43
THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1 , DAVID J. MIDDLETON 1 , THAWEESAKDI BOONKERD 2 & SOMRAN SUDDEE 3 ABSTRACT. There have been many changes to family and genus delimitation in Thai ferns since the publication of volume 3 (Pteridophytes) of the Flora of Thailand. These changes are discussed and current names for all taxa, including those recently described or recorded for Thailand, are presented. A new combination is made in Hymenasplenium. KEY WORDS: Ferns, Flora of Thailand, nomenclature, Thailand. INTRODUCTION There are approximately 670 taxa of ferns in Thailand comprising about 5–7% of the total vascular flora (Middleton, 2003). The ferns were published in volume 3 of the Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989) and 596 taxa were recognised as occurring in Thailand (excluding the lycophytes: Lycopodiaceae, Selaginellaceae and Isoetaceae). A number of papers have since been published in which new taxa have been described or new records have been added to the pteridophyte diversity of Thailand (Mitsuta, 1985; Iwatsuki et al., 1998; Parris, 1998a; Hovenkamp et al., 1998; Nooteboom, 1998; Boonkerd & Nooteboom, 2001; Boonkerd & Pollawatn 2000, 2002a, 2002b, 2006; Lindsay & Middleton, 2004, 2009c; Lindsay et al., 2004, 2008; Suksathan, 2004; Boonkerd et al., 2004; Boonkerd, 2006). Of particular significance amongst these are Boonkerd & Pollawatn (2000), which is a list of all the pteridophytes in Thailand (including an additional 27 taxa) with distribution maps and many photographs, and Boonkerd et al. (2004), in which many more species were added. A total of 71 fern taxa have been newly recorded or newly described for Thailand since the publication of the Flora of Thailand accounts. This rate of addition of new taxa and new records to a recently completed Flora account reflects the fact that pteridophytes had previously often been neglected on traditional collecting expeditions (and were therefore not available to Tagawa & Iwatsuki) and is testament to the relatively poor state of our taxonomic knowledge of them compared to many angiosperm plant groups. The taxonomy of ferns in Thailand has also been subject to considerable flux due to differences of opinion over the delimitation of families, genera and species. One need only glance at the lists of synonymy to see that very many species have had homotypic combinations made in many different genera (e.g. several species of Selliguea also have combinations in Polypodium, Phymatopsis, Pleopeltis, Phymatodes, Crypsinus ________________________________________________________________________________________________________________________________________________________ 1 Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, UK. 2 Department of Botany, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. 3 The Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, Chatuchak, Bangkok 10900, Thailand.

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Page 1: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

THAI FOR. BULL. (BOT.) 37: 64–106. 2009.

Towards a stable nomenclature for Thai ferns

STUART LINDSAY1, DAVID J. MIDDLETON1, THAWEESAKDI BOONKERD2 & SOMRAN SUDDEE3

ABSTRACT. There have been many changes to family and genus delimitation in Thai ferns since the publication of volume 3 (Pteridophytes) of the Flora of Thailand. These changes are discussed and current names for all taxa, including those recently described or recorded for Thailand, are presented. A new combination is made in Hymenasplenium.

KEY WORDS: Ferns, Flora of Thailand, nomenclature, Thailand.

INTRODUCTION

There are approximately 670 taxa of ferns in Thailand comprising about 5–7% of the total vascular flora (Middleton, 2003). The ferns were published in volume 3 of the Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989) and 596 taxa were recognised as occurring in Thailand (excluding the lycophytes: Lycopodiaceae, Selaginellaceae and Isoetaceae). A number of papers have since been published in which new taxa have been described or new records have been added to the pteridophyte diversity of Thailand (Mitsuta, 1985; Iwatsuki et al., 1998; Parris, 1998a; Hovenkamp et al., 1998; Nooteboom, 1998; Boonkerd & Nooteboom, 2001; Boonkerd & Pollawatn 2000, 2002a, 2002b, 2006; Lindsay & Middleton, 2004, 2009c; Lindsay et al., 2004, 2008; Suksathan, 2004; Boonkerd et al., 2004; Boonkerd, 2006). Of particular significance amongst these are Boonkerd & Pollawatn (2000), which is a list of all the pteridophytes in Thailand (including an additional 27 taxa) with distribution maps and many photographs, and Boonkerd et al. (2004), in which many more species were added. A total of 71 fern taxa have been newly recorded or newly described for Thailand since the publication of the Flora of Thailand accounts. This rate of addition of new taxa and new records to a recently completed Flora account reflects the fact that pteridophytes had previously often been neglected on traditional collecting expeditions (and were therefore not available to Tagawa & Iwatsuki) and is testament to the relatively poor state of our taxonomic knowledge of them compared to many angiosperm plant groups.

The taxonomy of ferns in Thailand has also been subject to considerable flux due to differences of opinion over the delimitation of families, genera and species. One need only glance at the lists of synonymy to see that very many species have had homotypic combinations made in many different genera (e.g. several species of Selliguea also have combinations in Polypodium, Phymatopsis, Pleopeltis, Phymatodes, Crypsinus

________________________________________________________________________________________________________________________________________________________

1 Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, UK.2 Department of Botany, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand.3 The Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, Chatuchak, Bangkok 10900, Thailand.

Page 2: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

65TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

and Phymatopteris - see Lindsay & Middleton, 2009a). This indecision about the best placement of species in genera is also manifest at the family level where the limits of several families have changed considerably over time and between contemporary Flora projects, leading, in some cases, to the abandonment of families altogether in some parts of Flora Malesiana (e.g. Kramer, 1971; Holttum, 1978, 1991) in favour of “groups” of genera.

In recent years phylogenetic techniques, particularly using molecular data, have been used to attempt to solve the many problems in fern familial and generic delimitation (see Smith et al., 2006, 2008 and references cited therein). This has led to a greater consensus on the delimitation of families and genera. We are eventually beginning to see the possibility of stability in the names applied to the ferns of Thailand.

THE TABLES

In this paper we apply the family and genus concepts summarised by Smith et al. (2006, 2008) to the ferns of Thailand and present the data in two tables. For Davalliaceae we follow the more recent generic concepts of Kato & Tsutsumi (2008). In Table 1 the families and genera recognised by Smith et al. (2006, 2008) and Kato & Tsutsumi (2008) are compared to the families and genera recognised in the Flora of Thailand accounts. In Table 2 all fern taxa now known to occur in Thailand are listed using the up-to-date family and genus concepts from Smith et al. (2006, 2008) and Kato & Tsutsumi (2008), with occasional modifications from other recent works (e.g. Parris, 2007). These are compared to the names used for the same taxon from the Flora of Thailand and from Boonkerd & Pollawatn (2000). For species delimitation we follow Tagawa & Iwatsuki (1979, 1985, 1988, 1989), unless other pieces of taxonomic work have superseded these publications (e.g. Hovenkamp et al., 1998), or we have come to our own differing opinion in specific groups. In some cases the authorship of names is not the same as has been previously published. We have checked the problematic ones and only the correct authorships are used without highlighting the discrepancies. Likewise the spelling of specific epithets not in conformity with the International Code of Botanical Nomenclature (McNeill et al., 2006) has been corrected in all columns without comment. We include taxa which have been published as occurring in Thailand. We also include a small number of taxa which we provisionally record for Thailand from our own observations but which still await formal publication. We acknowledge that this list might be incomplete judging by the rate at which taxa have been newly recorded from Thailand in the last 20 years. However, this list has been compiled to facilitate the development of an on-line multi-access key to the ferns of Thailand (Lindsay & Middleton, 2009b) and new taxa can easily be added to this.

The “nine new records” in Mitsuta (1985) were not included in the Flora of Thailand. Some of them are, however, attributable to species already known from Thailand. We have included a genuine additional record (Monachosorum henryi Christ) and as yet unconfirmed new records (Thelypteris laxa (Franch. & Sav.) Ching and Diplazium doerderleinii (Luerss.) Makino) in Table 2. The new record Diplazium okinawaense Tagawa has been synonymised to Diplazium virescens Kunze following Shieh et al. (1994). We have redetermined the other material cited in Mitsuta (1985) to taxa already reported in the Flora of Thailand, thus: Colysis flexiloba (Christ) Ching var. undulato-repanda (C.Chr.) Ching to Leptochilus ellipticus (Thunb.) Noot.; Crypsinus trisectus

Page 3: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

66 THaI FOreST BuLLeTIN (BOTaNY) 37

(Baker) Tagawa to Selliguea hirsuta (Baker) S.Linds.; Microlepia trichoclada Ching to Microlepia herbacea Ching & C.Chr. ex Tardieu & C.Chr.; Thelypteris angulariloba Ching to Thelypteris hirsutipes (C.B.Clarke) Ching; and Thelypteris esquirolii (Christ) Ching to Cyclosorus falcilobus (Hook.) Panigrahi.

Major changes in family delimitations

Table 1 summarises the changes in family delimitation between the Flora of Thailand accounts and those currently recognised and lists the genera in each of the families. These changes include losing some familiar families altogether, such as Parkeriaceae (now in Pteridaceae) and Athyriaceae (now mostly in Woodsiaceae), and changing the delimitation of others quite radically, such as Polypodiaceae (which now includes Grammitidaceae) and Dryopteridaceae (which has lost Tectaria and its relatives and gained genera from Lomariopsidaceae, Davalliaceae and Athyriaceae). Pteridaceae has also undergone major changes and is now much larger with five subfamilies which do not correspond well to former families; e.g. Pteridaceae subfamily Pteridoideae includes only Pteris from the previously delimited Pteridaceae, with all of the other genera previously having been placed in Parkeriaceae.

