Work Package 5 - MUSE · Work Package 5 Characterization of zoocoenoses ... Parole chiave: Insetti, foreste alpine, biomonitoraggio, bioindicazione, Trentino, Italia

Work Package 5

Characterization of zoocoenoses in correlation to environmental factors

Studio delle zoocenosi in correlazione ai principali fattori ambientali

167Studi Trent. Sci. Nat., Acta Biol., 81 (2004), Suppl. 1: 167-217Studi Trent. Sci. Nat., Acta Biol., 81 (2004), Suppl. 1: 167-217 ISSN 0392-0542© Museo Tridentino di Scienze Naturali, Trento 2005

1. INTroduCTIoN

Bioindication is based on the principle that orga-nisms or groups of organisms are strongly dependent on specific environmental factors and respond to changes in their habitat. This correlation is an ideal indication of changes in certain biotic and abiotic parameters (Schubert 1991; Carl 1996). The following basic assumptions must be considered in the choice of an organism as a bioindicator: the effect of an environmental factor should be recognised in the

Ecological assessment in alpine forest ecosystems: bioindication with insects (Auchenorrhyncha, Coleoptera (Staphylinidae), Lepidoptera)

Michael Carl1, Peter HueMer2*, adriano ZaNeTTI3 & Cristina SalVadorI4

1Institut für umweltforschung, Gollenbergstraße 12, G-82299 Türkenfeld, Germany2Tiroler landesmuseum Ferdinandeum, Naturwissenschaftliche Sammlungen, Feldstraße 11a, a-6020 Innsbruck, austria3Museo Civico di Storia Naturale, lungadige Porta Vittoria 9, I-37129 Verona, Italy4IaSMa research Center, Natural resources department, Via e. Mach 1, I-38010 San Michele all�adige (TN), ItalyVia e. Mach 1, I-38010 San Michele all�adige (TN), Italy*Corresponding author e-mail: p.huemer@tiroler-landesmuseum. at

SuMMarY - Ecological assessment in alpine forest ecosystems: bioindication with insects (Auchenorrhyncha, Coleop-tera (Staphylinidae), Lepidoptera)- The use of terrestrial arthropods as a management tool for monitoring status and change in forest ecosystems is critically evaluated. The suitability and value of three groups of insects as bioindicators (auchenorrhyncha, Coleoptera Staphylinidae, lepidoptera) was tested at two permanent monitoring areas situated in the southern alps (Trentino, Italy). one site is located at Pomarolo in a mixed thermophilous oak woodland (Quercetalia pubescentis), the other at Passo lavazè in a subalpine Norway spruce forest (Piceetalia excelsae). The fauna of both sites is dominated by a characteristic set of partially strongly specialized species. Prospects for bioindication using the groups examined are discussed. Their particular potential to monitor montane ecosystems and to assess a range of environmental problems, from pollution effects through to land use change up to long-term degradation and recovery of ecosystems, is highlighted. Indications for the choice of appropriate bioindicators are also provided.

rIaSSuNTo - Valutazione ecologica di ecosistemi forestali alpini: bioindicazione con gli insetti (Auchenorrhyncha, Coleoptera (Staphylinidae), Lepidoptera) - Nel presente lavoro sono trattati in modo sintetico problemi e vantaggi legati all�impiego di artropodi terrestri nell�ambito della gestione ambientale, quali strumenti di valutazione dello stato degli ecosistemi forestali e delle loro modificazioni. Il possibile utilizzo di tre gruppi di insetti come bioindicatori (auchenorrhyncha, Coleoptera Staphylinidae, lepidoptera) è stato testato in due siti forestali delle alpi meridionali (Trentino, Italia) sottoposti a monitoraggio integrato. la prima area è localizzata a Pomarolo in un querceto misto termofilo (Quercetalia pubescentis), la seconda in una pecceta subalpina (Piceetalia excelsae) a Passo lavazè. la fauna di entrambi i siti è dominata da una zoocenosi caratteristica costituita in larga misura da specie fortemente specializzate. Vengono presentate le prospettive per l�utilizzo dei gruppi indagati nel campo della bioindicazione, con particolare riferimento alle loro potenzialità nell�evidenziare gli effetti di varie perturbazioni ecologiche, quali inqui-namento atmosferico e cambiamenti d�uso del territorio, nonché le capacità di risposta e recupero degli ecosistemi. Sono inoltre fornite indicazioni per la scelta delle specie indicatrici più idonee.

Key words: Insects, alpine forest ecosystem, biomonitoring, bioindication, Trentino, ItalyParole chiave: Insetti, foreste alpine, biomonitoraggio, bioindicazione, Trentino, Italia

reaction of the bioindicator; the bioindicator should react as specifically as possible to a particular factor, to enable exclusion of other environmental influences; bioindicators are strongly related to specific habitats; as far as possible a simple method for registering the presence/absence of a bioindicator should be available.

The suitability of taxa as bioindicators therefore depends greatly on the problems to be monitored. Wit-hin the eFoMI project (ecological valuation of forest ecosystems by integrated monitoring) 4 taxonomic

168 Carl et al. Bioindication with insects

groups with a wide ecological amplitude have been selected as potential bioindicators; one of these groups (Collembola) will be dealt with elsewhere.

The authors responsible for the various taxa in this paper are M. Carl (auchenorrhyncha), P. Huemer (lepidoptera) and a. Zanetti (Coleoptera Staphylini-dae).

1.1. Auchenorrhyncha

The leafhopper fauna was examined at two moni-toring areas in alpine forest ecosystems in northern Italy. anthropogenic influence has led to air pollution at the examined sites (Tait 1995) which are the object of the monitoring programme. This research aimed to provide an inventory of the status of the monitor-ing sites as well as a comparative assessment of the lavazè, Pomarolo (Trentino), renon and Monticolo (South Tyrol) study areas using leafhoppers as bio-logical indicators. This was achieved by using, among other things:- comparison between the tested leafhopper fauna

and relevant environmental factors, location, and climatic factors;

- comparative assessment with earlier studies;- assessment of the species as specific and non-spe-

cific biological indicators with respect to selected environmental parameters;

- comparison of the observation areas with regard to their species inventory (dominance structure, Wain-stein- and evenness indexes, cluster analysis).

anthropogenic influence on air pollution in the eco-systems has been and is the object of numerous papers. different research projects have verified the influence of air pollution on leafhopper fauna (Heinrich 1984; Schäller 1985; Wiegand et al. 1994).

leafhoppers are present in almost all types of terrestrial habitats in the grass and canopy layers. The biocoenosis of auchenorrhyncha plant-sucking insects depends on the physiological state of their food plants (e.g. Prestidge & McNeill 1983), on climatic factors such as temperature, solar radiation, humidity etc. (e.g. Masters et al. 1998; Whittaker & Tribe 1998) as well as on the altitude of their habitat (Whittaker & Tribe 1996). achtziger (1996;1999), Niedringhaus (1999) and Bornholdt (2002) have demonstrated that leafhoppers are ideally suitable for bioindication and habitat assessment.

1.2. Coleoptera (Staphylinidae)

The use of staphylinid beetles as bioindicators in forest ecosystems has the advantage of offering a very wide ecological range, as they include species of the soil community, saproxylic species in trees and species linked to temporary microhabitats such as fungi, carrion and dung. Nevertheless, their use in bioindication was

until recently limited by poor knowledge of their taxonomy and autoecology. In more recent years this situation has improved as a result of several taxonomic reviews and ecological studies, some of which carried out in Italy. The rove-beetles in forest areas at Passo lavazè and Pomarolo were surveyed in 1992-2001 and the material obtained allowed us to outline in an exhaustive manner the characteristics of the community linked to the soil (Zanetti et al. 1997). The species linked to the forest vegetation, primarily the saproxylic species, were still little known and relevant information for bioindication was poor. The main target of research within the eFoMI project in 2003-2004 was the study of biodiversity levels for the Staphylinid fauna linked to trees in the mature spruce forest at the lavazè site and the developing broad-leaves wood at the Pomarolo site. a comparison with forest sites surveyed using the same sampling techniques will also be proposed in order to establish bioindication methods.

1.3. Lepidoptera

lepidoptera are a major group of mainly phyto-phagous insects with an exceptionally rich species diversity. In particular, forest ecosystems are of high value in terms of regional biodiversity (e.g. Hacker 1987; Hacker & Kolbeck 1996; Jaros & Spitzer 2002). due to their frequent host-specificity many species are excellent bioindicators for vegetation and climatic as-pects within forest ecosystems (albrecht 1990; Huemer 2002). a high proportion of most lepidoptera species inventories is more or less strongly specialized and usually restricted to only a few host-plant species of a single genus or family. Hence, the overall diversity is strongly linked to aspects of the vegetation and lepi-doptera can be used as bioindicators in terms of the composition, physiological state and development of plant communities. However, the presence/absence of species is also influenced by many other biotic factors (predators, parasites, physiology of host-plants etc.) and abiotic factors (climate, soil, exposure etc.). Within the two permanent monitoring areas chosen in the province of Trento, more than 200 species at lavazè and more than 700 at Pomarolo had already been recorded in the mid 1990s. It was planned to monitor any changes of species composition, with particular emphasis on specific indicators already identified during the 2001 and 2002 vegetation periods.

2. STudY areaS

2.1. Lavazè Pass

Geographical position/exposure: lavazè is situated close to the border of the provinces of Trento and Bol-zano at an altitude of around 1800 m; exposure mainly westerly or north-westerly at a slight incline.

169Studi Trent. Sci. Nat., Acta Biol., 81 (2004), Suppl. 1: 167-217

Geology: Bozner Quartzporphyr.average annual temperature: 3.9 °C; average annual

rainfall: 1100 mm (Salvadori & ambrosi 2005).Vegetation: sub-alpine spruce forest (order:

Piceetalia excelsae) with scattered Pinus cembra and rare Larix decidua. Species richness in terms of herbs is low and dominated by various grasses and ericaceae (Bolpagni et al. 2005).

2.2. Pomarolo

Geographical position/exposure: approx. 14 km SW of Trento at an altitude of around 700 m a.s.l.; exposure mainly south-easterly and easterly.

Geology: Cretaceous and Jurassic limestone.average annual temperature: 11.0 °C; average an-

nual rainfall: 1150 mm (Salvadori & ambrosi 2005).Vegetation: mixed termophilous oak forest (order:

Quercetalia pubescentis) with transformation to Quercus robur - Corylus avellana shrub-formations. Numerous thermophilous species such as Fraxinus ornus and Os-trya carpinifolia, but also small patches of xerophilous steppe vegetation (rigobello et al. 2005). In the northern part mesophilous plants with uniform Coryletum and a stock of Larix decidua and tall pine trees.

3. MeTHodS


The investigations were carried out from april 28, 2002 to october 19, 2003. Numerous collecting sites

were examined at lavazè and Pomarolo using pitfall traps, nets and suction-samplers as well as various tree traps. The collecting sites at the two monitoring areas were: l1-lavazè coniferous forest; l2-lavazè edge of the coniferous forest; l3-lavazè alpine pasture; l4-lavazè stream meadow; l5-lavazè forest road border; l6-lavazè southern lower situated slope; P1-Pomarolo meadow; P2-Pomarolo forest; P3-Pomarolo oak-forest trail border ; P4-Pomarolo forest road border; P5-Pomarolo steep rock. Sampling at the renon and Monticolo monitoring sites in the province of Bolzano had already been carried out during the 1996 and 2000 vegetation periods.

The leafhoppers were identified to species level. The taxonomical treatment mainly follows Bieder-mann & Niedringhaus (2004) as well as Holzinger et al. (2003).

The investigated areas were compared regarding their species inventory (evenness- and Wainstein-index). The multivariate method of the cluster analysis (single linkage method) was employed for the further evaluation of the data.


Sampling was carried out using window flight traps (three per site) and aerial pitfall traps (six per site) in the period May-october, 2003. The window flight traps were made up of two perpendicularly crossed Plexiglas panels (60 cm x 40 cm) connected below to a funnel and collecting jar. They were placed close to tree trunks. aerial pitfall traps consisted of plastic bottles (1.5 l) cut into two at the base of the upper

Site Kind of trap Symbol Tree species Height (m)

Pomarolo aerial pitfall trap Pb1 Pinus sylvestris 7Pomarolo aerial pitfall trap Pb2 Pinus sylvestris 7Pomarolo aerial pitfall trap Pb3 Pinus sylvestris 7Pomarolo aerial pitfall trap Qb1 Quercus cerris 7Pomarolo aerial pitfall trap Qb2 Quercus cerris 7Pomarolo aerial pitfall trap Qb3 Quercus cerris 7Pomarolo window flight trap Qa1 Quercus cerris 7Pomarolo window flight trap Qa2 Quercus cerris 3Pomarolo window flight trap Pa3 Pinus sylvestris 7

lavazè aerial pitfall trap Pb1 Picea abies 10lavazè aerial pitfall trap Pb2 Picea abies 10lavazè aerial pitfall trap Pb3 Picea abies 10lavazè aerial pitfall trap Cb1 Pinus cembra 10lavazè aerial pitfall trap Cb2 Pinus cembra 10lavazè aerial pitfall trap Cb3 Pinus cembra 10lavazè window flight trap Pa1 Picea abies 10lavazè window flight trap Pa2 Picea abies 3lavazè window flight trap Ca3 Pinus cembra 10

Tab. 1 - Traps at the monitoring sites lavazè and Pomarolo (Coleoptera, Staphylinidae).Tab. 1 - Tipi di trappole nei siti di monitoraggio di Lavazè e Pomarolo (Coleoptera, Staphylinidae).


conical section, which was inserted upside-down into the lower portion of the bottle, thus acting as a fun-nel. They were suspended from branches as high up as possible. More information on these techniques is given by Kaila (1993), Siitonen (1994) (window flight traps) and allemand & aberlenc (1991) (aerial pitfall traps). 18 traps were placed as shown in table 1 and the sampling data are given in table 2. The Staphylinid beetles collected were picked out from the samples and then identified at species level; the nomenclature follows löbl & Smetana (2004).

3.3. Lepidoptera

Sampling was carried out once a month and lasted from 27 June 2001 to 14 September 2001 at lavazè and from 14 March 2002 to 17 october 2002 at the Pomarolo site. Various methods having proved useful during earlier monitoring programs on lepidoptera were applied:

2 automatic light traps (eNTo-TeCH type; light source: 8W uV) for semi-quantitative registration of nocturnal lepidoptera. at Pomarolo one light trap was set up in the central part of the site, another in the southern part. at lavazè the light traps were situated in the north-eastern part of the monitoring site in the closed spruce forest and in the south-western part in the pasture and sub-alpine grassland, about 30 m outside the closed forest.

Illuminated white sheet (2x3 m; light source: HQl 125 W): semi-quantitative registration of nocturnal lepi-doptera. at Pomarolo the illuminated sheet was placed in the northernmost part of the site, while at lavazè it was placed in the southern part of the monitoring site, about 20 m outside the forest in the pastures and the side of the sheet facing the forest was illuminated.

1-2 light towers (light source: 15 W uV): semi-quantitative registration of nocturnal lepidoptera. at Pomarolo one light tower was used in the southern part of the site, the other east of the central section along the road. at lavazè a single light tower was placed in the central part of the monitoring site within the subalpine spruce forest.

Time registration: semi-quantitative registration of diurnal lepidoptera (butterflies and diurnal moths) for one hour (lavazè) and two hours (Pomarolo) respec-tively. at Pomarolo registration took place along the path at the site and along the road situated to the east, above the site. at lavazè time registration was under-taken outside the closed forest in an area of about 25 m, between the spruce wood and the pastures.

Visual registration: qualitative registration of larvae and/or leaf-miners, all over both sites.

The collected or observed specimens were identi-fied to species level; the systematic treatment mainly follows Huemer & Tarmann (1993). The similarity of species composition between both sites and during vari-ous sampling periods was measured using Sörensen´s co-efficient: CS (%) = 2G/S

a + S

B x 100 (G = number of

common species in sites a and B; Sa, S

B = total number

of species in sites a and B).Species diversity was also compared to that of

earlier samples, using the absolute species turnover rate (Turnover T

B). This measurement refers to the

percentage of exchanged species in relation to the initial species number (Hausmann 1991): T

B (%) =

(X1 + X

2) x 100/S

1 (S

1 = initial species number; X

1 =

number of newly recorded species; X2 = number of

missing species).The whole sample of species was attributed to

feeding categories as defined by Jaros & Spitzer (2002): monophagous (1) species feeding on a single species; monophagous (2) species feeding on a single genus; oligophagous (3) species feeding on a group of closely related genera; oligophagous (4) species feeding on a single family; polyphagous (5) species feeding on more than one family (woody plants); polyphagous (6) species feeding on more than one family (herbs); polyphagous (7) species feeding on more than one family (woody plants and herbs); mycophagous (8) species feeding on lichens, algae, fungi (rotten wood) or moss; detritophagous (9) species feeding on detritus (including keratophagous species). The monophagous (1) + (2) and oligophagous (3) + (4) feeding categories were classified. Furthermore, the whole sample of species was attributed to host-plant classes which are used to give a more precise picture in terms of specialization within feeding categories. Various host-plant groups forming natural entities for lepidoptera (deciduous trees/bushes, conifers, grass, herbs, lichens, mosses etc.) were defined as host-plant classes and their species composition was evaluated separately.

lavazè Pomarolo

--------

10/06/0324/06/0308/07/0322/07/0305/08/0319/08/0302/09/0316/09/0330/09/0314/10/03

--

09/04/0323/04/0307/05/0321/05/0304/06/0318/06/0302/07/0316/07/0331/07/0313/08/0327/08/0310/09/0324/09/0308/10/0322/10/03

Tab. 2 - Collection dates (Coleoptera, Staphylinidae).Tab. 2 - Date di campionamento (Coleoptera, Staphylini-dae).


4. reSulTS


2,732 individuals from 72 leafhopper species were collected at lavazè and Pomarolo from 2002 to 2003. a complete checklist of species detected in the study areas is reported in the appendix 1.

The number of species collected in the lavazè (39 species) and renon (30) sub-alpine research sites was insignificantly to considerably lower than the number of species collected at the Pomarolo (44) and Monticolo (45) lowland research sites. It must be considered that species numbers at renon and Monticolo are based on two collecting periods and that habitat l6, situated at a lower altitude, strongly differs from the lavazè Pass habitats. If the l6 (11) species are compared to the total number of species from lavazè, then the difference (28) becomes clearer in relation to the Pomarolo and Mon-ticolo sites, which are situated at a lower level. This is about the same number of species as at renon (30).

The Wainstein- and evenness indices are shown in table 3. The Wainstein indices of many compared habitats are remarkably low. This applies to compari-sons between monitoring sites as well as between habitats lying close together.

The evenness indices of Pomarolo are considerably higher than those of lavazè (not considering l6). due to the eudominance of one species and the low overall number of species, l1 shows the lowest evenness index of all surveyed habitats. The l6, P2, P3 and P4 habitats show the highest evenness indices of all.

The multivariate method of cluster analysis (shown in Figs 7, 8) was used for further evaluation of the data, results showing the following distance measures: The biggest distance is between P1 and the other habitats with approx. 80,000, followed by l3 which shows a distance of approx. 20,000 to the next (P5). all other habitats form relatively dense clusters with a distance of less than 10,000. P2/P3 and l4/l5 prove to be par-ticularly similar clusters.


1,395 specimens belonging to 15 species (11 in Pomarolo and 5 in lavazè) were collected (Tab. 4). The species are listed in appendix 2; records on the auto-ecology of species are also provided, with particular regard to informations useful in bioindication.

at Pomarolo most individuals were captured in pitfall traps, with only three specimens collected in window traps. on the contrary, at lavazè 43% of specimens were collected using window traps. In both localities a clear dominance of one species was noted (Aleochara sparsa at Pomarolo and Quedius mesomelinus skoraszewskyi at lavazè) and the Shannon index was therefore low (Pomarolo H

s = 1.011 and

lavazè Hs = 0.049).

4.3. Lepidoptera

4.3.1. Lavazè

272 species of lepidoptera were recorded between 1992 and 2001 (appendix 3). The species inventory of the 2001 sampling period included 106 species, belonging to 21 families.

The most species rich groups were Geometridae (32 spp.), Noctuidae (20 spp.), Crambidae (13 spp.), Tortricidae (10 spp.), Gelechiidae (6 spp.) and Nymphalidae (6 spp.) (24 spp.). The family structure is poor, but in a similar way to Pomarolo, Geometridae (30%) and Noctuidae (19%) are the most species rich groups though with a distinctly higher percentage. altogether 16 species were newly recorded at the site, whereas 182 species were already known from earlier years but not registered in 2001. Turnover rates are very high, ranging from 82.2% between the sampling periods of 2001 and 2000 and 76% between 2001 and

Collecting sites

Wainsteinindex

Collecting site

evennessindex

l1/P2 0 l1 0.33

l2/P3 0 l2 0.77

l3/P1 0 l3 0.69

l5/P4 6 l4 0.42

l6/P5 9 l5 0.77

l1/l2 31 l6 0.90

l1/l3 22 P1 0.67

l2/l3 50 P2 0.92

P1/P2 53 P3 0.84

P1/P3 42 P4 0.86

P1/P4 47 P5 0.75

l4/P4 3

Tab. 3 - Wainstein- and evenness indices of the monitoring sites lavazè and Pomarolo (auchenorrhyncha). Collecting sites: l1: lavazè coniferous forest; l2: lavazè edge of the coniferous forest; l3: lavazè alpine pasture; l4: lavazè stream meadow; l5: lavazè forest road border ; l6: lavazè southern lower situated slope; P1: Pomarolo meadow; P2: Pomarolo forest; P3: Pomarolo oak-forest trail border ; P4: Pomarolo forest road border; P5: Pomarolo steep rock.Tab. 3 - Indici di Wainstein e di evenness nei siti di moni-toraggio di Lavazè e Pomarolo (Auchenorrhyncha). Siti di campionamento: L1: Lavazè bosco di conifere; L2: Lavazè margine del bosco di conifere; L3: Lavazè pascolo alpino; L4: Lavazè prato umido; L5: Lavazè margine di strada forestale ; L6: Lavazè versante meridionale inferiore; P1: Pomarolo prato; P2: Pomarolo bosco misto; P3: Pomarolo fascia di margine di bosco di querce ; P4: Pomarolo margine di strada forestale; P5: Pomarolo parete rocciosa.


