TAXONOMIC RECORD OF THE OLIGOCENE BENTHIC … · 2018. 12. 14. · megasequence. According to Filipescu (2001) and Krézsek & Bally (2006), the deposition of the early Oligocene bituminous

ACTA PALAEONTOLOGICA ROMANIAE (2015) V. 11 (1), P. 25-62

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1Babeș-Bolyai University, Department of Geology, Str. Kogălniceanu 1, 400084, Cluj Napoca, Romania, [email protected], 25 [email protected]

TAXONOMIC RECORD OF THE OLIGOCENE BENTHIC FORAMINIFERA FROM

THE VIMA FORMATION (TRANSYLVANIAN BASIN, ROMANIA)

Szabolcs-Flavius Székely1* & Sorin Filipescu1

Received: 11 February 2015 / Accepted: 04 August 2015 / Published online: 26 August 2015

Abstract Samples collected from the Oligocene Vima Formation in the Transylvanian Basin revealed abundant and

diverse foraminifera assemblages. Among benthics, 139 taxa belonging to 76 genera, 43 families, 26 superfamilies,

11 suborders and 8 orders were identified. A large number of agglutinated and calcareous benthic foraminifera

species are reported and illustrated for the first time from the Oligocene of the Transylvanian Basin

Keywords: benthic foraminifera, taxonomy, Oligocene, Transylvanian Basin

INTRODUCTION

Regional geotectonic evolution during the Oligocene

produced major paleogeographic changes in Central and

Eastern Europe, mainly by the separation of a new marine

realm, the Paratethys (Laskarev, 1924; Rögl, 1998).

These new conditions influenced the development of

particular Oligocene sedimentary paleoenvironments in

the Transylvanian Basin. Krézsek & Bally (2006)

included the sedimentary formations in the Paleogene

megasequence. According to Filipescu (2001) and

Krézsek & Bally (2006), the deposition of the early

Oligocene bituminous shales of the Ileanda Formation

(Hofmann, 1879) continued with outer shelf to slope

Vima Formation (Lăzărescu, 1957, emend. Rusu, 1969)

and the equivalent inner shelf Buzaș Formation

(Dumitrescu, 1957).

Earlier studies focused on the taxonomy and

biostratigraphy of the foraminiferal assemblages in the

Vima Formation. Among first investigations from this

area belong to Majzon (1950a, 1950b). Diverse

planktonic and benthic foraminifera were documented by

Popescu (1971, 1972, 1975), Popescu & Iva (1971), and

Popescu & Brotea (1989). Our recent investigations in the

Vima Formation also revealed abundant and diverse

Oligocene planktonic and benthic foraminifera

communities. The aim of this contribution is to improve

the knowledge of the Oligocene benthic foraminifera

assemblages and to provide proper illustration of selected

species.

MATERIAL AND METHODS

One hundred and eleven samples were collected from

three outcrops (coordinates N47.41477, E23.82699;

N47.41356, E 23.82637; N47.41195, E 23.82692 - Fig. 1)

from one of the most representative sections of the Vima

Formation, located in Fântânele village (Maramureș

county). The studied deposits consist of mudstones and

sandstones (Fig. 2). The samples were processed

following standard micropaleontological methods.

Foraminifera were picked from the 63μm fraction.

Representative species were photographed using scanning

electron microscopes at the University of Fribourg and

Babeş-Bolyai University in Cluj-Napoca.

BENTHIC FORAMINIFERAL ASSEMBLAGES

Several types of assemblages were identified based on

quantitative analysis of the benthic community. These

were separated mainly according to the inferred

paleodepth and bottom water oxygenation, as follows:

1. Assemblages adapted to relatively well-oxygenated

environments affected by intervals of low-oxygen caused

by seasonal / interanual oscillations of oxygen content at

the sea floor, contain diverse benthic foraminiferal

assemblages, well represented by both epifaunal and

infaunal forms. Calcareous benthic species, such as

Cibicidoides acutus (Fig. 15.4), C. ungerianus filicosta

(Fig. 15.3), and C. oligocenicus (Fig. 15.6) are present in

moderate numbers and are known as intolerant to low-

oxygen conditions (Barmawidjaja et al., 1992; Rögl &

Spezzaferri, 2003). Additionaly, the co-occurrence of the

species Heterolepa praecincta (Figs. 17.1-17.2) and H.

eocaena (Figs. 17.3-17.4) suggest episodes with well-

oxygenated bottom waters (Murray, 2006). However,

intervals with fluctuating phytodetritus supply to the

seafloor (Gooday, 1993) could be suggested by the

presence of Epistominella molassica (Fig. 15.7),

Alabamina tangentialis (Figs. 16.8-16.9), and possibly

?Gyroidina constans (Fig. 15.2), while periodic high

primary productivity has been inferred based on the high

proportions of Uvigerina popescui (Rögl & Spezzaferri,

2003; Figs. 13.2-13.5). The above mentioned factors

triggered intervals with decreased oxygen content on the

substrate and the development of low oxygen tolerant

infaunal forms, such as Bolivina dilatata dilatata (Figs.

11.5-11.6), B. dilatata hyalina (Fig. 11.4), B. fastigia

(Fig. 11.2), and Bulimina schischkinskayae (Fig. 12.9).

Less representative benthic forms for these assemblages

are: Gyroidinoides octocameratus (Fig. 17.5),

Hansenisca soldanii (Figs. 17.7-17.8), Globocassidulina

subglobosa (Fig. 11.14), Spirorutilus carinatus (Fig. 5.1),

Stilostomella adolphina (Figs. 14.12-14.14), S. emaciata

(Figs. 14.15, 14.17, 14.18 & 14.21), and S. bradyi (Fig.

14.20).

mailto:[email protected]

Szabolcs-Flavius Székely & Sorin Filipescu

26

2. Assemblages adapted to low oxygen bottom water

conditions - are characterized by low species diversity.

The dominant species are Bulimina schischkinskayae,

Caucasina oligocaenica (Figs. 12.3-12.5), Bolivina

dilatata dilatata, B. dilatata hyalina, Fursenkoina

mustoni (Figs. 14.2-14.3) and F. halkyardi (Figs. 14.5-

14.10). Low oxygen bottom water conditions for this kind

of assemblages was suggested by Spezzaferri & Ćorić

(2001), Rögl & Spezzaferri (2003) and Murray (2006).

These co-occur with other less represented benthic

foraminifera such as the epifaunal-shallow infaunal

Haplophragmoides carinatus (Kaminski, Gradstein and

collaborators, 2005 - Figs. 4.5-4.9), Epistominella

molassica, Lenticulina inornata (Fig. 8.10) and

Lenticulina arcuatostriata (Fig. 8.9) indicating episodes

of phytodetritus supply and intervals with increased

dissolved oxygen content at the sea floor.

3. “Flysch-type” assemblages (Gradstein & Berggren,

1981) - populated the deeper upper slope environments.

These are completely dominated by agglutinated

foraminifera (mainly Haplophragmoides carinatus) or

mixed agglutinated (dominant tubular) and calcareous

benthic species. Agglutinated foraminifera are

represented by tubular forms (Bathysiphon taurinensis -

Fig. 3.1; Hyperammina elongata - Fig. 3.7; H. rugosa -

Fig. 3.6; Nothia excelsa - Figs. 3.4-3.5; Rhabdammina

spp.), uniserial Reophax (R. duplex - Fig. 3.12; R.

globosus - Fig. 3.14; R. subfusiformis - Fig. 3.15; R.

excentricus - Fig. 3.16; R. pilulifer - Fig. 4.2; R.

scorpiurus - Fig. 4.1), coiled Recurvoides spp., and

planispiral Reticulophragmium (R. amplectens - Fig. 6.1;

R. rotundidorsatum - Fig. 6.2; R. acutidorsatum - Fig.

6.3). Calcareous benthic foraminifera within the mixed

communities (Stilostomella adolphina, S. emaciata, S.

bradyi, Uvigerina gracilis - Fig. 13.8; Bolivina spp.,

Cibicidoides spp., Lenticulina inornata, L. limbosa - Fig.

8.7; Epistominella molassica and ?Gyroidina constans)

are less abundant in these assemblages.

BIOSTRATIGRAPHY

Although most of the identified planktonic species from

the Fântânele section have a large stratigraphic range, the

presence in the first outcrop of Chiloguembelina cubensis

associated with Paragloborotalia opima suggest the

Oligocene Zone O4 (late Rupelian, G. angulisuturalis /

C. cubensis Concurrent-range Zone following the

biozonation of Wade et al., 2011). The occurrence of

Paragloborotalia opima and Globigerina ciperoensis in

the third outcrop suggests a Late Oligocene age (early

Chattian, Zone O5 – P. opima Highest-occurrence Zone

according to Wade et al., 2011) for the younger

sediments. Based on these observations, the Rupelian -

Chattian boundary in the Fântânele section could be

placed within the second outcrop (Fig. 2).

SYSTEMATIC TAXONOMY

Identification of benthic foraminifera was mainly based

on the works of Bhatia (1955), Batjes (1958), Larsen &

Dinesen (1959), Kümmerle (1963), Grossheide & Trunkó

(1965), Popescu & Iva (1971), Popescu (1975), Sztrákos

Fig. 1 Simplified geological map of the studied area and location of sampled outcrops (modified after Beldean &

Filipescu, 2011; detailed map redrawn after Giușcă & Rădulescu, 1967).

Taxonomic record of the Oligocene benthic foraminifera from the Vima Formation (Transylvanian Basin, Romania)

27

(1979), Reiser (1987), Cicha et al. (1998), and Horváth

(2003). Classification of calcareous benthic foraminifera

is based on the suprageneric classification of Loeblich &

Tappan (1988), while the classification of Kaminski

(2004) was used for the agglutinated forms. Selected

illustrations for the plates represent certainly determined

foraminifera species. In some cases, specimens lacking

morphological features necessary to associate it with a

specific species are presented in the classification as

“sp.”. Synonymy list containing various foraminifera

species was compiled based on important works focused

on taxonomy (e.g. Rögl & Hansen, 1984; Papp &

Schmid, 1985; Horváth, 2003; Kaminski, Gradstein &

collaborators, 2005).

Class FORAMINIFEREA d’Orbigny, 1826

Subclass TEXTULARIIA Mikhalevich, 1980

Order ASTRORHIZIDA Lankester, 1885

Suborder ASTRORHIZINA Lankester, 1885

Family RHABDAMMINIDAE Brady, 1884

Subfamily BATHYSIPHONINAE Avnimelech, 1952

Fig. 2 Simplified lithology of the Fântânele section and position of the collected samples.


28

Genus Bathysiphon Sars, 1872

Bathysiphon taurinensis Sacco, 1893

Fig. 3.1a-b

Bathysiphon taurinensis Sacco, 1893, p. 168, pl. 11, fig. 2

(fide Ellis & Messina).

Bathysiphon taurinense Sacco - Popescu, 1975, p. 32, pl.

1, fig. 2.

Stratigraphical distribution in samples: Rupelian-

Chattian.

Genus Nothia Pflaumann, 1964

Nothia excelsa (Grzybowski, 1898) emend. Geroch &

Kaminski, 1992

Figs. 3.4-3.5

Dendrophrya excelsa Grzybowski, 1898, p. 272, pl. 10,

figs. 2-4 (fide Ellis & Messina).

Rhabdammina excelsa (Grzybowski) - Charnock &

Jones, 1990, pp. 151-153, pl. 1, figs. 26-27, pl. 13, fig.

24.

Nothia excelsa (Grzybowski) - Geroch & Kaminski,

1992, p. 255, pl. 1, figs. 1-4, pl. 2, figs. 1-11.


Chattian..

Nothia sp.

Figs. 3.2a-b, 3.3

Stratigraphical distribution in samples: Fig. 3.2 –

Chattian, Fig. 3.3 – Rupelian.

Suborder HIPPOCREPININA Saidova, 1981

Superfamily HIPPOCREPINACEA Rhumbler, 1895

Family HYPERAMMINIDAE Eimer & Fickert, 1899

Subfamily HYPERAMMININAE Eimer & Fickert,

1899

Genus Hyperammina Brady, 1878

Hyperammina elongata Brady, 1878

Fig. 3.7

Hyperammina elongata Brady, 1878, p. 433, pl. 1, figs.

16, 18-19.


Chattian.

Hyperammina rugosa Verdenius & van Hinte, 1983

Fig. 3.6a-b

Hyperammina rugosa Verdenius & van Hinte, 1983, p.

187, pl. 1, figs. 12-14.


Chattian.

Hyperammina sp.

Fig. 3.8

Stratigraphical distribution in samples: Rupelian.

Suborder AMMODISCINA Mikhalevich, 1980

Superfamily AMMODISCACEA Reuss, 1862

Family AMMODISCIDAE Reuss, 1862

Subfamily AMMODISCINAE Reuss, 1862

Genus Ammodiscus Reuss, 1862

Ammodiscus pennyi Cushman & Jarvis, 1928

Fig. 3.11

Ammodiscus pennyi Cushman & Jarvis, 1928, p. 87, pl.

12, figs. 4-5.


Subfamily USBEKISTANIINAE Vyalov, 1977

Genus Glomospira Rzehak, 1885

Glomospira charoides (Jones & Parker, 1860)

Fig. 3.9

Trochammina squamata Jones & Parker var. charoides

Jones & Parker, 1860, p. 304 (fide Ellis & Messina).

Trochammina charoides Jones & Parker - Carpenter et

al., 1862, p. 141, pl. 2, fig. 3 (fide Ellis & Messina).

Ammodiscus charoides (Jones & Parker) - Brady, 1884,

p. 334, pl. 38, figs. 10-16.

Glomospira charoides (Jones & Parker) var. corona

Cushman & Jarvis, 1928, p. 89, pl. 12, figs. 9-11.

Glomospira saturniformis Majzon, 1943, p. 231, pl. 5,

fig. 9.

Glomospira charoides (Jones & Parker) var. extendens

Emiliani, 1954, p. 133, pl. 13, fig. 14a-c.

Glomospira favilla Emiliani, 1954, p. 133, pl. 13, fig.

15a-c.

Glomospira charoides (Jones & Parker) var. leroyi Said

& Kenawy, 1956, p. 120, pl. 1, fig. 6.

Glomospira subcharoides Khalilov, 1959, p. 25, pl. 1,

fig. 1a-c.

Glomospira subcharoides var. sphaera Khalilov, 1959, p.