Major changes in generic delimitation

Table 2 lists all the currently used names for Thai ferns and compares them to Pteridophytes in Thailand (Boonkerd & Pollawatn, 2000) and Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989). A total of 191 taxa have a name which is now different to the name used in the Flora of Thailand accounts, most of these as a result of changes in generic delimitation. Changes in generic delimitation are particularly extensive in Davalliaceae, Hymenophyllaceae, Polypodiaceae and Thelypteridaceae. Well known genera such as Grammitis, Polypodium, Trichomanes and Vittaria are no longer recognised in Thailand. Table 2 also provides references to the literature on new taxa, new records and in cases where the taxon epithet is different to that used in the Flora of Thailand account (usually due to synonymy). If there is no reference given then the change is made here. Black dots are used to indicate a taxon which is under a different name to that used in the original Flora of Thailand account; circles are used to indicate a taxon which is new to Thailand, either as a newly described taxon or a new record.

ACKNOWLEDGEMENTS

We thank The Leverhulme Trust for financial support for this work; the Curators and Collection Managers of herbaria that have loaned material or welcomed us on visits (notably a, BK, BKF, BM, C, CMu, e, K, SING); the library staff of the royal Botanic Garden edinburgh for their help in locating or obtaining protologues and other references; and Christopher Fraser-Jenkins, John McNeill, Barbara Parris, alan Smith and John Thompson for advice and information.

Page 4: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

67TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

Tabl

e 1.

Com

paris

on o

f fam

ily d

elim

itatio

n an

d co

mpo

sitio

n be

twee

n Sm

ith e

t al.

(200

6, 2

008)

and

Flo

ra o

f Tha

iland

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9). N

ote

that

with

in P

terid

acea

e the

subf

amily

nam

e Vitt

ario

idea

e has

prio

rity

over

Adi

anto

idea

e (cf

. Sm

ith et

al.,

2006

, 200

8) an

d th

e sub

fam

ilial

nam

e Cry

ptog

ram

moi

deae

,

w

hich

did

not

exi

st fo

r Sm

ith e

t al.

(200

6, 2

008)

, has

now

bee

n pu

blis

hed

(Lin

dsay

& M

iddl

eton

, 200

9a).

Smith

et a

l. (2

006,

200

8) fa

mily

Gen

era

curr

ently

rec

ogni

zed

in T

haila

nd.

Tho

se in

bo

ld (2

6) w

ere

not r

ecog

nize

d in

Flo

ra o

f Tha

iland

Cor

resp

ondi

ng fa

mily

(or

fam

ilies

) in

Flor

a of

Tha

iland

(T

agaw

a &

Iwat

suki

, 197

9, 1

985,

198

8, 1

989)

ASP

LeN

IaC

eae

Aspl

eniu

m, H

ymen

aspl

eniu

mA

SPLe

NIa

Cea

e

BLe

CH

Na

Cea

eBl

echn

um, B

rain

ea, S

teno

chla

ena,

Woo

dwar

dia

BLe

CH

Na

Cea

e (B

lech

num

, Bra

inea

, Woo

dwar

dia)

, PT

ERID

AC

EAE

(Ste

noch

laen

a)

CIB

OTI

AC

EAE

Cib

otiu

mD

ICK

SON

IaC

eae

CYA

THEA

CEA

EC

yath

eaC

YATH

EAC

EAE

DAV

ALL

IAC

EAE

Ara

iost

egie

lla, D

aval

lia, D

aval

lode

s, H

umat

a, W

ibel

iaD

AVA

LLIA

CEA

E

DeN

NST

aeD

TIa

Cea

eD

enns

taed

tia, H

istio

pter

is, H

ypol

epis

, Mic

role

pia,

Pt

erid

ium

DeN

NST

aeD

TIa

Cea

e

DIP

TER

IDA

CEA

EC

heiro

pleu

ria,

Dip

teri

sD

IPTE

RID

AC

EAE

(Dip

teri

s), C

HEI

RO

PLEU

RIA

CEA

E (C

heiro

pleu

ria)

DRY

OPT

ERID

AC

EAE

Acro

phor

us, A

rach

niod

es, B

olbi

tis, C

teni

tis, C

yrto

miu

m,

Pera

nem

a, D

idym

ochl

aena

, Dry

opte

ris,

Elap

hogl

ossu

m,

Hyp

odem

atiu

m, L

euco

steg

ia, L

omag

ram

ma,

Pol

ystic

hum

, Te

rato

phyl

lum

DRY

OPT

ERID

AC

EAE

(Acr

opho

rus,

Arac

hnio

des,

Cte

nitis

, C

yrto

miu

m,

Did

ymoc

hlae

na, D

ryop

teri

s, Po

lyst

ichu

m),

ATH

YR

IAC

EAE

(Hyp

odem

atiu

m),

DAV

ALL

IAC

EAE

(Leu

cost

egia

), LO

Ma

rIO

PSID

aC

eae

(Bol

bitis

, Ela

phog

loss

um, L

omag

ram

ma,

Te

rato

phyl

lum

)

EQU

ISET

AC

EAE

Equi

setu

mEQ

UIS

ETA

CEA

E

GLe

ICH

eNIa

Cea

eD

icra

nopt

eris

, Dip

lopt

eryg

ium

, Gle

iche

nia,

Stic

heru

sG

LeIC

HeN

IaC

eae

HY

MeN

OPH

YLL

aC

eae

Abro

dict

yum

, Cal

listo

pter

is, C

epha

lom

anes

, Cre

pido

man

es,

Did

ymog

loss

um, H

ymen

ophy

llum

, Van

denb

osch

iaH

YM

eNO

PHY

LLa

Cea

e

LIN

DSa

eaC

eae

Lind

saea

, Sph

enom

eris

, Tap

eini

dium

LIN

DSa

eaC

eae

LOM

ar

IOPS

IDa

Cea

eC

yclo

pelti

s, Lo

mar

iops

is, N

ephr

olep

isLO

Ma

rIO

PSID

aC

eae

(Lom

ario

psis

), D

RYO

PTER

IDA

CEA

E (C

yclo

pelti

s), O

Lea

ND

ra

Cea

e (N

ephr

olep

is)

LYG

OD

IAC

EAE

Lygo

dium

SCH

IZA

EAC

EAE

MA

RAT

TIA

CEA

EAn

giop

teri

s, C

hris

tens

enia

, Ptis

ana

MA

RAT

TIA

CEA

E

Page 5: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

68 THaI FOreST BuLLeTIN (BOTaNY) 37

Smith

et a

l. (2

006,

200

8) fa

mily

Gen

era

curr

ently

rec

ogni

zed

in T

haila

nd.

Tho

se in

bol

d (2

6) w

ere

not r

ecog

nize

d in

Flo

ra o

f Tha

iland

Cor

resp

ondi

ng fa

mily

(or

fam

ilies

) in

Flor

a of

Tha

iland

(T

agaw

a &

Iwat

suki

, 197

9, 1

985,

198

8, 1

989)

MA

RSI

LEA

CEA

EM

arsi

lea

MA

RSI

LEA

CEA

E

MaT

ON

IaC

eae

Mat

onia

Abs

ent

OLe

aN

Dr

aC

eae

Ole

andr

aO

Lea

ND

ra

Cea

e

OPH

IOG

LOSS

AC

EAE

Botr

ychi

um, H

elm

inth

osta

chys

, Oph

iogl

ossu

mO

PHIO

GLO

SSA

CEA

E

OSM

uN

Da

Cea

eO

smun

daO

SMu

ND

aC

eae

PLA

GIO

GY

RIA

CEA

EPl

agio

gyri

aPL

AG

IOG

YR

IAC

EAE

POLY

POD

IAC

EAE

Acro

soru

s, Ag

laom

orph

a, A

rthr

omer

is, B

elvi

sia,

C

alym

mod

on, C

hris

tiopt

eris

, Chr

ysog

ram

miti

s, Ct

enop

tere

lla, D

asyg

ram

miti

s, D

ryna

ria, G

onio

phle

bium

, G

ymno

gram

miti

s, Le

cano

pter

is, L

emm

aphy

llum

, Le

piso

rus,

Lept

ochi

lus,

Loxo

gram

me,

Mic

roso

rum

, N

eoch

eiro

pter

is, O

reog

ram

miti

s, Pl

atyc

eriu

m,

Pros

aptia

, Pyr

rosi

a, R

adio

gram

miti

s, Sc

lero

glos

sum

, Se

lligu

ea, T

hem

eliu

m, T

omop

hyllu

m, X

ipho

pter

ella

POLY

POD

IAC

EAE

(Agl

aom

orph

a, A

rthr

omer

is,

Belv

isia

, Chr

istio

pter

is, C

olys

is, D

ryna

ria,

Lem

map

hyllu

m,

Lepi

soru

s, Le

ptoc

hilu

s, Lo

xogr

amm

e, M

icro

soru

m,

Neo

chei

ropt

eris

, Pla

tyce

rium

, Pyr

rosi

a, S

ellig

uea)

, D

AVA

LLIA

CEA

E (G

ymno

gram

miti

s), G

ra

MM

ITID

aC

eae

(Acr

osor

us, C

alym

mod

on, P

rosa

ptia

, Scl

erog

loss

um)