Species

Pom

arol

o

lav

azè

Bos

co F

onta

na

Nor

th. a

penn

in

Tarv

isio

eucoenic species in tree holes Euryusa optabilis xHapalaraea pygmaea xIschnoglossa ?prolixa xPhyllodrepa ioptera xPhyllodrepa melanocephala melanocephala xQuedius cruentus xQuedius maurus xStichoglossa semirufa xTotal 2 0 2 2

under barks Atheta picipes xLeptusa pulchella x xNudobius lentus xPhloeonomus punctipennis xPhloeopora corticalis xPhloeopora sp. xPhloeostiba plana xPlacusa atrata xPlacusa complanata xPlacusa depressa x x xPlacusa pumilio x x xPlacusa sp. x xPlacusa tachyporoides xTotal 3 1 4 5 8

nests in tree holes Haploglossa bernhaueri xfermented sap of trees Thamiaraea cinnamomea xnests of Hymenoptera Velleius dilatatus x

Total 2 1Total of eucoenic 5 1 8 8 8

Tycocoenic species in tree holes Coryphium angusticolle xQuedius mesomelinus mesomelinus x xQuedius mesomelinus skoraszewskyi x x

in nests Aleochara sparsa x x xAtheta harwoodi x x x xAtheta vaga x x x x x

Total tycocoenic 4 2 2 5 4

eucoenic + tycocoenic 9 3 10 13 12Number of xenocoenic 2 2 12 20 6

Tab. 4 - Captures of saproxylic species in northern Italian sites (Coleoptera, Staphylinidae).Tab. 4 - Catture di specie saproxiliche in aree dell’Italia settentrionale (Coleoptera, Staphylinidae).


remaining species are mycophagous or detritophagous. The diversity of species within feeding classes (Fig. 2a) reflects the importance of herbs (45 spp.), grass (26 spp.) and deciduous trees (25 spp.) as host-plants, but species feeding on conifers are also well represented (14 spp.). The number of taxa restricted to lichens and moss (6 spp.) is low.

4.3.2. Pomarolo

1037 species of lepidoptera were recorded between 1992 and 2002 (appendix 3). The inventory of the 2002 sampling period included 660 species belonging to 49 families. The most species rich groups are the Geome-tridae (125 spp.), Noctuidae (100 spp.), Tortricidae (87 spp.), Gelechiidae (41 spp.), Pyralidae (28 spp.), Cram-bidae (26 spp.) and Gracillariidae (24 spp.). The family structure is extraordinarily diverse and characteristic of Central europe. However, in most habitats Noctuidae exceed Geometridae in terms of number of species, even in some forests (e.g. Huemer & Tarmann 2001). altogether, 96 species were newly recorded at the site, whereas 281 species already known from earlier years

lavazè S1

X1

X2

TB %

2001/1992-95 217 28 137 76.0

Pomarolo S1

X1

X2

TB %

2002/1992-95 726 194 161 48.9

Tab. 5 - Species turnover rates 2001/02 - 2000 and 2001/02 - 1992/95 (lepidoptera). S

1= initial species number; X

1=

number of newly recorded species; X2= number of missing;

TB (%)= absolute turnover rate.

Tab. 5 - Turnover di specie 2001/02 - 2000 e 2001/02 - 1992/95 (Lepidoptera). S

1= numero iniziale di specie; X

1= numero

di specie nuove; X2= numero di specie mancanti; T

B (%)=

valore assoluto di turnover.

0%

10%

20%

30%

40%

50%

60%

70%

80%

90%

100%

2001 2000 1992-95Lavazé

polyphagous (6)polyphagous (5)oligophagous (4)oligophagous (3)monophagous (2)monophagous (1)

0%

10%

20%

30%

40%

50%

60%

70%

80%

90%

100%

2002 2000 1992-95Lavazé

polyphagous (6)polyphagous (5)oligophagous (4)oligophagous (3)monophagous (2)monophagous (1)

Fig. 1a - Proportion of species in feeding categories during various sampling periods (lavazè) (lepidoptera). (lepidoptera)..Fig. 1a - Distribuzione percentuale di specie in categorie trofiche nei diversi periodi di campionamento (Lavazè) (Le-pidoptera).

Fig. 1b - Proportion of species in feeding categories during various sampling periods (Pomarolo) (lepidoptera).Fig. 1b - Distribuzione percentuale di specie in categorie trofiche nei diversi periodi di campionamento (Pomarolo) (Lepidoptera).

1992-1995, respectively (Tab. 5). The proportion of species within feeding classes

proves that specialised species are of high importance for the faunal composition (Fig. 1a). Twenty-one spp. (19.9%) were classified as monophagous and a further 27 spp. (25.4%) as oligophagous, whereas about 44% of the species inventory (47 spp.) is polyphagous. The


were not registered. In comparison to 2000 a reduction of 8% in the species observed was noted, a percentage which is regarded as insignificant. Turnover rates are moderately high, ranging from 53.3% between the sample periods of 2002 and 2000 and 48.9% between 2002 and 1992-1995 respectively (Tab. 5).

The proportion of species in the various feeding class-es proves the importance of specialized species (Fig. 1b). 196 spp. (30.2%) were classified as monophagous and a further 159 spp. (24.5%) as oligophagous, whereas only about one third of the species inventory (237 spp. = 36.5%) is more or less unspecialized polyphagous. The remaining species are mycophagous, detritophagous or their feeding classes are unknown. The proportion of species within feeding classes reflects the importance of deciduous trees (289 spp.) and herbs (257 spp.) as host-plants, whereas species feeding on grass (49 spp.) or conifers (40 spp.) are less well represented. The number of taxa restricted to lichens and moss (21 spp.) and rotten wood (15 spp.) is moderately high.

The overall similarity between the species commu-

0%

20%

40%

60%

80%

100%

2001 2000 1992-95Lavazé

otherslichens, mossgrassherbsconifersdeciduous trees/shrubs

0%

20%

40%

60%

80%

100%

2002 2000 1992-95Pomarolo

othersrotten woodlichens, mossgrassherbsconifersdeciduous trees/shrubs

Fig. 2a - Proportion of species in host-plant classes (lavazè) (lepidoptera).Fig. 2a - Distribuzione percentuale di specie in base al tipo di pianta ospite (Lavazè) (Lepidoptera).

Fig. 2b - Proportion of species in host-plant classes (Pomarolo) (lepidoptera).Fig. 2b - Distribuzione percentuale di specie in base al tipo di pianta ospite (Pomarolo) (Lepidoptera).

lav95

lav00

lav01

Pom95

Pom00

Pom02

0.04 0.2 0.36 0.52 0.68 0.84 1

Fig. 3 - Similarity between sites/different sampling periods (lav95 = lavazè 1995 etc; Pom95 = Pomarolo 1995 etc) (based on uPGMa – Sörensen�s Coefficient) (lepidopte-ra). Fig. 3 - Similarità tra siti e periodi di campionamento (Lav95= Lavazè 1995 ecc.; Pom95= Pomarolo 1995 ecc.) (basato su UPGMA – Indice di Sörensen) (Lepi-doptera).


uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

Pom02Pom95Pom00lav01lav00lav95

0.04 0.2 0.36 0.52 0.68 0.84 1

uPGMa

Sörensen's Coefficient

Pom02Pom00Pom95

0.64 0.7 0.76 0.82 0.88 0.94 1

Sörensen's Coefficient

Sörensen's Coefficient Sörensen's Coefficient









a b

c d

e f

g h

Fig. 4 - Species similarity within feeding categories (lepidoptera).a: monophagous (1)+(2); b: oligopha-gous (3)+(4); c: polyphagous (5); d: polyphagous (6); e: polyphagous (7); f: mycophagous (8); g: detritophagous (9); h: monophagous (2) – Quercus.Fig. 4 - Similarità di specie all’interno delle categorie trofiche (Lepidoptera).a: monofagi (1)+(2); b: oligofagi (3)+(4); c: polifagi (5); d: polifagi (6); e: polifagi (7;, f: micofagi (8); g: detriti-vori (9); h: monofagi (2) – Quercus.

nities of lavazè and Pomarolo (Fig. 3) was calculated using Sörensen�s coefficient. It is very low and ranges between 5.8% and 14.4% of the species inventory in the different years of sampling. Within locations, species similarity is moderately high at Pomarolo with about 73% conformity during the last two sampling periods and not less than 67.3% over all periods. However, lavazè is quite different in this respect with only a moderately low similarity among sampling periods, ranging from 58.9% to 52.2%. These differences are mainly explained by species turnover processes.

Cluster analysis proves the relationship between feeding categories and species concordance (Fig. 4). It was not really surprising to find that the differences between the two sites are very high throughout the feeding categories.

Monophagous (1)+(2) (Fig. 4a): the species similar-

ity ranges between 61% - 65.9% (Pomarolo) and 36.1% - 50.6% (lavazè).

oligophagous (3)+(4) (Fig. 4b): the species simi-larity ranges between 68.9% - 73.5% (Pomarolo) and 44.4% - 54.5% (lavazè).

Polyphagous (5) (woody plants) (Fig. 4c): the spe-cies similarity ranges between 78.2% - 84.0% (Poma-rolo) and 48.6% - 64.0% (lavazè).

Polyphagous (6) (herbs) (Fig. 4d): the species simi-larity ranges between 63.1% - 74.2% (Pomarolo) and 49.4% - 62.4% (lavazè).

Polyphagous (7) (herbaceous and woody plants) (Fig. 4e): the species similarity ranges between 68.6% - 72.2% (Pomarolo) and 42.1% - 62.5% (lavazè).

Mycophagous (8) (Fig. 4f): the species similarity ranges between 77.1% - 81.1% (Pomarolo) and 58.3% - 66.7% (lavazè).


0

10

20

30

40

50

60

70

80

90

L1L2L3L4L5L6

%

species number1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 310

Fig. 5 - dominance structure of the auchenorrhyncha-coenosis of the mo-nitoring site lavazè (sub-sites compare Tab. 3).Fig. 5 - Struttura di dominanza della cenosi di Auchenorrinchi nell’area di monitoraggio di Lavazè (per le sottoaree vedere Tab. 3).

0

10

20

30

40

50

60

70

80

90

P1P2P3P4P5

species number

%

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 310

Fig. 6 - dominance structure of the au-chenorrhyncha-coenosis of the monito-ring site Pomarolo (sub-sites compare Tab. 3).Fig. 6 - Struttura di dominanza della cenosi di Auchenorrinchi nell’area di mo-nitoraggio di Pomarolo (per le sottoaree vedere Tab. 3).

detritophagous (9) (Fig. 4g): the species similarity ranges between 68.0% - 68.2% (Pomarolo) and 57.1% - 80.0% (lavazè).

Monophagous (10) – Quercus (Fig. 4h) (only Pomarolo): the species similarity ranges between 64.1% - 85.4% (Pomarolo). The similarity between species communities restricted to Quercus is exceptionally high with 85.4% similarity between 2000 and 2002.

5. dISCuSSIoN


The number of species collected and their vertical distribution prove the influence of altitude, climatic and botanic parameters in the form of a reduction of the species number and species diversity.- out of 111 species collected at the 4 monitoring

sites of Trentino-South Tyrol, 76 (68.5%) were found exclusively in one research site.

- out of the 111 species 25 (22.5%) were found in two research sites.

- only 10 species (9%) were found in three or four research sites.These results are impressive proof of the powerful

capacity of leafhopper fauna to adapt to habitats with varying environmental parameter sets. This demonstrates the excellent suitability of the leafhopper fauna for bioindication.

as regards the dominance structure of the leafhop-per fauna at lavazè (IT03) and Pomarolo (IT04) the following can be stated (Figs. 5, 6):

IT03 and IT04 show considerably different dominance structures of leafhopper fauna.

The proportion of eudominant or dominant species at IT03 is higher and the proportion of recedent and subrecedent species is lower than at the Pomarolo site.

exceptions are the following:- l3 (alpine pasture), without eudominant species and

with a higher proportion of recedent and subrece-dent species, and

- l6, without eudominant or dominant species and with a higher proportion of recedent and subrece-dent species.


The dominance structure at Pomarolo is relatively balanced, with very flat curves. None of the species are eudominant, three are dominant. The proportion of recedent and subrecedent species is fundamentally higher than at lavazè.

It will be necessary to examine whether these significant differences in the dominance structure are based on anthropogenic influence, such as air pollution or limiting parameters (altitude), or other factors of stress through further monitoring.

Wainstein indices: at lavazè, the similarities of habitats are altogether lower than at Pomarolo. The comparison of lavazè/Pomarolo habitats leads to no or nearly no similarity. These results can only be based on the strongly differing environmental parameter sets of the monitoring sites. If the habitats show a low similar-ity despite being situated close together, these results are clearly more remarkable and must also be based on strongly differing environmental parameter sets (see table 3 l1/l2, l1/l3). This again demonstrates the excellent suitability of leafhoppers for bioindication in alpine forest ecosystems.

With regard to evenness indices the following can be summarised:- the evenness indices at Pomarolo (P1 to P5) are al-

together higher than those of lavazè (l1 to l5, not considering l6). This is a strong indication of the reaction of the leafhopper fauna to the environmen-tal parameter set of a mountainous coniferous forest close to nature. However, anthropogenic influences such as air pollution cannot be excluded;

- the l6 lower altitude habitat is widely separated, having no or little ecological or climatic links to the pass habitats of lavazè;

- due to the eudominance of one species and the low species number, l1 shows the lowest evenness index of all habitats. This is again a strong indication of the reaction of the leafhopper fauna to the environ-mental parameter set of a mountainous coniferous forest close to nature;

- the l6, P2, P3 and P4 habitats show the highest

evenness indices of all the habitats. Such high evenness indices are very unusual even in habitats close to nature and are therefore particularly remark-able.The results of the cluster analysis can be sum-

marised as follows (Figs. 7, 8):- the distance measures and some of the clusters

are remarkable because they are partially in clear contradiction to the Wainstein indices. The reason for this is firstly the application of the “single link-age method” and secondly the fact that with the Wainstein index only two habitats are tested while the cluster analysis is comparing all the habitats examined with one another;

- P1 and l3 show a completely different structure for leafhopper fauna as compared to other habitats;

- habitat l6 is shown to be a distinctly separate lo-cation with no or little ecological or climatic links with the lavazè pass habitats because of a narrow cluster formation with P2, P3 and P4;

- l1, l2, l4 and l5 are habitats more or less associ-ated with woods. This leads to a narrower cluster formation in the comparison of all the habitats examined. This may be explained by more similar biotic and abiotic parameters in comparison to other habitats;

- the distance of P5 to cluster P2 - l1 is slightly larger than the distances within the dense P2 - l1 cluster. Furthermore there is an exclusive cluster formation within lavazè habitats. a difference in relation to the leafhopper fauna of all other habitats is suspected;

- as regards the cluster analysis of all four monitoring sites (Fig. 8) it can be stated that on the one hand subsisting clusters such as P2/P3 or l4/l5 are a strong indication of very similar biotic and abiotic parameters in comparison to other clusters. on the other hand a mixture of Bolzano province /Trento province clusters leads to the evaluation that some of the auchenorrhyncha biocoenosis must be more similar than within the monitoring sites.

Fig. 7 - Cluster analysis of the auche-norrhyncha-coenosis of the monitoring sites lavazè and Pomarolo (sub-sites compare Tab. 3).l= lavazè 2002 - 2003; P= Pomarolo 2002 - 2003.Fig. 7 - Analisi dei cluster della cenosi di Auchenorrinchi nei siti di monitoraggio di Lavazè e Pomarolo (per le sottoaree confrontare la Tab. 3).L= Lavazè 2002 - 2003; P= Pomarolo 2002 - 2003.



The results show that 72% of species and 92% of specimens collected at Pomarolo are more or less closely linked to trees, but 40% of species and only 0.3% of the specimens from Passo lavazè are saproxylic. These results have been compared with those from other North Italian localities recently investigated using the same methods (Tab. 4). at Bosco della Fontana, an ancient Quercus and Car-pinus wood in a flat area near Mantua (lombardy), 21 species, 10 of which saproxylic, were obtained using 20 aerial pitfall traps adopted from June to November (Tagliapietra & Zanetti 2002); 33 species, 14 of which were saproxylic, were collected at five sites in the Northern apennines (provinces of Parma and lucca) mostly in beech woods, using 18 window traps adopted from spring to autumn (Tagliapietra & Zanetti 2003); unpublished material relating to 15 window flight traps placed in the state forests of Tarvisio (Friuli, province of udine) for 5 months (June-November) contained 18 Staphylinid spe-cies, of 12 which which saproxylic. These numbers are not sufficient for differentiating taxocoenoses per se, and a critical analysis of the species is also necessary. The total number of species collected varies significantly, mostly because the number of xenocoenic species is very variable. This fact could depend on the structure of the forest and on the position of the traps. Saproxylic species occur in very similar numbers. They are brought together in two main groups in table 4, eucoenic species living in decaying trees only and tycocoenic species also living in other habitats. eucoenic species are mostly linked to two different microhabitats; the bark where they live mostly as predators following attack by

Scolytidae and other xylophagous insects, and the tree hollows where they can be associated with the nests of invertebrates (mostly Hymenoptera) and ver-tebrates (birds and mammals). The eucoenic species collected at Pomarolo are not highly specialized and are represented by species living under bark (Placusa spp. pl.) and by species with a wide ecological range which are not rare in tree hollows. More specialized species were sampled in the apennine and Tarvisio sites. The material from lavazè is rather enigmatic. It also demonstrates the unreliability of aerial traps in coniferous woods and could be related to the scarcity of tree hollows and to the presence of resinous sap.

5.3. Lepidoptera

5.3.1. Lavazè

Species diversity is low, mainly due to the modera-tely uniform vegetation. The site shows a significant reduction of overall species diversity compared to 1992-95 and 2000. Furthermore the turnover rate between 2000 to 2001 is extremely high (82%) and a comparison with earlier periods also shows an ex-tremely high turnover, standing at about 76% (Tab. 5). However, such turnover rates are quite characteristic of similar studies (e.g. Huemer & Tarmann 2001) and Hausmann (1991) notes a turnover rate of about 50% even at short recording intervals. When considering the autecology of single species, the turnover rates probably mainly refer to methodological problems and can be considered artificial, at least in most cases. Numerous decisive factors such as trapping intervals, seasonal periods, weather conditions, trap construction, identi-fication problems and others may reduce the number of recognized species. Trapping results in summer 2001

Fig. 8 - Cluster analysis of the auchenor-rhyncha-coenosis of the monitoring sites ritten, Montiggl, lavazè and Pomarolo (sub-sites compare Tab. 3).l= lavazè 2002 - 2003; P= Pomarolo 2002 - 2003; ra= ritten outside monitoring site; rV= ritten within monitoring site; Ma= Montiggl outside monitoring site; MV= Montiggl within monitoring site.Fig. 8 - Analisi dei cluster della cenosi di Auchenorrinchi nei siti di monitoraggio di Renon, Monticolo, Lavazè e Pomarolo (per le sottoaree confrontare la Tab. 3).L= Lavazè 2002 - 2003; P= Pomarolo 2002 - 2003; RA= Renon esterno area di monitoraggio; RV= Renon interno area di monitoraggio; MA= Monticolo esterno area di monitoraggio; MV= Monticolo interno area di monitoraggio.


were undoubtedly reduced by the exceptionally bad weather conditions on two evenings.

The proportional variation of species within feeding categories and host-plant classes shows only limited changes compared to the decline of the total number of taxa observed (Figs 1a, 2a). The proportion of species associated with various herbs has declined during the last few years, whereas that associated with grass has increased. Probably, this is due to recent intensifica-tion of pasture fertilisation in the southern part of the site. The proportion of species feeding on conifers is substantially constant, whereas the percentage of spe-cies numbers linked to deciduous trees/shrubs varies to a higher degree, probably because of less than ideal conditions.

5.3.2. Pomarolo

Species diversity is very high, due to the richness of flora and the diverse structure of microhabitats. The site shows no significant reduction of species diversity compared to 1992-95 and 2000, but high turnover rates can still be observed. The concordance of the species inventory between the various periods is rather low. at Pomarolo the turnover rates, in addition to methodological problems, mainly depend on the moderately high number of rare species and finally on a long-term change of habitats. In particular, the clearing of forests in the surrounding area significantly contributed to an increase in herbivorous species in 2000, which is now slowly declining due to succession processes. Furthermore, the study site itself is gradually changing, the forest becoming more closed-in due to the absence of human impact. However, the proportional variation within feeding categories and host-plant classes reflects only minor changes (Figs. 1b, 2b) during recent years. This is of particular interest when we take into consideration the high species turnover rates (Tab. 5). The percentage of species associated with various herbs has declined slowly, whereas species feeding on deciduous trees/bushes have increased.

The proportion of species in feeding categories and host plant-classes proved stable, though significantly different in both monitoring sites (Figs. 1, 2), and should be regarded as a key information about the stability of lepidoptera communities and their habitats. However, a minimum number of excursions using different me-thods is crucial to assess all host-plantclasses. The changes in host-plant composition can be assessed with about 5 excursions/year in Pomarolo and 3 excursions/year in lavazè during the main vegetation period. al-though the proportion of species in different host-plant classes showed only limited variation throughout the years, some interesting differences could be observed in the overall similarity within feeding-categories, which underlines a higher degree of variability (Fig. 4). The similarity of species communities is highest within the polyphagous species-community restricted to woody

plants at Pomarolo, with Sörensens´s coefficient reach-ing 84% between 2000 and 2002. Monophagous and oligophagous species-groups seem to be less suitable for long-term monitoring of forest ecosystems, due to high turnover rates, probably connected with ongoing succession processes in both study sites. Therefore, the species similarities are only average, with the excep-tion of some monophagous species on woody plants (e.g. Quercus/Fagaceae (Fig. 4h) and Picea/Pinaceae). The high similarity within the mycophagous feeding category in both sites is based on a very limited number of species, thus at high risk of considerable seasonal variability. This problem also applies to detritophagous feeding categories which show the highest similarities in lavazè.

6. CoNCluSIoNS


Censuses of the damage to woods in South Tyrol yielded indications that the degree of damage to trees could be linked to climatic factors as well as to proven changes in leafhopper fauna (Carl 2001).

The environmental factors controlling the individual and species numbers of the leafhopper fauna are multiple, complex and not fully understood. a link to climatic change has not been definitively proved but is very probable. The results of this study demonstrate, in harmony with other studies, the excellent suitability of leafhopper fauna for bioindication of changes in important environmental factors such as rainfall, temperature and air pollution.


The importance of the saproxylic fauna in forests has been highlighted mostly in recent years (Tagliapi-etra 2003), and the presence of species connected to particular arboreal niches in restricted areas is related to mature and long-standing situations. Some particu-larly specific Staphylinid species such as Trigonurus mellyi in the Northern apennines (Tagliapietra & Zanetti 2003) could be considered bioindicators of high value, like other saproxylic Coleoptera (Cucujus cinnaberinus, Osmoderma eremita) included in the appendix II Fauna - Flora Habitats directive (92/43/eeC May, 21 1992) of the european Council.

aerial traps are effective for a full investigation of Staphylinid taxocoenoses in forest sites, as they give a good support to information obtained by pitfall trap-ping on the surface of the soil. They work best of all in broad-leaved woods, while they are less effective in coniferous woods.

The data obtained from the Pomarolo site, which does not include very specialized saproxylic Staphy-linids, fits in well with the characteristics of the area,


which has a high diversity of trees but lacks large old trees with cavities. The records from the lavazè site provide little information.

Further investigation of Staphylinid beetles using aerial traps in Northern Italy is recommended, as this would allow a method to be established, which could provide more helpful information.