27, pl. 1, fig. 1d.

Glomospira charoides (Jones & Parker) - Barker, 1960,

pl. 38, figs. 10-16.

Glomospira charoides (Jones & Parker) - Geroch, 1960,

pp. 46-47, pl. 4, figs. 1,2,5.

Glomospira charoides (Jones & Parker) var. minima

Subbotina, 1960, p. 179, pl. 1, figs. 11a-12b.

Glomospira pileolus Subbotina, 1960, p. 179, pl. 1, figs.

13a-14c.

Glomospira charoides corona Cushman and Jarvis -

Hillebrandt, 1962, p. 25, pl. 2, fig. 24a-c.

Glomospira iranensis Kavary - Kavary & Frizzel, 1963,

p. 10, pl. 1, figs. 2-7.

Glomospira subarctica Chamney, 1969, p. 16, pl. 2, figs.

7-9.

Glomospira subarctica saturna Chamney, 1969, p. 17, pl.

2, figs. 10-12.

Glomospira praecharoides Soliman, 1972, p. 36, pl. 1,

figs. 1-2, pl. 2, fig. 1.

Usbekistania charoides antarctica Saidova, 1975, p. 71,

pl. 96, fig. 3.

Usbekistania charoides profunda Saidova, 1975, p. 71,

pl. 18, fig. 5.

Glomospira fulgida Maslun, 1987, p. 69, pl. 2, fig. 3.

Glomospira charoides charoides (Jones and Parker) -

Weidich, 1990, p. 83, pl. 34, figs. 7, 11.

Usbekistania charoides (Jones & Parker) - Jones, 1994,

p. 43, pl. 38, figs. 10-16.

Repmanina charoides (Jones & Parker) - Cicha et al.,

1998, p. 123, pl. 2, fig. 3.

Glomospira charoides (Jones & Parker) - Kaminski,

Gradstein and collaborators, 2005, p. 173, pl. 22, figs. 1-

16.


29

Fig. 3 All scales represent 100 μm. 1. Bathysiphon taurinensis Sacco. Same specimen, a. apertural view, b. lateral view, same

scale, sample F8, Rupelian. 2. Nothia sp. Same specimen, a. apertural view, b. lateral view, same scale, sample FB20, Chattian. 3.

Nothia sp. Sample F19, Rupelian. 4-5. Nothia excelsa (Grzybowski). 4. sample F4, Rupelian; 5. sample F22, Rupelian. 6.

Hyperammina rugosa Verdenius & Van Hinte. Same specimen, a. apertural view, b. lateral view, same scale, sample FB62,

Chattian. 7. Hyperammina elongata Brady. Sample F7, Rupelian. 8. Hyperammina sp. Sample F14, Rupelian. 9. Glomospira

charoides (Jones & Parker). Sample F32, Rupelian/Chattian. 10. “Glomospira” serpens (Grzybowski). Sample F20, Rupelian. 11.

Ammodiscus pennyi Cushman & Jarvis. Sample F15, Rupelian. 12. Reophax duplex Grzybowski. Sample F22, Rupelian. 13.

Hormosina trinitatensis Cushman & Renz. Sample F31, Rupelian/Chattian. 14. Reophax globosus Sliter. Sample FB62, Chattian.

15. Reophax subfusiformis Earland. Sample FB23, Chattian. 16. Reophax excentricus Cushman. Sample F1, Rupelian. 17.

Pseudonodosinella elongata (Grzybowski). Sample F13, Rupelian.


30


Chattian.

„Glomospira” serpens (Grzybowski, 1898)

Fig. 3.10

Ammodiscus serpens Grzybowski, 1898, p. 285, pl. 10,

fig. 31.

Glomospira serpens (Grzybowski) - Geroch, 1960, p. 47,

pl. 4, fig. 13.

Glomospira rostokiensis Mjatliuk, 1970, pp. 68-69, pl.

11, figs. 16a-18b.

?Glomospira tracta Saidova, 1975, p. 70, pl. 96, fig. 2.

Glomospirella serpens (Grzybowski) - Samuel, 1977, p.

30, pl. 4, fig. 2, pl. 22, fig. 4.

“Glomospira” serpens (Grzybowski) - Kaminski,

Gradstein and collaborators, 2005, p. 189, pl. 27, figs. 1a-

6b.


Order LITUOLIDA Lankester, 1885

Suborder HORMOSININA Mikhalevich, 1980

Superfamily HORMOSINELLACEA Rauser &

Reitlinger, 1986

Family HORMOSINELLIDAE Rauser & Reitlinger,

1986

Genus Subreophax Saidova, 1975

Subreophax scalaris (Grzybowski, 1896)

Fig. 4.3

Reophax guttifera Brady var. scalaria Grzybowski, 1896,

p. 277, pl. 8, fig. 26 (fide Ellis & Messina).

Reophax guttifera scalaris Grzybowski - Jurkiewicz,

1967, pp. 47-48, pl. 1, fig. 25.

Reophax scalaria Grzybowski - Samuel, 1977, pp. 35-36,

pl. 3, fig. 6a-b, pl. 19, fig. 4.

Reophax scalaris Grzybowski - Hemleben and Tröster,

1984, p. 521, pl. 2, figs. 10-11.

Subreophax scalaria (Grzybowski) - Kaminski et al.,

1988, p. 187, pl. 2, figs. 16-17.

Subreophax scalaris (Grzybowski) - Kuhnt, 1990, pl. 3,

figs. 4-5.


Superfamily HORMOSINACEA Haeckel, 1894

Family REOPHACIDAE Cushman, 1927

Genus Reophax de Montfort, 1808

Reophax duplex Grzybowski, 1896

Fig. 3.12

Reophax duplex Grzybowski, 1896, p. 276, pl. 8, figs. 23-

24 (fide Ellis & Messina).

Reophax paraduplex Mjatliuk, 1970, p. 65, pl. 9, figs. 1a-

3, pl. 10, fig. 2.


Chattian.

Reophax excentricus Cushman, 1910

Fig. 3.16

Reophax excentricus Cushman, 1910, p. 92, textfig. 134



Chattian.

Reophax globosus Sliter, 1968

Fig. 3.14

Reophax globosus Sliter, 1968, p. 43, pl. 1, fig. 12 (fide

Ellis & Messina).


Chattian.

Reophax pilulifer Brady, 1884

Fig. 4.2

Reophax pilulifer Brady, 1884, p. 292, pl. 30, figs. 18-20.


Chattian.

Reophax scorpiurus de Montfort, 1808

Fig. 4.1

Reophax scorpiurus de Montfort, 1808, p. 331, fig. p. 330



Chattian.

Reophax subfusiformis Earland, 1933 emend. Höglund,

1947

Fig. 3.15

Reophax subfusiformis Earland, 1933, p. 74, pl. 2, figs.

16-19 (fide Ellis & Messina).

Reophax subfusiformis Earland emend. Höglund, 1947, p.

82, pl. 9, figs. 1-4, pl. 26, figs. 1-36, pl. 27, figs. 1-19,

textfigs. 43-50.


Chattian.

Family HORMOSINIDAE Haeckel, 1894

Subfamily HORMOSININAE Haeckel, 1894

Genus Hormosina Brady, 1879

Hormosina trinitatensis Cushman and Renz, 1946

Fig. 3.13

Hormosina globulifera Brady var. trinitatensis Cushman

and Renz, 1946, p. 14, pl. l, figs. 15-19.

Reophax trinitatensis (Cushman & Renz) - Hanzliková,

1972, p. 37, pl. 3, fig. 10.

Hormosina trinitatensis Cushman & Renz – Kaminski et

al., 1988, pl. 3, fig. 1.


Chattian.

Genus Pseudonodosinella Saidova, 1970

Pseudonodosinella elongata (Grzybowski, 1898)

Fig. 3.17

Reophax elongata Grzybowski, 1898, p. 279, pl. 10, figs.

19-20.

?Reophax pyrifera Rhumbler, 1905, p. 103, textfig. 4.

?Hormosina mortenseni Hofker, 1972, p. 62, pl. 18, figs.

6-12.

Reophax elongatus Grzybowski - Morgiel and

Olszewska, 1981, p. 9, pl. 2, fig. 11.

Pseudonodosinella elongata (Grzybowski) - Kaminski,


9.


Suborder LITUOLINA Lankester, 1885

Superfamily LITUOLACEA de Blainville, 1827


31

Fig. 4 All scales represent 100 μm. 1. Reophax scorpiurus Montfort. Sample F38, Chattian. 2. Reophax pilulifer Brady. Sample

FB23, Chattian. 3. Subreophax scalaris (Grzybowski). Sample F25, Rupelian. 4. Ammobaculites agglutinans (d’Orbigny).

Sample FB62, Chattian. 5-9. Haplophragmoides carinatus Cushman & Renz. 5. sample F14, Rupelian; 6a-b. same specimen, a.

lateral view, b. peripherial view, same scale, sample FB61, Chattian; 7. sample F19, Rupelian; 8. sample F35, Rupelian/Chattian;

9. sample F2, Rupelian. 10-11. Haplophragmoides suborbicularis (Grzybowski). 10a-b. same specimen, a. lateral view, b.

peripherial view, same scale, sample F1, Rupelian; 11. sample F26, Rupelian. 12. Sculptobaculites barri Beckmann. Sample F24,

Rupelian. 13. Ammosphaeroidina pseudopauciloculata (Mjatliuk). Sample F28, Rupelian. 14. Recurvoides sp. Same specimen, a.

lateral view, b. lateral view, c. peripherial view, same scale, Sample FB20, Chattian.


32

Family HAPLOPHRAGMOIDIDAE Maync, 1952

Genus Haplophragmoides Cushman, 1910

Haplophragmoides carinatus Cushman & Renz, 1941

Figs. 4.5, 4.6a-b, 4.7-4.9

Haplophragmoides carinatus Cushman & Renz, 1941, pl.

1, fig. 1.


Chattian.

Haplophragmoides suborbicularis (Grzybowski, 1896)

emend. Kaminski, Gradstein and collaborators, 2005

Figs. 4.10a-b, 4.11

Cyclammina suborbicularis Grzybowski, 1896, p. 284,

pl. 9, figs. 5-6 (fide Ellis & Messina).

Haplophragmoides suborbicularis (Grzybowski) -

Geroch, 1960, pl. 5, fig. 1a-b.

Haplophragmoides suborbicularis suborbicularis

(Grzybowski) - Jurkiewicz, 1967, p. 77, pl. 4, figs. 12-13.

Cribrostomoides? ex. gr. suborbicularis (Grzybowski) -

Mjatliuk, 1970, p. 76, pl. 18, fig. 3.

Haplophragmoides (Cribrostomoides) suborbicularis

(Grzybowski) - Liszka & Liszkowa, 1981, p. 176, pl. 3,

figs. 2-3.

Haplophragmoides suborbicularis (Grzybowski) emend.

Kaminski, Gradstein & collaborators, 2005 - Kaminski,


4.


Chattian.

Family LITUOLIDAE de Blainville, 1827

Subfamily AMMOMARGINULINAE Podobina, 1978

Genus Ammobaculites Cushman, 1910

Ammobaculites agglutinans (d’Orbigny, 1846) emend.

Bartenstein, 1952

Fig. 4.4

Spirolina agglutinans d’Orbigny, 1846, p. 137, pl. 7, figs.

10-12.

Haplophragmium agglutinans (d’Orbigny) - Brady, 1884,

pl. 32, figs. 19-20, 24-26.

Ammobaculites agglutinans (d’Orbigny) - Bartenstein,

1952, p. 318, pl. 1, fig. 1, pl. 2, figs. 10-16.

Ammobaculites agglutinans (d’Orbigny) emend.

Bartenstein, 1952 - Kaminski, Gradstein and

collaborators, 2005, p. 324, pl. 70, figs. 1-8.

Stratigraphical distribution in samples: Chattian.

Genus Sculptobaculites Loeblich & Tappan, 1984

Sculptobaculites barri Beckmann, 1991

Fig. 4.12

Sculptobaculites barri Beckmann, 1991, p. 823, pl. 1,

figs. 6-9.


Superfamily RECURVOIDACEA Alekseychik-

Mitskevich, 1973

Family AMMOSPHAEROIDINIDAE Cushman, 1927

Subfamily AMMOSPHAEROIDININAE Cushman,

1927

Genus Ammosphaeroidina Cushman, 1910

Ammosphaeroidina pseudopauciloculata (Mjatliuk,

1966)

Fig. 4.13

Trochammina pauciloculata Brady - Grzybowski, 1896,

p. 283, pl. 8, figs. 51-52.

Cystamminella pseudopauciloculata Mjatliuk, 1966, p.

264, pl. 1, figs. 5-7, pl. 2, fig. 6, pl. 3, fig. 3.

Cystamminella ancestralis Mjatliuk, 1970, p. 103, pl. 30,

figs. 8-9.

Cystamminella elongata Mjatliuk, 1970, p. 104, pl. 31,

figs. 1, 3-5.

Cystamminella grzybowskii Mjatliuk, 1970, p. 105, pl.

10, figs. 19-21.

Cystamminella bytkovensis Mjatliuk, 1970, p. 106, pl. 28,

fig. 11, pl. 31, fig. 2.

Ammosphaeroidina pseudopauciloculata (Mjatliuk) -

Kaminski et al., 1988, p. 193, pl. 8, figs. 3-5.


Chattian.

Subfamily RECURVOIDINAE Alekseychik-

Mitskevich, 1973

Genus Recurvoides Earland, 1934

Recurvoides sp.

Fig. 4.14a-c


Suborder SPIROPLECTAMMININA Mikhalevich,

1992

Superfamily SPIROPLECTAMMINACEA Cushman,

1927

Family SPIROPLECTAMMINIDAE Cushman, 1927

Subfamily VULVULININAE Saidova, 1981

Genus Vulvulina d'Orbigny, 1826

Vulvulina haeringensis (Gümbel, 1868)

Fig. 5.4

Venilina haeringensis Gümbel, 1868, p. 649, pl. 2, fig.

84.

Schizophora haeringensis (Gümbel) - Hantken, 1875, p.

68, pl. 3, fig. 3.

Vulvulina haeringensis (Gümbel) - Hagn, 1956, p. 115,

pl. 9, figs. 7-8.


Chattian.

Vulvulina spinosa Cushman, 1927a

Figs. 5.3a-b, 5.6

Vulvulina spinosa Cushman, 1927a, pl. 23, fig. 1.