PSIL

OTA

CEA

EPs

ilotu

mPS

ILO

TAC

EAE

PTER

IDA

CEA

E su

bfam

. Che

ilant

hoid

eae

Che

ilant

hes,

Dor

yopt

eris

, Hem

ioni

tis, N

otho

laen

a,

Para

hem

ioni

tis, P

ella

eaPA

RK

ERIA

CEA

E

PTER

IDA

CEA

E su

bfam

. Cry

ptog

ram

moi

deae

Con

iogr

amm

ePA

RK

ERIA

CEA

E

PTER

IDA

CEA

E su

bfam

. Par

kerio

idea

eAc

rost

ichu

m, C

erat

opte

ris

PTER

IDA

CEA

E (A

cros

tichu

m),

PAR

KER

IAC

EAE

(Cer

atop

teri

s)

PTER

IDA

CEA

E su

bfam

. Pte

ridoi

deae

Ony

chiu

m, P

ityro

gram

ma,

Pte

ris,

Syng

ram

ma,

Tae

nitis

PAR

KER

IAC

EAE

(Ony

chiu

m, P

ityro

gram

ma,

Syn

gram

ma,

Ta

eniti

s), P

TER

IDA

CEA

E (P

teri

s)

PTER

IDA

CEA

E su

bfam

. Vitt

ario

idea

eAd

iant

um, A

ntro

phyu

m, H

aplo

pter

is, V

agin

ular

iaPA

RK

ERIA

CEA

E (A

dian

tum

), V

ITTA

RIA

CEA

E (A

ntro

phyu

m, V

agin

ular

ia, V

ittar

ia)

SaLV

INIa

Cea

eAz

olla

, Sal

vini

aSa

LVIN

IaC

eae

(Sal

vini

a), A

ZOLL

AC

EAE

(Azo

lla)

SCH

IZA

EAC

EAE

Schi

zaea

SCH

IZA

EAC

EAE

TEC

TAR

IAC

EAE

Arth

ropt

eris

, Het

erog

oniu

m, P

leoc

nem

ia, P

teri

drys

, Tec

tari

aD

RYO

PTER

IDA

CEA

E (H

eter

ogon

ium

, Ple

ocne

mia

, Pt

erid

rys,

Tect

aria

), O

Lea

ND

ra

Cea

e (A

rthr

opte

ris)

THEL

YPT

ERID

AC

EAE

Cycl

osor

us, M

acro

thel

ypte

ris, P

seud

ophe

gopt

eris,

The

lypt

eris

THEL

YPT

ERID

AC

EAE

WO

OD

SIA

CEA

EAt

hyri

um, C

orno

pter

is, D

epar

ia, D

ipla

zium

ATH

YR

IAC

EAE

Page 6: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

69TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

Tabl

e 2.

Com

paris

on o

f the

cur

rent

nam

es fo

r Tha

i fer

n sp

ecie

s w

ith th

e na

mes

use

d fo

r the

sam

e ta

xa in

Pte

ridop

hyte

s in

Tha

iland

(Boo

nker

d &

Pol

law

atn,

200

0) a

nd

Fl

ora

of T

haila

nd (T

agaw

a &

Iwat

suki

, 197

9, 1

985,

198

8, 1

989)

. Tw

o na

mes

for a

sing

le e

ntry

in th

e th

ird c

olum

n lin

ked

by a

‘/’ i

ndic

ate

that

a c

hang

e w

as m

ade

to

the

nam

e in

the

“add

ition

s an

d C

orre

ctio

ns”

sect

ion

in p

art 4

of v

olum

e 3

of th

e Fl

ora

of T

haila

nd a

ccou

nt. T

wo

or m

ore

nam

es fo

r a s

ingl

e en

try li

nked

by

a

‘+’ i

ndic

ate

that

thes

e ta

xa a

re n

ow tr

eate

d as

syn

onym

s. B

lack

dot

s in

dica

te s

peci

es w

hose

nam

es h

ave

chan

ged

sinc

e th

e pu

blic

atio

n of

the

Flor

a of

Tha

iland

or w

ere

chan

ged

in th

e “a

dditi

ons

and

Cor

rect

ions

” se

ctio

n. C

ircle

s in

dica

te s

peci

es th

at h

ave

been

dis

cove

red

or d

escr

ibed

sin

ce th

e pu

blic

atio

n of

the

Flor

a of

Thai

land

. Th

e re

fere

nce

besi

de a

nam

e re

fers

to th

e pl

ace(

s) o

f pub

licat

ion

of th

e ne

w ta

xon

or re

cord

, or t

he so

urce

of i

nfor

mat

ion

abou

t the

cha

nge

of a

nam

e: 1

Boo

nker

d (2

006)

; 2 B

oonk

erd

(in p

ress

); 3

Boo

nker

d &

Noo

tebo

om (2

001)

; 4 B

oonk

erd

& P

olla

wat

n (2

000)

; 5 B

oonk

erd

& P

olla

wat

n (2

002a

); 6

Boo

nker

d &

Polla

wat

n (2

002b

); 7

Boo

nker

d &

Pol

law

atn

(200

4); 8

Boo

nker

d &

Pol

law

atn

(200

6); 9

Boo

nker

d &

Suk

sath

an (

2009

); 10

Boo

nker

d et

al.

(200

4); 1

1 Fr

aser

-

Jenk

ins

(199

7); 1

2 H

olttu

m (

1982

); 13

Hol

ttum

(19

91);

14 H

olttu

m &

Grim

es (

1980

); 15

Hov

enka

mp

(198

6); 1

6 H

oven

kam

p et

al.

(199

8); 1

7 H

oven

kam

p

& M

iyam

oto

(200

5); 1

8 Iw

atsu

ki e

t al.

(199

8); 1

9 K

ato

& T

suts

umi (

2008

); 20

Khu

llar (

1994

); 21

Lin

dsay

& M

iddl

eton

(200

4); 2

2 Li

ndsa

y &

Mid

dlet

on (2

009c

);

23 L

inds

ay e

t al.

(200

4); 2

4 Li

ndsa

y et

al.

(200

8); 2

5 Lu

(199

9); 2

6 M

itsut

a (1

985)

; 27

Mur

dock

(200

8); 2

8 N

akai

ke (1

992)

; 29

Noo

tebo

om (1

997)

; 30

Noo

tebo

om

(1

998)

; 31

Parr

is (

1998

a); 3

2 Pa

rris

(19

98b)

; 33

Parr

is (

2007

); 34

Par

ris &

Lat

iff (

1997

); 35

Pet

chsr

i, B

oonk

erd

& B

aum

(20

09);

36 r

ödl-L

inde

r (1

990)

; 37

Su

ksat

han

(200

4); 3

8 Th

omps

on (2

008)

; 39

Zhan

g (1

999a

); 40

Zha

ng (1

999b

); 41

B. P

arris

, per

s. co

mm

.; 42

J. T

hom

pson

, per

s. co

mm

.; 43

Adi

antu

m p

hilip

pens

e

var.

subj

unon

icum

is re

porte

d fo

r Tha

iland

by

Boo

nker

d &

Pol

law

atn

(200

6) b

ut th

e com

bina

tion

has n

ot b

een

valid

ly p

ublis

hed

and

the s

tatu

s of t

he v

arie

ties o

f thi

s

spec

ies

need

furth

er re

sear

ch; 4

4 Pr

ovis

iona

l new

reco

rds,

curr

ently

und

er in

vest

igat

ion;

45

Hym

enas

plen

ium

exc

isum

(C.P

resl

) S.L

inds

., co

mb.

nov

. Bas

iony

m:

As

plen

ium

exc

isum

C.P

resl

, Epi

mel

. Bot

. 74

(185

1 [‘

1849

’]).

Alth

ough

the

nam

e H

ymen

aspl

eniu

m e

xcis

um h

as b

een

used

in r

ecen

t lite

ratu

re w

e ca

n fin

d no

evid

ence

that

the

com

bina

tion

has p

revi

ousl

y be

en v

alid

ly p

ublis

hed;

46

taxo

nom

ic c

hang

e by

cur

rent

aut

hors

; 47

Ebih

ara

et a

l. (2

006)

.

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

ASP

LE

NIA

CE

AE

Aspl

eniu

m a

ffine

Sw.

Aspl

eniu

m a

ffine

Sw.

Aspl

eniu

m a

ffine

Sw

.

Aspl

eniu

m a

ntro

phyo

ides

Chr

ist

Aspl

eniu

m a

ntro

phyo

ides

Chr

ist

Aspl

eniu

m a

ntro

phyo

ides

Chr

ist

Aspl

eniu

m b

atue

nse A

lder

w.As

plen

ium

bat

uens

e A

lder

w.As

plen

ium

bat

uens

e A

lder

w.

Aspl

eniu

m c

onfu

sum

Tar

dieu

& C

hing

Aspl

eniu

m c

onfu

sum

Tar

dieu

& C

hing

Aspl

eniu

m c

onfu

sum

Tar

dieu

& C

hing

Aspl

eniu

m c

rini

caul

e H

ance

Aspl

eniu

m c

rini

caul

e H

ance

Aspl

eniu

m c

rini

caul

e H

ance

○As

plen

ium

dec

resc

ens K

unze

44A

bsen

tA

bsen

t

Aspl

eniu

m d

elav

ayi (

Fran

ch.)

Cop

el.

Aspl

eniu

m d

elav

ayi (

Fran

ch.)

Cop

el.

Aspl

eniu

m d

elav

ayi (

Fran

ch.)

Cop

el.