6.3. Lepidoptera

Numerous species of lepidoptera are stenotopic and related to very specific habitats, particularly when larval habits are considered. Furthermore, the sampling of adul-ts is easy in many cases (not for some cryptical species or a number of microlepidoptera), though diversity indices are strongly affected by collecting methods and sample size. on the other hand, the reaction of species to a single environmental factor is extremely difficult to prove and in many cases unknown, a problem which applies to most organisms. Therefore, a monitoring programme with lepidoptera is not appropriate for precise bioindication without restrictions for the time being, though it may help to recognize slow (or fast) changes in the environ-mental quality of sites. Indeed, initial data sampled over several years demonstrate a slow change in the vegetation within the lavazè and Pomarolo monitoring sites when the host-plant relationships of the lepidoptera fauna as a whole are compared. even a very limited number of excursions indicates such changes in plant composi-tion (Niedermoser 2001; Huemer 2002) and also in the health of the host-plants (ellis et al. 2001). a complete species survey is important from the point of view of fauna and biogeography but did not prove necessary for bioindication.

The high number of stenotopic species observed both at lavazè and Pomarolo (Huemer 1997, 2002) is a further assumption for successful host-plant oriented bioindication. Changes in the vitality of host-plants are indicated e.g. by the fauna of highly specialized and frequently strictly monophagous leaf miners (el-lis et al. 2001) which could be particularly useful as bioindicators at the Pomarolo site in the future. With a very low number of leaf miners, lavazè is not suitable for bioindication within this ecological association, but at both sites several additional key species have been determined. To recapitulate, species belonging to communities of woody plants should be selected as bioindicators at both sites in the future, particularly if represented in high numbers in previous sampling sessions. assessment of additional taxa from all feed-ing classes would be helpful for monitoring short-term changes within the habitats.

aCKNoWledGeMeNTS

our special thanks go to Paolo ambrosi and Cristina Cont (Istituto agrario di San Michele all�adige, Trento)

for the support received during this study. a.Z. is also greatly indebted to andrea Tagliapietra (Verona) for bibliographic research and for helpful discussion of the results.

This work was supported by a funding from the autonomous Province of Trento (eFoMI project: ecological Valuation in alpine Forest ecosystems by Integrated Monitoring). earlier research activities at the renon and Monticolo monitoring sites were financed by the Forstwirtschaftsinspektorat of the autonomous Province of Bolzano and we would like to thank Stefano Minerbi (Bolzano) for his support.

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DelphaciDae

Kelisinae Anakelisia perspicillata (Boh.) (New for Pomarolo)Stenocraninae Stenocranus minutus (F.)Stirominae Eurysa lineataEurysa lineata (Perris) Eurybregma nigrolineataEurybregma nigrolineata Scott (New for lavazè) Stiroma affinisStiroma affinis Fieb. Ditropsis ��avipesDitropsis ��avipes (Sign.)delphacinae Acanthodelphax spinosus (Fieb.) Dicranotropis ��avipesDicranotropis ��avipes (Fieb.) Xanthodelphax ��aveolusXanthodelphax ��aveolus (Flor) (New for lavazè) Toya propinquaToya propinqua (Fieb.) Javesella discolor (Boh.) Javesella forcipata (Boh.) Ribautodelphax pungensRibautodelphax pungens (rib.) (New for Pomarolo)(New for Pomarolo)

issiDae

Issinae Issus coleoptratus (F.) Mycterodus orthocephalus Ferr. Hysteropterum schaefferiHysteropterum schaefferi Metc.

cicaDiDae

Cicadetta montana (Scop.)

cercopiDae

Cercopinae Cercopis sanguinolenta (Scop.) Cercopis vulnerata rossiaphrophorinae Neophilaenus infumatus (Hpt.) Neophilaenus lineatusNeophilaenus lineatus (l.) Aphrophora alniAphrophora alni (Fall.) Philaenus spumarius (l.)

MeMbraciDae

Centrotinae Centrotus cornutus (l.)

cicaDelliDae

Megophthalminae Megophthalmus scanicus (Fall.)Macropsinae Macropsis infuscataMacropsis infuscata (Sahlb.) (New for lavazè)(New for lavazè) Macropsis impura (Boh.) Macropsis fuscula (Zett.)agalliinae Dryodurgades dlabolai Wagn. (New for lavazè and Pomarolo)

appendix 1 - auchenorrhyncha: list of species recorded at lavazè and Pomarolo (2002-2003).Appendice 1 - Auchenorrhyncha: lista delle specie riscontrate a Lavazè e Pomarolo (2002-2003).

Idiocerinae Tremulicerus tremulae (estl.) (New for lavazè)(New for lavazè) Populicerus populi (l.)aphrodinae Aphrodes bicincta (Schr.) (New for Pomarolo) Planaphrodes nigrita (Kbm.) Anoscopus albifrons (l.)Cicadellinae Evacanthus acuminatus (F.) Evacanthus interruptus (l.) Cicadella viridisCicadella viridis (l.)Typhlocybinae Alebra albostriella (Fall.) Erythria manderstjernii (Kbm.) Emelyanoviana mollicula (Boh.) Forcipata obtusaForcipata obtusa Vidano (New for lavazè and Pomarolo) Empoasca apicalis (Flor) (New for lavazè) Empoasca decipiensEmpoasca decipiens Paoli Empoasca vitis (Göthe) Fagocyba cruenta (H.-S.) Ribautiana tenerrima (H.-S.) Eupteryx stachydearumEupteryx stachydearum (Hardy) Zygina suavis rey (New for Pomarolo) Arboridia parvulaArboridia parvula (Boh.) Arboridia pusilla (rib.) (New for Pomarolo)deltocephalinae Balclutha punctata (F.) Macrosteles alpinusMacrosteles alpinus (Zett.) Macrosteles ossiannilssoniMacrosteles ossiannilssoni lindb. (New for lavazè)(New for lavazè) Deltocephalus pulicaris (Fall.) Recilia schmidtgeniRecilia schmidtgeni (W.Wg.) Doratura stylata (Boh.) Selenocephalus griseus (F.) Allygidius abbreviatusAllygidius abbreviatus (leth.) Rhopalopyx adumbrata (c.Shlb.) (New for lavazè)(New for lavazè) Mocydia crocea (H.-S.) Mocydiopsis longicauda rem. Speudotettix subfusculus (Fall.) Thamnotettix confinis (Zett.) Thamnotettix dilutiorThamnotettix dilutior (Kbm.) Arocephalus longiceps (Kbm.) Psammotettix cephalotesPsammotettix cephalotes (H.-S.) Ebarrius interstinctusEbarrius interstinctus (Fieb.) Adarrus exornatus rib. Jassargus baldensis D. Giust. Jassargus repletus (Fieb.) (New for Pomarolo) Jassargus bisubulatus (Then) Verdanus abdominalisVerdanus abdominalis (F.)


appendix 2 - Coleoptera Staphylinidae: list of the collected species with autoecological records.Appendice 2 - Coleoptera Staphylinidae: lista delle specie con note autoecologiche.

Aleochara sparsa Heer, 1839 [Pomarolo] an european species living in nests of birds and sometimes in decaying materials.

Anthophagus omalinus arrowi Koch, 1933 [lavazè] Spread in Northern europe and on the mountains of Central europe, on the alps and the Carpathians,

montane and subalpine. adults on leaves of grasses and bushes, accidental in aerial traps.

Atheta harwoodi Williams, 1930 [Pomarolo] an european species inhabiting nests on trees and sometimes vegetal detritus.

Atheta vaga (Heer, 1839) [lavazè and Pomarolo] an european species inhabiting nests on trees and sometimes vegetal detritus.

Lordithon lunulatus (linné, 1760) [Pomarolo] an european mycetophylic species occasional in aerial traps.

Placusa atrata (Mannerheim, 1830) [Pomarolo] It is spread in europe and Siberia where lives under coniferous barks (pine, spruce).

Placusa depressa Maeklin, 1845 [lavazè] It is spread in europe and Siberia where lives under coniferous barks (pine, spruce, fir).

Placusa pumilio (Gravenhorst, 1802) [Pomarolo] a palaearctic species living under barks of broad-leaf trees.

Placusa tachyporoides (Waltl, 1838) [Pomarolo] a palaearctic species living under barks of recently felled trees (beech, oak, poplar, sometimes coniferous too).

Quedius cruentus (olivier, 1795) [Pomarolo] Spread in europe and Northafrica. It is an arboreal species living under decaying barks and in tree cavities, also

with bird nests, often on isolated plants (e. g. in the plantations along boulevards), but also in woods, mostly in the South.

Quedius maurus (Sahlberg, 1830) [Pomarolo] It is an european species living under barks and in cavities of broad-leaf trees, often in chestnut in the adige

valley.

Quedius mesomelinus mesomelinus (Marsham, 1802) [Pomarolo] a subcosmopolitan species with three subspecies in Italy, in Pomarolo the typical form is present. Phytodetricolous, often in subterranean cavities and in nests of small mammals. often in tree holes too.


Quedius mesomelinus skoraszewskyi Korge, 1960 [lavazè] It is a well differentiated subspecies, endemical of the eastern alps. In the dolomites it is normally montane

and subalpine. The exceptional abundance in the traps of lavazè site is hardly explicable because the species is not related to coniferous barks or holes. The specimens could be from not necessary arboreal nests of Vertebrate.

Quedius nemoralis nemoralis Baudi, 1848 [Pomarolo] an european species usually found in litter of submediterranean woods. occasional in aerial traps.

Quedius punctatellus (Heer, 1839) [lavazè] a subalpine species in Central europe, usually in litter and under stones. occasional in aerial traps.

(appendix 2 - continued)(Appendice 2 - continua)


appendix 3 - lepidoptera: Systematic and ecological species inventory.Appendice 3 - Lepidoptera: lista tassonomico-ecologica delle specie.

TaXoN - Name of taxon (family, genus, species, author and year of description). The systematic treatment mainly follows Huemer & Tarmann (1993).

Po = Pomarolola = lavazèx = records 2001/2002, 2000 and 1992-95

e – ecological amplitudeeu = eurytope: wide ecological amplitudest = stenotope: restricted ecological amplitude

BIoToPe1. Wetland biotopes 1 a. Biotopes beside lakes and along drains 1a B. Biotope along rivers and rivulets 1B C. Fens, marshland 1C2. Woodland biotopes 2 a. Floodplain forests, moorland forest 2a B. deciduous forests 2B a) Thermophilous forests 2Ba b) Mesophilous forests, not beech dominated 2Bb c) Beech dominated forests 2Bc C. Coniferous forest 2C a) Pine forests 2Ca b) Fir-tree dominated forests 2Cb c) Spruce forests 2Cc d) larch forests 2Cd e) Cembra pine forests 2Ce f) dwarf-pine stands 2Cf d. Mountainous green alder shrubs 2d e. Hedgerows and related biotopes 2e3. alpine biotopes inclusive rock and scree formations below the tree line 3 a. rock formations 3a B. Scree formations 3B C. Subalpine-alpine dwarf shrub zone 3CC. Subalpine-alpine dwarf shrub zone 3C d. alpine grassland 3d4. Biotopes with high anthropogenic impact 4 a. Meadows and pastures 4a a) Xerophilous meadows and pastures 4aa b) Mesophilous meadows and pastures 4ab c) Intensively used meadows and pastures 4ac B. Intensively used agricultural areas 4B C. ruderal biotopes 4C d. Buildings 4d e. Gardens, parks, groves 4e

F – FeedING CaTeGorIeS(mainly following the concept of JaroS & SPITZer (2002)1 = monophagous (1) – species feeding on a single species2 = monophagous (2) – species feeding on a single genus3 = oligophagous (3) – species feeding on a group of closely related genera4 = oligophagous (4) – species feeding on a single family5 = polyphagous (5) – species feeding on more than one family (woody plants)6 = polyphagous (6) – species feeding on more than one family (herbs)7 = polyphagous (7) – species feeding on more than one family (woody plant and herbs)8 = mycophagous (8) – species feeding on lichen, algae, fungi (rotten wood) or moss9 = detritophagous (9) – species feeding on detritus (including keratophagous species)

r – reMarKS

P! = possible bioindicator Pomarolo= possible bioindicator Pomarolol! = possible bioindicator lavazè



TaXoN Po02 Po00 Po95 la01 la00 la95 e BIoToPe F HoST-PlaNT r

MicropterigiDae

Micropterix aureatella (Scopoli, 1763) x x x eu 1dd,2a,2C 9 dead vegetable substances

Micropterix aruncella (Scopoli, 1763) x x x eu 1d,2a,2B,3d,4a 9 dead vegetable substances

Micropterix osthelderi Heath, 1976 x eu 2a,2B,4a 9 dead vegetable substances

Micropterix rothenbachii Frey, 1856 x x x st 2Ba 9 dead vegetable substances

eriocraniiDae

Dyseriocrania subpurpurella (Haworth, 1828) x st 2Ba 2 Quercus

hepialiDae

Triodia sylvina (l., 1761) x x x eu 4ac,4ad,4Ca 6 herbs

Korscheltellus lupulinus (l., 1758) x st 2B 6 herbs

Hepialus humuli (l., 1758) x eu 4ac,4ad,4Ca 6 herbs

nepticuliDae

Simplimorpha promissa (Staudinger, 1870) x x x st 2Ba 2 Cotinus coggyria

Stigmella microtheriella (Stainton, 1854) x x x st 2a,2B,2ea 4 Corylaceae: Corylus, Ostrya, Carpinus

Stigmella prunetorum (Stainton, 1855) x x x eu 2B,2ea,4e 2 Prunus (esp. P. spinosa)

Stigmella aceris (Frey, 1857) x x x st 2Ba,2Bb,2ea 2 Acer

Stigmella catharticella (Stainton, 1853) x st 2aa,2Ba,2Bb,2ea 1 Rhamnus cathartica

Stigmella anomalella (Göze, 1783) x x x eu 2,4e 2 Rosa

Stigmella johanssonella last. & last., 1997 x x x st 2Ba,2ea 1 Ostrya

Stigmella hahniella (Wörz, 1937) x x st 2Ba 1 Sorbus torminalis

Stigmella desperatella (Frey, 1856) x x x eu 2a,2B,4e 3 rosaceae (Malus, Pyrus)

Stigmella hybnerella (Hübner, 1796) x x x eu 2a,2B,2ea,4e 2 Crataegus

?Stigmella mespilicola (Frey, 1856) x x x st 2Ba 3 rosaceae (Sorbus, Amelanchier)

Stigmella salicis (Stainton, 1854) x x eu 2 2 Salix caprea, S. cinerea etc.

Stigmella trimaculella (Haworth, 1828) x st 2a,2Bb 2 Populus nigra, P. canadensis

Stigmella assimilella (Zeller, 1848) x x st 2a,2B,2ea 1 Populus tremula

Stigmella plagicolella (Stainton, 1854) x eu 2a,2B,2ea,4e 2 Prunus

Stigmella aurella (Fabricius, 1775) x x st 2Ba,2ea 2 Rubus

Stigmella hemargyrella (Kollar, 1832) x st 2Bc 1 Fagus

Stigmella svenssoni (Johansson, 1971) x st 2Bc 2 Quercus

Stigmella atricapitella (Haworth, 1828) x st 2Ba,4F 2 Quercus

Stigmella samiatella (Zeller, 1839) x st 2Ba,4F 3 Fagaceae: Quercus, Castanea

Stigmella sp. x st 2Ba 2 Quercus

Stigmella sp. x st 2Ba 2 Crataegus

Trifurcula cryptella (Stainton, 1856) x x st 2Ba,4aa 3 Fabaceae

Trifurcula moravica last. & last., 1994 x st 2Ba 1 Lembotropis nigricans

Bohemannia pulverosella (Stainton, 1849) x x x eu 2Ba,2e,4Ba,4e 3 rosaceae (Malus, Pyrus, Prunus)

Ectoedemia atrifrontella (Stainton, 1851) x st 2Ba 2 Quercus

Ectoedemia argyropeza (Zeller, 1839) x st 2B,2ea 1 Populus tremula

Ectoedemia albifasciella (Heinemann, 1871) x x x st 2Ba 2 Quercus

Ectoedemia angulifasciella (Stainton, 1849) x x st 2Ba,2ea 2 Rosa

Ectoedemia atricollis (Stainton, 1857) x x x eu 2Ba,2e,4e 3 wooden rosaceae




Ectoedemia arcuatella (H.-S., 1855) x x x eu 2a,2Bb,2Bc,2C 3 rosaceae (Fragaria, Potentilla)

Ectoedemia mahalebella (Klimesch, 1936) x x x st 2Ba 1 Prunus mahaleb

heliozeliDae

Antispila treitschkiella (F. v. r., 1843) x x x st 2Ba,2ea 2 Cornus

aDeliDae

Nematopogon swammerdamella (l., 1758) x x x x x eu 2 6 herbs, decaying leaves

Nematopogon robertella (Clerck, 1759) x x x st 2Cc 9 dead spruce needles

Nemophora metallica (Poda, 1761) x x eu 1dc,4aa,4ab 3 dipsacaceae: Scabiosa, Knautia, ?Succisa

Nemophora minimella (d. & Schiff., 1775) x st 1dc,4aa 3 dipsacaceae: Scabiosa, Succisa

Adela reaumurella (l., 1758) x x x st 2Ba 6 herbs

Adela croesella (Scopoli, 1763) x eu 2a,2B 9 ?dead vegetable substances

incurvariiDae

Incurvaria masculella (d. & Schiff., 1775) x st 2Ba 5 deciduous trees/shrubs

Paraclemensia cyanella (Zeller, 1850) x x x st 2Ba 1 Acer campestre

tisheriiDae

Tischeria ekebladella (Bjerkander, 1795) x x x st 2Ba 3 Fagaceae: Quercus, Castanea

Tischeria dodonaea Stainton, 1858 x st 2Ba 2 Quercus

Tischeria decidua Wocke, 1876 x x st 2Ba 2 Quercus

Emmetia marginea (Haworth, 1828) x x x st 2B,2ea 2 Rubus

Emmetia angusticolella (dup., 1843) x x x st 2Ba,2ea 2 Rosa

psychiDae

Dahlica triquetrella (Hübner, 1813) bisex. x eu 2 8 algae

Taleporia tubulosa (retzius, 1783) x x eu 2 8 lichens, withered leaves

Bijugis bombycella (d. & Schiff., 1775) x x x eu 1d,2 6 herbs, grass

Sterrhopterix fusca (Haworth, 1809) x x x eu 1dd,2a,2B 5 deciduous trees/shrubs

Apterona helicoidella (Vallot, 1827) x st 4aa 6 herbs

tineiDae

Euplocamus anthracinalis (Scopoli, 1763) x st 2Ba 8 dead wood/fungi

Cephimallota angusticostella (Zeller, 1839) x x x st 2B 8 dead wood/fungi

Infurcitinea albicomella (H.-S., 1851) x st 2Ba,?3a,?3B 8 lichens

Infurcitinea finalis Gozmány, 1959 x x st 3a,3B 8 lichens

Nemapogon ruricolella (Stainton, 1849) x x st 2Ba 8 dead wood/fungi

Nemapogon clematella (Fabricius, 1781) x eu 2a,2B,2C 8 dead wood/fungi

Triaxomera fulvimitrella (Sodoffsky, 1830) x st 2Ba 8 dead wood/fungi

Monopis laevigella (d. & Schiff., 1775) x x eu 2a,2B,2C 9 dead organic matter

Monopis obviella (d. & Schiff., 1775) x x x eu 2a,2B,2C 9 dead organic matter

Tinea semifulvella Haworth, 1828 x st 2a,2B 9 birdsńests

Tinea trinotella Thunberg, 1794 x x st 2 9 birdsńests

DouglasiiDae

Tinagma signatum Gaedike, 1991 x x st 3C,3d 3 ?Potentilla, ?Geum




bucculatriciDae

Bucculatrix demaryella castaneae Klimesch, 1950

x x st 2Ba,4F 1 Castanea sativa

Bucculatrix ulmella Zeller, 1848 x x x st 2Ba,4F 3 Fagaceae: Quercus, Castanea

Bucculatrix bechsteinella (Be. & Scharfenberg, 1805)

x x st 2Ba,2ea 2 Crataegus

Bucculatrix frangutella (Goeze, 1783) x x x eu 1dc,2a,2B,2ea 2 Rhamnus

gracillariiDae

Caloptilia syringella (Fabricius, 1794) x x x eu 2,4e 3 oleaceae: Fraxinus, Ligustrum

Caloptilia alchimiella (Scopoli, 1763) x x st 2Ba,4F 2 Quercus

Caloptilia robustella Jäckh, 1972 x x x st 2Ba 2 Quercus

Caloptilia sp. x x eu 2,4e 1 Acer pseudoplatanus

Aspilapteryx limosella (dup., 1843) x x st 4aa 2 Teucrium

Eucalybites auroguttella (Stephens, 1835) x x eu 2B,2eb,2ec,4ab 2 Hypericum

Micrurapteryx kollariella (Zeller, 1839) x x x st 2Ba 3 Fabaceae

Parectopa robiniella Clemens, 1863 x x st 2aa,2Ba,4e 1 Robinia

Parornix anglicella (Stainton, 1850) x x x eu 2 3 rosaceae: Prunus, Crateagus, Amelanchier

Parornix carpinella (Frey, 1861) x x x st 2Ba 3 Corylaceae: Carpinus, Ostrya

Parornix devoniella (Stainton, 1850) x x x st 2B,2ea 1 Corylus

Parornix scoticella (Stainton, 1850) x x x eu 2a,2B,2C,2ea,3C 3 rosaceae (esp. Sorbus)

Parornix torquilella (Zeller, 1850) x st 2Bb 1 Prunus spinosa

Callisto denticulella (Thunberg, 1794) x x x eu 2a,2B,2ea,4B,4e 2 Malus

Phyllonorycter robiniella (Clemens, 1859) x x st 2as,2Ba,4e 1 Robinia

Phyllonorycter sp. x st 2a 2 Populus (P. canadensis)

Phyllonorycter sagitella (Bjerkander, 1790) x x x st 2B,2ea 1 Populus tremula

Phyllonorycter corylifoliella (Hübner, 1796) x x x eu 2,4Ba,4e 5 rosaceae, Betula

Phyllonorycter quercifoliella (Zeller, 1839) x x st 2Ba 2 Quercus

Phyllonorycter parisiella (Wocke, 1848) x st 2Ba 2 Quercus

Phyllonorycter pomonella (Zeller, 1846) x x x st 2B,2ea 2 Prunus (esp. P. spinosaP. spinosa)

Phyllonorycter cerasicolella (H.-S., 1855) x x x eu 2B,2ea,4Ba,4e 2 Prunus (esp. P. avium)

Phyllonorycter lantanella (Schrank, 1802) x x x st 2B,2ea 2 Viburnum

Phyllonorycter hilarella (Zetterstedt, 1839) x x eu 2 2 Salix

Phyllonorycter oxyacanthae (Frey, 1856) x x x st 2B,2ea 3 rosaceae (esp. Crataegus)

Phyllonorycter sorbi (Frey, 1855) x eu 2B,2ea 3 rosaceae (esp. SorbusSorbus)

Phyllonorycter cydoniella (d. & Schiff., 1775) x st 2Ba 3 rosaceae: Amelanchier, Pyrus

Phyllonorycter junoniella (Zeller, 1846) x x st 1da,2Cc,2Ce,2Cf 1 Vaccinium vitis-idaea

Phyllonorycter schreberella (Fabricius, 1781) x st 2B 2 Ulmus

Phyllonorycter nicellii (Stainton, 1851) x st 2B,2ea 1 Corylus

Phyllonorycter coryli (Nicelli, 1851) x x x st 2B,2ea 4 Corylaceae

Phyllonorycter aemula Trib., de. & Hue., 1996

x x x st 2Ba 1 Ostrya carpinifolia

Phyllonorycter maestingella (Müller, 1794) x x x st 2Bc 1 Fagus




Phyllonorycter harrisella (l., 1761) x x st 2Ba 2 Quercus

Phyllonorycter roboris (Zeller, 1839) x x x st 2Ba 2 Quercus

Phyllonorycter saportella (dup., 1840) x st 2Ba 2 Quercus

Phyllonorycter nigrescentella (Bouché, 1834) x x x st 2Ba,2eb,4aa 3 Fabaceae

Phyllonorycter fraxinella (Zeller, 1846) x x x st 2Ba 1 Genista germanica

Phyllonorycter sp. x st 2Ba 3 Fabaceae

Phyllonorycter acerifoliella (Zeller, 1839) x x st 2Ba 1 Acer campestre

Phyllonorycter geniculella (ragonot, 1874) x x x eu 2 1 Acer pseudoplatanus

yponoMeutiDae

Yponomeuta evonymella (l., 1758) x x st 2a,2ea 3 rosaceae: esp. Prunus padus

Yponomeuta padella (l., 1758) x x x x eu 2a,2B,2C 3 rosaceae: Prunus, Crataegus

Yponomeuta irrorella (Hübner, 1796) x st 2B,2ea 1 Euonymus

Yponomeuta plumbella (d. & Schiff., 1775) x st 2aa,2B,2ea 1 Euonymus

Yponomeuta sedella Treitschke, 1832 x x st 3a 2 Sedum

Parahyponomeuta egregiella (dup., 1838) x st 2Ba,2Ca 1 Erica

Swammerdamia pyrella (Villers, 1789) x eu 2a,2B,2C 3 wooden rosaceae

Paraswammerdamia nebulella (Goeze, 1783) x st 2Ba,2ea 3 wooden rosaceae

Cedestis gysseleniella Zeller, 1839 x st 2Ca,2Cd,2Cf,4e 2 Pinus

Cedestis subfasciella (Stephens, 1834) x x st 2Ca,2Cd,2Cf 2 Pinus

Ocnerostoma piniariellum Zeller, 1847 x eu 2C 2 Pinus

Prays fraxinella (Bjerkander, 1784) x x x eu 2a,2B 2 Fraxinus

Argyresthia amiantella (Zeller, 1847) x x x st 2Cc 1 Picea abies l!