?Vulvulina jarvisi Cushman, 1932, p. 84, pl. 10, fig. 20.

Spiroplectammina spinosa (Cushman) - Parisi &

Coccioni, 1988, p. 105, pl. 1, fig. 7.


Suborder VERNUILININA Mikhalevich & Kaminski,

2004

Superfamily VERNEUILINACEA Cushman, 1911

Family PROLIXOPLECTIDAE Loeblich & Tappan,

1985

Genus Karrerulina Finlay, 1940

Karrerulina conversa (Grzybowski, 1901)


33

Fig. 5 All scales represent 100 μm. 1. Spirorutilus carinatus (d’Orbigny). Same specimen, a. apertural view, b. lateral

view, same scale, sample F4, Rupelian. 2. Semivulvulina pectinata (Reuss). Same specimen, a. apertural view, b. lateral

view, same scale, sample F15, Rupelian. 3,6. Vulvulina spinosa Cushman. 3a-b. same specimen, a. apertural view, b.

lateral view, same scale, sample F2, Rupelian; 6. sample F7, Rupelian. 4. Vulvulina haeringensis (Gümbel). Sample F7,

Rupelian. 5. Semivulvulina sagittula (Defrance). Sample F3, Rupelian. 7-8. Karrerulina conversa (Grzybowski). 7.

sample F7, Rupelian; 8. sample F25, Rupelian. 9. Popovia plana (Bykova). Same specimen, a. lateral view, b. peripherial

view, c. lateral view, same scale, sample FB20, Chattian.


34

Figs. 5.7-5.8

Gaudryina conversa Grzybowski, 1901, p. 254, pl. 8,


Gaudryina apicularis Cushman, 1911, p. 69, fig. 110.

Karreriella (Karrerulina) apicularis (Cushman) - Finlay,

1940, p. 450.

Karreriella (Karrerulina) aegra Finlay, 1940, p. 451, pl.

62, figs. 21-22.

Karreriella (Karrerulina) bortonica Finlay, 1940, ibid. p.

451, pl. 62, figs. 18-20.

Plectina apicularis (Cushman) - Phleger et al., 1953, p.

27, pl. 5, fig. 10.

Plectina conversa (Grzybowski) - Geroch, 1960, p. 59,

pl. 6, fig. 7.

Karreriella indigena Mjatliuk, 1970, p. 116, pl. 34, figs.

10-14.

Karreriella pokornyi Mjatliuk, 1970, p. 117-118, pl. 34,

figs. 15-17.

Karreriella conversa (Grzybowski) - Webb, 1975, p. 835,

pl. 3, fig. 12.

Karreriella apicularis (Cushman) - Pflum & Frerichs,

1976, pl. 1, fig. 2.

Karrerulina conversa (Grzybowski) - King, 1989, p. 456,

pl. 9.1, figs. 23-24.

Gerochammina conversa (Grzybowski) - Kaminski &

Geroch, 1993, p. 279, pl. 13, figs. 5-11.


Chattian.

Order LOFTUSIIDA Kaminski & Mikhalevich, 2004

Suborder LOFTUSIINA Kaminski & Mikhalevich,

2004

Superfamily LOFTUSIACEA Brady, 1884

Family CYCLAMMINIDAE Marie, 1955

Subfamily ALVEOLOPHRAGMIINAE Saidova, 1981

Genus Popovia Suleymanov, 1965

Popovia plana (Bykova, 1939)

Fig. 5.9a-c

Alveolophragmium planum Bykova, 1939, p. 20, pl. 1,

fig. 10, textfig. 1. (fide Ellis & Messina).

Popovia plana (Bykova) - Suleymanov, 1965, p. 48.


Chattian.

Genus Reticulophragmium Maync, 1955

Reticulophragmium acutidorsatum (Hantken, 1868)

Fig. 6.3

Haplophragmium acutidorsatum Hantken, 1868, p. 82,

pl. 1, fig. 1 (fide Ellis & Messina).

? Cyclammina appeninica Emiliani, 1954, p. 117, pl. 21,

figs. 18-19.

Cyclammina acutidorsata (Hantken) - Lühr, 1962, pl. 1,

fig. 2.

Alveolophragmium acutidorsatum (Hantken) - Popescu,

1972, p. 302, pl. 1, figs. 1-2, pl. 2, fig. 16, pl. 5, fig. 31,

pl. 6, fig. 43, pl. 7, figs. 45, 48.

Haplophragmium acutidorsatum Hantken - Gellai-Nagy,

1988, pl. 1, figs. 1-3.

Reticulophragmium acutidorsatum (Hantken) - King,

1989, p. 458, pl. 9.2, fig. 15.

Cyclammina (Cyclammina) acutidorsata (Hantken) -

Charnock & Jones, 1990, p. 175, pl. 7, figs. 3-4, pl. 18,

fig. 2.

Reticulophragmium acutidorsatum (Hantken) -

Kaminski, Gradstein and collaborators, 2005, p. 490, pl.

122, figs. 1-7.


Chattian.

Reticulophragmium amplectens (Grzybowski, 1898)

Fig. 6.1

Cyclammina amplectens Grzybowski, 1898, p. 292, pl.

12, figs. 1-3.

Cyclammina samanica Berry, 1928, p. 393, textfig. 5a-b.

Cyclammina grangei Finlay, 1940, p. 448, pl. 62, figs. 3–

4.

Cyclammina subcarpatica Majzon, 1943, p. 156, pl. 2,

fig. 8a–b.

Reticulophragmium amplectens (Grzybowski) - King,

1989, p. 458, pl. 9.2, figs. 16-18.

Cyclammina (Reticulophragmium) amplectens

(Grzybowski) - Charnock & Jones, 1990, p. 176, pl. 8,

figs. 1-5, pl. 18, fig. 3.

Reticulophragmium amplectens (Grzybowski) - Kaminski

& Geroch, 1993, pp. 266-267, pl. 11, figs. 5-7.


Reticulophragmium rotundidorsatum (Hantken, 1875)

Fig. 6.2

Haplophragmium rotundidorsatum Hantken, 1875, p. 12,

pl. 1, fig. 2 (fide Ellis & Messina).

Cyclammina rotundidorsata (Hantken) - Cicha &

Zapletalová, 1963, p. 103, abh. 14.

Haplophragmoides rotundidorsatum (Hantken) - Braga et

al., 1975, p. 102.

Alveolophragmium rotundidorsatum (Hantken) -

Popescu, 1972, p. 304, pl. 1, figs. 3-4, pl. 2, fig. 15, pl. 4,

fig. 20, pl. 5, fig. 37, pl. 7, figs. 47,49.

Cyclammina (Reticulophragmium) rotundidorsatum

(Hantken) - Charnock & Jones, 1990, pl. 7, figs. 13-15,

pl. 19, fig. 1.

Reticulophragmium rotundidorsatus (Hantken) -

Schröder-Adams & McNiel, 1994, pl. 8, figs. 5-7.

Reticulophragmium rotundidorsatum (Hantken) - Cicha

et al., 1998, pl. 5, fig. 5.


Chattian.

Order TEXTULARIIDA Delage & Hérouard, 1896

Suborder TEXTULARIINA Delage & Hérouard, 1896

Superfamily EGGERELLACEA Cushman, 1937

Family EGGERELLIDAE Cushman, 1937

Subfamily EGGERELLINAE Cushman, 1937

Genus Karreriella Cushman, 1933

Karreriella chilostoma (Reuss, 1852)

Fig. 6.4

Textularia chilostoma Reuss, 1852, p. 18, textfigs. a-b.

Gaudryina chilostoma (Reuss) - Reuss, 1865, p. 120, pl.

1, fig. 5.

Gaudryina pupoides var. chilostoma Reuss - Brady,

1884, p. 379, pl. 46, figs. 5-6.


35

Fig. 6 All scales represent 100 μm. 1. Reticulophragmium amplectens (Grzybowski). Sample F7, Rupelian. 2.

Reticulophragmium rotundidorsatum (Hantken). Sample F28, Rupelian. 3. Reticulophragmium acutidorsatum (Hantken).

Sample F2, Rupelian. 4. Karreriella chilostoma (Reuss). Sample F18, Rupelian. 5. Valvulina sp. Same specimen, a.

umbilical view, b. spiral view, same scale, sample F2, Rupelian. 6. Karreriella hantkeniana Cushman. Sample F4,

Rupelian. 7. Karreriella gaudryinoides (Fornasini). Sample F15, Rupelian. 8-10. Valvulina flexilis Cushman & Renz. 8.

sample F8, Rupelian; 9-10. sample F31, Rupelian/Chattian.


36

Gaudryina pupoides Brady - Parker & Jones, 1887, p.

279, pl. 42, figs. 7-9.

Karreriella chilostoma (Reuss) - Cushman, 1937, p. 126,

pl. 15, figs. 1-8.

Karreriella siphonella (Reuss) var. chilostoma (Reuss) -

Batjes, 1958, p. 100, pl. 1,

fig. 8.

Siphotextularia chilostoma (Reuss) - Schudack &

Nuglisch, 2004, p. 78, pl. 8, figs. 156-157.


Chattian.

Karreriella gaudryinoides (Fornasini, 1885)

Fig. 6.7

Clavulina gaudryinoides Fornasini, 1885, p. 8, pl. 6, figs.

3-9.

Gaudryina gaudryinoides (Fornasini) - Cole, 1927, p. 13.

Karreriella gaudryinoides (Fornasini) - Cicha et al.,

1998, p. 108, pl. 8, figs. 10-12.


Karreriella hantkeniana Cushman, 1936

Fig. 6.6

Karreriella hantkeniana Cushman, 1936, p. 36, pl. 5, fig.

19.


Chattian.

Family VALVULINIDAE Berthelin, 1880

Subfamily VALVULININAE Berthelin, 1880

Genus Valvulina d'Orbigny, 1826

Valvulina flexilis Cushman & Renz, 1941

Figs. 6.8-6.10

Valvulina flexilis Cushman & Renz, 1941, p. 7, pl. 1, figs.

16-17.

Dorothia flexilis (Cushman & Renz) - Popescu, 1975, p.

40, pl. 6, figs. 2-4, pl. 7, figs. 3-5.


Chattian.

Valvulina sp.

Fig. 6.5a-b


Superfamily TEXTULARIACEA Ehrenberg, 1838

Family KAMINSKIIDAE Neagu, 1999

Genus Spirorutilus Hottinger, Halicz & Reiss, 1990

Spirorutilus carinatus (d'Orbigny, 1846)

Fig. 5.1a-b

Textularia carinata d'Orbigny, 1846, p. 247, pl. 14, figs.

32-34.

Textularia lacera Reuss, 1851, p. 84, pl. 6, figs. 52-53.

Textularia attenuata Reuss, 1851, p. 84, pl. 6, fig. 54.

Textilaria carinata d’Orbigny - Reuss, 1865, p. 474.

Textularia pseudocarinata Cushman, 1921, p. 121, pl. 22,

figs. 5, 15-16.

Spiroplectammina carinata (d'Orbigny) - Weber, 1939, p.

17-20.

Spiroplectammina carinata (d'Orbigny) - Dam &

Reinhold, 1942, p. 42, pl. 6, fig. 2.

Bolivinopsis carinata (d’Orbigny) - Pokorny, 1958, p.

193, fig. 111.

Spirorutilis carinatus (d'Orbigny) - Hofker, 1976, p. 69.

Spirorutilis carinatus (d’Orbigny) - Banner & Pereira,

1981, p. 102.

Spiroplectinella carinata (d’Orbigny) - Papp & Schmid,

1985, p. 86, pl. 80, figs. 1-4.

Spirorutilus carinatus (d’Orbigny) - Hottinger et al.,

1990, p. 67, pl. 1, figs. 6-7.


Chattian.

Family TEXTULARIIDAE Ehrenberg, 1838

Subfamily TEXTULARIINAE Ehrenberg, 1838

Genus Semivulvulina Finlay, 1939

Semivulvulina pectinata (Reuss, 1850)

Fig. 5.2a-b

Textularia pectinata Reuss, 1850, p. 381, pl. 49, figs. 2-3.

Semivulvulina pectinata pectinata (Reuss) - Cicha &

Zapletalová, 1965, p. 130, abb. 22.

Semivulvulina pectinata (Reuss) - Rögl & Cicha, 1973, p.

301, pl. 1, fig. 2, pl. 4, figs. 20-21.


Chattian.

Semivulvulina sagittula (Defrance, 1824)

Fig. 5.5

Textularia sagittula Defrance, 1824, p. 117, pl. 13, fig. 5


Textulaire sagittule Defrance - Blainville, 1825, p. 370,

pl. 5, fig. 5.

Textularia sagitata Defrance - Jones et al. 1866, pl. 3,

figs. 7-9.

Textilaria sagittula Defrance - Jones & Parker, 1871, pp.

511, 564.

Spiroplectammina sagittula (Defrance) - Hofker, 1960, p.

237, pl. A, fig. 17.

Semivulvulina sagittula (Defrance) - Popescu, 1975, p.

39, pl. 4, fig. 5.

Spiroplectinella sagittula (Defrance) - Schiebel, p. 26, pl.

6, fig. 14.


Order MILIOLIDA Lankester 1885

Superfamily MILIOLACEA Ehrenberg, 1839

Family SPIROLOCULINIDAE Wiesner, 1920

Genus Spiroloculina d’Orbigny, 1826

Spiroloculina canaliculata d’Orbigny, 1846

Fig. 7.2

Spiroloculina canaliculata d’Orbigny, 1846, p. 269, pl.

16, figs. 10-12.


Spiroloculina sp.

Fig. 7.1


Family HAUERINIDAE Schwager, 1876

Subfamily HAUERININAE Schwager, 1876

Genus Cycloforina Łuczkowska, 1972


37

Fig. 7 All scales represent 100 μm. 1. Spiroloculina sp. Sample F9, Rupelian. 2. Spiroloculina canaliculata d’Orbigny.

Sample F5, Rupelian. 3,5. Cycloforina ludwigi (Reuss). 3. sample F14, Rupelian; 5a-c. same specimen, a. lateral view,

b. apertural view, c. lateral view, same scale, sample FB21, Chattian. 4. Quinqueloculina sp. Sample F17, Rupelian. 6.

Sigmoilinita tenuis (Czjzek). Same specimen, a. apertural view, b. lateral view, same scale, sample F5, Rupelian. 7.