Aspl

eniu

m e

nsifo

rme

Wal

l. ex

Hoo

k. &

Gre

v.As

plen

ium

ens

iform

e W

all.

ex H

ook.

& G

rev.

Aspl

eniu

m e

nsifo

rme

Wal

l. ex

Hoo

k. &

Gre

v.

Aspl

eniu

m e

xigu

um B

edd.

Aspl

eniu

m e

xigu

um B

edd.

Aspl

eniu

m e

xigu

um B

edd.

○As

plen

ium

finl

ayso

nian

um W

all.

ex H

ook.

10A

bsen

tA

bsen

t

Aspl

eniu

m g

revi

llei W

all.

ex H

ook.

& G

rev.

Aspl

eniu

m g

revi

llei W

all.

ex H

ook.

& G

rev.

Aspl

eniu

m g

revi

llei W

all.

ex H

ook.

& G

rev.

Page 7: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

70 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

○As

plen

ium

gri

ffith

ianu

m H

ook.

44A

bsen

tA

bsen

t

○As

plen

ium

gue

inzi

anum

Met

t. ex

Kuh

n.9

Abs

ent

Abs

ent

Aspl

eniu

m h

umbe

rtii

Tard

ieu

Aspl

eniu

m h

umbe

rtii

Tard

ieu

Aspl

eniu

m h

umbe

rtii

Tard

ieu

○As

plen

ium

inae

quila

tera

le W

illd.

2A

bsen

tA

bsen

t

Aspl

eniu

m in

terj

ectu

m C

hris

tAs

plen

ium

inte

rjec

tum

Chr

ist

Aspl

eniu

m in

terj

ectu

m C

hris

t

○As

plen

ium

kha

sian

um S

ledg

e.44

Abs

ent

Abs

ent

●As

plen

ium

laci

niat

um D

.Don

.20As

plen

ium

var

ians

Wal

l. ex

Hoo

k. &

Gre

v.As

plen

ium

var

ians

Wal

l. ex

Hoo

k. &

Gre

v.

Aspl

eniu

m lo

ngis

sim

um B

lum

eAs

plen

ium

long

issi

mum

Blu

me

Aspl

eniu

m lo

ngis

sim

um B

lum

e

Aspl

eniu

m m

acro

phyl

lum

Sw

.As

plen

ium

mac

roph

yllu

m S

w.As

plen

ium

mac

roph

yllu

m S

w.

Aspl

eniu

m n

idus

L. v

ar. n

idus

As

plen

ium

nid

us L

. var

. nid

usAs

plen

ium

nid

us L

. var

. nid

us

Aspl

eniu

m n

idus

L. v

ar. m

usifo

lium

(Met

t.) C

.Chr

.As

plen

ium

nid

us L

. var

. mus

ifoliu

m (M

ett.)

C.C

hr.

Aspl

eniu

m n

idus

L. v

ar. m

usifo

lium

(Met

t.) C

.Chr

.

Aspl

eniu

m n

itidu

m S

w.

Aspl

eniu

m n

itidu

m S

w.As

plen

ium

niti

dum

Sw.

Aspl

eniu

m n

orm

ale

D.D

on

Aspl

eniu

m n

orm

ale

D.D

onAs

plen

ium

nor

mal

e D

.Don

Aspl

eniu

m p

arad

oxum

Blu

me

Aspl

eniu

m p

arad

oxum

Blu

me

Aspl

eniu

m p

arad

oxum

Blu

me

Aspl

eniu

m p

ellu

cidu

m L

am.

Aspl

eniu

m p

ellu

cidu

m L

am.

Aspl

eniu

m p

ellu

cidu

m L

am.

Aspl

eniu

m p

erak

ense

C.G

.Mat

thew

& C

hris

tAs

plen

ium

per

aken

se C

.G.M

atth

ew &

Chr

ist

Aspl

eniu

m p

erak

ense

C.G

.Mat

thew

& C

hris

t

Aspl

eniu

m p

hylli

tidis

D.D

on. s

ubsp

. phy

llitid

is

Aspl

eniu

m p

hylli

tidis

D.D

on. s

ubsp

. phy

llitid

isAs

plen

ium

phy

llitid

is D

.Don

. sub

sp. p

hylli

tidis

Aspl

eniu

m p

hylli

tidis

D.D

on. s

ubsp

. m

ales

icum

Hol

ttum

As

plen

ium

phy

llitid

is D

.Don

. sub

sp.

mal

esic

um H

olttu

mAs

plen

ium

phy

llitid

is D

.Don

. sub

sp.

mal

esic

um H

olttu

m

●As

plen

ium

pol

yodo

n G

.For

st.34

Aspl

eniu

m fa

lcat

um L

am.

Aspl

eniu

m fa

lcat

um L

am.

Aspl

eniu

m ro

ckii

C.C

hr.

Aspl

eniu

m ro

ckii

C.C

hr.

Aspl

eniu

m ro

ckii

C.C

hr.

Aspl

eniu

m sa

lignu

m B

lum

e As

plen

ium

salig

num

Blu

me

Aspl

eniu

m sa

lignu

m B

lum

e

Aspl

eniu

m sc

orte

chin

ii B

edd.

As

plen

ium

scor

tech

inii

Bed

d.As

plen

ium

scor

tech

inii

Bed

d.

Aspl

eniu

m si

amen

se T

agaw

a &

K.Iw

ats.

Aspl

eniu

m si

amen

se T

agaw

a &

K.Iw

ats.

Aspl

eniu

m si

amen

se T

agaw

a &

K.Iw

ats.

Aspl

eniu

m si

mon

sian

um H

ook.

As

plen

ium

sim

onsi

anum

Hoo

k.As

plen

ium

sim

onsi

anum

Hoo

k.

Aspl

eniu

m te

neru

m G

.For

st.

Aspl

eniu

m te

neru

m G

.For

st.

Aspl

eniu

m te

neru

m G

.For

st.

Page 8: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

71TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Aspl

eniu

m te

nuifo

lium

D.D

on

Aspl

eniu

m te

nuifo

lium

D.D

onAs

plen

ium

tenu

ifoliu

m D

.Don

Aspl

eniu

m th

unbe

rgii

Kun

ze

Aspl

eniu

m th

unbe

rgii

Kun

zeAs

plen

ium

thun

berg

ii K

unze

●As

plen

ium

trun

catu

m B

lum

e.46

Aspl

eniu

m c

auda

tum

auc

t. no

n G

.For

st.

Aspl

eniu

m c

auda

tum

auc

t. no

n G

.For

st.

●As

plen

ium

vitt

aefo

rme

Cav

.34As

plen

ium

squa

mul

atum

Blu

me

Aspl

eniu

m sq

uam

ulat

um B

lum

e

Aspl

eniu

m y

oshi

naga

e M

akin

o As

plen

ium

yos

hina

gae

Mak

ino

Aspl

eniu

m y

oshi

naga

e M

akin

o

●H

ymen

aspl

eniu

m a

poga

mum

(N.M

urak

. & H

atan

.) N

akai

keAs

plen

ium

apo

gam

um N

.Mur

ak. &

Hat

an.

Aspl

eniu

m u

nila

tera

le a

uct.

non

Lam

. /

Aspl

eniu

m a

poga

mum

N.M

urak

. & H

atan

.

●H

ymen

aspl

eniu

m c

heilo

soru

m (K

unze

ex

Met

t.) T

agaw

aAs

plen

ium

che

iloso

rum

Kun

ze e

x M

ett.

Aspl

eniu

m c

heilo

soru

m K

unze

ex

Met

t.

Hym

enas

plen

ium

exc

isum

(C.P

resl

) S.L

inds

.45As

plen

ium

exc

isum

C.P

resl

Aspl

eniu

m e

xcis

um C

.Pre

sl

○H

ymen

aspl

eniu

m in

than

onen

se N

.Mur

ak. &

J.Yo

koy.

18H

ymen

aspl

eniu

m in

than

onen

se N

.Mur

ak. &

J.Yo

koy.

Abs

ent

●H

ymen

aspl

eniu

m o

bscu

rum

(Blu

me)

Tag

awa

Aspl

eniu

m o

bscu

rum

Blu

me

Aspl

eniu

m o

bscu

rum

Blu

me

BL

EC

HN

AC

EA

E

Blec

hnum

finl

ayso

nian

um W

all.

ex H

ook.

& G

rev.

Bl

echn

um fi

nlay

soni

anum

Wal

l. ex

Hoo

k. &

Gre

v.Bl

echn

um fi

nlay

soni

anum

Wal

l. ex

Hoo

k. &

Gre

v.

Blec

hnum

indi

cum

Bur

m.f.

Bl

echn

um in

dicu

m B

urm

.f.Bl

echn

um in

dicu

m B

urm

.f.

Blec

hnum

ori

enta

le L

. Bl

echn

um o

rien

tale

L.

Blec

hnum

ori

enta

le L

.

Brai

nea

insi

gnis

(Hoo

k.) J

.Sm

. Br

aine

a in

sign

is (H

ook.

) J.S

m.

Brai

nea

insi

gnis

(Hoo

k.) J

.Sm

.

Sten

ochl

aena

pal

ustr

is (B

urm

.f.) B

edd.

St

enoc

hlae

na p

alus

tris

(Bur

m.f.

) Bed

d.St

enoc

hlae

na p

alus

tris

(Bur

m.f.

) Bed

d.

○W

oodw

ardi

a ha

rlan

dii H

ook.

4,7

Woo

dwar

dia

harl

andi

i Hoo

k.A

bsen

t

●W

oodw

ardi

a ja

poni

ca (L

.f.) S

m.