Argyresthia abdominalis Zeller, 1839 x x st 2C 2 Juniperus

Argyresthia retinella Zeller, 1839 x st 2a 2 Betula, ?Salix

Argyresthia sorbiella (Treitschke, 1833) x x st ?2Ba,2C 2 Sorbus aucuparia, Sorbus aria

Argyresthia albistria (Haworth, 1828) x st 2Ba,2ea 1 Prunus spinosa

Argyresthia spinosella Stainton, 1849 x st 2Ba 1 Prunus spinosa

Argyresthia pruniella (Clerck, 1759) x x eu 2a,2B,4Ba,4e 2 Prunus

Argyresthia bonnetella (l., 1758) x x x st 2Ba,2ea 2 Crataegus

Argyresthia conjugella Zeller, 1839 x x eu 2,4e 3 rosaceae: Sorbus, Malus

ypsolophiDae

Ypsolopha mucronella (Scopoli, 1763) x eu 2a,2B 2 Euonymus

Ypsolopha dentella (Fabricius, 1775) x st 2a,2B,2ea,4e 2 Lonicera

Ypsolopha falcella (d. & Schiff., 1775) x st 2a,2B,2ea,4e 2 Lonicera

Ypsolopha scabrella (l., 1761) x x st 2Ba 3 rosaceae: Malus, Crataegus

Ypsolopha horridella (Treitschke, 1835) x st 2Ba 3 rosaceae: Malus, Crataegus, Prunus

Ypsolopha lucella (Fabricius, 1775) x st 2Ba 2 Quercus

Ypsolopha sylvella (l., 1767) x x st 2Ba 2 Quercus

Ypsolopha parenthesella (l., 1761) x x x eu 2a,2B,2C,2ea,4e 5 deciduous trees/shrubs

Ypsolopha ustella (Clerck, 1759) x st 2Ba 5 deciduous trees/shrubs

plutelliDae

Plutella xylostella (l., 1758) x x x x x eu 1d,2,4 4 Cruciferae

Digitivalva arnicella (Heyden, 1863) x x st 3d 1 Arnica montana




Acrolepia assectella (Zeller, 1839) x x x eu 1dc,2e,4a,4e 2 Allium

glyphipterigiDae

Glyphipterix simpliciella (Stephens, 1834) x x x eu 1d,2B,2e,4a 3 Poaceae

lyonetiiDae

Leucoptera coronillae (Hering, 1933) x x x st 2Ba 2 Cytisus

Lyonetia clerkella (l., 1758) x eu 2a,2B,2C,4e 5 rosaceae, Betulaceae

coleophoriDae

Coleophora albella (Thunberg, 1788) x x st 2Ba,4aa 3 Caryophyllaceae: Lychnis, Silene

Coleophora lutipennella (Zeller, 1838) x x x st 2Ba 2 Quercus

Coleophora gryphipennella (Hübner, 1796) x x eu 2B,2C,2d,2ea 2 Rosa

Coleophora ��avipennella (dup., 1843) x x x st 2Ba 2 Quercus

Coleophora milvipennis Zeller, 1839 x x x st 2aa,2Ba,2Bb 5 deciduous trees/shrubs

Coleophora coracipennella (Hübner, 1796) x eu 2B,2ea,4Ba,4e 3 rosaceae: Crataegus, Prunus

Coleophora fuscocuprella H.-S., 1855 x st 2B,2ea 1 Corylus

Coleophora lusciniaepennella (Treitschke, 1833)

x st 2a 2 Salix

Coleophora glitzella Hoofmann, 1869 x st 2Cc,3C 1 Vaccinium vitis-idaea

Coleophora violacea (Ström, 1783) x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Coleophora lineolea (Haworth, 1828) x st 2Ba,2Bb,4a 3 labiatae: Ballota, Stachys, Betonica

Coleophora hemerobiella (Scopoli, 1763) x x x eu 2a,2B,2ea,4e 3 wooden rosaceae

Coleophora onobrychiella Zeller, 1849 x st 4aa 2 Astragalus

Coleophora colutella (Fabricius, 1794) x x st 2Ba,2ea 3 Fabaceae

Coleophora discordella Zeller, 1849 x st 2Ba,2eb,4aa 3 Fabaceae

Coleophora bilineatella Zeller, 1849 x st 2Ba 3 Fabaceae: Genista, Sarothamnus

Coleophora deauratella lienig & Zeller, 1846 x x x eu 4a,4C 2 Trifolium

Coleophora mayrella (Hübner, 1813) x x eu 4a,4C 2 Trifolium

Coleophora kuehnella (Goeze, 1783) x x x st 2Ba,4F 2 Quercus

Coleophora ibipennella Zeller, 1849 x st 2Ba 2 Quercus

Coleophora currucipennella Zeller, 1839 x st 2Ba 2 Quercus

Coleophora virgatella Zeller, 1849 x st 2eb,4aa 2 Salvia

Coleophora serpylletorum Hering, 1889 x x st 4aa 2 Thymus

Coleophora auricella (Fabricius, 1794) x x x st 2Ba,4aa 3 labiatae: Stachys, Teucrium

Coleophora vibicella (Hübner, 1813) x st 2Ba,2ea 2 Genista

Coleophora lixella Zeller, 1849 x st 4aa 6 Poaceae, labiatae: Thymus

Coleophora ornatipennella (Hübner, 1796) x x x eu 4a 6 Poaceae, labiatae

Coleophora hartigi Toll, 1944 x st 2Ba,2ea 2 Genista

Coleophora laricella (Hübner, 1817) x x x eu 2C,4e 1 Larix

Coleophora otidipennella (Hübner, 1817) x x st 2 2 Luzula

Coleophora alticolella Zeller, 1849 x st 1B,1C,1d 2 Juncus

Coleophora taeniipennella H.-S., 1855 x st 1C 2 Juncus

Coleophora sylvaticella Wood, 1892 x x st 2 1 Luzula sylvatica

Coleophora obscenella H.-S., 1855 x eu 2,3C,4a,4C 2 Solidago




Coleophora versurella Zeller, 1849 x st 2eb,4ab,4Cd 4 Chenopodiaceae

Coleophora vestianella (l., 1758) x x st 4C 2 Atriplex

Coleophora galbulipennella Zeller, 1838 eu 2 2 Silene

Coleophora setarii Wocke, 1877 x x st 2Ba,4aa 2 Artemisia

Coleophora trochilella (dup., 1843) x eu 3a,3d,4aa,4ab 3 asteraceae

Coleophora follicularis (Vallot, 1802) x x x eu 1Bb,1d,2a 3 asteraceae: esp. Eupatorium

Coleophora nubivagella Zeller, 1849 x eu 3d,4a 3 Caryophyllaceae

Coleophora dianthi H.-S., 1855 x x st 4aa 2 Dianthus

Coleophora silenella H.-S., 1855 x st 2eb,4aa,4ab 2 Silene nutans, S. vulgaris

Coleophora paripennella Zeller, 1839 x x eu 2eb,4a,4Cc,4Cd 3 asteraceae: esp. Centaurea

Pseudatemelia latipennella (Jäckh, 1959) x x x eu 2 9 ?decaying leaves, ?lichens

Pseudatemelia synchrozella (Jäckh, 1959) x eu 2 9 ?decaying leaves

Pseudatemelia josephinae (Toll, 1956) x eu 2 9 ?decaying leaves

elachistiDae

Spuleria ��avicaput (Haworth, 1828) x st 2Ba,2ea 2 Crataegus

Heinemannia festivella (d. & Schiff., 1775) x x st 2Ba,4aa 3 Fabaceae

Blastodcana atra (Haworth, 1828) x x st 2Ba,2ea,4e 3 wooden rosaceae

Tetanocentria ochraceella rebel, 1903 x x x st 2Ba 8 ?dead wood/fungi

Elachista regificella Sircom, 1849 x x st 2B,2e 3 Poaceae

Elachista gleichenella (Fabricius, 1781) x x st 1Bb,1d,2ab 6 Juncaceae, Cyperaceae

Elachista atricomella Stainton, 1849 x x eu 1d,2eb,4a 3 Poaceae

Elachista bifasciella Treitschke, 1833 x x x st 2C,3C 3 Poaceae

Elachista humilis Zeller, 1850 x x x eu 1d,2 3 Poaceae

Elachista sp. x ? ?2Ba,4aa 3 ?Poaceae

Elachista dispilella Zeller, 1839 x x x st 4aa 3 Poaceae: Festuca, Corynephora

Elachista chrysodesmella Zeller, 1850 x st 4aa,4ab 3 Poaceae

Elachista atrisquamosa Staudinger, 1880 x x x st 4aa 3 Poaceae

Elachista subalbidella Schläger, 1847 x st 3d 3 Poaceae

Elachista cingillella (H.-S., 1855) x x eu 2a,2B 2 ?Milium

Elachista adscitella Stainton, 1851 x eu 1d,2ea,4a 3 Poaceae

Elachista cinereopunctella (Haworth, 1828) x st 2Ba 6 Cyperaceae, Poaceae

Elachista juliensis (Frey, 1870) x st 4aa 2 ?Carex

Cephalispheira ferrugella (d. & Schiff., 1775) x st 2Ba 2 Campanula

Anchinia daphnella (d. & Schiff., 1775) x x st 2C 1 Daphne mezereum

Ethmia terminella Fletcher, 1938 x st 2Ba,2eb,4aa,4Cd 2 Echium

Agonopterix hippomarathri (Nickerl, 1864) x x st 4aa 3 apiaceae

Agonopterix selini (Heinemann, 1870) x x x st 1d 2 Peucedanum

Agonopterix kaekeritziana (l., 1767) x x x eu 1dc,4aa,4ab 2 Centaurea

Agonopterix laterella (d. & Schiff., 1775) x st 4aa,4ab 1 Centaurea cyanus

Agonopterix ocellana (Fabricius, 1775) x eu 2a,2B 2 Salix

Agonopterix nervosa (Haworth, 1811) x eu 2B,2e 3 apiaceae

Agonopterix scopariella (Heinemann, 1870) x x st 2Ba 3 Fabaceae: esp. SarothamnusSarothamnus

Agonopterix cnicella (Treitschke, 1832) x st 4aa 3 asteraceae: esp. EryngiumEryngium




Agonopterix senecionis (Nickerl, 1864) x x st 4aa 3 asteraceae

Agonopterix parilella (Treitschke, 1835) x st 2Ba,4aa 1 Peucedanum oroselinum

Agonopterix purpurea (Haworth, 1811) x st 2Ba,4aa,4ab 3 apiaceae

Agonopterix pallorella (Zeller, 1839) x x st 4aa,4ab 3 asteraceae

Agonopterix arenella (d. & Schiff., 1775) x eu 2B,2e,4aa 3 asteraceae

Agonopterix pupillana (Wocke, 1887) x x x st 4aa 1 Dictamnus albus

Agonopterix rotundella (douglas, 1846) x st 2Ba,2e,4aa 3 apiaceae

Agonopterix ciliella (Stainton, 1849) x eu 2B,2e,4a,4C 3 apiaceae

Agonopterix nodi��orella (Milliére, 1866) x st 4aa 1 Ferula nodi��ora

Levipalpus hepatariellus (lienig & Zeller, 1846)

x st 3C,3d

Depressaria depressana (Fabricius, 1775) x st 4aa 3 apiaceae

Depressaria pimpinellae Zeller, 1839 x x x st 2Ba,2eb,4aa 2 Pimpinella

Depressaria pulcherrimella Stainton, 1849 x st 2Ba,2eb,4aa 2 Pimpinella

Depressaria olerella Zeller, 1854 x st 4aa 1 Achillea millefolium

Horridopalpus dictamnellus (Treitschke, 1835) x st 4aa 1 Dictamnus albus

Semioscopis steinkellneriana (d. & Schiff., 1775)

x st 2Ba 3 rosaceae: Sorbus, Prunus, Crataegus

Telechrysis tripuncta (Haworth, 1828) x x st 2B,2C 8 ?dead wood/fungi

chiMabachiDae

Diurnea fagella (d. & Schiff., 1775) x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Diurnea lipsiella (d. & Schiff., 1775) x eu 2a,2B,2C,2e,4e 5 deciduous trees/shrubs

carciniDae

Carcina quercana (Fabricius, 1775) x x x eu 2aa,2B,4e 5 deciduous trees/shrubs

oecophoriDae

Denisia stipella (l., 1758) x x x eu 2 8 dead wood/fungi

Denisia nubilosella (H.-S., 1854) x st 2C 8 ?dead wood/fungi

Denisia similella (Hübner, 1796) x eu 2B,2C 8 dead wood/fungi

Goidanichiana jourdheuillella (ragonot, 1875)

x st 2Ba 8 dead wood/fungi

Batia unitella (Hübner, 1796) x x eu 2a,2B 8 dead wood/fungi

Batia lambdella (donovan, 1793) x x st 2B 8 dead wood/fungi

Batia internella Jäckh, 1972 x x x st 2Ba 8 dead wood/fungi

Metalampra italica Baldizzone, 1977 x x st 2Ba 8 dead wood/fungi, esp. Quercus

Borkhausenia fuscescens (Haworth, 1828) x eu 2B,2d 9 decaying leaves, birdsńests

Oecophora bractella (l., 1758) x x eu 2B 8 dead wood/fungi

Harpella forficella (Scopoli, 1763) x x x eu 2a,2B 8 dead wood/fungi

Hofmannophila pseudospretella (Stainton, 1849)

x eu 2,4d,4e 9 dead vegetable substances, birdsńests etc

Aplota palpella (Haworth, 1828) x st 2B 8 moss, lichens

Protasis punctella (o.G.Costa, 1836) x x st 2Ba,4aa

Pleurota bicostella (Clerck, 1759) x eu 2Ca,3C 3 ericaceae: Erica, Calluna

Pleurota pungitiella (H.-S., 1854) x x x st 4aa




syMMociDae

Symmoca caliginella Mann, 1867 x x x st 3a 8 ?moss

Symmoca signatella H.-S., 1854 x x x st 2Ba 8 ?moss

Apatema mediopallidum Walsingham, 1900 x x st 2Ba,2ea 9 dead vegetable substances

Apatema apolausticum Gozmány, 1996 x x st 2Ba 9 ?dead vegetable substances

lecithoceriDae

Homaloxestis briantiella (Turati, 1879) x x x st 2Ba 8 dead wood/fungi (?Quercus pubescens)

scythriDiDae

Scythris obscurella (Scopoli, 1763) x x st 4aa,4ab

Scythris punctivittella (o.G. Costa, 1836) x st 4aa

Scythris laminella (d. & Schiff., 1775) x st 4aa,4ab 2 Hieracium

blatobasiDae

Blastobasis phycidella (Zeller, 1839) x x x st 2Ba 8 ?rotten wood/fungi

Blastobasis huemeri Sinev, 1994 x x x st 2Ba 8 ?rotten wood/fungi

MoMphiDae

Mompha locupletella (d. & Schiff., 1775) x x x st 2e 2 Epilobium

Mompha miscella (d. & Schiff., 1775) x eu 3d,4aa,4ab 2 Helianthemum

Mompha epilobiella (d. & Schiff., 1775) x eu 1d,2a,2B 2 Epilobium (esp. E. hirsutum)

cosMopterigiDae

Pancalia leuwenhoekella (l., 1761) x x st 2Ba,2eb,4aa,4ab 2 Viola

Cosmopterix orichalcea Stainton, 1861 x x st 1d,2aa,2B 3 Poaceae: Phalaris, Phragmites, Milium

Stagmatophora heydeniella (F. v. r., 1838) x x x st 1dc,2Ba,4aa,4ab 3 labiatae: Betonica, Stachys

Vulcaniella extremella (Wocke, 1871) x st 2Ba,4aa 3 labiatae: Salvia, Prunella

gelechiDae

Xystophora pulveratella (H.-S., 1854) x st 4aa 3 Fabaceae

Isophrictis anthemidella (Wocke, 1871) x x x st 4aa,4ab 3 asteraceae

Metzneria metzneriella (Stainton, 1851) x x x st 4aa 1 Centaurea nigra

Metzneria neuropterella (Zeller, 1839) x st 4aa 2 Centaurea

Metzneria aprilella (H.-S., 1854) x st 2Ba,4aa 3 asteraceae

Apodia bifractella (dup., 1843) x st 2Ba,2ea,2eb,4aa 2 Inula

Monochroa nomadella (Zeller, 1868) x st 2Ba

Eulamprotes libertinella (Zeller, 1872) x x eu 4aa,4ab

Eulamprotes unicolorella (duponchel, 1843) x x eu 2e,4aa,4ab

Eulamprotes atrella (d. & Schiff., 1775) x x eu 2eb,4a,4C 2 Hypericum

Bryotropha similis (Stainton, 1854)) x x eu 2,4C 8 moss

Bryotropha senectella (Zeller, 1839) x st 1Bc,2Ba,4aa 8 moss

Recurvaria nanella (d. & Schiff., 1775) x x x eu 2a,2B,2e,4Ba,4e 3 rosaceae: Malus, Prunus

Recurvaria leucatella (Clerck, 1759) x x eu 2a,2B,2e,4Ba,4e 3 rosaceae: Crataegus, Prunus

Exoteleia dodecella (l., 1758) x x eu 2C 2 Pinus

Stenolechia gemmella (l., 1758) x x x st 2Ba,4F 2 Quercus




Parachronistis albiceps (Zeller, 1839) x x x st 2ab,2Ba,2Bb,2ea 1 Corylus

Teleiodes italica Huemer, 1992 x x st 2Ba 3 ?rosaceae

Teleiodes wagae (Nowicki, 1861) x x x st 2Ba 3 Corylaceae: Corylus, Ostrya

Teleiodes saltuum (Zeller, 1878) x st 2Cd 1 Larix

Teleiodes luculella (Hübner, 1813) x x x st 2aa,2Ba 5 deciduous trees/shrubs: esp. Quercus

Teleiodes sequax (Haworth, 1828) x x st 4aa,4ab 2 Helianthemum

Pseudotelphusa scalella (Scopoli, 1763) x x x st ?2Ba,4aa 6 ?herbs

Carpatolechia decorella (Haworth, 1811) x x x st 2Ba,4F 5 deciduous trees/shrubs: esp. Cotinus

Pseudotelphusa paripunctella (Thunberg, 1794)

x x x eu 2a,2B,2ea 5 deciduous trees/shrubs: Quercus, Hippophae

Pseudotelphusa tessella (l., 1758) x x st 2Ba,2ea 1 Berberis

Altenia scriptella (Hübner, 1796) x x st 2Ba 1 Acer campestre

Gelechia scotinella H.-S., 1854 x x st 2Ba,2ea,4e 3 wooden rosaceae

Gelechia sabinella Zeller, 1839 x x st 2Ca,4e 2 Juniperus

Gelechia nigra (Haworth, 1828) x eu 2a,2B 4 Salicaceae

Psoricoptera gibbosella (Zeller, 1839) x x x st 2aa,2Ba 2 Quercus

Mirificarma lentiginosella (Zeller, 1839) x x st 2Ba 3 Fabaceae: Genista, Chamaecytisus, Laburnum

Mirificarma maculatella (Hübner, 1796) x x st 2Ba 2 Coronilla emerus, C. varia

Mirificarma cytisella (Treitschke, 1833) x st 2Ba,4aa 3 Fabaceae: Genista, Laburnum, Lembotropis

Chionodes tragicella (Heyden, 1865) x x st 2Cc,2Cd,2Ce 1 Larix

Chionodes perpetuella (H.-S., 1854) x x st 3B

Chionodes electella (Zeller, 1839) x x eu 2C 1 Picea

Aroga ��avicomella (Zeller, 1839) x st 2Ba 1 Prunus spinosa

Neofaculta ericetella (Geyer,, 1832) x st 2Ca,4aa 4 ericaceae

Neofaculta infernella (H.-S., 1854) x x x x x x eu 2,3C 4 ericaceae

Prolita sexpunctella (Fabricius, 1794) x x x eu 1da,2Ca,3C,3d 7 herbs, deciduous shrubs

Prolita solutella (Zeller, 1839) x st 2Ba,2ea 3 Fabaceae: Genista, Sarothamnus

Athrips mouffetella (l., 1758) x st 2B 2 Lonicera

Athrips amoenella (Frey, 1882) x st 2Ba,4aa 2 ?Vicia

Scrobipalpa acuminatella (Sircom, 1850) x x eu 3C,3d,4ab,4Cd 3 asteraceae: Carduus, Cirsium

Caryocolum tischeriella (Zeller, 1839) x x x st 2Ba,2Ca,2ea,4aa 1 Silene nutans

Caryocolum leucomelanella (Zeller, 1839) x x x st 3a,4aa,4ab 3 Caryophyllaceae: Dianthus

Caryocolum leucothoracellum (Klimesch, 1953)

x x x st 4aa 2 Dianthus

Caryocolum moehringiae (Klimesch, 1954) x st 2B,2C 2 Moehringia

Stomopteryx ��avipalpella Jäckh, 1959 x st 2Ba,4aa 3 Fabaceae

Syncopacma sangiella (Stainton, 1863) x st 2Ba,3a,4aa 1 Lotus corniculatus

Syncopacma coronillella (Treitschke, 1833) x st 2Ba,3a,4aa 3 Fabaceae

Syncopacma cincticulella (Bruand, 1850) x st 2Ba,4aa,4ab 3 Fabaceae

Syncopacma cinctella (Clerck, 1759) x x st 2Ba,3a,4aa 3 Fabaceae

Syncopacma taeniolella (Zeller, 1839) x x x st 2Ba,4aa,4ab 3 Fabaceae

Aproaerema anthyllidella (Hübner, 1813) x x eu 2Ba,2e,3a,4a 3 Fabaceae




Anacampsis populella (Clerck, 1759) x eu 2a,2B 4 Salicaceae

Anacampsis quercella (Chrétien, 1907) x x x st 2Ba 2 Quercus P!