Sigmoilinita tenuissima (Reuss). Same specimen, a. apertural view, b. lateral view, same scale, sample F5, Rupelian. 8-

9. Laevidentalina elegans (d’Orbigny). 8. sample F7, Rupelian; 9. sample F15, Rupelian. 10. Laevidentalina communis

(d’Orbigny). Sample F18, Rupelian. 11. Nodosaria soluta (Reuss). Sample F3, Rupelian. 12. Neugeborina longiscata

(d’Orbigny). Sample F12, Rupelian.


38

Cycloforina ludwigi (Reuss, 1866)

Figs. 7.3, 7.5a-c

Quinqueloculina ludwigi Reuss, 1866, p. 126, pl. 1, fig.

12.

Cycloforina ludwigi (Reuss) - Łuczkowska, 1972, p. 365.


Chattian.

Genus Quinqueloculina d’Orbigny, 1826

Quinqueloculina sp.

Fig. 7.4


Subfamily SIGMOILINITINAE Luczkowska, 1974

Genus Sigmoilinita Seiglie, 1965

Sigmoilinita tenuis (Czjzek, 1848)

Fig. 7.6a-b

Quinqueloculina tenuis Czjzek, 1848, p. 149, pl. 13, figs.

31-34.

Spiroloculina berchtoldsdorfensis Karrer, 1877, p. 375,

pl. 16a, fig. 10.

Spiroloculina tenuis (Czjzek) - Brady, 1884, p. 152, pl.

10, figs. 7-8, 11.

Sigmoilina tenuis (Czjzek) - Marks, 1951, p. 39, pl. 5, fig.

7.

“Sigmoilina” tenuis (Czjzek) - Barker, 1960, pl. 10, figs.

7-8, 11.

Sigmoilinita tenuis (Czjzek) - Seiglie, 1965, p. 72.

Spirosigmoilina tenuis (Czjzek) - Sztrákos, 1979, pl. 7,

fig. 13.


Chattian.

Sigmoilinita tenuissima (Reuss, 1867)

Fig. 7.7a-b

Spiroloculina tenuissima Reuss, 1867, p. 55, pl. 1, fig.

11.

Sigmoilina tenuissima (Reuss) - Serova, 1955, p. 326, pl.

12, figs. 5-6.

Sigmoilinita tenuissima (Reuss) - Łuczkowska, 1974, p.

149, pl. 16, figs. 4-6.


Order LAGENIDA Lankester, 1885

Suborder LAGENINA Delage & Hérouard, 1896

Superfamily NODOSARIACEA Ehrenberg, 1838

Family NODOSARIIDAE Ehrenberg, 1838

Subfamily NODOSARIINAE Ehrenberg, 1838

Genus Dentalina Risso, 1826

Dentalina contorta (Hantken, 1868)

Fig. 8.2

Nodosaria (Dentalina) contorta Hantken, 1868, p. 159,

pl. 16, fig. 2.

Dentalina contorta (Hantken) - Hantken, 1875, p. 36, pl.

4, fig. 5.

Dentalina contorta (Hantken) - Cicha et al., 1998, p. 93,

pl. 21, fig. 1.


Genus Laevidentalina Loeblich & Tappan, 1986

Laevidentalina communis (d’Orbigny, 1826)

Fig. 7.10

Dentalina communis d’Orbigny, 1826, p. 254, no. 35.

Nodosarina communis (d’Orbigny) - Goës, 1894, pl. 2,

fig. 36.

Nodosaria communis (d’Orbigny) - Cushman, 1923, p.

75, pl. 12, figs. 3-4, 15-17.

Enantiodentalina communis (d’Orbigny) - Giannini &

Tavani, 1960, p. 46, pl. 6, fig. 15.

Dentalia communis (d’Orbigny) - Le Roy, 1964, p. 23, pl.

15, fig. 28.

Laevidentalina communis (d’Orbigny) - Cicha et al.,

1998, p. 109, pl. 21, fig. 3.


Laevidentalina elegans (d’Orbigny, 1846)

Figs. 7.8-7.9

Dentalina elegans d’Orbigny, 1846, p. 45, pl. 1, figs. 52-

56.

Siphonodosaria elegans (d’Orbigny) - Stainforth, 1952,

p. 12.

Stilostomella elegans (d’Orbigny) - Stainforth, 1952, p.

136.

Stilostomella elegans (d’Orbigny) - Popescu, 1975, p. 66,

pl. 17, fig. 8.

Dentalina elegans d’Orbigny - Papp & Schmid, 1985, p.

28, pl. 10, figs. 1-5.

Laevidentalina elegans (d’Orbigny) - Cicha et al., 1998,

p. 109, pl. 21, figs. 6-7


Genus Neugeborina Popescu, 1998

Neugeborina longiscata (d’Orbigny, 1846)

Fig. 7.12

Nodosaria longiscata d’Orbigny, 1846, p. 32, pl. 1, fig.

10-12.

Nodosaria irregularis d’Orbigny, 1846, p. 32, pl. 1, figs.

13-14.

Nodosaria orbignyana Neugeboren, 1852, p. 40, pl. 1,

fig. 15.

Nodosaria buchiana Neugeboren, 1852, p. 41, pl. 1, fig.

16.

Nodosaria ackneriana Neugeboren, 1852, p. 41, pl. 1,

figs. 17-18.

Nodosaria roemeriana Neugeboren, 1852, p. 42, pl. 1,

fig. 19.

Nodosaria nodifera Neugeboren, 1852, p. 42, pl. 1, figs.

20-21.

Nodosaria capillaris Neugeboren, 1852, p. 50, pl. 1, figs.

22-24.

Nodosaria exilis Neugeboren, 1852, p. 51, pl. 1, figs. 25-

26.

Nodosaria c1avaeformis Neugeboren, 1852, p. 53, pl. 1,

fig. 38.

Nodosaria culmen Costa, 1856, p. 158, pl. 13, fig. 15.

Nodosaria arundinea Schwager, 1866, p. 211, pl. 5, figs.

43-45.

Nodosaria (Dentalina) longiscata (d’Orbigny) -

Schuhert, 1899, p. 219.

Lagenodosaria longiscata (d’Orbigny) - Colom, 1956, pl.

8, figs. 1-4.


39

Fig. 8 All scales represent 100 μm. 1. Pyramidulina latejugata (Gümbel). Sample FB20, Chattian. 2. Dentalina contorta

(Hantken). Sample F12, Rupelian. 3. Amphimorphina haueriana Neugeboren. Sample F4, Rupelian. 4. Dimorphina sp. Sample

F12, Rupelian. 5. Lenticulina vortex (Fichtel & Moll). Sample F15, Rupelian. 6. Neolenticulina peregrina (Schwager). Sample

FB21, Chattian.7. Lenticulina limbosa (Reuss). Same specimen, a. lateral view, b. peripherial view, c. lateral view, same scale,

sample FB21, Chattian. 8. Lenticulina gibba (d’Orbigny). Sample F11, Rupelian. 9. Lenticulina arcuatostriata (Hantken). Same

specimen, a. lateral view, b. peripherial view, same scale, sample FB20, Chattian. 10. Lenticulina inornata (d’Orbigny). Same

specimen, a. lateral view, b. peripherial view, same scale, sample FB20, Chattian. 11. Lenticulina calcar (Linné). Sample F15,

Rupelian. 12. Saracenaria senni Hedberg. Same specimen, a. lateral view, b. apertural view, same scale, sample F3, Rupelian.

13. Saracenaria propinqua (Hantken). Same specimen, a. lateral view, b. apertural view, same scale, sample F5, Rupelian.


40

Stilostomella longiscata (d’Orbigny) - Hausmann, 1964,

p. 380, pl. 3, fig. 11.

Chrysalogonum longiscatum (d’Orbigny) - Sztrákos,

1979, pl. 9, fig. 16.

Siphonodosaria longiscata (d’Orbigny) - Perembo, 1995,

pl. 9, fig. 20.

Neugeborina longiscata (d’Orbigny) - Cicha et al., 1998,

p. 113, pl. 21, fig. 12.


Chattian.

Genus Nodosaria Lamarck, 1812

Nodosaria soluta (Reuss, 1851)

Fig. 7.11

Dentalina soluta Reuss, 1851, p. 60, pl. 3, fig. 4a-b.

Nodosaria (Dentalina) soluta (Reuss) - Andreae, 1884, p.

201, pl. 10, fig. 8.

Nodosaria soluta (Reuss) - Batjes, 1958, p. 114, pl. 3,

figs. 17-18.


Genus Pyramidulina Fornasini, 1894

Pyramidulina latejugata (Gümbel, 1868)

Fig. 8.1

Nodosaria latejugata Gümbel, 1868, p. 619, pl. 1, fig. 32.

Pyramidulina latejugata (Gümbel) - Cicha et al., 1998, p.

121, pl. 21, fig. 13.


Chattian.

Subfamily PLECTOFRONDICULARIINAE Cushman, 1927

Genus Amphimorphina Neugeboren, 1850

Amphimorphina haueriana Neugeboren, 1850

Fig. 8.3

Amphimorphina haueriana Neugeboren, 1850, p. 127, pl.

4, figs. 13-14.

Nodosaria compressiuscula Neugeboren, 1852, p. 59, pl.

1, figs. 54-56.

Plectofrondicularia trinitatensis Cushman & Jarvis,

1929, p. 11, pl. 2, fig. 16.

Amphimorphina miocenica Cushman, 1948, p. 220, pl.

15, figs. 16-18.

Amphimorphinella butonensis Keyzer var. compressa

Keyzer, 1953, p. 276, pl. 3, figs. 7, 9-16.


Chattian.

Family VAGINULINIDAE Reuss, 1860

Subfamily LENTICULININAE Chapman, Parr &

Collins, 1934

Genus Dimorphina d'Orbigny, 1826

Dimorphina sp.

Fig. 8.4


Genus Lenticulina Lamarck, 1804

Lenticulina arcuatostriata (Hantken, 1868)

Fig. 8.9a-b

Cristellaria (Robulina) arcuato striata Hantken, 1868, p.

93, pl. 2, fig. 20 (fide Ellis & Messina).

Robulina arcuato-striata (Hantken) - Hantken, 1875, p.

56, pl. 7, fig. 2.

Robulus arcuatostriatus (Hantken) - Cuvillier & Szakall,

1949, p. 51, pl. 23, fig. 9.

Robulus arcuato-striatus (Hantken) - Ruiz de Caona &

Colom, 1950, p. 402, textfig. 6, nr. 1-9.

Lenticulina arcuatostriata (Hantken) - Braga et al., 1975,

p. 104.

Lenticulina arcuato-striata (Hantken) - Grünig, 1985, p.

261, pl. 4, fig. 19.

Robulina arcuatostriata (Hantken) - Gellai-Nagy, 1988,

pl. 8, figs. 1-3.

Lenticulina arcuatostriata (Hantken) - Cicha et al., 1998,

p. 109, pl. 24, figs. 1-2.


Chattian.

Lenticulina calcar (Linné, 1758)

Fig. 8.11

Nautilus calcar Linné, 1758, p. 709 (fide Ellis &

Messina).

Nautilus calcar Linné, 1758 var. α Fichtel & Moll, 1798,

p. 71, pl. 11, figs. a-c.

Robulina calcar (Linné) - d’Orbigny, 1846, p. 99, pl. 4,

fig. 20.

Cristellaria calcar (Linné) - Parker & Jones, 1860, tab. p.

100, p. 111.

Nodosarina calcar (Linné) - Goës, 1882, pl.3, figs. 54-

56.

Robulus calcar (Linné) - Dieci, 1959, p. 28, pl. 2, figs. 8-

10.

Lenticulina calcar (Linné) - Rögl, 1968, p. 75, pl. 5, fig.

1.


Chattian.

Lenticulina gibba (d’Orbigny, 1839)

Fig. 8.8

Cristellaria gibba d’Orbigny, 1839, p. 40, pl. 7, figs. 20-


Robulus gibbus (d’Orbigny) - Bermúdez, 1949, p. 126,

pl. 7, figs. 53-54.

Lenticulina gibba (d’Orbigny) - Dieci, 1959, p. 36, pl. 3,

figs. 7-8.

Lenticulina (Lenticulina) gibba (d’Orbigny) - Kiesel,

1962, p. 28, pl. 4, fig. 6.


Chattian.

Lenticulina inornata (d’Orbigny, 1846)

Fig. 8.10a-b

Robulina inornata d’Orbigny, 1846, p. 102, pl. 4, figs.

27-28.

Cristellaria crassa d’Orbigny, 1846, p. 90, pl. 4, figs. 1-

3.

Robulina simplex d’Orbigny, 1846, p. 103, pl. 4, figs. 27-

28.

Robulina austriaca d’Orbigny, 1846, p. 103, pl. 5, figs. 1-

2.


41

Fig. 9 All scales represent 100 μm. 1. Saracenaria arcuata (d’Orbigny). Same specimen, a. lateral view, b. apertural

view, same scale, sample F3, Rupelian. 2-3. Frondovaginulina tenuissima (Hantken). 2. sample F17, Rupelian; 3. sample

F28, Rupelian. 4. Percultazonaria fragaria (Gümbel). Same specimen, a. apertural view, b. lateral view, same scale,

sample FB20, Chattian. 5-6. Amphicoryna spinicosta (d’Orbigny). 5. sample F13, Rupelian; 6. same specimen, a.

apertural view, b. lateral view, same scale, sample FB21, Chattian. 7. Amphicoryna armata (Neugeboren). Sample F21,

Rupelian. 8. Amphicoryna crassa (Hantken). Sample F10, Rupelian. 9. Marginulina behmi (Reuss). Same specimen, a.

apertural view, b. lateral view, c. peripherial view, same scale, sample FB21, Chattian. 10. Hemirobulina recta

(Hantken). Sample F12, Rupelian. 11. Hemirobulina hantkeni (Bandy). Same specimen, a. apertural view, b. lateral view,

same scale, sample FB21, Chattian.


42

Robulina intermedia d’Orbigny, 1846, p. 104, pl. 5, figs.

3-4.

Cristellaria inornata (d’Orbigny) - Reuss, 1865, p. 144.

Robulus inornatus (d’Orbigny) - Agip, 1957, pl. 10, fig.

4.

Lenticulina inornata (d’Orbigny) - Rögl, 1968, p. 75, pl.

2, fig. 7a-b.