Woo

dwar

dia

japo

nica

(L.f.

) Sm

.W

oodw

ardi

a co

chin

-chi

nens

is C

hing

CIB

OT

IAC

EA

E

Cib

otiu

m b

arom

etz (

L.) J

.Sm

. C

ibot

ium

bar

omet

z (L.

) J.S

m.

Cib

otiu

m b

arom

etz (

L.) J

.Sm

.

Page 9: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

72 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

CYA

TH

EA

CE

AE

Cya

thea

bor

neen

sis C

opel

. C

yath

ea b

orne

ensi

s Cop

el.

Cya

thea

bor

neen

sis C

opel

.

Cya

thea

chi

nens

is C

opel

. C

yath

ea c

hine

nsis

Cop

el.

Cya

thea

chi

nens

is C

opel

.

Cya

thea

con

tam

inan

s (W

all.

ex H

ook.

) Cop

el.

Cya

thea

con

tam

inan

s (W

all.

ex H

ook.

) Cop

el.

Cya

thea

con

tam

inan

s (W

all.

ex H

ook.

) Cop

el.

Cya

thea

gig

ante

a (W

all.

ex H

ook.

) Hol

ttum

C

yath

ea g

igan

tea

(Wal

l. ex

Hoo

k.) H

olttu

mC

yath

ea g

igan

tea

(Wal

l. ex

Hoo

k.) H

olttu

m

○C

yath

ea g

labr

a (B

lum

e) C

opel

.44A

bsen

tA

bsen

t

○C

yath

ea h

ymen

odes

Met

t.44A

bsen

tA

bsen

t

Cya

thea

late

bros

a (W

all.

ex H

ook.

) Cop

el.

Cya

thea

late

bros

a (W

all.

ex H

ook.

) Cop

el.

Cya

thea

late

bros

a (W

all.

ex H

ook.

) Cop

el.

○C

yath

ea m

oluc

cana

R.B

r.10A

bsen

tA

bsen

t

Cya

thea

pod

ophy

lla (H

ook.

) Cop

el.

Cya

thea

pod

ophy

lla (H

ook.

) Cop

el.

Cya

thea

pod

ophy

lla (H

ook.

) Cop

el.

Cya

thea

spin

ulos

a W

all.

ex H

ook.

C

yath

ea sp

inul

osa

Wal

l. ex

Hoo

k.C

yath

ea sp

inul

osa

Wal

l. ex

Hoo

k.

DAV

AL

LIA

CE

AE

●Ar

aios

tegi

ella

fabe

rian

a (C

.Chr

.) M

.Kat

o &

Tsu

tsum

i Ar

aios

tegi

a fa

beri

ana

(C.C

hr.)

Chi

ngAr

aios

tegi

a fa

beri

ana

(C.C

hr.)

Chi

ng

Dav

allia

pet

elot

ii Ta

rdie

u &

C.C

hr.

Dav

allia

pet

elot

ii Ta

rdie

u &

C.C

hr.

Dav

allia

pet

elot

ii Ta

rdie

u &

C.C

hr.

Dav

allia

solid

a (G

.For

st.)

Sw.

Dav

allia

solid

a (G

.For

st.)

Sw.

Dav

allia

solid

a (G

.For

st.)

Sw.

Dav

allia

tric

hom

anoi

des B

lum

e va

r. tr

icho

man

oide

s D

aval

lia tr

icho

man

oide

s Blu

me

var.

tric

hom

anoi

des

Dav

allia

tric

hom

anoi

des B

lum

e va

r. tr

icho

man

oide

s

Dav

allia

tric

hom

anoi

des B

lum

e va

r. lo

rrai

nii

(Han

ce) H

olttu

m

Dav

allia

tric

hom

anoi

des B

lum

e va

r. lo

rrai

nii

(Han

ce) H

olttu

mD

aval

lia tr

icho

man

oide

s Blu

me

var.

lorr

aini

i (H

ance

) Hol

ttum

○D

aval

lode

s bor

neen

se (H

ook.

) Cop

el.30

Abs

ent

Abs

ent

○D

aval

lode

s hym

enop

hyllo

ides

(Blu

me)

M.K

ato

&

Tsut

sum

i30,1

9A

bsen

tA

bsen

t

●D

aval

lode

s im

bric

atum

(Chi

ng) M

.Kat

o &

Tsu

tsum

iAr

aios

tegi

a im

bric

ata

Chi

ngAr

aios

tegi

a im

bric

ata

Chi

ng

Dav

allo

des m

embr

anul

osum

(Hoo

k.) C

opel

. D

aval

lode

s mem

bran

ulos

um (H

ook.

) Cop

el.

Dav

allo

des m

embr

anul

osum

(Hoo

k.) C

opel

.

Page 10: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

73TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●D

aval

lode

s pse

udoc

ysto

pter

is (K

unze

) M.K

ato

&

Tsut

sum

i Ar

aios

tegi

a ps

eudo

cyst

opte

ris (

Kun

ze) C

opel

.Ar

aios

tegi

a ps

eudo

cyst

opte

ris (

Kun

ze) C

opel

.

●D

aval

lode

s pul

chra

(D.D

on) M

.Kat

o &

Tsu

tsum

iAr

aios

tegi

a pu

lchr

a (D

.Don

) Cop

el.

Arai

oste

gia

pulc

hra

(D.D

on) C

opel

.

Dav

allo

des v

isci

dulu

m (K

uhn)

Ald

erw

. D

aval

lode

s vis

cidu

lum

(Kuh

n) A

lder

w.D

aval

lode

s vis

cidu

lum

(Kuh

n) A

lder

w.

Hum

ata

angu

stat

a (W

all.

ex H

ook.

& G

rev.

) J.S

m.

Hum

ata

angu

stat

a (W

all.

ex H

ook.

& G

rev.

) J.S

m.

Hum

ata

angu

stat

a (W

all.

ex H

ook.

& G

rev.

) J.S

m.

/ Pac

hypl

euri

a an

gust

ata

(Wal

l. ex

Hoo

k. &

G

rev.

) C.P

resl

●H

umat

a co

rnic

ulat

a (T

.Moo

re) M

.Kat

o &

Tsu

tsum

iD

aval

lia c

orni

cula

ta T

.Moo

reD

aval

lia c

orni

cula

ta T

.Moo

re

Hum

ata

hete

roph

ylla

(Sm

.) D

esv.

H

umat

a he

tero

phyl

la (S

m.)

Des

v.H

umat

a he

tero

phyl

la (S

m.)

Des

v. /

Dav

allia

het

erop

hylla

Sm

.

Hum

ata

pect

inat

a (S

m.)

Des

v.

Hum

ata

pect

inat

a (S

m.)

Des

v.H

umat

a pe

ctin

ata

(Sm

.) D

esv.

/ Pa

chyp

leur

ia p

ectin

ata

(Sm

.) C

.Pre

sl

Hum

ata

repe

ns (L

.f.) D

iels

Hum

ata

repe

ns (L

.f.) J

.Sm

ex

Die

lsH

umat

a re

pens

(L.f.

) J.S

m e

x D

iels

/ Pa

chyp

leur

ia re

pens

(L.f.

) M.K

ato

Hum

ata

vest

ita (B

lum

e) T

.Moo

reH

umat

a ve

stita

(Blu

me)

T.M

oore

Hum

ata

vest

ita (B

lum

e) T

.Moo

re /

Pach

yple

uria

ves

tita

(Blu

me)

C.P

resl

●W

ibel

ia d

entic

ulat

a ( B

urm

.f.) M

.Kat

o &

Tsu

tsum

iD

aval

lia d

entic

ulat

a (B

urm

.f.) M

ett.

ex K

uhn

Dav

allia

den

ticul

ata

(Bur

m.f.

) Met

t. ex

Kuh

n

●W

ibel

ia d

ivar

icat

a (B

lum

e) M

.Kat

o &

Tsu

tsum

iD

aval

lia d

ivar

icat

a B

lum

eD

aval

lia d

ivar

icat

a B

lum

e

○W

ibel

ia e

mbo

lost

egia

(Cop

el.)

M.K

ato

& T

suts

umi10

,19

Abs

ent

Abs

ent

DE

NN

STA

ED

TIA

CE

AE

Den

nsta

edtia

scab

ra (W

all.

ex H

ook.

) T.M

oore

D

enns

taed

tia sc

abra

(Wal

l. ex

Hoo

k.) T

.Moo

reD

enns

taed

tia sc

abra

(Wal

l. ex

Hoo

k.) T

.Moo

re

His

tiopt

eris

inci

sa (T

hunb

.) J.S

m.

His

tiopt

eris

inci

sa (T

hunb

.) J.S

m.

His

tiopt

eris

inci

sa (T

hunb

.) J.S

m.

Hyp

olep

is p

unct

ata

(Thu

nb.)

Met

t. ex

Kuh

n H

ypol

epis

pun

ctat

a (T

hunb

.) M

ett.

ex K

uhn

Hyp

olep

is p

unct

ata

(Thu

nb.)

Met

t. ex

Kuh

n

Mic

role

pia

calv

esce

ns (W

all.

ex H

ook.

) C.P

resl

M

icro

lepi

a ca

lves

cens

(Wal

l. ex

Hoo

k.) C

.Pre

slM

icro

lepi

a ca

lves

cens

(Wal

l. ex

Hoo

k.) C

.Pre

sl

Mic

role

pia

firm

a M

ett.

ex K

uhn

Mic

role

pia

firm

a M

ett.

ex K

uhn

Mic

role

pia

firm

a M

ett.

ex K

uhn

Mic

role

pia

herb

acea

Chi

ng &

C.C

hr. e

x Ta

rdie

u &

C.C

hr.