Mesophleps silacella (Hübner, 1796) x eu 2Ba,2eb,4aa,4ab 2 Helianthemum

Uncustriodonta trinotella (H.-S., 1856) x st 2Ba,4aa 2 Erysimum

Anarsia lineatella (Zeller, 1839) x st 4Ba,4e 2 Prunus

Hypatima rhomboidella (l., 1758) x x eu 2a,2B 5 deciduous trees/shrubs

Nothris verbascella (d. & Schiff., 1775) x st 4aa,4ab,4Cc,4Cd 2 Verbascum

Nothris lemniscella (Zeller, 1839) x st 4aa 2 Globularia

Dichomeris marginella (Fabricius, 1781) x st 2Ba 1 Juniperus communis

Dichomeris ustalella (Fabricius, 1794) x x x st 2ab,2Ba 5 deciduous trees/shrubs: Carpinus, Betula, Salix

Dichomeris derasella (d. & Schiff., 1775) x x x st 2Ba,2ea 1 Prunus spinosa

Dichomeris limosella (Schläger, 1849) x st 4aa 3 Fabaceae

Dichomeris alacella (Zeller, 1839) x x x st 2a 8 lichens

Brachmia dimidiella (d. & Schiff., 1775) x st 4aa 4 Poaceae

Helcystogramma lutatella (H.-S., 1854) x x x st 4aa 4 Poaceae

Helcystogramma rufescens (Haworth, 1828) x x eu 1d,2,4aa,4ab 4 Poaceae

Acompsia cinerella (Clerck, 1759) x x x eu 2,4a 8 moss

Acompsia tripunctella (d. & Schiff., 1775) x x x eu 1d,3C,3d 8 ?moss

cossiDae

Zeuzera pyrina (l., 1761) x x eu 2a,2d,4e 5 deciduous trees/shrubs

sesiiDae

Chamaesphecia empiformis (esper, 1783) x eu 2eb,4aa 2 Euphorbia

zygaeniDae

Zygaena carniolica (Scopoli, 1763) x st 4aa 3 Fabaceae

Zygaena romeo dup., 1835 x x st 2Ba,4aa,4ab 3 Fabaceae: Vicia, Lathyrus

Zygaena transalpina (esper, 1782) x x x eu 1d,2Ba,2e,4a 3 Fabaceae

Zygaena filipendulae (l., 1758) x x x eu 1,2,3,4 2 Lotus

Zygaena lonicerae (Scheven, 1777) x x x st 2Ba,2e,4aa 2 Lotus

Zygaena purpuralis (Brünnich, 1763) x st 4aa 2 Thymus

Adscita albanica (Naufock, 1929) x x x st ?2Ba 1 Geranium sanguineum P!

liMacoDiDae

Apoda limacodes (Hufnagel, 1766) x x x eu 2B 5 deciduous trees/shrubs: Quercus, Fagus

tortriciDae

Isotrias rectifasciana (Haworth, 1811) x x x x eu 2 5 deciduous trees/shrubs

Cochylimorpha hilarana (H.-S., 1851) x x st 4aa 1 Artemisia campestris

Cochylimorpha jucundana (Treitschke, 1835) x x x st 4aa 2 Artemisia

Cochylimorpha straminea (Haworth, 1811) x st 4aa 2 Centaurea

Agapeta zoegana (l., 1767) x x x st 4aa,4ab 6 herbs

Eupoecilia ambiguella (Hübner, 1796) x eu 2Ba,2eb,4Bb,4e 5 deciduous trees/shrubs: Vitis, Frangula, Lonicera




Aethes williana (Brahm, 1791) x st 4aa 6 apiaceae, asteraceae

Aethes tesserana (d. & Schiff., 1775) x st 4aa,4ab 3 asteraceae

Aethes francillana (Fabricius, 1794) x st 4aa 3 apiaceae

Cochylidia rupicola (Curtis, 1834) x x x st 1Bb,1dc,2a,2e 1 Eupatorium

Cochylidia heydeniana (H.-S., 1851) x st 4aa 3 asteraceae

Cochylis hybridella (Hübner, 1813) x x x st 2eb,4aa,4ab,4C 3 asteraceae: Picris, Crepis

Cochylis pallidana Zeller, 1847 x x x x st ?2eb,2ec,4ab ? ?Jasione

Falseuncaria ruficiliana (Haworth, 1811) x x eu 1d,2e,3d,4a 6 herbs

Tortrix viridana l., 1758 x x st 2Ba,4F 5 deciduous trees/shrubs: esp. QuercusQuercus P!

Aleimma loe��ingiana (l., 1758) x x x st 2Ba 2 Quercus

Acleris forsskaleana (l., 1758) x eu 2a,2B,2ea 2 Acer

Acleris sparsana (d. & Schiff., 1775) x x eu 2aa,2B,4F 5 deciduous trees/shrubs: esp. QuercusQuercus

Acleris rhombana (d. & Schiff., 1775) x eu 2a,2B,2C,2ea 5 deciduous trees/shrubs

Acleris schalleriana (l., 1761) x x x eu 2 2 Viburnum

Acleris variegana (d. & Schiff., 1775) x x x eu 2 5 deciduous trees/shrubs: esp. wooden rosaceae

Acleris permutana (dup., 1836) x x x st 2Ba 3 rosaceae (Rosa, Prunus)

Acleris cristana (d. & Schiff., 1775) x x x st 2Ba,2ea 3 rosaceae (Crataegus, Prunus)

Acleris literana (l., 1758) x x st 2Ba 2 Quercus

Acleris emargana (Fabricius, 1775) x eu 2a,2B 5 deciduous trees/bushes: esp. SalicaceaeSalicaceae

Doloploca punctulana (d. & Schiff., 1775) x x eu 2a,2B,2ea 5 deciduous trees/shrubs

Tortricodes alternella (d. & Schiff., 1775) x x st 2Ba,4F 2 Quercus

Eana argentana (Clerck, 1759) x x x x x eu 1dc,3C,3d 6 herbs, grass

Eana osseana (Scopoli, 1763) x x x eu 1dc,3C,3d 6 herbs, grass

Eana canescana (Guenée, 1845) x x st 3a,4aa 6 ?herbs

Eana penziana (Thunberg & Becklin, 1791) x x x eu 3a,3B,3C,3d 6 ?Poaceae: esp. Festuca

Eana incanana (Stephens, 1852) x st 2Ba,2ea 6 herbs

Cnephasia incertana (Treitschke, 1835) x eu 1,2,3,4 7 herbs, deciduous trees/shrubs, conifers

Cnephasia stephensiana (doubleday, 1849) x eu 1,2,3,4 7 herbs, deciduous trees/shrubs, conifers

Cnephasia alticolana (H.-S., 1851) x eu 3B,3C,4a 6 herbs

Cnephasia asseclana (d. & Schiff., 1775) x x eu 1,2,3,4 7 herbs, deciduous trees/shrubs,conifers

Cnephasia communana (H.-S., 1851) x eu 1,2,3,4 6 herbs

Sparganothis pilleriana (d. & Schiff., 1775) x eu 1d,2,4Bb 7 herbs, deciduous trees/shrubs

Eulia ministrana (l., 1758) x x x x x eu 2B,3C 5 deciduous trees/shrubs

Pseudargyrotoza conwagana (Fabricius, 1775) x x x eu 2a,2B,2e,4e 3 oleaceae: Ligustrum, Fraxinus

Epagoge grotiana (Fabricius, 1781) x x x eu 2B 5 deciduous trees/shrubs: esp. QuercusQuercus, Rubus

Paramesia gnomana (Clerck, 1759) x eu 2a,2B,2eb 7 deciduous trees/shrubs, herbs

Philedone gerningana (d. & Schiff., 1775) x eu 2a,2B,4aa 6 herbs

Capua vulgana (Frölich, 1828) x x x eu 2a,2B,2ea 5 deciduous trees/shrubs

Archips oporana (l., 1758) x x x eu 2C 5 conifers

Archips podana (Scopoli, 1763) x x x eu 2,4e 5 deciduous trees/shrubs, conifers

Archips xylosteana (l., 1758) x x x st 2B 5 deciduous trees/shrubs: esp. Quercus

Archips rosana (l., 1758) x eu 2B 5 deciduous trees/shrubs: esp. RosaceaeRosaceae




Argyrotaenia ljungiana (Thunberg, 1797) x x eu 2,4e 7 esp. deciduous trees/shrubs, herbs

Choristoneura hebenstreitella (Müller, 1764) x x x st 2Ba,2ea 5 deciduous trees/shrubs: esp. QuercusQuercus P!

Ptycholomoides aeriferanus (H.-S., 1851) x x x st 2C 1 Larix

Ptycholoma lecheana (l., 1758) x x x eu 2a,2B,2ea 5 deciduous trees/shrubs

Pandemis corylana (Fabricius, 1794) x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Pandemis cerasana (Hübner, 1786) x x eu 2a,2B,2ea,4e 7 deciduous trees/shrubs, herbs

Pandemis cinnamomeana (Treitschke, 1830) x x x eu 2,4e 5 deciduous trees/shrubs, conifers

Pandemis heparana (d. & Schiff., 1775) x x x eu 2a,2B,2ea,4e 7 deciduous trees/shrubs, herbs

Syndemis musculana (Hübner, 1799) x x x eu 2 5 deciduous trees/shrubs, conifers

Lozotaenia forsterana (Fabricius, 1781) x st 3C 5 deciduous trees/shrubs: esp. Vaccinium

Dichelia histrionana (Frölich, 1828) x st 2C 4 Pinaceae

Clepsis rurinana (l., 1758) x x x eu 2aa,2B 5 deciduous trees/shrubs

Bactra lancealana (Hübner, 1799) x x st 1C,1d 6 Juncaceae, Cyperaceae

Endothenia gentianaeana (Hübner, 1799) x st 2Ba,2e 6 herbs

Endothenia marginana (Haworth, 1811) x eu 1d,2a,2B 6 Scrophulariaceae, labiatae

Endothenia sp. x st 2Ba,2e

Eudemis profundana (d. & Schiff., 1775) x x x st 2Ba 2 Quercus

Pseudosciaphila branderiana (l., 1758) x x st 2ab 1 Populus tremula

Apotomis inundana (d. & Schiff., 1775) x st 2ab 1 Populus tremula

Orthotaenia undulana (d. & Schiff., 1775) x eu 2 7 herbs, deciduous trees/shrubs

Hedya dimidioalba (retzius, 1783) x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs: esp. rosaceaerosaceae

Metendothenia atropunctana (Zetterstedt, 1839)

x x eu 2,3C 5 deciduous trees/shrubs: esp. BetulaBetula, Salix

Celypha striana (d. & Schiff., 1775) x x x eu 1dc,2eb,4aa,4ab 3 asteraceae

Celypha ��avipalpana (H.-S., 1851) x x eu 2e,4aa,4ab,4C 6 herbs

Celypha lacunana (d. & Schiff., 1775) x x x x eu 1d,2,3C,4 7 herbs, deciduous trees/shrubs, conifers

Phiaris metallicana (Hübner, 1799) x x x st 3C 6 herbs l!

Phiaris schulziana (Fabricius, 1776) x st 1da,2C,3C 3 ericaceae

Phiaris bipunctana (Fabricius, 1794) x st 2C,3C 2 Vaccinium

Stictea mygindiana (d. & Schiff., 1775) x st 3C 3 ericaceae: esp. Vaccinium

Olethreutes arcuella (Clerck, 1759) x x x st 2B 8 dead wood/fungi

Piniphila bifasciana (Haworth, 1811) x x x st 2Ca,2Cd 2 Pinus

Pseudohermenias abietana (Fabricius, 1787) x st 2Ca,2Cb 3 Pinaceae: Pinus, Abies

Lobesia bicinctana (dup., 1844) x x st 4aa 2 Allium

Thiodia torridana (lederer, 1859) x x st 2Ba,4aa 6 herbs

Rhopobota ustomaculana (Curtis, 1831) st 2Cc,2Ce,3C 1 Vaccinium vitis-idaea

Rhopobota naevana (Hübner, 1817) x x eu 2,3C,4e 5 deciduous trees/shrubs

Spilonota ocellana (d. & Schiff., 1775) x x x eu 2 5 deciduous trees/shrubs

Spilonota laricana (Heinemann, 1863) x x x st 2C 3 Pinaceae: Larix, rarely Picea

Epinotia solandriana (l., 1758) x x eu 2a,2B,2d,2ea 5 deciduous trees/shrubs

Epinotia festivana (Hübner, 1799) x x x st 2Ba 2 Quercus

Epinotia abbreviana (Fabricius, 1794) x st 2aa,2B 5 ulmaceae, rarely aceraceae

Epinotia subocellana (donovan, 1806) x st 1dd,2ab 2 Salix

Epinotia tenerana (d. & Schiff., 1775) x x x st 2a 5 deciduous trees/shrubs, esp. Alnus

Epinotia tedella (Clerck, 1759) x x x x x x eu 2C 5 Conifers, esp. Picea abies l!

Epinotia thapsiana (Zeller, 1847) x x x st 4aa 3 apiaceae




Epinotia granitana (H.-S., 1851) x st 2Cb,2Cc,2Ce 1 Picea

Epinotia nanana (Treitschke, 1835) x st 2C 2 Pinus

Zeiraphera ratzeburgiana (Saxesen, 1840) x st 2C 3 Pinaceae

Zeiraphera isertana (Fabricius, 1794) x x x st 2Ba,4F 2 Quercus

Zeiraphera griseana (Hübner, 1799) x x x x st 2Cd 3 Pinaceae, esp. Larix

Phaneta pauperana (dup., 1843) x st 2Ba,2ea 2 Rosa

Eucosma cana (Haworth, 1811) x x x eu 4a,4C 3 asteraceae: Cirsium, Carduus

Eucosma hohenwartiana (d. & Schiff., 1775) x x x st 1db,4aa 3 asteraceae: Centaurea, Serratula

Eucosma fulvana (Stephens, 1834) x st 4aa 1 Centaurea scabiosa

Eucosma scutana (Constant,, 1893) x x st 1dc,2Ba,4aa 1 Serratula

Eucosma campoliliana (d. & Schiff., 1775) x x eu 2eb,4ab,4C 1 Senecio jacobaea

Eucosma albidulana (H.-S., 1851) x st 4aa 1 ?Serratula

Eucosma conterminana (H.-S., 1851) x x st 2Ba,2eb,2ec 2 Solidago

Eucosma pupillana (Clerck, 1759) x st 4aa 1 Artemisia absinthium

Eucosma lugubrana (Treitschke, 1830) x st 4aa ? unknown

Gypsonoma dealbana (Frölich, 1828) x x eu 2a,2B 5 deciduous trees/shrubs

Gypsonoma sociana (Haworth, 1811) x x st 2a 2 Salix

Epiblema foenella (l., 1758) x st 4aa,4ab,4Cc 2 Artemisia

Epiblema obscurana (H.-S., 1851) x st 4aa 2 Inula

Notoelia cynosbatella (l., 1758) x x x eu 2B,2ea,4e 2 Rosa

Notocelia uddmanniana (l., 1758) x x st 2a,2B,2e 2 Rubus

Notocelia roborana (d. & Schiff., 1775) x x x x st 2Ba,2ea,4e 2 ?Rosa

Notocelia rosaecolana (doubleday, 1850) x eu 2a,2B 2 Rosa

Rhyacionia buoliana (d. & Schiff., 1775) x x st 2Ca,2Cd 2 Pinus

Rhyacionia pinicolana (doubleday, 1849) x x x st 2Ca,2Cd 2 Pinus

Rhyacionia pinivorana (lienig & Zeller, 1846)

x x x st 2Ca,2Cd 2 Pinus

Enarmonia formosana (Scopoli, 1763) x x eu 2aa,2B,2ea,4e 3 rosaceae: Malus, Prunus

Ancylis unguicella (l., 1758) x x x eu 2C,3C 3 ericaceae: Calluna, Erica

Ancylis laetana (Fabricius, 1775) x x x st 2a 2 Populus, esp. P. tremula

Ancylis obtusana (Haworth, 1811) x eu 1dd,2a,2B,2ea 5 rosaceae, rhamnaceae

Ancylis unculana (Haworth, 1811) x x eu 1dd,2a,2B,2ea 4 rhamnaceae

Ancylis comptana (Frölich, 1828) x eu 1d,2e,3C,3d 7 herbs, deciduous shrubs

Ancylis myrtillana (Treitschke, 1830) x st 3C 2 Vaccinium

Ancylis apicella (d. & Schiff., 1775) x x x eu 1d,2B,2e,3a 4 rhamnaceae

Ancylis badiana (d. & Schiff., 1775) x x x x x eu 2B,2e,3C,3d 3 Fabaceae

Ancylis mitterbacheriana (d. & Schiff., 1775) x x x st 2Ba,2Bc,4F 3 Fagaceae: Quercus, Castanea

Grapholita fissana (Frölich, 1828) x x st 2B,2e 2 Vicia

Grapholita caecana (Schläger, 1847) x st 1d,4aa 2 Onobrychis

Grapholita jungiella (Clerck, 1759) x x x eu 1dc,2B,2e 3 Fabaceae: Lathyrus, Vicia

Grapholita tenebrosana (dup., 1843) x st 2Ba,2ea 2 Rosa

Cydia succedana (d. & Schiff., 1775) x x x eu 2Ba,2e,4aa,4ab 3 Fabaceae

Cydia milleniana (adamczewski, 1967) x st 2Cd,2Ce 1 Larix




Cydia pomonella (l., 1758) x eu 2a,2B,4Ba,4e,4F 5 rosaceae, Juglandaceae, Fagaceae

Cydia penkleriana (d. & Schiff., 1775) x x x st 2Ba,4F 4 Fagaceae: Quercus, Castanea (fruits)

Cydia fagiglandana (Zeller, 1841) x x x eu 2Ba,2Bc,4F 4 Fagaceae: Fagus, Quercus (fruits)

Pammene fasciana (l., 1761) x st 2Ba,4F 4 Fagaceae: Quercus, Castanea (fruits)

Pammene gallicolana (lienig & Zeller, 1846) x x st 2Ba 2 Quercus, galls of Cynpiidae

Pammene germmana (Hübner, 1799) eu 2B,2ea,4Ba,4e 3 rosaceae: Prunus, Crataegus

Dichrorampha aeratana (Pierce & Metcalfe, 1915)

x eu 2e,4aa,4Cc,4Cd 2 asteraceae: Chrysanthemum

Dichrorampha petiverella (l., 1758) x x eu 2e,4aa,4Cc,4Cd 3 asteraceae: Achillea, Chrysanthemum

eperMeniiDae

Ochromolopis ictella (Hübner, 1813) x x st 3B,4aa,4ab 2 Thesium

Epermenia pontificella (Hübner, 1796) x x x st 3B,4Cd 2 Thesium

Epermenia scurella (Stainton, 1851) x st 2d,3C,3d 3 apiaceae

Epermenia aequidentella (Hofmann, 1867) x x eu 2e,3C,4aa,4ab 3 apiaceae

alucitiDae

Pterotopteryx zonodactyla Zeller, 1847 x st 2Ba

pterophoriDae

Geina didactyla (l., 1758) x st 2e,4ab 2 Geum

Cnaemidophorus rhododactyla (d. & Schiff., 1775)

x x st 2Ba,2ea 2 Rosa

Platyptilia farfarella Zeller, 1867 x x st 2e,4Cc 2 Senecio

Platyptilia gonodactyla (d. & Schiff., 1775) x eu 1Bb,2eb,2ec,4Cd 3 asteraceae: Tussilago, Petasites

Gillmeria pallidactyla (Haworth, 1811) x st 4ab 3 asteraceae: esp. Achillea

Amblyptilia punctidactyla (Haworth, 1811) x x x eu 1d,2,4ab,4C 6 herbs

Stenoptilia pterodactyla (l., 1761) x eu 1d 2 ?Veronica

Stenoptilia bipunctidactyla (Scopoli, 1763) x x x eu 1dc,2eb,4aa,4ab 3 ?dipsacaceae

Stenoptilia annadactyla Sutter, 1988 x st 4aa 1 Scabiosa columbaria

Stenoptilia coprodactyla (Stainton, 1851) x x x st 3C,3d,4ab 2 Gentiana

Marasmarcha lunaedactyla (Haworth, 1811) x x st 2Ba,4aa 2 Ononis

Adaina microdactyla (Hübner, 1813) x x st 1Bb,2a,2ec 2 Eupatorium

Odeaematophorus carphodactylus (Hübner, 1813)

x st 1dc,4aa,4ab 2 Inula

Oidaematophorus osteodactylus (Zeller, 1841) x eu 2d,2e,3C,4ab 3 asteraceae

Oidaematophorus tephradactylus (Hübner, 1813)

x st 2C,2e,3C 3 asteraceae

Oidaematophorus lithodactylus (Treitschke, 1833)

x x st 1dc,4aa,4ab 3 asteraceae: Inula, Pulicaria

Pterophorus pentadactylus (l., 1758) x eu 1d,2e,4C,4e 4 Convolvulaceae

Merrifieldia leucodactyla (d. & Schiff., 1775) x x eu 3C,3d,4a 3 labiatae

Merrifieldia tridactyla (l., 1758) x x x st 4aa 3 Thymus

Emmelina monodactyla (l., 1758) x x x eu 1d,2,4e 4 Convolvulus

pyraliDae

Aphomia sociella (l., 1758) x x x eu 2 9 bumble bees wax

Hypsopygia costalis (Fabricius, 1775) x x x eu 2,4d,4e 9 dead vegetable substances




Synaphe punctalis (Fabricius, 1775) x st 2a,2e 8 moss

Actenia brunnealis (Treitschke, 1829) x st 4aa 6 herbs

Orthopygia glaucinalis (l., 1758) x eu 2,4d,4e 9 dead vegetable substances

Pyralis farinalis (l., 1758) x eu 2B,2e,4d,4e 9 dead vegetable substances

Pyralis regalis (d. & Schiff., 1775) x x x st 2Ba 9 dead vegetable substances

Endotricha ��ammealis (d. & Schiff., 1775) x x x eu 1dd,2 9 dead vegetable substances etc.