Chattian.

Lenticulina limbosa (Reuss, 1863)

Fig. 8.7a-c

Cristellaria (Robulina) limbosa Reuss, 1863, p. 55, pl. 6,

fig. 69 (fide Ellis & Messina).

Cristellaria limbosa (Reuss) - Nuttall, 1932, p. 11.

Rubulus limbosus (Reuss) - Ruiz de Gaona & Colom,

1950, p. 405, fig. 5/1-14.

Robulus limbosus (Reuss) - Hagn, p. 127, pl. 11, fig. 2.

Lenticulina limbosa (Reuss) - Mathelin & Sztrákos, 1993,

p. 75, pl. 8, fig. 6


Chattian.

Lenticulina vortex (Fichtel & Moll, 1798)

Fig. 8.5

Nautilus vortex Fichtel & Moll, 1798, p. 33, pl. 2, fig. d-i.

Nautilus calcar Linné, 1758 var. η Fichtel & Moll, 1798,

p. 76, pl. 12, fig. g-h.

Polystomella vortex (Fichtel & Moll) - Di Blainville,

1824, p. 183.

Robulina vortex (Fichtel & Moll) - d’Orbigny, 1826, p.

288, no. 4.

Robulina imperatoria d’Orbigny, 1846, p. 104, pl. 5, figs.

5-6.

Cristellaria vortex (Fichtel & Moll) - Parker & Jones,

1860, p. 113.

Cristellaria vortex (Fichtel & Moll) - Brady, 1884, p.

1548, pl. 69, figs. 14-16.

Robulus vortex (Fichtel & Moll) - Cushman, 1946, p. 6,

pl. 1, figs. 3-4.

Lenticulina (Lenticulina) vortex (Fichtel & Moll) - Butt,

1966, p. 41, pl. 2, fig. 1.

Lenticulina vortex (Fichtel & Moll) - Rögl, 1968, p. 75.

Lenticulina vortex (Fichtel & Moll) - Popescu, 1975, p.

55, pl. 30, fig. 30, pl. 31, fig. 1.

Lenticulina vortex (Fichtel & Moll) - Rögl & Hansen,

1984, p. 30, pl. 2, figs. 3-4, textfig. 8.


Genus Neolenticulina McCulloch, 1977

Neolenticulina peregrina (Schwager, 1866)

Fig. 8.6

Cristellaria peregrina Schwager, 1866, p. 245, pl. 7, fig.

89.

Lenticulina peregrina (Schwager) – AGIP Mineraria,

1957, pl. 11, fig. 2.

Neolenticulina peregrina (Schwager) - Loeblich &

Tappan, 1987, p. 406, pl. 447, figs. 9-12, 16.


Chattian.

Genus Percultazonaria Loeblich & Tappan, 1986

Percultazonaria fragaria (Gümbel, 1868)

Fig. 9.4a-b

Marginulina fragaria Gümbel, 1868, p. 635, pl. 1, fig.

58a-c.

Cristellaria fragaria (Gümbel) - Hantken, 1875, pl. 6,

figs. 1-3.

Lenticulina fragaria (Gümbel) - Subbotina, 1947, p. 85,

pl. 8, figs. 6-7.

Marginulinopsis fragaria (Gümbel) - Hagn, 1956, p. 134,

pl. 12, figs. 3, 8, 10.

Percultazonaria fragaria (Gümbel) - Mathelin &

Sztrákos, 1993, p. 76, pl. 9, figs. 1-2.

Vaginulinopsis fragaria (Gümbel) - Ozsvárt, 2007, p. 56,

pl. 6, fig. 1.


Genus Saracenaria Defrance, 1824

Saracenaria arcuata (d’Orbigny, 1846)

Fig. 9.1a-b

Cristellaria arcuata d’Orbigny, 1846, p. 87, pl. 3, figs.

34-36.

Lenticulina (Sarazenaria) arcuata (d’Orbigny) -

Hausmann, 1964, p. 352, pl. 5, fig. 2.

Saracenaria arcuata (d’Orbigny) - Rögl & Cicha, 1973,

pl. 5, fig. 28.


Chattian.

Saracenaria propinqua (Hantken, 1875)

Fig. 8.13a-b

Cristellaria propinqua Hantken, 1875, p. 52, pl. 5, fig. 4.

Robulus propinquus (Hantken) - Cushman, 1935, p. 16,

pl. 5, fig. 1.

Saracenaria propinqua (Hantken) - Majzon, 1962, pl.

33(5), fig. 4.

Marginulinopsis propinqua (Hantken) - Hagn et al.,

1981, p. 96.

Saracenaria propinqua (Hantken) - Cicha et al., 1998, p.

125, pl. 24, fig. 9.


Chattian.

Saracenaria senni Hedberg, 1937

Fig. 8.12a-b

Saracenaria senni Hedberg, 1937, p. 674, pl. 90, fig. 18a-

b.


Chattian.

Subfamily PALMULINAE Saidova, 1981

Genus Frondovaginulina Schubert, 1912

Frondovaginulina tenuissima (Hantken, 1875)

Figs. 9.2-9.3

Frondicularia tenuissima Hantken, 1875, p. 43, pl. 13,

fig. 11a-b.

Flabelinella tenuissima (Hantken) - Hagn, 1952, p. 156,

pl. 4, fig. 20.

Flabellina tenuissima (Hantken) - Hagn, 1956, p. 139.

Citharinella tenuissima (Hantken) - Popescu, 1975, p. 49,

pl. 36, fig. 1, pl. 37, figs. 3-4.


43

Fig. 10 All scales represent 100 μm. 1. Vaginulinopsis cumulicostatus (Gümbel). Same specimen, a. apertural view, b.

lateral view, same scale, sample F10, Rupelian. 2. Vaginulinopsis mexicana (Nuttall). Same specimen, a. apertural view,

b. lateral view, same scale, sample F10, Rupelian. 3. Lagena tenuis (Bornemann). Sample FB17, Chattian. 4.

Pseudofissurina kapellensis (Spiegler). Same specimen, a. apertural view, b. lateral view, same scale, sample FB17,

Chattian. 5. Favulina hexagona (Williamson). Sample F26, Rupelian. 6. Glandulina ovula d’Orbigny. Sample FB20,

Chattian. 7. Bolivina beyrichi carinata Hantken. Same specimen, a. apertural view, b. lateral view, same scale, sample

F21, Rupelian. 8. Bolivina beyrichi beyrichi Reuss. Sample F9, Rupelian. 9. Bolivina plicatella Cushman. Sample FB17,

Chattian. 10-11. Bolivina crenulata Cushman. 10. sample F33, Rupelian/Chattian; 11a-b. same specimen, a. apertural

view, b. lateral view, same scale, FB01, Chattian.


44

Plectofrondicularia tenuissima (Hantken) - Foraminiferi

padani, AGIP, 1982, pl. 24, fig. 8.

Frondovaginulina tenuissima (Hantken) - Mathelin &

Sztrákos, 1993, p. 76, pl. 9, fig. 8.

Palmula tenuissima (Hantken) - Cimerman et al., 2006, p.

24, pl. 5, figs. 8-9.


Chattian.

Subfamily MARGINULININAE Wedekind, 1937

Genus Amphicoryna Schlumberger, 1881

Amphicoryna armata (Neugeboren, 1852)

Fig. 9.7

Nodosaria armata Neugeboren, 1852, p. 56, pl. 1, fig. 44.

Amphicoryna armata (Neugeboren) - Popescu, 1975, p.

47, pl. 19, figs. 6-8.


Amphicoryna crassa (Hantken, 1868)

Fig. 9.8

Nodosaria crassa Hantken, 1868, p. 86, pl. 1, fig. 15.

Amphicoryna crassa (Hantken) - Sztrákos, 1979, p. 63,

pl. 10, figs. 1-2.


Chattian.

Amphicoryna spinicosta (d’Orbigny, 1846)

Figs. 9.5, 9.6a-b

Nodosaria spinicosta d’Orbigny, 1846, p. 37, pl. 1, figs.

32-33.

Amphicoryna spinicosta (d’Orbigny) - Sztrákos, 1979, pl.

10, figs. 3-4.


Chattian.

Genus Hemirobulina Stache, 1864

Hemirobulina hantkeni (Bandy, 1949)

Fig. 9.11a-b

Marginulina subbullata Hantken, 1875, p. 46, pl. 4, figs.

9-10, pl. 5, fig. 9.

Marginulina hantkeni Bandy, 1949, p. 46, pl. 6, fig. 9


Hemirobulina hantkeni (Bandy) - Cicha et al., 1998, p.

107, pl. 25, figs. 9-10.


Chattian.

Hemirobulina recta (Hantken, 1875)

Fig. 9.10

Marginulina recta Hantken, 1875, p. 47, pl. 4, fig. 15.

Hemirobulina recta (Hantken) - Horváth, 2003, p. 22, pl.

5, fig. 18.


Genus Marginulina d’Orbigny, 1826

Marginulina behmi (Reuss, 1866)

Fig. 9.9a-c

Cristellaria (Marginulina) behmi Reuss, 1866, p. 138, pl.

2, fig. 37.

Cristellaria behmi Reuss - Gümbel, 1868, p. 55, pl. 1, fig.

61.

Marginulina behmi (Reuss) - Hantken, 1875, p. 48, pl. 5,

figs. 1-2.

Marginulinopsis behmi (Reuss) - Cimerman et al., 2006,

p. 24, pl. 5, fig. 1.


Chattian.

Genus Vaginulinopsis Silvestri, 1904

Vaginulinopsis cumulicostatus (Gümbel, 1868)

Fig. 10.1a-b

Cristellaria cumulicostata Gümbel, 1868, p. 638, pl. 1,

fig. 67.

Vaginulinopsis cumulicostata (Gümbel) - Hagn, 1956, p.

135, pl. 13, fig. 3.

Vaginulopsis cumulicostatus (Gümbel) - Mathelin &

Sztrákos, 1993, p. 76, pl. 10, fig. 12.


Vaginulinopsis mexicana (Nuttall, 1932)

Fig. 10.2a-b

Vaginula elegans d’Orbigny var. mexicana Nuttall, 1932,

p. 16, pl. 13, figs. 12, 16.

Vaginulina mexicana Nuttall - Nuttall et al., 1941, p. 336,

pl. 9, fig. 4

Vaginulinopsis mexicana (Nuttall) - Poag, 2012, p. 109,

pl. 9, figs. 13-15.


Family LAGENIDAE Reuss, 1862

Genus Lagena Walker & Jacob, 1798

Lagena tenuis (Bornemann, 1855)

Fig. 10.3

Ovulina tenuis Bornemann, 1855, p. 317, pl. 12, fig. 3.

Lagena tenuis (Bornemann) - Reuss, 1862, p. 325, pl. 3,

figs. 30-39.


Chattian.

Family ELLIPSOLAGENIDAE Silvestri, 1923

Subfamily OOLININAE Loeblich & Tappan, 1961

Genus Favulina Patterson & Richardson, 1987

Favulina hexagona (Williamson, 1848)

Fig. 10.5

Entosolenia squamosa (Montagu) var. hexagona

Williamson, 1848, p. 20, pl. 2, fig. 23 (fide Ellis &

Messina).

Lagena hexagona (Williamson) - Brady, 1884, p. 472, pl.

58, figs. 32-33.

Oolina hexagona (Williamson) - Barker, 1960, p. 120, pl.

58, figs. 32-33.

Favulina hexagona (Williamson) - Cicha et al., 1998, p.

96, pl. 28, fig. 11.


Subfamily PARAFISSURININAE Jones, 1984

Genus Pseudofissurina Jones, 1984

Pseudofissurina kapellensis (Spiegler, 1974)


45

Fig. 11 All scales represent 100 μm. 1. Bolivina scalprata retiformis Cushman. Same specimen, a. apertural view, b.

lateral view, same scale, sample F11, Rupelian. 2. Bolivina fastigia Cushman. Same specimen, a. apertural view, b.

lateral view, same scale, sample F11, Rupelian. 3. Bolivina dilatata maxima Cicha & Zapletalová. Same specimen, a.

apertural view, b. lateral view, same scale, sample FB14, Chattian. 4. Bolivina dilatata hyalina Hofmann. Same

specimen, a. apertural view, b. lateral view, same scale, sample FB01, Chattian. 5-6. Bolivina dilatata dilatata Reuss.

5. sample F27, Rupelian; 6. sample F33, Rupelian/Chattian. 7. Bolivina antiqua d’Orbigny. Sample F23, Rupelian. 8.

Bolivina tereta (Cushman). Sample F38, Chattian. 9. Bolivina semistriata semistriata Hantken. Same specimen, a.

apertural view, b. lateral view, same scale, sample F15, Rupelian. 10. Bolivina molassica Hofmann. Same specimen, a.

apertural view, b. lateral view, same scale, sample F23, Rupelian. 11. Bolivina semistriata nobilis Hantken. Same

specimen, a. apertural view, b. lateral view, same scale, sample F4, Rupelian. 12. Bolivina pokornyi serrata Cicha &

Zapletalová. Same specimen, a. apertural view, b. lateral view, same scale, sample F13, Rupelian. 13. Cassidulinoides

bodeni Hagn. Same specimen, a. lateral view, b. apertural view, same scale, sample F20, Rupelian. 14.

Globocassidulina subglobosa (Brady). Sample F12, Rupelian. 15. Loxostomoides zsigmondyi (Hantken). Sample F20,

Rupelian. 16. Globocassidulina globosa (Hantken). Sample F23, Rupelian.


46

Fig. 10.4a-b

Fissurina kapellensis Spiegler, 1974, p. 60, pl. 1, figs. 16-

17.

Pseudofissurina kapellensis (Spiegler) - De Man, 2006, p.

275, pl. 6, fig. 15.


Family GLANDULINIDAE Reuss, 1860

Subfamily GLANDULININAE Reuss, 1860

Genus Glandulina d’Orbigny, 1839

Glandulina ovula d’Orbigny, 1846

Fig. 10.6

Glandulina ovula d’Orbigny, 1846, p. 29, pl. 1, figs. 6-7.

Glandulina ovula d’Orbigny - Papp & Schmid, 1985, p.

21, pl. 2, figs. 1-9.