Mic

role

pia

herb

acea

Chi

ng &

C.C

hr. e

x Ta

rdie

u &

C.C

hr.

Mic

role

pia

herb

acea

Chi

ng &

C.C

hr. e

x Ta

rdie

u &

C.C

hr.

Page 11: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

74 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Mic

role

pia

hook

eria

na (W

all.

ex H

ook.

) C

.Pre

sl

Mic

role

pia

hook

eria

na (W

all.

ex H

ook.

) C

.Pre

slM

icro

lepi

a ho

oker

iana

(Wal

l. ex

Hoo

k.)

C.P

resl

Mic

role

pia

kurz

ii (C

.B.C

lark

e) B

edd.

M

icro

lepi

a ku

rzii

(C.B

.Cla

rke)

Bed

d.M

icro

lepi

a ku

rzii

(C.B

.Cla

rke)

Bed

d.

Mic

role

pia

plat

yphy

lla (D

.Don

) J.S

m.

Mic

role

pia

plat

yphy

lla (D

.Don

) J.S

m.

Mic

role

pia

plat

yphy

lla (D

.Don

) J.S

m.

Mic

role

pia

pube

rula

Ald

erw.

M

icro

lepi

a pu

beru

la A

lder

w.M

icro

lepi

a pu

beru

la A

lder

w.

Mic

role

pia

ridl

eyi C

opel

. M

icro

lepi

a ri

dley

i Cop

el.

Mic

role

pia

ridl

eyi C

opel

.

Mic

role

pia

spel

unca

e (L

.) T.

Moo

re

Mic

role

pia

spel

unca

e (L

.) T.

Moo

reM

icro

lepi

a sp

elun

cae

(L.)

T.M

oore

Mic

role

pia

stri

gosa

(Thu

nb.)

C.P

resl

M

icro

lepi

a st

rigo

sa (T

hunb

.) C

.Pre

slM

icro

lepi

a st

rigo

sa (T

hunb

.) C

.Pre

sl

Mic

role

pia

taiw

ania

na T

agaw

a M

icro

lepi

a ta

iwan

iana

Tag

awa

Mic

role

pia

taiw

ania

na T

agaw

a

Mic

role

pia

trap

ezifo

rmis

(Rox

b.) K

uhn

Mic

role

pia

trap

ezifo

rmis

(Rox

b.) K

uhn

Mic

role

pia

trap

ezifo

rmis

(Rox

b.) K

uhn

○M

onac

hoso

rum

hen

ryi C

hris

t26

Abs

ent

Abs

ent

○Pt

erid

ium

aqu

ilinu

m (L

.) K

uhn

subs

p. ja

poni

cum

(N

akai

) Á.L

öve

& D

.Löv

e4,6,

38,4

2Pt

erid

ium

aqu

ilinu

m (L

.) K

uhn

var.

latiu

scul

um

(Des

v.) U

nder

w. e

x A

.Hel

ler

Abs

ent

●Pt

erid

ium

aqu

ilinu

m (L

.) K

uhn

subs

p. w

ight

ianu

m

(J.a

gard

h) W

.C.S

hieh

38,4

2Pt

erid

ium

aqu

ilinu

m (L

.) K

uhn

var.

wig

htia

num

(J

.aga

rdh)

r.M

.Try

onPt

erid

ium

aqu

ilinu

m (L

.) K

uhn

subs

p.

aqui

linum

var

. wig

htia

num

(J.a

gard

h)

r.M

.Try

on

●Pt

erid

ium

sem

ihas

tatu

m (W

all.

ex J.

aga

rdh)

S.

B.A

ndre

ws42

Pter

idiu

m a

quili

num

(L.)

Kuh

n va

r. ya

rrab

ense

D

omin

Pter

idiu

m a

quili

num

(L.)

Kuh

n su

bsp.

ca

udat

um v

ar. y

arra

bens

e D

omin

DIP

TE

RID

AC

EA

E

Che

iropl

euri

a bi

cusp

is (B

lum

e) C

.Pre

sl

Che

iropl

euri

a bi

cusp

is (B

lum

e) C

.Pre

slC

heiro

pleu

ria

bicu

spis

(Blu

me)

C.P

resl

Dip

teri

s con

juga

ta R

einw

. D

ipte

ris c

onju

gata

Rei

nw.

Dip

teri

s con

juga

ta R

einw

.

DR

YO

PTE

RID

AC

EA

E

●Ac

roph

orus

nod

osus

C.P

resl

Acro

phor

us st

ipel

latu

s T.M

oore

, nom

. nud

.Ac

roph

orus

stip

ella

tus T

.Moo

re, n

om. n

ud.

Arac

hnio

des a

ssam

ica

(Kuh

n) O

hwi

Arac

hnio

des a

ssam

ica

(Kuh

n) O

hwi

Arac

hnio

des a

ssam

ica

(Kuh

n) O

hwi

Page 12: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

75TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Arac

hnio

des c

aval

erii

(Chr

ist)

Ohw

i Ar

achn

iode

s cav

aler

ii (C

hris

t) O

hwi

Arac

hnio

des c

aval

erii

(Chr

ist)

Ohw

i

Arac

hnio

des c

hine

nsis

(Ros

enst

.) C

hing

Ar

achn

iode

s chi

nens

is (R

osen

st.)

Chi

ngAr

achn

iode

s chi

nens

is (R

osen

st.)

Chi

ng

○Ar

achn

iode

s con

iifol

ia (T

.Moo

re) C

hing

8,10

Abs

ent

Abs

ent

Arac

hnio

des h

enry

i (C

hris

t) C

hing

Ar

achn

iode

s hen

ryi (

Chr

ist)

Chi

ngAr

achn

iode

s hen

ryi (

Chr

ist)

Chi

ng

Arac

hnio

des s

peci

osa

(D.D

on) C

hing

Ar

achn

iode

s spe

cios

a (D

.Don

) Chi

ngAr

achn

iode

s spe

cios

a (D

.Don

) Chi

ng

Arac

hnio

des s

pect

abili

s (C

hing

) Chi

ng

Arac

hnio

des s

pect

abili

s (C

hing

) Chi

ngAr

achn

iode

s spe

ctab

ilis (

Chi

ng) C

hing

○Ar

achn

iode

s sub

refle

xipi

nna

(Oga

ta) C

hing

4Ar

achn

iode

s sub

refle

xipi

nna

(Oga

ta) C

hing

A

bsen

t

Bolb

itis a

ngus

tipin

na (H

ayat

a) H

.Ito

Bolb

itis a

ngus

tipin

na (H

ayat

a) H

.Ito

Bolb

itis a

ngus

tipin

na (H

ayat

a) H

.Ito

Bolb

itis a

ppen

dicu

lata

(Will

d.) K

.Iwat

s. su

bsp.

ap

pend

icul

ata

Bolb

itis a

ppen

dicu

lata

(Will

d.) K

.Iwat

s. su

bsp.

ap

pend

icul

ata

Bolb

itis a

ppen

dicu

lata

(Will

d.) K

.Iwat

s. su

bsp.

ap

pend

icul

ata

Bolb

itis c

opel

andi

i Chi

ng e

x C

.Chr

. & T

ardi

eu

Bolb

itis c

opel

andi

i Chi

ng e

x C

.Chr

. & T

ardi

euBo

lbiti

s cop

elan

dii C

hing

ex

C.C

hr. &

Tar

dieu

Bolb

itis c

osta

ta (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis c

osta

ta (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis c

osta

ta (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis d

eltig

era

(Bed

d.) C

.Chr

. Bo

lbiti

s del

tiger

a (B

edd.

) C.C

hr.

Bolb

itis d

eltig

era

(Bed

d.) C

.Chr

.

Bolb

itis h

eter

oclit

a (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis h

eter

oclit

a (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis h

eter

oclit

a (C

.Pre

sl) C

hing

ex

C.C

hr.

Bolb

itis h

ooke

rian

a K

.Iwat

s. Bo

lbiti

s hoo

keri

ana

K.Iw

ats.

+ Bo

lbiti

s app

endi

cula

ta

(Will

d.) K

.Iwat

s. su

bsp.

viv

ipar

a (B

uch.

-Ham

. ex

Hoo

k.)

Hen

nipm

an

Bolb

itis h

ooke

rian

a K

.Iwat

s.

Bolb

itis s

calp

tura

ta (F

ée) C

hing

Bo

lbiti

s sca

lptu

rata

(Fée

) Chi

ngBo

lbiti

s sca

lptu

rata

(Fée

) Chi

ng

Bolb

itis s

inen

sis (

Bak

er) K

.Iwat

s. va

r. si

nens

is

Bolb

itis s

inen

sis (

Bak

er) K

.Iwat

s. va

r. si

nens

isBo

lbiti

s sin

ensi

s (B

aker

) K.Iw

ats.

var.

sine

nsis

Bolb

itis s

inen

sis (

Bak

er) K

.Iwat

s. va

r. co

stul

ata

(Hoo

k.) T

agaw

a &

K.Iw

ats.

Bolb

itis s

inen

sis (

Bak

er) K

.Iwat

s. va

r. co

stul

ata

(Hoo

k.) T

agaw

a &

K.Iw

ats.