Cryptoblabes bistriga (Haworth, 1811) x x eu 2a,2B 5 deciduous trees/shrubs: Betula, Alnus, Quercus

Oncocera semirubella (Scopoli, 1763) x x x st 2Ba,4aa,4ab 3 Fabaceae

Laodamia faecella (Zeller, 1839) x st 4aa

Pempelia palumbella (d. & Schiff., 1775) x x x st 2Ba,2Ca,4aa 6 herbs, esp. ericaceae

Pempelia obductella (Zeller, 1839) x st 2Ba,2eb,4aa 3 labiatae: Satureja, Mentha

Pempelia formosa (Haworth, 1811) x st 2Ba 2 Ulmus

Salebriopsis albicilla (H.-S., 1849) x x st 2aa,2B 5 deciduous trees/shrubs, esp. TiliaTilia

Sciota rhenella (Zincken, 1818) x st 2a,2ec 2 Populus

Selagia argyrella (d. & Schiff., 1775) x x x st 2Ca,4aa 1 Calluna

Selagia spadicella (Hübner, 1796) x x x st 2Ca,4aa 6 herbs: esp. Calluna, Teucrium

Phycita roborella (d. & Schiff., 1775) x x x st 2aa,2Ba,4F 5 Fagaceae: Quercus, rosaceae: Malus P!

Dioryctria abietella (d. & Schiff., 1775) x x x x x st 2Ca,2Cb,2Cc 2 Pinus

Dioryctria sylvestrella (ratzeburg, 1840) x x st 2Ca 2 Pinus

Hypochalcia ahenella (d. & Schiff., 1775) x x eu 1dc,2eb,4aa,4ab 6 herbs

Hypochalcia bruandella (Guenée, 1845) x x st 4aa

Elegia fallax (Staudinger, 1881) x x st 2Ba 2 Quercus

Elegia similella (Zincken, 1818) x x x st 2Ba 2 Quercus

Pyla fusca (Haworth, 1811) x x x st 2Cc,2Ce,2Cd,3C 3 ericaceae (esp. Erica)

Etiella zinckenella (Treitschke, 1832) x x x st 2Ba,4aa 3 Fabaceae

Pempeliella ornatella (d. & Schiff., 1775) x st 4aa,4Cd 2 Thymus

Pempeliella dilutella (d. & Schiff., 1775) x eu 2Ba,2eb,4aa,4ab 6 herbs

Nephopterix angustella (Hübner, 1796) x st 2B,2ea 1 Euonymus

Acrobasis glaucella Staudinger, 1859 x x x st 2Ba 2 Quercus

Acrobasis consociella (Hübner, 1813) x x x st 2Ba 2 Quercus

Conobathra tumidana (d. & Schiff., 1775) x x st 2Ba 2 Quercus

Conobathra repandana (Fabricius, 1798) x st 2Ba 2 Quercus

Trachycera advenella (Zincken, 1818) x x x st 2Ba,2ea 3 wooden rosaceae

Trachycera suavella (Zincken, 1818) x st 2Ba,2ea 3 wooden rosaceae

Trachycera marmorea (Zeller, 1848) x st 2Ba,2ea 3 wooden rosaceae

Myelopsis tetricella (d. & Schiff., 1775) x x x st 2Ba,2ea 5 deciduous trees/shrubs: Salix, ?Quercus

Assara terebrella (Zincken, 1818) x x x st 2C 1 Picea abies

Euzophera bigella (Zeller, 1848) x x eu 2a,2B,2ea 5 deciduous trees/shrubs, conifers

Nyctegretis lineana (Scopoli, 1786) x st 4aa,4Cc,4Cd 6 herbs: Ononis, Trifolium, Artemisia

Ancylosis cinnamomella (dup., 1836) x x x st 3a,4aa 6 herbs

Homoeosoma inustellum ragonot, 1884 x st 4aa

Phycitodes binaevella (Hübner, 1813) x x eu 1d,2Ba,4aa,4ab 3 asteraceae




Phycitodes albatella (ragonot, 1887) x x x x st 4aa 3 asteraceae

Vitula biviella (Zeller, 1848) x st 2Ca 2 Pinus

Ephestia elutella (Hübner, 1796) x x x eu 2,4d 9 dead organic matter

Ephestia parasitella (Staudinger, 1859) x eu ?2,4d 9 dead vegetable substances

Ephestia welseriella (Zeller, 1848) x x x st 2Ba,4aa 9 dried fruits

craMbiDae

Euchromius ocellea (Haworth, 1811) x migrating species 9 dead vegetable substances

Chrysoteuchia culmella (l., 1758) x x eu 1dc,2e,4a,4e 4 Poaceae

Crambus ericella (Hübner, 1813) x x x st 1d 3 ericaceae l!

Crambus pratella (l., 1758) x x eu 1dc,2e,4aa,4ab 4 Poaceae

Crambus lathoniellus (Zincken, 1817) x x x x x x eu 1dc,2e,4a,4e 4 Poaceae

Crambus perlella (Scopoli, 1763) x eu 1dc,2e,4a,4e 4 Poaceae

Agriphila tristella (d. & Schiff., 1775) x x eu 1d,4a 4 Poaceae

Agriphila inquinatella (d. & Schiff., 1775) x eu 4aa,4ab 4 Poaceae

Agriphila straminella (d. & Schiff., 1775) x x x eu 1dc,4aa,4ab 4 Poaceae

Catoptria myella (Hübner, 1796) x x x eu 1Bc,2ea,4aa,4Cd 8 moss

Catoptria speculalis Hübner, 1825 x x st 3C,3d 8 ?moss

Catoptria pyramidella (Treitschke, 1832) x x st 3C,3d 8 moss

Catoptria conchella (d. & Schiff., 1775) x x x st 3C,3d 8 ?moss

Catoptria mytilella (Hübner, 1805) x x x st 2Ba,? 8 moss

Catoptria pinella (l., 1758) x x x eu 2,4a 6 grass

Catoptria falsella (d. & Schiff., 1775) x x x x eu 1d,2,4d,4e 8 moss

Thisanotia chrysonuchella (Scopoli, 1763) x x x st 4aa,4ab 4 Poaceae

Scoparia italica Turati, 1919 x x eu 2 8 moss

Scoparia subfusca Haworth, 1811 x eu 2 8 moss

Eudonia lacustrata (Panzer, 1804) x x x x x x eu 1dd,2,4d,4e 8 moss

Eudonia murana (Curtis, 1827) x x x st 2C,3a 8 moss l!

Eudonia petrophila (Standfuss, 1848) x st 2C,?3a 8 moss

Eudonia truncicolella (Stainton, 1849) x st 2C 8 moss

Eudonia mercurella (l., 1758) x x x eu 2a,2B 8 moss

Eudonia sudetica (Zeller, 1839) x x x st 3B 8 moss

Evergestis aenealis (d. & Schiff., 1775) x x eu 4aa,4ab 3 Cruciferae

Evergestis sophialis (Fabricius, 1787) x x x st 3a,3B 3 Cruciferae

Cynaeda dentalis (d. & Schiff., 1775) x x x st 4aa,4Cc,4Cd 3 Boraginaceae: Echium, Anchusa

Metaxmeste phrygialis (Hübner, 1796) x eu 3a 6 ?herbs

Metaxmeste schrankiana (Hochenwarth, 1785) x eu 3a 6 ericaceae, ?herbs

Harpadispar diffusalis (Guenée, 1854) x x st 2Ba,4aa 6 herbs

Pyrausta purpuralis (l., 1758) x eu 2eb,4aa,4ab,4C 4 labiatae

Pyrausta despicata (Scopoli, 1763) x eu 2eb,4a,4C 2 Plantago

Pyrausta aerealis (Hübner, 1793) x eu 3C,3d 6 herbs

Loxostege virescalis (Guenée, 1854) x st 4aa 2 Artemisia

Ecpyrrhorrhoe rubiginalis (Hübner, 1796) x x st 2eb,4aa 3 labiatae




Sitochroa palealis (d. & Schiff., 1775) x st 4aa,4ab 4 apiaceae

Sitochroa verticalis (l., 1758) x x eu 1d,2B,2e,4a 6 herbs

Microstega pandalis (Hübner, 1825) x x eu 1d,2a,2B,2e,4a 6 herbs

Ostrinia nubilalis (Hübner, 1796) x eu 4a,4Bc 6 herbs, esp. Zea

Eurrhypara hortulata (l., 1758) x eu 2eb,4C 6 herbs: esp. Urtica

Perinephela lancealis (d. & Schiff., 1775) x x eu 1d,2a,2eb 6 herbs: esp. Stachys, Senecio

Phlyctaenia coronata (Hufnagel, 1767) x st 2Ba,2ea 5 deciduous trees/shrubs

Mutuuraia terrealis (Treitschke, 1829) x x x x eu 2eb,4aa,4C 3 asteraceae: Solidago, Aster

Anania funebris (Ström, 1768) x x eu 3d,4aa,4ab 6 herbs

Anania verbascalis (d. & Schiff., 1775) x x x eu 1dc,2a,2B,2e,4a 6 herbs

Ebulea crocealis (Hübner, 1796) x x x st 4aa,4ab 3 asteraceae: Inula, Pulicaria

Obsibotys fuscalis (d. & Schiff., 1775) x x x x eu 1d,2a,2B,2e 6 herbs

Udea lutealis (Hübner, 1809) x eu 3C,4ab 6 herbs

Udea inquinatalis (lienig & Zeller, 1846) x st 3Ca 5 deciduous shrubs: Betula, Salix, Vaccinium

Udea prunalis (d. & Schiff., 1775) x eu 2 7 herbs, deciduous trees/shrubs

Udea olivalis (d. & Schiff., 1775) x x x eu 1,2,4 6 herbs

Udea nebulalis (Hübner, 1796) x x st 3C,3d 6 herbs

Udea decrepitalis (H.-S., 1847) x x st 3C 6 herbs

Udea cyanalis (la Harpe, 1855) x x ? ? ? unknown

Udea uliginosalis (Stephens, 1829) x x x eu 3C,3d 6 herbs

Udea ferrugalis (Hübner, 1796) x x x migrating species 6 herbs

Mecyna ��avalis (d. & Schiff., 1775) x x x st 2Ba,4aa 6 herbs

Nomophila noctuella (d. & Schiff., 1775) x x migrating species 6 herbs, grass

Dolicharthria punctalis (d. & Schiff., 1775) x st 2Ba,2ea 6 herbs

Metasia ophialis (Treitschke, 1829) x x st ?2Ba,4aa ? unknown

Pleuroptya ruralis (Scopoli, 1763) x x x eu 2eb,4C,4e 1 Urtica

Pleuroptya balteata (Fabricius, 1798) x x st 2Ba,4aa 3 apiaceae

Agrotera nemoralis (Scopoli, 1763) x x x st 2Ba 5 deciduous trees/shrubs: esp. CarpinusCarpinus

thyriDiDae

Thyris fenestrella (Scopoli, 1763) x st 2ea 1 Clematis vitalba

lasiocaMpiDae

Malacosoma neustrium (l., 1758) x x x eu 2Ba,4Ba,4e 5 deciduous trees/shrubs

Malacosoma alpicolum Staudinger, 1870 x x eu 3C,3d,4a 7 herbs, deciduous trees/shrubs

Trichiura crataegi (l., 1758) x x eu 2,3,4 5 deciduous trees/shrubs

Lasiocampa quercus (l., 1758) x x x x eu 1da,2e,3C 7 deciduous trees/shrubs, Vaccinium, Rubus

Lasiocampa trifolii (d. & Schiff., 1775) x x st 4aa,4ab 6 herbs, grass

Macrothylacia rubi (l., 1758) x x x x x x eu 2,4 6 herbs

Cosmotriche lobulina (d. & Schiff., 1775) x x st 2Cc,2Ce 1 ?Pinus cembra

Phyllodesma tremulifolia (Hübner, 1810) x x x eu 2 5 deciduous trees/shrubs

Odonestis pruni (l., 1758) x eu 2a,4Ba 5 deciduous trees/shrubs

Dendrolimus pini (l., 1758) x x x eu 2C 3 Pinaceae: Pinus (Pices, Abies)




enDroMiDiDae

Endromis versicolora (l., 1758) x st 1dd,2a 5 deciduous trees/shrubs

sphingiDae

Agrius convolvuli (l., 1758) x x migrating species 2 Convolvulus

Sphinx ligustri (l., 1758) x eu 2a,2ea,4e 5 deciduous trees/shrubs: esp. oleaceae

Hyloicus pinastri (l., 1758) x x eu 2C 5 conifers

Mimas tiliae (l., 1758) x eu 2,4e 5 deciduous trees/shrubs

Laothoe populi (l., 1758) x x x eu 2 4 Salicaceae: Populus, Salix

Hemaris fuciformis (l., 1758) x eu 2 2 Lonicera

Macroglossum stellatarum (l., 1758) x x migrating species 2 Galium

Hyles livornica (esper, 1780) x migrating species 6 herbs: esp. Galium, Linaria

Deilephila elpenor (l., 1758) x eu 2,4e 6 herbs

Deilephila porcellus (l., 1758) x x x eu 2,4 6 herbs

saturniiDae

Saturnia pyri (d. & Schiff., 1775) x x eu 2Ba,4Ba,4e 5 deciduous trees/shrubs

Saturnia pavoniella (Scopoli, 1763) x x x eu 1d,2 5 deciduous trees/shrubs,rosaceae

Aglia tau (l., 1758) x x x eu 2Bc 5 deciduous trees/shrubs, esp. Fagus

hesperiiDae

Thymelicus lineolus (ochsenheimer, 1808) x st 2Ba,4aa 4 Poaceae

Erynnis tages (l., 1758) x x x eu 2e,4a,4C 3 Fabaceae

Pyrgus malvoides (elwes & edwards, 1897) x x st 4aa,4ab 2 Potentilla

Pyrgus serratulae (rambur, 1840) x st 3a,4aa,4ab 2 Potentilla

Pyrgus fritillarius (Poda, 1761) x x st 4aa 6 herbs

Ochlodes venatus faunus Turati, 1905 x x x eu 1d,2e,4a 4 Poaceae

papilioniDae

Papilio machaon l., 1758 x eu 1,4 3 apiaceae

Iphiclides podalirius (l., 1758) x x x st 2Ba,2ea 3 wooden rosaceae

pieriDae

Leptidea sinapis (l., 1758) x x x eu 2e,4a 3 Fabaceae

Colias hyale (l., 1758) x eu 4a,4C 2 Medicago

Colias alfacariensis ribbé, 1905 x x st 4aa,4ab 1 Hippocrepis comosa

Gonepteryx rhamni (l., 1758) x x eu 2a,2B,2e,4e 3 rhamnaceae

Pieris brassicae (l., 1758) x eu 2e,4Bc,4C,4e 4 Cruciferae

Pieris rapae (l., 1758) x x x eu 2e,4a,4Bc,4e 4 Cruciferae

Pieris napi (l., 1758) x x x x x eu 2e,4a,4Bc,4e 4 Cruciferae

Anthocharis cardamines (l., 1758) x x x eu 2e,4B,4C,4e 3 Cruciferae: Cardamine

nyMphaliDae

Limenitis camilla (l., 1764) x x eu 2a,2B 3 Caprifoliaceae

Nymphalis polychloros (l., 1758) x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Nymphalis antiopa (l., 1758) x eu 2a,2B,2ea 5 deciduous trees/shrubs




Vanessa atalanta (l., 1758) x x eu migrating species 1 Urtica

Aglais urticae (l., 1758) x x eu 2e,4C,4e 1 Urtica

Cynthia cardui (l., 1758) x x eu migrating species 6 herbs

Polygonia c-album (l., 1758) x x eu 2e,4C,4e 7 deciduous trees/shrubs, herbs

Argynnis paphia (l., 1758) x x x x st 2e 2 Viola

Argynnis aglaja (l., 1758) x eu 1dc,2e,4ab 6 herbs: Viola, Polygonum

Argynnis niobe (l., 1758) x eu 1d,4 2 Viola

Boloria pales (d. & Schiff., 1775) x st 3d 2 Viola

Boloria euphrosyne (l., 1758) x x eu 2ec 2 Viola

Melitaea cinxia (l., 1758) x x x st 4aa,4ab 6 herbs: Plantago, Hieracium

Melitaea didyma (esper, 1779) x x x st 4aa,4ab 6 herbs: Plantago, Centaurea

Mellicta athalia (rottemburg, 1775) x x x eu 1d,2e,4ab 6 herbs

Melanargia galathea (l., 1758) x x x st 1dc,4aa,4ab 4 Poaceae P!

Satyrus ferula (Fabricius, 1793) x x x st 2Ba,4aa 1 Festuca ovina P!

Hipparchia statilinus (Hufnagel, 1766) x x st 2Ba,2Ca,4aa 4 Poaceae

Minois dryas (Scopoli, 1763) x x x st 1dc,4aa 4 Poaceae

Erebia euryale (esper, 1805) x x x st 2C 4 Poaceae

Erebia aethiops (esper, 1777) x x x eu 2 4 Poaceae

Erebia medusa (d. & Schiff., 1775) x x x eu 1dc,2ec 4 Poaceae

Erebia cassioides (reiner & Hochenwarth, 1793)

x x x st 3d 4 Poaceae

Maniola jurtina (l., 1758) x x x eu 1dc,2e,4a 4 Poaceae

Coenonympha gardetta (Prunner, 1798) x x eu 3C,3d,4a 4 Poaceae

Pararge aegeria (l., 1758) eu 2a,2B,2C 6 Cyperaceae, Poaceae

Lasiommata megera (l., 1767) x x x st 3a 4 Poaceae

rioDiniDae

Hamearis lucina (l., 1758) x x st 2eb,2ec 2 Primula

lycaeniDae

Callophrys rubi (l., 1758) x x x eu 2Ca,2e,3C,4ab 7 herbs, deciduous shrubs

Neozephyrus quercus (l., 1758) x x x st 2Ba 2 Quercus

Cupido minimus (Fueslly, 1775) x x x eu 2e,3C,4ab,4ad 3 Fabaceae

Celastrina argiolus (l., 1758) x eu 2 5 deciduous trees/shrubs

Scolitantides orion (Pallas, 1771) x x x st 3a 2 Sedum

Glaucopsyche alexis (Poda, 1761) x x st 2Ba,4aa 3 Fabaceae

Plebejus idas (l., 1761) x st 2Ba,4aa 3 Fabaceae, Hippophaes

Aricia agestis (d. & Schiff., 1775) x st 2Ba,2eb 6 herbs: Geranium, Helianthemum

Polyommatus semiargus (rottemburg, 1775) x eu 2B,4aa,4ab 2 ?Trifolium pratense

Polyommatus thersites thersites (Cantener, 1834) x x st 4aa 1 Onobrychis viciifolia

Polyommatus coridon (Poda, 1761) x st 2Ba,4aa,4ab 3 Fabaceae

Polyommatus bellargus (rottemburg, 1775) x x x st 2Ba,4aa,4ab 3 Fabaceae

Polyommatus icarus (rottemburg, 1775) x x eu 2Ba,2e,4a,4C,4e 3 Fabaceae




DrepaniDae

Watsonalla binaria (Hufnagel, 1769) x x x eu 2B 5 deciduous trees/shrubs: esp. QuercusQuercus

Watsonalla cultraria (Fabricius, 1775) x eu 2B 1 Fagus

Drepana falcataria (l., 1758) x eu 2,4e 5 deciduous trees/shrubs

Sabra harpagula (esper, 1786) x st 2Ba 5 ?Fagaceae: Quercus, ?Tiliaceae

Cilix glaucata (Scopoli, 1763) x st 2Ba,2ea 3 wooden rosaceae

Thyatira batis (l., 1758) x x eu 2,4e 2 Rubus

Tethea ocularis (l., 1767) x eu 2a,4e,4F 2 Populus

Tethea or (Goeze, 1781) x x x x eu 2a,4e,4F 4 Salicaceae: Populus, Salix

Ochropacha duplaris (l., 1761) x x x st 1dd,2a 5 deciduous trees/shrubs

Polyploca ridens (Fabricius, 1787) x x x st 2Ba 2 Quercus

geoMetriDae

Alsophila aescularia (d. & Schiff., 1775) x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Pseudoterpna pruinata (Hufnagel, 1767) x x x st 2Ba,2Ca,2ea,4aa 3 Fabaceae: Genista, Sarothamnus

Geometra papilionaria (l., 1758) x eu 2a,2B 4 Betulaceae

Comibaena bajularia (d. & Schiff., 1775) x x st 2Ba 2 Quercus P!