Order BULIMINIDA Fursenko, 1958

Suborder ROTALIINA Delage & Hérouard, 1896

Superfamily BOLIVINACEA Glassner, 1937

Family BOLIVINIDAE Glassner, 1937

Genus Bolivina d’Orbigny, 1839

Bolivina antiqua d’Orbigny, 1846

Fig. 11.7

Bolivina antiqua d’Orbigny, 1846, p. 240, pl. 14, figs. 11-

13.


Chattian.

Bolivina beyrichi beyrichi Reuss, 1851

Fig. 10.8

Bolivina beyrichi Reuss, 1851, p. 83, pl. 6, fig. 51 (fide

Ellis & Messina).

Bolivina beyrichi beyrichi Reuss - Hofmann, 1967, p.

137, pl. 3, figs. 1-4.

Bolivina beyrichi Reuss - Popescu, 1975, p. 73, pl. 43,

figs. 1-2.


Chattian.

Bolivina beyrichi carinata Hantken, 1875

Fig. 10.7a-b

Bolivina beyrichi Reuss var. carinata Hantken, 1875, pl.

7, fig. 12.

Bolivina beyrichi carinata Hantken - Lindenberg, 1966,

p. 96, abb. 4/4.

Bolivina beyrichi carinata Hantken - Cicha et al., 1998,

p. 83, pl. 44, fig. 3.


Chattian.

Bolivina crenulata Cushman, 1936

Figs. 10.10, 10.11a-b

Bolivina crenulata Cushman, 1936, p. 50, pl. 7, fig. 13a-

b.

Bolivina crenulata crenulata Cushman - Hofmann, 1967,

p. 146, pl. 1, figs. 4-6.


Chattian.

Bolivina dilatata dilatata Reuss, 1850

Figs. 11.5-11.6

Bolivina dilatata Reuss, 1850, p. 381, pl. 48, fig. 15a-c


Bolivina dilatata dilatata Reuss - Cicha & Zapletalová,

1963, p. 131, textfigs. 11a-f.


Chattian.

Bolivina dilatata hyalina Hofmann, 1967

Fig. 11.4a-b

Bolivina dilatata hyalina Hofmann, 1967, p. 156, pl. 1,

figs. 7-9.


Chattian.

Bolivina dilatata maxima Cicha & Zapletalová, 1963

Fig. 11.3a-b

Bolivina dilatata maxima Cicha & Zapletalová, 1963, p.

136, textfig. 15.


Bolivina fastigia Cushman, 1936

Fig. 11.2a-b

Bolivina fastigia Cushman, 1936, p. 51, pl. 7, fig. 17 (fide

Ellis & Messina).


Chattian.

Bolivina molassica Hofmann, 1967

Fig. 11.10a-b

Bolivina molassica Hofmann, 1967, p. 158, pl. 4, figs. 9-

11.


Chattian.

Bolivina plicatella Cushman, 1930

Fig. 10.9

Bolivina plicatella Cushman, 1930, p. 46, pl. 8, fig. 10.


Bolivina pokornyi serrata Cicha & Zapletalová, 1963

Fig. 11.12a-b

Bolivina pokornyi serrata Cicha & Zapletalová, 1963, p.

144, textfig. 19.


Chattian.

Bolivina scalprata retiformis Cushman, 1936

Fig. 11.1a-b

Bolivina scalprata Schwager var. retiformis Cushman,

1936, p. 53, pl. 7, fig. 19.


Bolivina semistriata semistriata Hantken, 1875

Fig. 11.9a-b

Bolivina semistriata Hantken, 1875, p. 95, pl. 15, fig. 4.

Bolivina semistriata semistriata Hantken - Cicha et al.,

1998, p. 85, pl. 44, fig. 6.


Chattian.

Bolivina semistriata nobilis Hantken, 1875

Fig. 11.11a-b

Bolivina nobilis Hantken, 1875, p. 65, pl. 14, fig. 4.


47

Fig. 12 All scales represent 100 μm. 1,8. Bulimina coprolithoides Andreae. 1a-b. same specimen, a. apertural view, b.

lateral view, same scale, sample F14, Rupelian; 8. sample F33, Rupelian/Chattian. 2. Bulimina elongata d’Orbigny.

Same specimen, a. apertural view, b. lateral view, same scale, sample F17, Rupelian. 3-5. Caucasina oligocaenica

Khalilov. 3a-b. same specimen, a. apertural view, b. lateral view, same scale, sample F14, Rupelian; 4. sample F3,

Rupelian; 5. sample F12, Rupelian. 6-7. Bulimina alsatica Cushman & Parker. 6a-b. same specimen, a. apertural view, b.

lateral view, same scale, sample F7, Rupelian; 7. sample F5, Rupelian. 9. Bulimina schischkinskayae Samoylova. Sample

F27, Rupelian. 10. Bulimina alazanensis Cushman. Sample F18, Rupelian. 11. Bulimina striata striata d’Orbigny. Same

specimen, a. apertural view, b. lateral view, same scale, sample F7, Rupelian. 12. Bulimina arndti Hagn. Same specimen,

a. apertural view, b. lateral view, same scale, FB19, Chattian. 13. Praeglobobulimina pupoides (d’Orbigny). Sample F25,

Rupelian.


48

Bolivina semistriata nobilis Hantken - Cicha et al., 1998,

p. 85, pl. 44, fig. 7.


Chattian.

Bolivina tereta (Cushman, 1936)

Fig. 11.8

Loxostoma teretum Cushman, 1936, p. 60, pl. 8, fig. 14.

Loxostomum teretum (Cushman) - Batjes, 1958, p. 133,

pl. 5, fig. 17.

Bolivina tereta (Cushman) - Hofmann, 1967, p. 179, pl.

4, figs. 7-8.


Genus Loxostomoides Reiss, 1957

Loxostomoides zsigmondyi (Hantken, 1868)

Fig. 11.15

Nodosaria (Dentalina) zsigmondyi Hantken, 1868, p. 87,

pl. 1, fig. 12.

Dentalina zsigmondyi (Hantken ) - Hantken, 1875, p. 32,

pl. 12, fig. 17.

Rectobolivina zsigmondyi (Hantken) - Popescu & Iva,

1971, p. 44, pl. 5, fig. 6, pl. 6, fig. 2.

Loxostomoides zsigmondyi (Hantken) - Cicha et al., 1998,

p. 111, pl. 44, figs. 12-13.

Zsigmondiella zsigmondyi (Hantken) - Popescu & Crihan,

2005, p. 385, pl. 4, figs. 2-3.


Superfamily CASSIDULINACEA d’Orbigny, 1839

Family CASSIDULINIDAE d'Orbigny, 1839

Subfamily CASSIDULININAE d’Orbigny, 1839

Genus Cassidulinoides Cushman, 1927

Cassidulinoides bodeni Hagn, 1952

Fig. 11.13a-b

Cassidulinoides bodeni Hagn, 1952, p. 181, pl. 2, fig. 12.


Chattian.

Genus Globocassidulina Voloshinova, 1960

Globocassidulina globosa (Hantken, 1875)

Fig. 11.16

Cassidulina globosa Hantken, 1875, p. 54, pl. 16, fig. 2.

Globocassidulina globosa (Hantken) - Popescu & Iva,

1971, p. 12, pl. 7, figs. 2-3.

Globocassidulina globosa (Hantken) - Cicha et al., 1998,

p. 102, pl. 45, figs. 12-14.


Chattian.

Globocassidulina subglobosa (Brady, 1881)

Fig. 11.14

Cassidulina subglobosa Brady, 1881, p. 60, pl. 54, fig.

17a-c.

Globocassidulina subglobosa (Brady) - Sztrákos, 1979,

pl. 27, fig. 16.


Chattian.

Superfamily BULIMINACEA Jones, 1875

Family BULIMINIDAE Jones, 1875

Genus Bulimina d’Orbigny, 1826

Bulimina alazanensis Cushman, 1927b

Fig. 12.10

Bulimina alazanensis Cushman, 1927b, p. 161, pl. 25,

fig. 4


Bulimina alsatica Cushman & Parker, 1937

Figs. 12.6a-b, 12.7

Bulimina alsatica Cushman & Parker, 1937, p. 39, pl. 4,

figs. 6-7.


Chattian.

Bulimina arndti Hagn, 1952

Fig. 12.12a-b

Bulimina arndti Hagn, 1952, p. 168, pl. 2, fig. 15.


Chattian.

Bulimina coprolithoides Andreae, 1884

Figs. 12.1a-b, 12.8

Bulimina coprolithoides Andreae, 1884, p. 305, pl. 6, fig.

4.

Caucasina coprolithoides (Andreae) - Le Calvez, 1966,

p. 408, pl. 1, figs. 7-8.

Bulimina coprolithoides Andreae - Cicha et al., 1998, p.

86, pl. 47, fig. 1.


Chattian.

Bulimina elongata d’Orbigny, 1846

Fig. 12.2a-b

Bulimina elongata d’Orbigny, 1846, p. 187, pl. 11, figs.

19-20.

Caucasina elongata (d’Orbigny) - Loeblich & Tappan,

1964, p. 82, pl. 2, figs. 14-15.

Bulimina elongata elongata d’Orbigny - Didkowski &

Satanovskaja, 1970, p. 129, pl. 77, fig. 2.

Baggatella elongata (d’Orbigny) - Popescu, 1975, p. 71,

pl. 49, fig. 1.


Chattian.

Bulimina schischkinskayae Samoylova, 1947

Fig. 12.9

Bulimina schischkinskye Samoylova, 1947, p. 82, pl. 10


Caucasina schischkinskayae (Samoylova) - Khalilov,

1958, p. 4.

Baggatella schischkinskayae (Samoylova) - Popescu &

Iva, 1971, p. 43, pl. 11, figs. 1-5.

Bulimina schischkinskayae Samoylova - Cicha et al.,

1998, p. 87, pl. 47, figs. 2-4.


Chattian.

Bulimina striata striata d’Orbigny, 1843

Fig. 12.11a-b

Bulimina striata d’Orbigny in Guérin-Méneville, 1843, p.

18, pl. 3, fig. 16.


49

Fig. 13 All scales represent 100 μm.1. Uvigerina moravia Boersma. Same specimen, a. apertural view, b. lateral view,

same scale, sample F4, Rupelian. 2-5. Uvigerina popescui Rögl. 2a-b. same specimen, a. lateral view, b. lateral view,

same scale, sample F6, Rupelian; 3a-b. same specimen, a. apertural view, b. apertural view, same scale, sample F6,

Rupelian; 4a-b. same specimen, a. apertural view, b. lateral view, same scale, sample FB20, Chattian; 5a-b. same

specimen, a. apertural view, b. lateral view, same scale, sample F5, Rupelian. 6. Uvigerina vicksburgensis Cushman &

Ellisor. Same specimen, a. apertural view, b. lateral view, same scale, sample F14, Rupelian. 7. Uvigerina steyri Papp.

Same specimen, a. apertural view, b. lateral view, same scale, sample FB21, Chattian. 8. Uvigerina gracilis Reuss. Same

specimen, a. apertural view, b. lateral view, same scale, sample FB21, Chattian. 9. Angulogerina globosa (Stoltz). Same

specimen, a. apertural view, b. lateral view, same scale, sample F5, Rupelian. 10,13. Angulogerina angulosa

(Williamson). 10a-b. same specimen, a. apertural view, b. lateral view, same scale, sample F14, Rupelian; 13a-b. same

specimen, a. apertural view, b. lateral view, same scale, sample F9, Rupelian.11. Uvigerina tenuistriata schlichti

Schudack & Nuglisch. Sample F3, Rupelian. 12. Uvigerina gracilis var. oligocaenica (Andreae). Sample F34,

Rupelian/Chattian.


50

Bulimina buchiana d’Orbigny, 1846, p. 186, pl. 11, figs.

15-18.

Bulimina costata d’Orbigny, 1852, p. 194.

Bulimina inflata Seguenza var. mexicana Cushman,

1922, p. 95, pl. 21, fig. 2.

Bulimina striata d’Orbigny var. mexicana Cushman,

1940, p. 16, pl. 3, fig. 9.

Bulimina australis Vella, 1957, p. 32, pl. 8, figs. 168-169.

Bulimina striata striata d’Orbigny - Cicha et al., 1998, p.

87, pl. 48, figs. 1-3.


Chattian.

Genus Praeglobobulimina Hofker, 1951

Praeglobobulimina pupoides (d’Orbigny, 1846)

Fig. 12.13

Bulimina pupoides d’Orbigny, 1846, p. 185, pl. 11, figs.

11-12.

Praeglobobulimina pupoides (d’Orbigny) - Foraminiferi

Padani, AGIP, 1982, p. 122, pl. 6, fig. 12.

Protoglobobulimina pupoides (d’Orbigny) - Loeblich &

Tappan, 1988, p. 522, pl. 572, figs. 1-6.


Chattian.

Family UVIGERINIDAE Haeckel, 1894

Subfamily UVIGERININAE Haeckel, 1894

Genus Uvigerina d’Orbigny, 1826

Uvigerina moravia Boersma, 1984

Fig. 13.1a-b

Uvigerina moravia Boersma, 1984, p. 114, textfig. p.

114, figs. 1-4.


Chattian.

Uvigerina popescui Rögl, 1998

Figs. 13.2a-b, 13.3a-b, 13.4a-b, 13.5a-b

Uvigerina popescui Rögl, 1998, p. 74, pl. 52, figs. 1-3.


Chattian.

Uvigerina steyri Papp, 1975

Fig. 13.7a-b

Uvigerina steyri steyri Papp, 1975, p. 282, pl. 1, figs. 5-6.

Uvigerina steyri Papp - Cicha et al., 1998, p. 135, pl. 50,

figs. 10-12.


Chattian.

Uvigerina vicksburgensis Cushman & Ellisor, 1931

Fig. 13.6a-b

Uvigerina vicksburgensis Cushman & Ellisor, 1931, p.

54, pl. 7, fig. 7.


Uvigerina gracilis Reuss, 1851

Fig. 13.8a-b

Uvigerina gracilis Reuss, 1851, p. 77, pl. 5, fig. 39.

Angulogerina gracilis (Reuss) - Batjes, 1958, p. 134, pl.

6, figs. 1-2.

Trifarina gracilis (Reuss) - Sztrákos, 1979, pl. 20, fig. 10.

Uvigerina gracilis Reuss - Cicha et al., 1998, p. 133, pl.

49, fig. 11.


Chattian.

Uvigerina gracilis var. oligocaenica (Andreae, 1894)

Fig. 13.12

Uvigerina oligocaenica Andreae, 1894, p. 50, fig. 1.