Bolb

itis s

inen

sis (

Bak

er) K

.Iwat

s. va

r. co

stul

ata

(Hoo

k.) T

agaw

a &

K.Iw

ats

Bolb

itis s

inua

ta (C

.Pre

sl) H

enni

pman

Bo

lbiti

s sin

uata

(C.P

resl

) Hen

nipm

anBo

lbiti

s sin

uata

(C.P

resl

) Hen

nipm

an

Bolb

itis t

onki

nens

is (C

.Chr

. ex

Chi

ng) K

.Iwat

s.Bo

lbiti

s ton

kine

nsis

(C.C

hr. e

x C

hing

) K.Iw

ats.

Bolb

itis t

onki

nens

is (C

.Chr

. ex

Chi

ng) K

.Iwat

s

Bolb

itis v

irens

(Wal

l. ex

Hoo

k. &

Gre

v.) S

chot

t va

r. vi

rens

Bolb

itis v

irens

(Wal

l. ex

Hoo

k. &

Gre

v.) S

chot

t va

r. vi

rens

Bolb

itis v

irens

(Wal

l. ex

Hoo

k. &

Gre

v.) S

chot

t va

r. vi

rens

Bolb

itis v

irens

(Wal

l. ex

Hoo

k. &

Gre

v.) S

chot

t va

r. co

mpa

cta

Hen

nipm

anBo

lbiti

s vire

ns (W

all.

ex H

ook.

& G

rev.

) Sch

ott

var.

com

pact

a H

enni

pman

Bolb

itis v

irens

(Wal

l. ex

Hoo

k. &

Gre

v.) S

chot

t va

r. co

mpa

cta

Hen

nipm

an

Page 13: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

76 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Cte

nitis

dum

rong

ii Ta

gaw

a &

K.Iw

ats.

Cte

nitis

dum

rong

ii Ta

gaw

a &

K.Iw

ats.

Cte

nitis

dum

rong

ii Ta

gaw

a &

K.Iw

ats.

●C

teni

tis su

bobs

cura

(Chr

ist)

Hol

ttum

C

teni

tis su

bobs

cura

(Chr

ist)

Hol

ttum

Cte

nitis

man

nii a

uct.

non

(C.H

ope)

Chi

ng /

Cte

nitis

subo

bscu

ra (C

hris

t) H

olttu

m

Cte

nitis

vili

s (K

unze

) Chi

ng

Cte

nitis

vili

s (K

unze

) Chi

ngC

teni

tis v

ilis (

Kun

ze) C

hing

Cyr

tom

ium

fort

unei

J.Sm

. C

yrto

miu

m fo

rtun

ei J.

Sm.

Cyr

tom

ium

fort

unei

J.Sm

.

Did

ymoc

hlae

na tr

unca

tula

(Sw.

) J.S

m.

Did

ymoc

hlae

na tr

unca

tula

(Sw.

) J.S

m.

Did

ymoc

hlae

na tr

unca

tula

(Sw.

) J.S

m.

Dry

opte

ris c

ochl

eata

(D.D

on) C

.Chr

. D

ryop

teri

s coc

hlea

ta (D

.Don

) C.C

hr.

Dry

opte

ris c

ochl

eata

(D.D

on) C

.Chr

.

○D

ryop

teri

s diff

ract

a (B

aker

) C.C

hr.10

Abs

ent

Abs

ent

Dry

opte

ris g

ymno

phyl

la (B

aker

) C.C

hr.

Dry

opte

ris g

ymno

phyl

la (B

aker

) C.C

hr.

Dry

opte

ris g

ymno

phyl

la (B

aker

) C.C

hr.

●D

ryop

teri

s has

selti

i (B

lum

e) C

.Chr

. Ar

achn

iode

s has

selti

i (B

lum

e) C

hing

Arac

hnio

des h

asse

ltii (

Blu

me)

Chi

ng

Dry

opte

ris h

ende

rson

ii (B

edd.

) C.C

hr.

Dry

opte

ris h

ende

rson

ii (B

edd.

) C.C

hr.

Dry

opte

ris h

ende

rson

ii (B

edd.

) C.C

hr.

Dry

opte

ris h

irtip

es (B

lum

e) K

untz

e D

ryop

teri

s hir

tipes

(Blu

me)

Kun

tze

Dry

opte

ris h

irtip

es (B

lum

e) K

untz

e

Dry

opte

ris i

nteg

rilo

ba C

.Chr

. D

ryop

teri

s int

egri

loba

C.C

hr.

Dry

opte

ris i

nteg

rilo

ba C

.Chr

.

Dry

opte

ris n

eoas

sam

ensi

s Chi

ng

Dry

opte

ris n

eoas

sam

ensi

s Chi

ngD

ryop

teri

s neo

assa

men

sis C

hing

Dry

opte

ris n

eoch

ryso

com

a C

hing

D

ryop

teri

s neo

chry

soco

ma

Chi

ngD

ryop

teri

s neo

chry

soco

ma

Chi

ng

Dry

opte

ris p

olita

Ros

enst

. D

ryop

teri

s pol

ita R

osen

st.

Dry

opte

ris p

olita

Ros

enst

.

Dry

opte

ris p

oros

a C

hing

D

ryop

teri

s por

osa

Chi

ngD

ryop

teri

s por

osa

Chi

ng

Dry

opte

ris p

seud

ospa

rsa

Chi

ng

Dry

opte

ris p

seud

ospa

rsa

Chi

ngD

ryop

teri

s pse

udos

pars

a C

hing

Dry

opte

ris r

heop

hila

Mits

uta

ex D

arna

edi,

M.K

ato

& K

.Iwat

s. D

ryop

teri

s rhe

ophi

la M

itsut

a ex

Dar

naed

i, M

.Kat

o &

K.Iw

ats.

Dry

opte

ris r

heop

hila

Mits

uta

ex D

arna

edi,

M.K

ato

& K

.Iwat

s.

Dry

opte

ris s

cotti

i (B

edd.

) Chi

ng

Dry

opte

ris s

cotti

i (B

edd.

) Chi

ngD

ryop

teri

s sco

ttii (

Bed

d.) C

hing

Dry

opte

ris s

pars

a (D

.Don

) Kun

tze

Dry

opte

ris s

pars

a (D

.Don

) Kun

tze

Dry

opte

ris s

pars

a (D

.Don

) Kun

tze

Dry

opte

ris s

ubtr

iang

ular

is (C

.Hop

e) C

.Chr

. D

ryop

teri

s sub

tria

ngul

aris

(C.H

ope)

C.C

hr.

Dry

opte

ris s

ubtr

iang

ular

is (C

.Hop

e) C

.Chr

.

Elap

hogl

ossu

m a

ngul

atum

(Blu

me)

T.M

oore

El

apho

glos

sum

ang

ulat

um (B

lum

e) T

.Moo

reEl

apho

glos

sum

ang

ulat

um (B

lum

e) T

.Moo

re

Elap

hogl

ossu

m d

umro

ngii

Taga

wa

& K

.Iwat

s. El

apho

glos

sum

dum

rong

ii Ta

gaw

a &

K.Iw

ats.

Elap

hogl

ossu

m d

umro

ngii

Taga

wa

& K

.Iwat

s.

Elap

hogl

ossu

m m

alay

ense

Hol

ttum

El

apho

glos

sum

mal

ayen

se H

olttu

mEl

apho

glos

sum

mal

ayen

se H

olttu

m

Page 14: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

77TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Elap

hogl

ossu

m m

argi

natu

m (F

ée) T

.Moo

re

Elap

hogl

ossu

m m

argi

natu

m (F

ée) T

.Moo

reEl

apho

glos

sum

mar

gina

tum

(Fée

) T.M

oore

Elap

hogl

ossu

m m

elan

ostic

tum

(Blu

me)

T.M

oore

El

apho

glos

sum

mel

anos

tictu

m (B

lum

e) T

.Moo

reEl

apho

glos

sum

mel

anos

tictu

m (B

lum

e) T

.Moo

re

Elap

hogl

ossu

m st

ellig

erum

(Wal

l. ex

Bak

er)

T.M

oore

ex

als

ton

& B

onne

rEl

apho

glos

sum

stel

liger

um (W

all.

ex B

aker

) T.

Moo

re e

x a

lsto

n &

Bon

ner

Elap

hogl

ossu

m st

ellig

erum

(Wal

l. ex

Bak

er)

T.M

oore

ex

als

ton

& B

onne

r

Elap

hogl

ossu

m su

belli

ptic

um R

osen

st.

Elap

hogl

ossu

m su

belli

ptic

um R

osen

st.

Elap

hogl

ossu

m su

belli

ptic

um R

osen

st.

Elap

hogl

ossu

m y

oshi

naga

e (Y

atab

e) M

akin

o El

apho

glos

sum

yos

hina

gae

(Yat

abe)

Mak

ino

Elap

hogl

ossu

m y

oshi

naga

e (Y

atab

e) M

akin

o

Hyp

odem

atiu

m c

rena

tum

(For

ssk.

) Kuh

n H

ypod

emat

ium

cre

natu

m (F

orss

k.) K

uhn

Hyp

odem

atiu

m c

rena

tum

(For

ssk.

) Kuh

n

Hyp

odem

atiu

m g

land

ulos

o-pi

losu

m (T

agaw

a)

Ohw

i H

ypod

emat

ium

gla

ndul

oso-

pilo

sum

(Tag

awa)

O

hwi

Hyp

odem

atiu

m g

land

ulos

o-pi

losu

m (T

agaw

a)

Ohw

i

Leuc

oste

gia

imm

ersa

(Wal

l. ex

Hoo

k.) C

.Pre

sl

Leuc

oste

gia

imm

ersa

(Wal

l. ex

Hoo

k.) C

.Pre

slLe

ucos

tegi

a im

mer

sa (W

all.

ex H

ook.