Thetidia smaragdaria (Fabricius, 1787) x st 2Ba,4aa 3 asteraceae

Hemithea aestivaria (Hübner, 1789) x x x eu 2B,4e,4F 5 deciduous trees/shrubs

Thalera fimbrialis (Scopoli, 1763) x eu 1d,2e,4ac 6 herbs

Hemistola biliosata (Villers, 1789) x x x eu 2e,4e 2 Clematis

Cyclophora annulata (Schulze, 1775) x x st 2Ba 1 Acer campestre

Cyclophora quercimontaria (Bastelberger, 1897)

x st 2Ba 2 Quercus

Cyclophora punctaria (l., 1758) x x eu 2Ba,2ea,4e,4F 2 Quercus

Cyclophora linearia (Hübner, 1799) x x x eu 2B 4 Fagaceae: Fagus, Quercus

Phaiogramma etruscaria (Zeller, 1849) x st 2Ba,2e 7 herbs, deciduous shrubs

Jodis lactearia (l., 1758) x x eu 2B,2e,4e 5 deciduous trees/shrubs

Scopula imitaria (Hübner, 1799) x st 2Ba,4aa 7 herbs, deciduous shrubs

Scopula nigropunctata (Hufnagel, 1767) x eu 2B,2eb,4aa,4ab 6 herbs

Scopula ornata (Scopoli, 1763) x x eu 2B,2eb,4aa,4ab 6 herbs

Scopula marginepunctata (Goeze, 1781) x x x eu 3B,4aa 6 herbs: esp. Sedum

Scopula incanata (l., 1758) x x eu 3B,4aa 6 herbs

Scopula ternata (Schrank, 1802) x x x eu 1da,2Cc,2Ce,3C 2 Vaccinium

Scopula ��oslactata (Haworth, 1809) x x x eu 1d,2 7 herbs, deciduous trees/shrubs

Glossotrophia confinaria (H.-S., 1847) x st 4aa 6 herbs

Idaea ochrata (Scopoli, 1763) x x st 4aa 6 herbs, Poaceae

Idaea muricata (Hufnagel, 1767) x x st 1d,4aa 6 herbs

Idaea filicata (Hübner, 1799) x st 4aa 6 herbs

Idaea rusticata (d. & Schiff., 1775) x st 4aa 9 withered leaves

Idaea moniliata (d. & Schiff., 1775) x x x st 3B,4aa 6 herbs

Idaea obsoletaria (rambur, 1833) st 2Ba,4aa 6 herbs

Idaea biselata (Hufnagel, 1767) x x x eu 2a,2B 9 withered and decaying leaves

Idaea dilutaria (Hübner, 1799) x eu 2B,4aa 9 dead vegetable substances




Idaea humiliata (Hufnagel, 1767) x x x eu 2Ba,4aa,4ab 6 herbs: esp. Ononis

Idaea pallidata (d. & Schiff., 1775) x eu 2,4a,4B 9 withered leaves

Idaea contiguaria (Hübner, 1799) x eu 3a,3B 6 herbs: esp. Sedum

Idaea aversata (l., 1758) x x x x eu 2B,2e,4e 9 withered leaves

Idaea rubraria (Staudinger, 1871) x x x st 2Ba,3aa,4aa 6 herbs

Idaea degeneraria (Hübner, 1799) x x x st 2Ba,2e 7 herbs, Frangula

Idaea straminata (Borkhausen, 1794) x x eu 2Ba,2e,4aa 6 herbs

Idaea deversaria (H.-S., 1847) x st 2Ba,2eb,2ec 6 herbs

Rhodostrophia vibicaria (Clerck, 1759) x x x eu 2Ba,2ea,3B,4aa 7 herbs, deciduous shrubs

Rhodostrophia calabra (Petagna, 1787) x x x st 2Ba,2ea 3 Fabaceae: Genista, Sarothamnus

Cataclysme riguata (Hübner, 1813) x x x st 4aa 6 herbs

Scotopteryx moeniata (Scopoli, 1763) x x x st 2Ca,2ea,2eb 3 Fabaceae: Genista, Sarothamnus

Scotopteryx diniensis Neuburger, 1906 x x x st 2Ba 3 Fabaceae

Scotopteryx chenopodiata (l., 1758) x x x eu 1Bb,4ac,4C 3 Fabaceae

Scotopteryx luridata (Hufnagel, 1767) x x x st 2Ba,2Ca,2e 3 Fabaceae

Xanthorhoe designata (Hufnagel, 1767) x x x x eu 1,2a 4 Cruciferae

Xanthorhoe munitata (Hübner, 1809) x x eu 3 6 herbs

Xanthorhoe spadicearia (d. & Schiff., 1775) x x eu 2,3C,4 6 herbs

Xanthorhoe ferrugata (Clerck, 1759) x eu 1Bb,1d,2,4C,4e 6 herbs

Xanthorhoe montanata (d. & Schiff., 1775) x x x eu 2,3C 6 herbs

Xanthorhoe ��uctuata (l., 1758) x x eu 2,4C,4e 6 herbs

Xanthorhoe incursata (Hübner, 1813) x st 2Cc,2Ce 2 Vaccinium

Catarhoe rubidata (d. & Schiff., 1775) x st 2Bb,4a,4C 3 rubiaceae

Catarhoe cuculata (Hufnagel, 1767) x x x eu 1,2B,2e,4e,4F 2 Galium

Epirrhoe alternata (o.F.Müller, 1764) x x x eu 1d,2e,4C,4d,4e 2 Galium

Epirrhoe rivata (Hübner, 1813) x eu 2a,2B,2e 2 Galium

Epirrhoe galiata (d. & Schiff., 1775) x x x x eu 2B,2C,3B,4ab 2 Galium

Camptogramma bilineatum (l., 1758) x x x eu 2,4 6 herbs

Entephria nobiliaria (H.-S., 1852) x st 3a,3B 2 Saxifraga

Entephria ��avicinctata (Hübner, 1813) x x eu 3a,3B 6 herbs: esp. Saxifraga

Entephria caesiata (d. & Schiff., 1775) x x x eu 1da,2B,2C,3C 7 herbs, deciduous shrubs l!

Earophila badiata (d. & Schiff., 1775) x eu 1d,2a,2B 2 Rosa

Anticlea derivata (d. & Schiff., 1775) x eu 1d,2a,2B 2 Rosa

Mesoleuca albicillata (l., 1758) x x x eu 2,4e 2 Rubus

Lampropteryx suffumata (d. & Schiff., 1775) x x eu 2Bc,2Cc 2 Galium

Cosmorhoe ocellata (l., 1758) x x x x eu 1,2,4ab,4e 2 Galium

Nebula salicata (d. & Schiff., 1775) x x x eu 2ab,3a,3B 2 Galium

Nebula tophaceata (d. & Schiff., 1775) x eu 2,3a 6 herbs: esp. Galium

Eulithis prunata (l., 1758) x eu 2B,2ea,4e 5 deciduous trees/shrubs: esp. RibesRibes

Eulithis populata (l., 1758) x x x eu 2C,3C 5 deciduous trees/shrubs: esp. Vaccinium l!

Eulithis pyraliata (d. & Schiff., 1775) x st 1d,2e 2 Galium

Ecliptopera silaceata (d. & Schiff., 1775) x x x eu 2a,2d,2e 6 herbs: Epilobium, Impatiens

Chloroclysta siterata (Hufnagel, 1767) x x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs




Chloroclysta miata (l., 1758) x x eu 2a,2Cc,2d 5 deciduous trees/shrubs, Vaccinium

Chloroclysta citrata (l., 1761) x x eu 2,3C,4e 7 herbs, deciduous trees/shrubs

Chloroclysta truncata (Hufnagel, 1767) x x x eu 2,3C,4e 7 herbs, deciduous trees/shrubs

Cidaria fulvata (Forster, 1771) x x x eu 2,4e 2 Rosa

Pennithera firmata (Hübner, 1822) x st 2Ca 1 Pinus sylvestris

Thera obeliscata (Hübner, 1787) x x x st 2Ca 1 Pinus sylvestris

Thera variata (d. & Schiff., 1775) x x x x x x eu 2C 5 conifers l!

Thera cembrae Kitt, 1912 x x st 2Ce 1 Pinus cembra

Thera stragulata (Hübner, 1809) x x st 2Cc 1 Picea abies

Thera cognata (Thunberg, 1792) x x x x eu 2C,3C 1 Juniperus communis l!

Electrophaes corylata (Thunberg, 1792) x x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs

Colostygia aptata (Hübner, 1813) x x x x x eu 2,3a,3B,3C 2 Galium

Colostygia olivata (d. & Schiff., 1775) x x eu 2B,2C,3a 6 herbs

Colostygia pectinataria (Knoch, 1781) x eu 2 6 herbs

Hydriomena furcata (Thunberg, 1784) x x eu 2 5 deciduous trees/shrubs: Salix, Vaccinium

Hydriomena impluviata (d. & Schiff., 1775) x eu 2a,2B,2d 5 deciduous trees/shrubs

Horisme vitalbata (d. & Schiff., 1775) x x x eu 2ea,2ec,4d,4e 2 Clematis

Horisme tersata (d. & Schiff., 1775) x x x x x eu 2a,2B,2e,4e 2 Clematis

Horisme radicaria (la Harpe, 1855) x st 2aa,2B 2 Clematis

Horisme aemulata (Hübner, 1813) x x x x x st 3a 2 Clematis

Melanthia procellata (d. & Schiff., 1775) x x x st 2ea,2ec,4e 2 Clematis

Pareulype berberata (d. & Schiff., 1775) x x x eu 2Ba,2Ca,2ea,3B 1 Berberis

Hydria cervinalis (Scopoli, 1763) x eu 2Ca,2ea,4e 1 Berberis

Hydria undulata (l., 1758) x eu 1dd,2 5 deciduous trees/shrubs

Triphosa dubitata (l., 1758) x x x x eu 2,4e 5 deciduous trees/shrubs: esp. Rhamnus

Philereme vetulata (d. & Schiff., 1775) x eu 2 4 rhamnaceae

Euphyia biangulata (Haworth, 1809) x st 2 3 Caryophyllaceae: esp. StellariaStellaria

Euphya adumbraria (H.-S., 1852) x st 3a 3 ?Caryophyllaceae

Euphya scripturata (Hübner, 1799) x st 3a,3B 6 herbs

Epirrita dilutata (d. & Schiff., 1775) x x x eu 2a,2B,2e,4F 5 deciduous trees/shrubs

Epirrita autumnata (Borkhausen, 1794) x x eu 1dd,2a,2B,2Cd 5 deciduous trees/shrubs

Operophtera brumata (l., 1758) x x eu 2B,4Ba,4e,4F 5 deciduous trees/shrubs

Mesotype parallelolineata (retzius, 1783) x eu 2 6 herbs

Mesotype verberata (Scopoli, 1763) x x x eu 2e,3,4ab,4ad 6 herbs

Perizoma alchemillata (l., 1758) x x x x x eu 2e,4e 6 herbs: esp. Galeopsis

Perizoma hydrata (Treitschke, 1829) x x st 3a,3B 2 Silene

Perizoma minorata (Treitschke, 1828) x x st 2e,4ab 2 Euphrasia

Perizoma blandiata (d. & Schiff., 1775) x st 1db,2e,4ab 2 Euphrasia

Perizoma albulata (d. & Schiff., 1775) x eu 1d,2a,2e,4ab 2 Rhinanthus

Perizoma ��avofasciata (Thunberg, 1792) x st 2a,2B,2e 2 Silene

Perizoma obsoletata (H.-S., 1838) x st 4ab 2 Gentiana

Perizoma incultaria (H.-S., 1848) x st 3a 1 Primula auricula

Martania taeniata (Stephens, 1831) x x x st 2C 6 herbs




Eupithecia tenuiata (Hübner, 1813) eu 1d,2a,2e 1 Salix caprea

Eupithecia haworthiata doubleday, 1856 x x x st 2e 2 Clematis

Eupithecia plumbeolata (Haworth, 1809) x x x eu 2e,4aa,4ab 3 Scrophularicaeae

Eupithecia abietaria (Goeze, 1781) x x x st 2Cc 5 conifers

Eupithecia analoga europaea lempke, 1969 x st 2Cc 1 Picea abies

Eupithecia venosata (Fabricius, 1787) x eu 2,3B 1 Silene vulgaris

Eupithecia gueneata Mabille, 1862 x st 2Ba,4aa 1 Pimpinella saxifraga

Eupithecia intricata arceuthata (Freyer, 1842) x x x x x eu 2Ca,2e,3C,4e 2 Juniperus

Eupithecia veratraria H.-S., 1848 x st 3a,4ab 1 Veratrum

Eupithecia absinthiata (Clerck, 1759) x x eu 2B,2e 4 asteraceae

Eupithecia vulgata (Haworth, 1809) x eu 2B,2e,4e 6 herbs: decaying leaves

Eupithecia icterata (Villers, 1789) x x x x eu 2,4e 4 asteraceae

Eupithecia impurata (Hübner, 1813) x x st 3a,3B 1 Campanula rotundifolia

Eupithecia semigraphata Bruand, 1851 x x st 4aa 3 labiatae: Origanum, Thymus

Eupithecia indigata (Hübner, 1813) x eu 2C 2 Pinus

Eupithecia druentiata dietze, 1902 x x st 4aa 1 Artemisia alba

Eupithecia distinctaria H.-S., 1861 x st 4aa 3 labiatae: Origanum, Thymus

Eupithecia gemellata H.-S., 1861 x x x st 3a,4aa 1 Petrohagia saxifraga

Eupithecia pimpinellata (Hübner, 1813) x eu 1d,2,4ab 4 apiaceae

Eupithecia abbreviata Stephens, 1831 x x st 2Ba 2 Quercus

Eupithecia dodoneata Guenée, 1857 x x x st 2Ba,2Bc,4F 2 Quercus

Eupithecia pusillata (d. & Schiff., 1775) x x x eu 2,4aa 2 Juniperus

Eupithecia ericeata (rambur, 1833) x x st 2Ba 2 Juniperus

Eupithecia lanceata (Hübner, 1825) x x st 2Cc 1 Picea abies

Eupithecia lariciata (Freyer, 1842) x x x x x x eu 2Cd,4e 1 Larix

Eupithecia tantillaria Boisduval, 1840 x x x x x eu 2Cc,2Cd 3 Pinaceae: Picea, Larix

Gymnoscelis rufifasciata (Haworth, 1809) x x x eu 2 2 Clematis

Chloroclystis v-ata (Haworth, 1809) x x x eu 1Bb,2B,2e 6 herbs

Chesias rufata (Fabricius, 1775) x st 2Ba 3 Fabaceae: Genista, Sarothamnus

Aplocera plagiata (l., 1758) x eu 2B,2C,2e,4ab 2 Hypericum

Aplocera praeformata (Hübner, 1826) x x x eu 2e,4a 2 Hypericum

Euchoeca nebulata (Scopoli, 1763) x x st 1dd,2a 4 Betulaceae: Alnus, rarely Betula

Asthena albulata (Hufnagel, 1767) x x x eu 2B,2ea 5 deciduous trees/shrubs P!

Asthena anseraria (H.-S., 1855) x st 2aa,2Ba,2ea 1 Cornus sanguinea

Hydrelia ��ammeolaria (Hufnagel, 1767) x x eu 1dd,2a,2B 5 deciduous trees/shrubs

Minoa murinata (Scopoli, 1763) x x eu 2e,4aa,4Bb,4C 1 Euphorbia cyparissias

Lobophora halterata (Hufnagel, 1767) x x eu 2a,2B,2ea,4e,4F 5 deciduous trees/shrubs: esp. PopulusPopulus

Trichopteryx carpinata (Borkhausen, 1794) x x eu 2,4e 5 deciduous trees/shrubs

Lomaspilis marginata (l., 1758) x x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Ligdia adustata (d. & Schiff., 1775) x x x eu 2,4e 2 Euonymus

Stegania trimaculata (Villers, 1789) x st 2a 2 Populus

Heliomata glarearia (d. & Schiff., 1775) x x x st 4aa 3 Fabaceae

Macaria notata (l., 1758) x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs




Macaria alternata (d. & Schiff., 1775) x x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs

Macaria liturata (Clerck, 1759) x x x x eu 2Ca,2Cc 5 conifers

Chiasmia clathrata (l., 1758) x x x eu 2e,4a,4Ce 4 Fabaceae

Petrophora chlorosata (Scopoli, 1763) x x eu 1dd,2,3Ba 1 Pteridium aquilinum

Anagoga pulveraria (l., 1758) x x x eu 2 5 deciduous trees/shrubs: esp. LoniceraLonicera

Plagodis dolabraria (l., 1758) x x x eu 2,4Ba,4e 5 deciduous trees/shrubs

Opisthograptis luteolata (l., 1758) x x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs

Pseudopanthera macularia (l., 1758) x x x eu 2,4aa,4ab,4e 6 herbs

Apeira syringaria (l., 1758) x x eu 2B,2C 5 deciduous trees/shrubs: esp. LoniceraLonicera

Selenia dentaria (Fabricius, 1775) x x x x eu 2,4e 5 deciduous trees/shrubs

Selenia lunularia (Hübner, 1788) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Selenia tetralunaria (Hufnagel, 1767) x x eu 2,4e,4F 5 deciduous trees/shrubs

Odontopera bidentata (Clerck, 1759) x x x x x eu 2,4e,4F 5 deciduous trees/shrubs

Crocallis tusciaria (Borkhausen, 1793) x st 2Ba 5 deciduous trees/shrubs

Crocallis elinguaria (l., 1758) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Colotois pennaria (l., 1761) x x x eu 2a,2B,2e,4F 5 deciduous trees/shrubs

Angerona prunaria (l., 1758) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Lycia hirtaria (Clerck, 1759) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Biston stratarius (Hufnagel, 1767) x x eu 1dd,2,4e 5 deciduous trees/shrubs

Biston betularia (l., 1758) x x x eu 1dd,2,4e,4F 5 deciduous trees/shrubs

Agriopis marginaria (Fabricius, 1777) x x eu 2,4e,4F 5 deciduous trees/shrubs

Erannis defoliaria (Clerck, 1759) x x x eu 2,4Ba,4e,4F 5 deciduous trees/shrubs

Nychiodes obscuraria (Villers, 1789) x x st 2Ba 1 Prunus spinosa

Menophra abruptaria (Thunberg, 1792) x x x st 2Ba,2ea 7 deciduous trees/shrubs, herbs

Peribatodes rhomboidarius (d. & Schiff., 1775)

x x x eu 2,4e,4F 7 deciduous trees/shrubs, herbs

Peribatodes secundarius (d. & Schiff., 1775) x x x x eu 2Ca,2Cb,2Cc 5 conifers

Cleora cinctaria (d. & Schiff., 1775) x eu 2 7 herbs, deciduous trees/shrubs

Alcis repandatus (l., 1758) x x x x x x eu 1d,2,4e,4F 7 deciduous trees/shrubs, conifers, herbs

Hypomecis punctinalis (Scopoli, 1763) x x x eu 2,4e,4F 5 deciduous trees/shrubs, conifers

Fagivorina arenaria (Hufnagel, 1767) x x st 2B 5 deciduous trees/shrubs

Ectropis crepuscularia (d. & Schiff., 1775) x x x eu 2 7 herbs, deciduous trees/shrubs, conifers

Paradarisa consonaria (Hübner, 1799) x x st 2B 5 deciduous trees/shrubs

Parectropis similaria (Hufnagel, 1767) x st 2Ba,?4F 5 deciduous trees/shrubs

Ematurga atomaria (l., 1758) x x eu 2,4a 6 herbs

Tephronia sepiaria (Hufnagel, 1767) x st 2Ba,2ea,4e 8 lichens

Cabera pusaria (l., 1758) x x x eu 1dd,2 5 deciduous trees/shrubs P!

Cabera exanthemata (Scopoli, 1763) x x eu 1dd,2 5 deciduous trees/shrubs

Lomographa bimaculata (Fabricius, 1775) eu 2a,2B,2C,2ea,4e 5 deciduous trees/shrubs

Lomographa temerata (d. & Schiff., 1775) x x x eu 1dd,2 5 deciduous trees/shrubs

Campaea margaritata (l., 1767) x x x x x eu 2B,4e,4F 5 deciduous trees/shrubs: esp. Fagus

Hylaea fasciaria (l., 1758) x x x eu 2C 5 conifers

Gnophos furvatus (d. & Schiff., 1775) x x x st 2Ba,3B,4aa 6 herbs P!




Gnophos obfuscatus (d. & Schiff., 1775) x eu 3C 6 herbs

Kemptrognophos ambiguata (dup., 1830) x x eu 2Ca,3a,3B 6 herbs

Rhopalogrophos glaucinaria (Hübner, 1799) x eu 3a 6 herbs

Euchrogrophos variegata (dup., 1830) x x x eu 3a,4aa 6 herbs

Elophos dilucidarius (d. & Schiff., 1775) x x eu 2C,3C 6 herbs

Elophos vittarius mendicarius (H.-S., 1852) x x eu 2C,3C 6 herbs, Vaccinium

Perconia strigillaria (Hübner, 1787) x x x eu 2 6 herbs: Ericaceae, Sarothamnus

notoDontiDae

Phalera bucephala (l., 1758) x x x eu 2,4e 5 deciduous trees/shrubs

Cerura vinula (l., 1758) x x eu 1dd,2a 4 Salicaceae

Stauropus fagi (l., 1758) x x x eu 2a,2B 5 deciduous trees/shrubs

Peridea anceps (Goeze, 1781) x x x eu 2aa,2Ba 5 deciduous trees/shrubs: esp. QuercusQuercus P!

Notodonta dromedarius (l., 1767) x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Notodonta ziczac (l., 1758) x eu 2,4e,4F 5 deciduous trees/shrubs

Ochrostigma velitaris (Hufnagel, 1766) x x st 2Ba 2 Quercus

Drymonia dodonaea (d. & Schiff., 1775) x x x eu 2B,4F 5 deciduous trees/shrubs: esp. QuercusQuercus P!

Drymonia ruficornis (Hufnagel, 1766) x eu 2B,4F 5 deciduous trees/shrubs: esp. QuercusQuercus

Pheosia tremula (Clerck, 1759) x x x eu 2,4e 5 deciduous trees/shrubs: Populus, Salix, Betula

Pterostoma palpinum (Clerck, 1759) x x eu 2,4e 5 deciduous trees/shrubs

Ptilodon capucina (l., 1758) x x x eu 2,4e 5 deciduous trees/shrubs

Ptilodontella cucullina (d. & Schiff., 1775) x x st 2aa,2Ba,2ea 2 Acer

Spatalia argentina (d. & Schiff., 1775) x x x st 2Bb,4F 2 Quercus

Clostera curtula (l., 1758) x st 1d,2a 3 Salicaceae: Populus, Salix

Clostera pigra (Hufnagel, 1766) x x st 1d,2a 3 Salicaceae: Populus, Salix

Thaumetopoea processionea (l., 1759) x x x st 2Ba,4F 2 Quercus P!

Traumatocampa pityocampa (d. & Schiff., 1775)

x x x st 2Cc 2 Pinus P!

lyMantriiDae

Calliteara pudibunda (l., 1758) x x x eu 2Bc,2e 5 deciduous trees/shrubs

Orgyia antiqua (l., 1758) x x x eu 2a,2B,2ea,4e,4F 5 deciduous trees/shrubs, rarely conifers

Lymantria dispar (l., 1758) x x x eu 2,4e 5 deciduous trees/shrubs, rarely conifers

Lymantria monacha (l., 1758) x x x eu 2a,2C 5 conifers, deciduous trees/shrubs

Arctornis l-nigrum (Müller, 1764) x x x st 2B 5 deciduous trees/shrubs

arctiiDae

Nudaria mundana (l., 1761) x st 3a 8 lichens, moss, decaying leaves, herbs

Atolmis rubricollis (l., 1758) x x x eu 2a,2B,2C 8 lichens

Lithosia quadra (l., 1758) x x x eu 1d,2,4e 8 lichens

Eilema deplana (esper, 1787) x x x x eu 2 8 lichens

Eilema lurideola (Zincken, 1817) x x x eu 2 8 lichens

Eilema complana (l., 1758) x x x x eu 2B,2e,4aa 8 lichens

Eilema caniola (Hübner, 1808) x x x eu 3B,4a,4e 8 lichens P!




Eilema palliatella (Scopoli, 1763) x st 3B,4aa 8 lichens

Eilema sororcula (Hufnagel, 1766) x x x eu 2 8 lichens

Setina irrorella (l., 1758) x st 3a,3B,3C,3d 8 lichens

Setema cereola (Hübner, 1803) x st 3C,3d 8 lichens

Coscinia cribraria (l., 1758) x x eu 2Ca,3B,4aa 6 herbs

Chelis maculosa (Gerning, 1780) x st 2Ba,4aa 6 herbs

Watsonarctia casta (esper, 1785) x x x eu 2Ba,4aa 6 herbs

Phragmatobia fuliginosa (l., 1758) x eu 1,2,3,4 6 herbs

Spilosoma luteum (Hufnagel, 1766) x x eu 1,2,3,4 6 herbs

Diaphora mendica (Clerck, 1759) x x x eu 2,4 6 herbs

Diacrisia sannio (l., 1758) x x x eu 2,4 6 herbs

Hyphoraia testudinaria (Geoffroy, 1785) x x x eu 3B,4aa 6 herbs

Arctia villica (l., 1758) x x x eu 2Ba,3B,4aa,4ab 6 herbs

Euplagia quadripunctaria (Poda, 1761) x x x eu 2 5 deciduous trees/shrubs

Amata phegea (l., 1758) x x x eu 2Ba,2B,4aa 6 herbs P!

noctuiDae

Orectis proboscidata (H.-S., 1851) x x x st 3B 2 Sedum

Trisateles emortualis (d. & Schiff., 1775) x x x st ?2Ba,2Bc,4F 9 decaying leaves (Quercus)

Eutelia adulatrix (Hübner, 1813) x x x st 2Ba 3 anacardiaceae: Rhus, Pistacia

Paracolax tristalis (Fabricius, 1794) x x x eu 1aa,2B 9 withered and decaying leaves P!