Angulogerina oligocaenica (Andreae) - Cushman &

Edwards, 1938, p. 86, pl. 15, figs. 8-11.

Uvigerina gracilis (Reuss) var. oligocaenica (Andreae) -

Batjes, 1958, p. 135, pl. 6, fig. 3.


Chattian.

Uvigerina tenuistriata schlichti Schudack & Nuglisch,

2007

Fig. 13.11

Uvigerina tenuistriata schlichti Schudack & Nuglisch,

2007, p. 41, pl. 14, figs. 245-250.


Subfamily ANGULOGERININAE Galloway, 1933

Genus Angulogerina Cushman, 1927

Angulogerina angulosa (Williamson, 1858)

Figs. 13.10a-b, 13.13a-b

Uvigerina angulosa Williamson, 1858, p. 67, pl. 5, fig.

140.

Angulogerina angulosa (Williamson) - Cushman, 1949,

p. 111, pl. 6, figs. 9-10.

Trifarina angulosa (Williamson) - Popescu, 1979, p. 36,

pl. 21, fig. 6.

Angulogerina angulosa (Williamson) - Loeblich &

Tappan, 1987, pl. 574, figs. 5-9.


Angulogerina globosa (Stoltz, 1925)

Fig. 13.9a-b

Uvigerina tenuistriata d’Orbigny var. globosa Stoltz,

1925, p. 130, fig. 2 (fide Ellis & Messina).

Trifarina globosa (Stoltz) - Sztrákos, 1979, p. 67, pl. 20,

figs. 13-14.

Angulogerina globosa (Stoltz) - Mathelin & Sztrákos,

1993, p. 48, pl. 34, fig. 20.


Chattian.

Superfamily FURSENKOINACEA Loeblich &

Tappan, 1961

Family FURSENKOINIDAE Loeblich & Tappan, 1961

Genus Fursenkoina Loeblich & Tappan, 1961

Fursenkoina halkyardi (Cushman, 1936)

Figs. 14.5a-b, 14.6a-b, 14.7-14.10

Virgulina schreibersiana Halkyard, 1918, p. 48, pl. 8, fig.


Virgulina halkyardi Cushman, 1936, p. 47, pl. 7. Fig. 5.

Fursenkoina halkyardi (Cushman) - Popescu & Iva,

1971, pl. 12, fig. 3.


Chattian.


51

Fig. 14 All scales represent 100 μm. 1. Fursenkoina schreibersiana (Czjzek). Same specimen, a. apertural view, b. lateral view,

same scale, sample F14, Rupelian. 2,3. Fursenkoina mustoni (Andreae). 2a-b. same specimen, a. apertural view, b. lateral view,

same scale, sample F3, Rupelian; 3. sample F3, Rupelian. 4. Fursenkoina sp. Same specimen, a. apertural view, b. lateral view,

same scale, sample F3, Rupelian. 5-10. Fursenkoina halkyardi (Cushman). 5. same specimen, a. apertural view, b. lateral view,

same scale; 6. same specimen, a. apertural view, b. lateral view; same scale. 5-10. sample FB01, Chattian. 11. Pleurostomella

incrassata Hantken. Same specimen, a. lateral view, b. apertural view, same scale, sample F5, Rupelian. 12-14. Stilostomella

adolphina (d’Orbigny). 12. sample FB20, Chattian; 13. sample F14, Rupelian; 14. sample F18, Rupelian. 15, 17, 18, 21.

Stilostomella emaciata (Reuss). 15. sample F9, Rupelian; 17. sample F9, Rupelian; 18. sample F21, Rupelian; 21. sample F25,

Rupelian. 16. Siphonodosaria consobrina (d’Orbigny). Sample F3, Rupelian. 19. Stilostomella pauperata (d’Orbigny). Sample

F21, Rupelian. 20. Stilostomella bradyi (Cushman). Sample F5, Rupelian.


52

Fursenkoina mustoni (Andreae, 1884)

Figs. 14.2a-b, 14.3

Virgulina mustoni Andreae, 1884, p. 254, pl. 11, fig. 4.

Fursenkoina mustoni (Andreae) - Reiser, 1987, p. 80, pl.

7, figs. 4, 9.


Chattian.

Fursenkoina schreibersiana (Czjzek, 1848)

Fig. 14.1a-b

Virgulina schreibersiana Czjzek, 1848, p. 147, pl. 13,


Stainforthia schreibersiana (Czjzek) - Kümmerle, 1963,

p. 40, pl. 5, fig. 10a-b.

Fursenkoina schreibersiana (Czjzek) - Belford, 1966, p.

136, pl. 9, figs. 18-21.


Chattian.

Fursenkoina sp.

Fig. 14.4a-b


Superfamily DELOSINACEA Parr, 1950

Family CAUCASINIDAE Bykova, 1959

Subfamily CAUCASININAE Bykova, 1959

Genus Caucasina Khalilov, 1951

Caucasina oligocaenica Khalilov, 1951

Figs. 12.3a-b, 12.4-12.5

Caucasina oligocaenica Khalilov, 1951, p. 58, pl. 4, fig.


Baggatella oligocaenica (Khalilov) - Popescu & Iva,

1971, p. 43, pl. 10, figs. 2-7.


Chattian.

Superfamily PLEUROSTOMELLACEA Reuss, 1860

Family PLEUROSTOMELLIDAE Reuss, 1860

Subfamily PLEUROSTOMELLINAE Reuss, 1860

Genus Pleurostomella Reuss, 1860

Pleurostomella incrassata Hantken, 1883

Fig. 14.11a-b

Pleurostomella incrassata Hantken, 1883, p. 25, pl. 1,

figs. 4, 7.

Pleurostomella alternans Schwager var. moravica

Schubert, 1900, p. 157, pl. 2, figs. 5a-c.

Pleurostomella velascoensis Cushman, 1926, p. 590, pl.

16, fig. 4.

Pleurostomella paleocenica Cushman, 1947, p. 86, pl.

18, figs. 14-15.


Superfamily STILOSTOMELLACEA Finlay, 1947

Family STILOSTOMELLIDAE Finlay, 1947

Genus Siphonodosaria Silvestri, 1924

Siphonodosaria consobrina (d’Orbigny, 1846)

Fig. 14.16

Dentalina consobrina d’Orbigny, 1846, p. 46, pl. 2, figs.

1-3.

Dentalina verneuilii d’Orbigny, 1846, p. 48, pl. 2, figs. 7-

8.

Dentalina aequalis Karrer, 1864, p. 74, pl. 16, fig. 1.

Nodosaria consobrina (d’Orbigny) - Reuss & Schlicht,

1870, p. 473/29, 31-32, pl. 9, figs. 1, 8, 22, pl. 10, figs. 1,

25.

Nodosaria japonica Cushman, 1913, p. 57, pl. 28, fig. 4.

Ellipsonodosaria verneuilii (d’Orbigny) var. emaciata

Palmer & Bermúdez, 1941, p. 189.

Siphonodosaria consobrina (d’Orbigny) - Stainforth,

1952, p. 12.

Stilostomella consobrina (d’Orbigny) - Stainforth, 1952,

p. 146.

Stilostomella consobrina (d’Orbigny) - Beckmann, 1953,

p. 370, pl. 21, figs. 24-25.


Chattian.

Genus Stilostomella Guppy, 1894

Stilostomella adolphina (d’Orbigny, 1846)

Figs. 14.12-14.14

Dentalina adolphina d’Orbigny, 1846, p. 51, pl. 2, figs.

18-20.

Dentalina ornata Neugeboren, 1856, pl. 4, figs. 9a-b.

Siphonodosaria adolphina (d’Orbigny) - Subbotina,

1953, p. 180, pl. 6, figs. 1-2.

Stilostomella adolphina (d’Orbigny) - Rögl, 1968, p. 80,

pl. 3, fig. 12.

Nodosaria adolphina (d’Orbigny) - Krayeva & Zerneckij,

1969, p. 41, pl. 14, fig. 6.

Nodogenerina adolphina (d’Orbigny) - Mathelin &

Sztrákos, 1993, pl. 35, fig. 17.

Caveastomella adolphina (d’Orbigny) – Hayward et al.,

2012, p. 163, pl. 16, figs. 1-6.


Chattian.

Stilostomella bradyi (Cushman, 1927c)

Fig. 14.20

Sagrina virgula Brady, 1879, p. 275, pl. 8, figs. 19-21.

Nodogenerina bradyi Cushman, 1927c, p. 89, pl. 76, fig.


Stilostomella bradyi (Cushman) - Barker, 1960, pl. 76,

fig. 8.

Nodogenerina virgula (Brady) - Jones, 1994, pl. 76, fig.

8.

Siphonodosaria bradyi (Cushman) - Hayward &

Kawagata, 2005, pl. 1, fig. 31.


Chattian.

Stilostomella emaciata (Reuss, 1851)

Figs. 14.15, 14.17, 14.18, 14.21

Dentalina emaciata Reuss, 1851, p. 63, pl. 3, fig. 9.

Nodosaria emaciata (Reuss) - Batjes, 1958, p. 116, pl. 3,

figs. 20-21.

Stilostomella emaciata (Reuss) - Sztrákos, 1979, pl. 19,

fig. 14.

Stilostomella emaciata (Reuss) - Cicha et al., 1998, p.

128, pl. 56, figs. 10-11.


Chattian.


53

Fig. 15 All scales represent 100 μm.1. Neoeponides schreibersi (d’Orbigny). Sample F4, Rupelian. 2. ?Gyroidina constans

(Reiser). Same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, sample F14, Rupelian. 3.

Cibicidoides ungerianus filicosta (Hagn). Same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale,

sample F14, Rupelian. 4. Cibicidoides acutus (Samoylova). Sample F10, Rupelian. 5. Cibicides amphysiliensis (Andreae).

Same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, FB18, Chattian. 6. Cibicidoides oligocenicus

(Samoylova). Same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, sample F14, Rupelian. 7.

Epistominella molassica (Hagn). Same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, sample F14,

Rupelian. 8. Fontbotia wuellerstorfi (Schwager). Same specimen, a. umbilical view, b. spiral view, same scale, sample F11,

Rupelian. 9. Epistominella exigua (Brady). Sample F14, Rupelian.


54

Stilostomella pauperata (d’Orbigny, 1846)

Fig. 14.19

Dentalina pauperata d’Orbigny, 1846, p. 46, pl. 1, figs.

57-58.

Siphonodosaria pauperata (d’Orbigny) - Stainforth,

1952, p. 13.

Stilostomella pauperata (d’Orbigny) - Stainforth, 1952,

p. 146.

Stilostomella pauperata (d’Orbigny) - Sztrákos, 1979, pl.

19, fig. 13.


Order ROTALIIDA Lankester, 1885

Superfamily DISCORBACEA Ehrenberg, 1838

Family DISCORBIDAE Ehrenberg, 1838

Genus Neoeponides Reiss, 1960

Neoeponides schreibersi (d’Orbigny, 1846)

Fig. 15.1

Rotalina schreibersi d’Orbigny, 1846, p. 154, pl. 8, figs.

4-6.

Eponides schreibersi (d’Orbigny) - Kaasschieter, 1961, p.

210, pl. 11, figs. 14-15.

Neoeponides schreibersi (d’Orbigny) - Popescu, 1975, p.

96.


Superfamily DISCORBINELLACEA Sigal, 1952

Family PARRELLOIDIDAE Hofker, 1956

Genus Cibicidoides Thalmann, 1939

Cibicidoides acutus (Samoylova, 1947)

Fig. 15.4

Cibicides acutus Samoylova, 1947, p. 97, textfigs. 40-42


Cibicidoides acutus (Samoylova) - Popescu & Iva, 1971,

p. 48, pl. 9, fig. 1.


Cibicidoides oligocenicus (Samoylova, 1947)

Fig. 15.6a-c

Cibicides dutemplei (d’Orbigny) var. oligocenica

Samoylova, 1947, p. 96, figs. 34-36.

Cibicides oligocenicus (Samoylova) - Pozaryska, 1977 p.

39, pl. 6. figs. 1–6; pl. 7, fig. 3a–c; pl. 13, figs. 8–9.

Cibicidoides oligocenicus (Samoylova) - Cicha et al.,

1998, p. 90, pl. 62, figs. 1-3.


Cibicidoides ungerianus filicosta (Hagn, 1952)

Fig. 15.3a-c

Cibicides ungerianus (d’Orbigny) var. filicosta Hagn,

1952, p. 186, pl. 1, fig. 4.

Cibicides ungerianus filicosta Hagn - Hagn, 1960, pl. 7,

fig. 9.

Cibicidoides ungerianus filicosta (Hagn) - Reiser, 1987,

p. 117, pl. 19, figs. 4, 7-8.


Chattian.

Family PSEUDOPARRELLIDAE Voloshinova, 1952

Subfamily PSEUDOPARRELLINAE Voloshinova,

1952

Genus Epistominella Husezima & Maruhasi, 1944

Epistominella exigua (Brady, 1884)

Fig. 15.9

Pulvinulina exigua Brady, 1884, p. 696, pl. 103, figs. 13-

14.

Pseudoparrella exigua (Brady) - Phleger & Parker, 1951,

p. 28, pl. 15, fig. 6.

Pulvinulinella exigua (Brady) - Hofker, 1951, p. 322,

textfigs. 219-221.

Epistominella exigua (Brady) - Barker, 1960, pl. 103,

figs. 13-14.

Alabamina exigua (Brady) - Popescu, 1975, p. 102, pl.

83, fig. 5.

Alabaminoides exiguus (Brady) - Jones, 1994, p. 103, pl.

103, figs. 13-14.

Pseudoparrella exigua (Brady) - Loeblich & Tappan,

1994, p. 146, pl. 307, figs. 1-7.


Chattian.

Epistominella molassica (Hagn, 1952)

Fig. 15.7a-c

Pseudoparella molassica Hagn, 1952, p. 180, pl. 2, fig.

13.

Epistominella molassica (Hagn) - Reiser, 1987, p. 100,

pl. 11, figs. 14-15, pl. 12, fig. 1.


Chattian.

Superfamily PLANORBULINACEA Schwager, 1877

Family PLANULINIDAE Bermúdez, 1952

Genus Planulina d’Orbigny, 1826

Planulina compressa (Hantken, 1875)

Fig. 16.1a-c

Truncatulina compressa Hantken, 1875, p. 62, pl. 8, fig.

8.

Planulina compressa (Hantken) - Hagn, 1956, p. 179, pl.

17, fig. 5a-b.


Chattian.

Planulina costata (Hantken, 1875)

Fig. 16.2a-b

Truncatulina costata Hantken, 1875, p. 63, pl. 9, fig. 2.

Planulina costata (Hantken) - Hagn, 1956, p. 178, pl. 17,

figs. 3a-b, 4a-b.


Chattian.

Family CIBICIDIDAE Cushman, 1927

Subfamily CIBICIDINAE Cushman, 1927

Genus Cibicides de Montfort, 1808

Cibicides amphysiliensis (Andreae, 1884)

Fig. 15.5a-c

Truncatulina amphysiliensis Andreae, 1884, p. 255, pl.

10, fig. 30.

Cibicides amphysiliensis (Andreae) - Cicha et al., 1998,

p. 90, pl. 72, figs. 14-17.


Chattian.


55

Fig. 16 All scales represent 100 μm. 1. Planulina compressa (Hantken). Same specimen, a. umbilical view, b. peripherial view, c.

spiral view, same scale, sample FB20, Chattian. 2. Planulina costata (Hantken). Same specimen, a. umbilical view, b. spiral view,

same scale, sample F12, Rupelian. 3. Asterigerinoides guerichi (Franke). Same specimen, a. umbilical view, b. spiral view, same

scale, sample F18, Rupelian. 4. Nonion commune (d’Orbigny). Same specimen, a. lateral view, b. peripherial view, same scale,

sample F4, Rupelian. 5. Pullenia salisburyii Stewart & Stewart. Same specimen, a. lateral view, b. peripherial view, same scale,

sample F10, Rupelian. 6. Chilostomella cylindroides Reuss. Sample F34, Rupelian/Chattian. 7. Quadrimorphina petrolei

(Andreae). Same specimen, a. umbilical view, b. spiral view, same scale, sample F18, Rupelian. 8-9. Alabamina tangentialis

(Clodius). 8a-b. same specimen, a. umbilical view, b. spiral view, same scale, sample FB01, Chattian; 9a-b. same specimen, a.

umbilical view, b. spiral view, same scale, sample FB21, Chattian. 10. Anomalinoides granosus (Hantken). Same specimen, a.

umbilical view, b. spiral view, same scale, sample F15, Rupelian


56

Genus Fontbotia Gonzales-Donoso & Linares, 1970

Fontbotia wuellerstorfi (Schwager, 1866)

Fig. 15.8a-b

Anomalina wuellerstorfi Schwager, 1866, p. 258, pl. 7,

figs. 105-107.

Truncatulina wuellerstorfi (Schwager) - Brady, 1884, pp.

627, 662, pl. 93, figs. 8-9.

Planorbulina wuellerstorfi (Schwager) - Goës, 1894, p.

89, pl. 15, fig. 777.

Planulina wuellerstorfi (Schwager) - Cushman, 1931, p.

110, pl. 19, figs. 5-6.

Cibicides wuellerstorfi (Schwager) - Leroy, 1941, p. 46,

pl. 1, figs. 27-29.

Fontbotia wuellerstorfi (Schwager) - Gonzales-Donoso &

Linares, 1970, p. 238, pl. 1, fig. 4.

Cibicidoides wuellerstorfi (Schwager) - Mackensen et al.,

1993, p. 57, pl. 2, figs. 4-5.


Chattian.

Superfamily ASTERIGERINACEA d'Orbigny, 1839

Family ASTERIGERINATIDAE Reiss, 1963

Genus Asterigerinoides Bermúdez, 1952

Asterigerinoides guerichi (Franke, 1912)

Fig. 16.3a-b

Discorbina guerichi Franke, 1912, p. 29, fig. 8.

Asterigerina guerichi (Franke) - Dam & Reinhold, 1942,

p. 90, pl. 7, fig. 1.

Asterigerinoides guerichi (Franke) - Hofker, 1959, p.

255, abb. 17.

Asterigerina guerichi guerichi (Franke) - Ellerman, 1960,

p. 687, pl. 53, fig. 17.


Superfamily NONIONACEA Schultze, 1854

Family NONIONIDAE Schultze, 1854

Subfamily NONIONINAE Schultze, 1854

Genus Nonion de Montfort, 1808

Nonion commune (d’Orbigny, 1826)

Fig. 16.4a-b

Nautilus scapha Fichtel & Moll, 1798, p. 105, pl. 19, fig.

d-f.

Cristellaria scapha (Fichtel & Moll) - Lamarck, 1822, p.

608.

Nonionina communis d’Orbigny, 1826, p. 128, no. 20.

Polystomella (Nonionina) scapha (Fichtel & Moll) -

Parker & Jones, 1860, p. 102.

Nonion scaphum (Fichtel & Moll) - Cushman, 1939, p.

20, pl. 5, fig. 18-21.

Nonion commune (d’Orbigny) - Cushman, 1939, p. 10,

pl. 3, fig. 2.

Nonion boueanus (d’Orbigny) - Voloshinova, 1952, p.

18, pl. 1, figs. 3-5.

Florilus communis (d’Orbigny) - Cicha & Zapletalová,

1967, p. 136.

Florilus boueanus (d’Orbigny) - Didkowski &

Satanovskaja, 1970, p. 102, pl. 65, fig. 3


Subfamily PULLENIINAE Schwager, 1877

Genus Pullenia Parker & Jones,1862

Pullenia salisburyii Stewart & Stewart, 1930

Fig. 16.5a-b

Pullenia salisburyii R. E & K. C. Stewart, 1930, p. 72, pl.

8, fig. 2 (fide Ellis & Messina).


Chattian.

Superfamily CHILOSTOMELLACEA Brady, 1881

Family CHILOSTOMELLIDAE Brady, 1881

Subfamily CHILOSTOMELLINAE Brady, 1881

Genus Chilostomella Reuss, 1849

Chilostomella cylindroides Reuss, 1851

Fig. 16.6

Chilostomella cylindroides Reuss, 1851, p. 80, pl. 6, fig.

43.


Chattian.

Family QUADRIMORPHINIDAE Saidova, 1981

Genus Quadrimorphina Finlay, 1939

Quadrimorphina petrolei (Andreae, 1884)

Fig. 16.7a-b

Pulvinulina petrolei Andreae, 1884, p. 125, pl. 8, fig. 15.

Valvulineria petrolei (Andreae) - Batjes, 1958, p. 146, pl.

8, fig. 1.

Quadrimorphina petrolei (Andreae) - Cicha et al., 1998,

p. 121, pl. 66, figs. 16-18.


Chattian.

Family ALABAMINIDAE Hofker, 1951

Genus Alabamina Toulmin, 1941

Alabamina tangentialis (Clodius, 1922)

Figs. 16.8a-b, 16.9a-b

Pulvinulina tangentialis Clodius, 1922, p. 138, pl. 1, fig.

14.

Alabamina tangentialis (Clodius) - Batjes, 1958, p. 155,

pl. 8, fig. 7.


Family HETEROLEPIDAE Gonzales-Donoso, 1969

Genus Anomalinoides Brotzen, 1942

Anomalinoides granosus (Hantken, 1875)

Fig. 16.10a-b

Truncatulina granosa Hantken, 1875, p. 65, pl. 10, fig. 2.

Anomalina granosa (Hantken) - Sztrákos, 1979, pl. 29,

fig. 9.

Anomalinoides granosus (Hantken) - Cicha et al., 1998,

p. 81, pl. 69, figs. 14-18.


Chattian.

Genus Heterolepa Franzenau, 1884

Heterolepa eocaena (Gümbel, 1868)

Figs. 17.3a-b, 17.4a-b

Rotalia eocaena Gümbel, 1868, p. 650, pl. 2, fig. 87.


57

Fig. 17 All scales represent 100 μm. 1-2. Heterolepa praecincta (Karrer). 1a-c. same specimen, a. umbilical view, b.

peripherial view, c. spiral view, same scale, sample F14, Rupelian; 2a-c. same specimen, a. umbilical view, b. peripherial

view, c. spiral view, same scale, sample FB20, Chattian. 3-4. Heterolepa eocaena (Gümbel). 3a-b. same specimen, a.

umbilical view, b. spiral view, same scale, sample F12, Rupelian; 4a-b. same specimen, a. umbilical view, b. spiral view,

same scale, sample FB21, Chattian. 5. Gyroidinoides octocameratus (Cushman & Hanna). Same specimen, a. umbilical

view, b. peripherial view, c. spiral view, same scale, sample FB21, Chattian. 6. Gyroidinoides mamillatus (Andreae). Same

specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, sample F5, Rupelian. 7-8. Hansenisca soldanii

(d’Orbigny). 7a-c. same specimen, a. umbilical view, b. peripherial view, c. spiral view, same scale, sample F14, Rupelian; 8.

sample F25, Rupelian.


58

Cibicides eocaenus (Gümbel) - Hagn, 1956, p. 180, pl.

17, figs. 8-9.

Heterolepa eocaena (Gümbel) - Grünig, 1985, p. 275, pl.

11, figs. 1-3.

Cibicidoides eocaenus (Gümbel) - Van Morkhoven et al.,

1986, p. 256, pl. 86A, figs. 1–4; pl. 86B, figs. 1–2; pl.

86C, 1–3 (cum syn).


Chattian.

Heterolepa praecincta (Karrer, 1868)

Figs. 17.1a-c, 17.2a-c

Rotalina praecincta Karrer, 1868, p. 189, pl. 5, fig. 7.

Truncatulina praecincta (Karrer) - Brady, 1884, p. 667,

pl. 95, figs. 1-3.

Cibicides dutemplei (d’Orbigny) var. praecinctus

(Karrer) - Batjes, 1958, p. 150, pl. 9,

fig.10.

Heterolepa praecincta (Karrer) - Popescu, 1975, p. 105,

pl. 85, fig. 3.

Cibicidoides (?) praecinctus (Karrer) - McCulloch, 1977,

p. 446, pl. 152, fig. 11.

Cibicides praecinctus (Karrer) - Tu & Zheng, 1991, p.

186, pl. 9, fig. 3.

Neoeponides praecinctus (Karrer) - Jones, 1994, p. 99, pl.

95, figs. 1-3.

Heterolepa praecincta (Karrer) - Loeblich & Tappan,

1994, p. 163, pl. 360, figs. 1-10.

Cibicidoides praecinctus (Karrer) - De Man, 2006, p.

291, pl. 12, fig. 1.


Chattian.

Family GAVELINELLIDAE Hofker, 1956

Subfamily GYROIDINOIDINAE Saidova, 1981

Genus Gyroidinoides Brotzen, 1942

Gyroidinoides octocameratus (Cushman and Hanna,

1927)

Fig. 17.5a-c

Gyroidina soldanii d’Orbigny var. octocamerata

Cushman and Hanna, 1927, p. 223, pl. 14, figs. 16-18.

Gyroidina soldanii d’Orbigny octocamerata Cushman &

Hanna - Willems, 1972, p. 39, pl. 5, fig. 8.

Gyroidinoides octocameratus (Cushman and Hanna) -

Cicha et al., 1998, p. 105, pl. 71,

figs. 20-22.


Chattian.

Gyroidinoides mamillatus (Andreae, 1884)

Fig. 17.6a-c

Rotalia girardana Reuss var. mamillata Andreae, 1884,

p. 142, pl. 9, fig. 4.

Gyroidina mamillata (Andreae) - Dam & Reinhold, 1942,

p. 87, pl. 6, fig. 6.

Gyroidina soldanii d’Orbigny var. mamillata Andreae -

Batjes, 1958, p. 147, pl.7,

fig. 15.

Rotaliatina bulimoides mamillata (Andreae) - Ellerman,

1960, p. 675.

Gyroidina soldanii mamillata (Andreae) - King, 1989, p.

478, pl. 9.7, fig. 17.

Gyroidinoides mamillatus (Andreae) - Cicha et al., 1998,

p. 105, pl. 72, figs. 1-3.


Chattian.

Subfamily GAVELINELLINAE Hofker, 1956

Genus Gyroidina d’Orbigny, 1826

?Gyroidina constans (Reiser, 1987)

Fig. 15.2a-c

Eponides constans Reiser, 1987, p. 98, pl. 11, figs. 13,

16-17.

Gyroidina ? constans (Reiser) - Cicha et al., 1998, p. 105,

pl. 58, figs. 4-6.


Chattian.

Genus Hansenisca Loeblich and Tappan, 1987

Hansenisca soldanii (d’Orbigny, 1826)

Figs. 17.7a-c, 17.8

Gyroidina soldanii d’Orbigny, 1826, p. 112, no. 5.

Rotalina soldanii (d’Orbigny) - d’Orbigny, 1846, p. 155,

pl. 8, figs. 10-12.

Rotalia soldanii (d’Orbigny) - Andreae, 1884, p. 155-

156, 257, 262, pl. 9, fig. 3.

Gyroidinoides soldanii (d’Orbigny) - Braga et al., 1975,

p. 109, pl. 6, figs. 10-11.

Gyroidinoides soldanii (d’Orbigny) - Popescu, 1975, p.

103, pl. 84, fig. 2a-c.

Hansenisca soldanii (d’Orbigny) - Loeblich & Tappan,

1987, pl. 719, figs. 5-9.


Chattian.

ACKNOWLEDGEMENTS

We thank the reviewers Katarína Holcová, Ileana-Monica

Crihan and Gheorghe Popescu for the review and

comments on the manuscript.

This paper is a result of a doctoral research made possible

by the financial support of the Sectoral Operational

Programme for Human Resources Development 2007-

2013, co-financed by the European Social Fund, under

the project POSDRU/159/1.5/S/133391 - “Doctoral and

postdoctoral excellence programs for training highly

qualified human resources for research in the fields of

Life Sciences, Environment and Earth”. Additional

financial support was provided by the Grzybowski

Foundation (the Brian J. O’Neill Memorial Grant-in-

Aid), Sciex (Grant No. 12.116), and S.N.G.N Romgaz

(Contract 18/2011).

The authors are grateful for Christoph Neururer

(University of Fribourg, Switzerland) for technical help

in photographing the foraminifera.

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Documents

TAXONOMIC RECORD OF THE OLIGOCENE BENTHIC … · 2018. 12. 14. · megasequence. According to Filipescu (2001) and Krézsek & Bally (2006), the deposition of the early Oligocene bituminous