Herminia tarsicrinalis (Knoch, 1782) x x eu 1aa,2B 9 withered and decaying leaves

Herminia grisealis (Fabricius, 1775) x x eu 1aa,2B 9 withered and decaying leaves

Pechipogo strigilata (l., 1758) x x x eu 2a,2B 5 deciduous trees/shrubs

Zanclognatha lunalis (Scopoli, 1763) x x eu 2Ba,2e,4aa 9 withered and decaying leaves

Zanclognatha zelleralis (Wocke, 1850) x x eu 2Ba,2e,4aa 9 withered and decaying leaves

Zanclognatha tarsipennalis (Treitschke, 1835) x x eu 1aa,2B 9 withered and decaying leaves

Schrankia costaestrigalis (Stephens, 1834) x st 1d,2a 6 herbs

Schrankia taenialis (Hübner, 1809) x x st 2Ba 3 Poaceae

Hypena crassalis (Fabricius, 1787) x x x st 1da,2C 2 ericaceae: Vaccinium, Calluna

Phytometra viridaria (Clerck, 1759) x x x st 1dc,2,4a 2 Polygala

Scoliopteryx libatrix (l., 1758) x eu 2a,4e 4 Salicaceae

Catocala fraxini (l., 1758) x x st 2aa,2B,4e 2 Populus

Minucia lunaris (d. & Schiff., 1775) x x x st 2Ba 2 Quercus

Dysgonia algira (l., 1767) x x st 2Ba,4aa 2 Rubus

Lygephila viciae (Hübner, 1822) x x x eu 2eb,4aa,4ab,4C 3 Fabaceae

Lygephila craccae (d. & Schiff., 1775) x x eu 2eb,4aa,4ab,4C 3 Fabaceae

Catephia alchymista (d. & Schiff., 1775) x st 2Ba 2 Quercus

Euclidia glyphica (l., 1758) x x x eu 1d,2,4 3 Fabaceae

Meganola togatulalis (Hübner, 1796) x st 2Ba 2 Quercus

Nola confusalis (H.-S., 1847) x x x eu 2B 5 deciduous trees/shrubs

Nola cicatricalis (Treitschke, 1835) x st 2B 8 lichens

Nola cristatula (Hübner, 1793) x st 1d,4aa 3 labiatae: Mentha, Teucrium

Nola subchlamydula Staudinger, 1871 x x st 4aa 6 herbs

Nycteola revayana (Scopoli, 1772) x x x x st 2aa,2Ba,4F 2 Quercus




Pseudoips prasinana (l., 1758) x x x st 2Ba,2Bc 2 Quercus

Bena bicolorana (Fuessly, 1775) x x x st 2aa,2Bc 5 deciduous trees/shrubs: esp. Fagus, Quercus

Panthea coenobita (esper, 1785) x x x st 2Cb,2Cc 3 Pinaceae

Colocasia coryli (l., 1758) x x x eu 2aa,2B,2ea 5 deciduous trees/shrubs P!

Diloba coeruleocephala (l., 1758) x x x eu 2a,2B,4Ba 5 deciduous trees/shrubs

Moma alpium (osbeck, 1778) x x x st 2aa,2Ba,2Bc 5 deciduous trees/shrubs

Acronicta psi (l., 1758) x x x eu 2a,2B,2ea,4e 5 deciduous trees/shrubs

Acronicta megacephala (d. & Schiff., 1775) x x eu 2a,2B,4e 4 Salicaceae: Populus, Salix

Acronicta rumicis (l., 1758) x x eu 2,4 7 herbs, deciduous shrubs

Craniophora ligustri (d. & Schiff., 1775) x x x eu 2,4e 4 oleaceae: Fraxinus, Ligustrum

Cryphia algae (Fabricius, 1775) x x x st 2a,2B,4e 8 lichens P!

Cryphia domestica (Hufnagel, 1766) x x st 3a,4d 8 lichens

Cryphia muralis (Forster, 1771) x st 3a,4d 8 lichens

Emmelia trabealis (Scopoli, 1763) x st 2e,4a,4C 2 Convolvulus

Phyllophila obliterata (rambur, 1833) x x x st 4aa 2 Artemisia

Lithacodia pygarga (Hufnagel, 1766) x x x eu 2 4 Poaceae

Eublemma purpurina (d. & Schiff., 1775) x st 4aa 2 Cirsium

Eublemma polygramma (dup., 1836) x st 2Ba,?3B,?4aa ? unknown

Metachrostis dardouini (Boisduval, 1840) x x st 2Ba 2 Anthericum

Panchrysia v-argenteum (esper, 1798) x st 2eb,2ec 2 Thalictrum

Diachrysia chrysitis (l., 1758) x eu 1Bb,1d,2,4 6 herbs

Macdunnoughia confusa (Stephens, 1850) migrating species 6 herbs

Autographa gamma (l., 1758) x x x x migrating species 6 herbs

Autographa pulchrina (Haworth, 1809) x eu 2a,2eb,3d 6 herbs

Autographa jota (l., 1758) x eu 2C,2d,2e,3C 6 herbs

Autographa bractea (d. & Schiff., 1775) x eu 1Bb,2a,2eb,3d 6 herbs

Syngrapha interrogationis (l., 1758) x st 1da,2Cc,2Ce,3C 2 Vaccinium

Syngrapha ain (Hochenwarth, 1785) x st 2Cd 1 Larix

Chrysodeixis chalcites (esper, 1789) x migrating species 6 herbs

Abrostola triplasia (l., 1758) x eu 2a,2d,2e,4C,4e 1 Urtica

Abrostola asclepiadis (d. & Schiff., 1775) x x x eu 2a,2B,2C,2e 1 Cynanchum

Abrostola agnorista dufay, 1956 x x ? ? ? unknown

Cucullia lactucae (d. & Schiff., 1775) x eu 3B,4C 3 asteraceae: Prenanthes, Sonchus

Shargacucullia lychnitis (rambur, 1833) x st 3Ba,4aa 2 Verbascum

Amphipyra pyramidea (l., 1758) x x x eu 2a,2B,2e,4e,4F 5 deciduous trees/shrubs

Amphipyra berbera svenssoni Fletcher, 1968 x eu ? 5 deciduous trees/shrubs

Amphipyra tragopoginis (Clerck, 1759) x x x eu 2,4e 5 deciduous trees/shrubs

Amphpyra tetra (Fabricius, 1787) x st 2Ba,? 6 herbs

Heliothis peltigera (d. & Schiff., 1775) x migrating species 6 herbs

Pyrrhia umbra (Hufnagel, 1766) x x eu 1Bb,2,4 6 herbs

Elaphria venustula (Hübner, 1790) x x x st 2Ba,2ea,4aa 6 herbs, grass

Acosmetia caliginosa (Hübner, 1813) x st 4aa 1 Serratula tinctoria




Platypterigea kadenii (Freyer, 1836) x st 3a,4aa 6 herbs

Paradrina ��avirena (Guenée, 1852) x x x st 3a,4aa 6 herbs

Hoplodrina octogenaria (Goeze, 1781) x x x eu 2,4 6 herbs

Hoplodrina blanda (d. & Schiff., 1775) x eu 2B,2e,4 6 herbs

Hoplodrina respersa (d. & Schiff., 1775) x x x st 2Ba,2Ca,3B,4aa 6 herbs

Hoplodrina ambigua (d. & Schiff., 1775) x x eu 4aa,4Cc,4Cd,4Ce 6 herbs

Athetis furvula (Hübner, 1808) x x st 4aa 6 herbs

Rusina ferruginea (esper, 1785) x x eu 2 6 herbs

Polyphaenis sericata (esper, 1787) x x x st 2Ba 5 deciduous trees/shrubs: Cornus, Lonicera

Talpophila matura (Hufnagel, 1766) x eu 1d,2Ba,4aa,4ab 3 Poaceae

Euplexia lucipara (l., 1758) x eu 2 6 herbs

Phlogophora meticulosa (l., 1758) x migrating species 7 herbs, deciduous shrubs

Hyppa rectilinea (esper, 1788 x x eu 1da,2C 7 herbs, deciduous shrubs

Auchmis detersa (esper, 1791) x eu 2,4aa 1 Berberis

Chloantha hyperici (d. & Schiff., 1775) x st 4aa,4ab 2 Hypericum

Methorasa latreillei (dup., 1827) x x x st 2Ba,2Bc 3 ferns P!

Ipimorpha subtusa (d. & Schiff., 1775) x x x x st 1dd,2a,4e 2 Populus

Mesogona acetosellae (d. & Schiff., 1775) x x st 2Ba,2ea,4aa 7 herbs, deciduous trees/shrubs

Cosmia trapezina (l., 1758) x x x x x eu 2a,2B,2e,4e,4F 5 deciduous trees/shrubs

Xanthia aurago (d. & Schiff., 1775) x x x st 2B 7 deciduous trees/shrubs, herbs

Xanthia icteritia (Hufnagel, 1766) x eu 1dd,2 7 Salicaceae, later herbs

Xanthia fulvago (Clerck, 1759) x st 2Ba 5 deciduous trees/shrubs

Agrochola lota (Clerck, 1759) x x eu 1dd,2a,2Bb 5 deciduous trees/shrubs

Agrochola macilenta (Hübner, 1809) x x eu 2aa,2B,4e 7 deciduous trees/shrubs, later herbs

Agrochola nitida (d. & Schiff., 1775) x eu 2,4ac,4ad 6 herbs: esp. Primula

Agrochola helvola (l., 1758) x eu 2aa,2B,2ea,4e 5 deciduous trees/shrubs: esp. Quercus, herbs

Eupsilia transversa (Hufnagel, 1766) x x x eu 2a,2B,2e,4e 5 deciduous trees/shrubs

Jodia croceago (d. & Schiff., 1775) x x st 2Ba 2 Quercus

Conistra vaccinii (l., 1761) x x x eu 2,4e 7 deciduous trees/shrubs, herbs

Conistra rubiginosa (Scopoli, 1763) x eu 2a,2B,2e,4e 7 deciduous trees/shrubs, herbs

Conistra rubiginea (d. & Schiff., 1775) x x x eu 2,4e,4F 7 deciduous trees/shrubs, herbs

Conistra erytrocephala (d. & Schiff., 1775) x x x st 2Ba,2Bc,4F 5 deciduous trees/shrubs: esp. QuercusQuercus

Orbona fragariae (Vieweg, 1790) x st 2B 6 herbs

Episema glaucina (esper, 1789) x st 2Ba,4aa 3 liliaceae

Asteroscopus sphinx (Hufnagel, 1766) x eu 2aa,2Ba,2ea,4F 5 deciduous trees/shrubs

Brachylomia viminalis (Fabricius, 1777) x x x eu 2,4e 2 Salix

Lithophane hepatica (Clerck, 1759) x eu 2,4e 5 deciduous trees/shrubs

Lithophane ornitopus (Hufnagel, 1766) x x x eu 2,4e 5 deciduous trees/shrubs

Allophyes oxyacanthae (l., 1758) x x eu 2aa,2Bb,2Ca,2ea 3 wooden rosaceae

Griposia aprilina (l., 1758) x x st 2Ba,4F 2 Quercus

Trigonophora ��ammea (esper, 1785) x st ? 2 Ranunculus repens, R. ficaria

Polymixis gemmea (Treitschke, 1825) x eu 1d,2,4ac 4 Poaceae




Blepharita satura (d. & Schiff., 1775) x x eu 2 7 herbs, deciduous trees/shrubs

Mniotype adusta (esper, 1790) x x x eu 1d,2 7 herbs, deciduous trees/shrubs

Apamea monoglypha (Hufnagel, 1766) x x x x x x eu 2,4a,4e 4 Poaceae

Apamea sublustris (esper, 1788) x x x eu 1d,2a,2e,4ac 4 Poaceae

Apamea crenata (Hufnagel, 1766) x x x x eu 2,3d,4a 4 Poaceae

Apamea sordens (Hufnagel, 1766) x eu 1d,2,4a,4C 4 Poaceae

Apamea lateritia (Hufnagel, 1766) x eu 2,4aa 4 Poaceae

Apamea furva (d. & Schiff., 1775) x x eu 2C,3,4a 4 Poaceae

Apamea maillardi (Geyer,, 1834) x x eu 3B,3d 4 Poaceae

Apamea rubrirena (Treitschke, 1825) x eu 3B,3C,3d 4 Poaceae

Oligia strigilis (l., 1758) x x x x eu 2,4a 4 Poaceae

Oligia latruncula (d. & Schiff., 1775) x x x x x eu 2,4a 4 Poaceae

Oligia dubia (Heydemann, 1942) x st 4aa 4 Poaceae

Mesoligia furuncula (d. & Schiff., 1775) x x eu 4aa,4ab 4 Poaceae

Mesoligia literosa (Haworth, 1809) x eu 2Ca,4aa,4ab 4 Poaceae

Mesapamea didyma (esper, 1788) x x eu 2,4a 4 Poaceae

Luperina testacea (d. & Schiff., 1775) x eu 4aa,4ab,4Cd 4 Poaceae

Charanyca trigrammica (Hufnagel, 1766) x eu 1d,2e,4a,4e 7 herbs, grass, deciduous shrubs

Lacanobia w-latinum (Hufnagel, 1766) x x x x eu 2,4 7 herbs, deciduous trees/shrubs

Lacanobia thalassina (Hufnagel, 1766) x x x eu 2,4e 7 herbs, deciduous trees/shrubs

Lacanobia contigua (d. & Schiff., 1775) x x eu 2 7 herbs, deciduous trees/shrubs

Lacanobia suasa (d. & Schiff., 1775) x eu 2,4 6 herbs

Hada plebeja (l., 1761) x x x eu 2,3,4 6 herbs

Lasionycta calberlai (Staudinger, 1883) x st 2Ba,2ea 2 Clematis

Hadena luteago (d. & Schiff., 1775) x x st 3B,4aa 2 Silene

Hadena compta (d. & Schiff., 1775) x x x eu 2eb,3B,4aa 3 Caryophyllaceae

Hadena confusa (Hufnagel, 1766) x eu 2eb,3B,4aa 3 Caryophyllaceae

Hadena filograna (esper, 1788) x x st 3B,4aa 3 Caryophyllaceae

Hadena albimacula (Borkhausen, 1792) x eu 2Ba,3B,4aa 3 Caryophyllaceae

Hadena magnolii (Boisduval, 1829) x x x st 3B,4aa 3 Caryophyllaceae

Hadena perplexa (d. & Schiff., 1775) x x eu 2,3B,3C,4a 3 Caryophyllaceae

Sideridis lampra (Schawerda, 1913) x x x st 2Ba,4aa 6 grass, herbs: apiaceae

Heliophobus reticulata (Goeze, 1781) x x x eu 3B,4aa,4ab,4Cd 3 Caryophyllaceae

Caramica pisi (l., 1758) x x x eu 2,4 6 herbs: esp. Gartenpflanzen

Mamestra brassicae (l., 1758) x eu 2,4 6 herbs

Papestra biren (Goeze, 1781) x x eu 1da,2C,3C 6 herbs

Polia bombycina (Hufnagel, 1766) x eu 2,4e 7 herbs, deciduous trees/shrubs

Polia nebulosa (Hufnagel, 1766) x eu 2 7 herbs, deciduous trees/shrubs

Leucania comma (l., 1761) x x x eu 4 6 grass, herbs l!

Mythimna conigera (d. & Schiff., 1775) x x eu 2,4 6 grass, herbs

Mythimna ferrago (Fabricius, 1787) x x x eu 2,4 6 grass

Mythimna albipuncta (d. & Schiff., 1775) x eu 2,4 6 grass

Mythimna vitellina (Hübner, 1808) x migrating species 6 grass




Mythimna impura (Hübner, 1808) x x st 1af,1db,1dc 3 Poaceae: Molinia, Phragmites

Mythimna l-album (l., 1767) x x x eu 1,2,4 6 grass

Mythimna andereggii (Boisduval, 1840) x x eu 3 6 grass

Mythimna scirpi (dup., 1836) x x x st 2Ba,4aa 6 grass P!

Orthosia incerta (Hufnagel, 1766) x eu 2,4e,4F 5 deciduous trees/shrubs

Orthosia gothica (l., 1758) x x x x eu 2,4e,4F 7 deciduous trees/shrubs, herbs

Orthosia cruda (d. & Schiff., 1775) x x eu 2a,2B,2ea,4F 7 deciduous trees/shrubs, rarely herbs

Orthosia cerasi (Fabricius, 1775) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Orthosia munda (d. & Schiff., 1775) x x x eu 2,4e,4F 5 deciduous trees/shrubs

Panolis ��ammea (d. & Schiff., 1775) x x st 2Ca 5 Pinaceae: Pinus, rarely Picea

Egira conspicillaris (l., 1758) x x x eu 2B,4aa 7 herbs, deciduous trees/shrubs

Cerapteryx graminis (l., 1758) x x x eu 3d,4a 6 grass l!

Tholera cespitis (d. & Schiff., 1775) x eu 2,4a 6 grass

Neuronia decimalis (Poda, 1761) x eu 4a 6 grass

Pachetra saggitigera (Hufnagel, 1766) x x x eu 2,4e,4aa,4ab 6 herbs, grass

Eriopygodes imbecilla (Fabricius, 1794) x x st 3d,4a 6 herbs, grass

Lasionhada proxima (Hübner, 1809) x x eu 2,3,4 6 herbs

Axylia putris (l., 1761) x x x eu 2,4 6 herbs, grass

Ochropleura plecta (l., 1761) x eu 4a,4e 6 herbs, grass

Diarsia mendica (Fabricius, 1775) x x eu 1dd,2B,2C,2d,3C 6 herbs

Diarsia brunnea (d. & Schiff., 1775) x eu 2 6 herbs, grass

Diarsia rubi (Vieweg, 1790) x x st 1Bb,1d,2a 6 herbs, grass

Noctua pronuba (l., 1758) x x x eu 1,2,3,4 6 herbs, grass

Noctua fimbriata (Schreber, 1759) x x x eu 2,4a,4e 7 herbs, deciduous trees/shrubs

Noctua tirrenica Biebinger, Speidel & Hanik, 1983

x x ? ? 6 ?herbs

Noctua orbona (Hufnagel, 1766) x st 2aa,2B 6 herbs, grass

Noctua comes Hübner, 1813 x x x eu 2,4e 6 herbs

Noctua janthe (Borkhausen, 1792) x x eu 1dd,2a,2e,4e 6 herbs

Epilecta linogrisea (d. & Schiff., 1775) x x x st 2Ca,2eb 6 herbs, grass

Lycophotia porphyrea (d. & Schiff., 1775) x x x eu 1da,2Ca 1 Calluna

Chersotis ocellina (d. & Schiff., 1775) x st 3C,3d 6 herbs

Chersotis multangula (Hübner, 1803) x x eu 3B,4aa 2 Galium

Chersotis cuprea (d. & Schiff., 1775) x x eu 2d,3d,4ac 6 herbs

Margasotis margaritacea (Villers, 1789) x x st 3B 3 rubiaceae: Asperula, Galium

Rhyacia simulans (Hufnagel, 1766) x st 2Ca,4aa,4Cd 6 herbs, grass

Epipsilia latens (Hübner, 1809) x st 3B,4aa 6 herbs, grass

Eurois occulta (l., 1758) x st 1da,2C,3C 6 herbs, Vaccinium etc.

Spaelotis senna contorta rebel & Zerny, 1932 x st 4aa,4Cd 2 ?Artemisia

Graphiphora augur (Fabricius, 1775) x x eu 1d,2,3C,4 7 herbs, deciduous trees/shrubs

Eugnorisma depuncta (l., 1761) x eu 2eb 6 herbs

Xestia rhaetica (Staudinger, 1871) x st 1da,3C 1 Vaccinium myrtillus

Xestia viridescens (Turati, 1919) x x st 1da,3C 6 herbs, grass




Xestia c-nigrum (l., 1758) x x eu 1,2,3,4 6 herbs

Xestia ditrapezium (d. & Schiff., 1775) x x eu 2a,2B,2eb,4e 6 herbs

Xestia triangulum (Hufnagel, 1766) x eu 2a,2B,2eb,4e 6 herbs

Xestia baja (d. & Schiff., 1775) x eu 2a,2B,2eb,4e 6 herbs

Xestia rhomboidea (esper, 1790) x x x eu 2a,2B,2eb,4e 6 herbs, grass

Xestia castanea neglecta (Hübner, 1803) x x x st 1da,2Ca,4aa 6 herbs, esp. Calluna

Xestia xanthographa (d. & Schiff., 1775) x x eu 1d,2,4e 6 herbs, grass

Eugraphe sigma (d. & Schiff., 1775) x x x eu 2,4e 7 herbs, deciduous trees/shrubs

Cerastis rubricosa (d. & Schiff., 1775) x eu 1d,2a,2e 6 herbs

Anaplectoides prasina (d. & Schiff., 1775) x x eu 1dd,2 6 herbs

Peridroma saucia (Hübner, 1808) x migrating species 6 herbs

Euxoa recussa (Hübner, 1817) x x eu 3d 4 Poaceae

Euxoa nigricans (l., 1761) x eu 2eb,4a,4C 6 herbs, grass

Euxoa obelisca (d. & Schiff., 1775) x st 4aa 6 herbs, grass

Crassagrotis crassa (Hübner, 1803) st 4aa,4C 6 herbs, grass

Agrotis ipsilon (Hufnagel, 1766) x x x migrating species 6 herbs, grass

Agrotis trux Hübner, 1824 x st 2Ba,4aa 6 herbs, grass

Agrotis exclamationis (l., 1758) x x x x x eu 4a,4B,4C,4e 6 herbs, grass

Agrotis clavis (Hufnagel, 1766) x x x eu 3d,4aa,4ab 6 herbs, grass

Agrotis segetum (d. & Schiff., 1775) x x x eu 4a,4Bc,4C,4e 6 herbs, grass

Agrotis simplonia (Geyer, 1832) x x eu 3a,3B 6 herbs, grass

Agrotis cinerea (d. & Schiff., 1775) x x x x eu 4aa,4ab,4ad 6 herbs, grass

Documents

Work Package 5 - MUSE · Work Package 5 Characterization of zoocoenoses ... Parole chiave: Insetti, foreste alpine, biomonitoraggio, bioindicazione, Trentino, Italia