) C.P

resl

Lom

agra

mm

a gr

osso

serr

ata

Hol

ttum

Lo

mag

ram

ma

gros

sose

rrat

a H

olttu

mLo

mag

ram

ma

gros

sose

rrat

a H

olttu

m

●Pe

rane

ma

aspi

dioi

des (

Blu

me)

Met

t. D

iaca

lpe

aspi

dioi

des B

lum

eD

iaca

lpe

aspi

dioi

des B

lum

e

Poly

stic

hum

atte

nuat

um T

agaw

a &

K.Iw

ats.

Poly

stic

hum

atte

nuat

um T

agaw

a &

K.Iw

ats.

Poly

stic

hum

atte

nuat

um T

agaw

a &

K.Iw

ats.

Poly

stic

hum

bia

rist

atum

(Blu

me)

T.M

oore

Po

lyst

ichu

m b

iari

stat

um (B

lum

e) T

.Moo

rePo

lyst

ichu

m b

iari

stat

um (B

lum

e) T

.Moo

re

Poly

stic

hum

exi

miu

m (K

uhn)

C.C

hr.

Poly

stic

hum

exi

miu

m (K

uhn)

C.C

hr.

Poly

stic

hum

exi

miu

m (K

uhn)

C.C

hr.

Poly

stic

hum

lind

saei

foliu

m S

cort.

ex

Rid

l. Po

lyst

ichu

m li

ndsa

eifo

lium

Sco

rt. e

x. R

idl.

Poly

stic

hum

lind

saei

foliu

m S

cort.

ex

Rid

l.

Poly

stic

hum

pro

lifica

ns A

lder

w.

Poly

stic

hum

pro

lifica

ns A

lder

w.Po

lyst

ichu

m p

rolifi

cans

Ald

erw.

○Po

lyst

ichu

m p

seud

otsu

s-si

men

se C

hing

8A

bsen

tA

bsen

t

○Po

lyst

ichu

m sc

ario

sum

(rox

b.) C

.V.M

orto

n8,10

Abs

ent

Abs

ent

Poly

stic

hum

sem

ifert

ile (C

.B.C

lark

e) C

hing

Po

lyst

ichu

m se

mife

rtile

(C.B

.Cla

rke)

Chi

ngPo

lyst

ichu

m se

mife

rtile

(C.B

.Cla

rke)

Chi

ng

Poly

stic

hum

teng

gere

nse

Ros

enst

. Po

lyst

ichu

m te

ngge

rens

e R

osen

st.

Poly

stic

hum

teng

gere

nse

Ros

enst

.

Tera

toph

yllu

m a

cule

atum

(Blu

me)

Met

t. ex

Kuh

n Te

rato

phyl

lum

acu

leat

um (B

lum

e) M

ett.

ex K

uhn

Tera

toph

yllu

m a

cule

atum

(Blu

me)

Met

t. ex

Kuh

n

Tera

toph

yllu

m lu

dens

(Fée

) Hol

ttum

Te

rato

phyl

lum

lude

ns (F

ée) H

olttu

mTe

rato

phyl

lum

lude

ns (F

ée) H

olttu

m

EQ

UIS

ETA

CE

AE

○Eq

uise

tum

diff

usum

D.D

on4

Equi

setu

m d

iffus

um D

.Don

Abs

ent

Page 15: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

78 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●Eq

uise

tum

ram

osis

sim

um D

esf.

subs

p. d

ebile

(Rox

b. e

x Va

uche

r) H

auke

Eq

uise

tum

deb

ile R

oxb.

ex

Vauc

her

Equi

setu

m d

ebile

Rox

b. e

x Va

uche

r

GL

EIC

HE

NIA

CE

AE

Dic

rano

pter

is c

urra

nii C

opel

. D

icra

nopt

eris

cur

rani

i Cop

el.

Dic

rano

pter

is c

urra

nii C

opel

.

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

linea

ris

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

linea

ris

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

linea

ris

○D

icra

nopt

eris

line

aris

(Bur

m.f)

Und

erw.

va

r. m

onta

na H

olttu

m4

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

mon

tana

Hol

ttum

Abs

ent

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

subp

ectin

ata

(Chr

ist)

Hol

ttum

D

icra

nopt

eris

line

aris

(Bur

m.f.

) Und

erw.

va

r. su

bpec

tinat

a (C

hris

t) H

olttu

mD

icra

nopt

eris

line

aris

(Bur

m.f.

) Und

erw.

va

r. su

bpec

tinat

a (C

hris

t) H

olttu

m

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

tetr

aphy

lla (r

osen

st.)

Nak

ai

Dic

rano

pter

is li

near

is (B

urm

.f.) U

nder

w.

var.

tetr

aphy

lla (r

osen

st.)

Nak

aiD

icra

nopt

eris

line

aris

(Bur

m.f.

) Und

erw.

va

r. te

trap

hylla

(ros

enst

.) N

akai

Dic

rano

pter

is sp

ecio

sa (C

.Pre

sl) H

olttu

m

Dic

rano

pter

is sp

ecio

sa (C

.Pre

sl) H

olttu

mD

icra

nopt

eris

spec

iosa

(C.P

resl

) Hol

ttum

Dic

rano

pter

is sp

lend

ida

(Han

d.-M

azz.

) Tag

awa

Dic

rano

pter

is sp

lend

ida

(Han

d.-M

azz.

) Tag

awa

Dic

rano

pter

is sp

lend

ida

(Han

d.-M

azz.

) Tag

awa

●D

iplo

pter

ygiu

m b

lotia

num

(C.C

hr.)

Nak

aiG

leic

heni

a bl

otia

na C

.Chr

.G

leic

heni

a bl

otia

na C

.Chr

.

●D

iplo

pter

ygiu

m lo

ngis

sim

um (B

lum

e) N

akai

Gle

iche

nia

long

issi

ma

Blu

me

Gle

iche

nia

long

issi

ma

Blu

me

●D

iplo

pter

ygiu

m n

orri

sii (

Met

t.) N

akai

Gle

iche

nia

norr

isii

Met

t.G

leic

heni

a no

rris

ii M

ett.

Gle

iche

nia

mic

roph

ylla

R.B

r. G

leic

heni

a m

icro

phyl

la R

.Br.

Gle

iche

nia

mic

roph

ylla

R.B

r.

○St

iche

rus h

irtu

s (B

lum

e) C

hing

10A

bsen

tA

bsen

t

●St

iche

rus t

runc

atus

(Will

d.) N

akai

Gle

iche

nia

trun

cata

(Will

d.) S

pren

g.G

leic

heni

a tr

unca

ta (W

illd.

) Spr

eng.

HY

ME

NO

PHY

LL

AC

EA

E

●Ab

rodi

ctyu

m id

oneu

m (C

.V.M

orto

n)

Ebih

ara

& K

.Iwat

s.47

Cep

halo

man

es g

emm

atum

(J.S

m. e

x B

aker

) K

.Iwat

s. M

acro

glen

a ge

mm

ata

(J.S

m. e

x B

aker

) Cop

el.

/ Cep

halo

man

es g

emm

atum

(J.S

m. e

x B

aker

) K

.Iwat

s.

●Ab

rodi

ctyu

m o

bscu

rum

(Blu

me)

Ebi

hara

& K

.Iwat

s. va

r. ob

scur

um

Cep

halo

man

es o

bscu

rum

(Blu

me)

K.Iw

ats.

var.

obsc

urum

Sele

node

smiu

m o

bscu

rum

(Blu

me)

Cop

el.

Page 16: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

79TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●Ab

rodi

ctyu

m o

bscu

rum

(Blu

me)

Ebi

hara

& K

.Iwat

s. va

r. si

amen

se (C

hris

t) K

.Iwat

s.C

epha

lom

anes

obs

curu

m (B

lum

e) K

.Iwat

s. va

r. si

amen

se (C

hris

t) K

.Iwat

s.Se

leno

desm

ium

obs

curu

m (B

lum

e) C

opel

., pr

o pa

rte /

Cep

halo

man

es o

bscu

rum

(Blu

me)

K

.Iwat

s. va

r. si

amen

se (C

hris

t) K

.Iwat

s.

●Ab

rodi

ctyu

m p

lum

a (H

ook.

) Ebi

hara

& K

.Iwat

s.47

Cep

halo

man

es m

eifo

lium

(Bor

y ex

Will

d.) K

.Iwat

s.M

acro

glen

a m

eifo

lia (B

ory

ex W

illd.

) Cop

el.

/ Cep

halo

man

es m

eifo

lium

(Bor

y ex

Will

d.)

K.Iw

ats.

Cal

listo

pter

is a

piifo

lia (C

.Pre

sl) C

opel

. C

epha

lom

anes

api

ifoliu

m (C

.Pre

sl) K

.Iwat

s.C

allis

topt

eris

api

ifolia

(C.P

resl

) Cop

el. /

C

epha

lom

anes

api

ifoliu

m (C

.Pre

sl) K

.Iwat

s.

Cep

halo

man

es ja

vani

cum

(Blu

me)

C.P

resl

C

epha

lom

anes

java

nicu

m (B

lum

e) C

.Pre

slC

epha

lom

anes

java

nicu

m (B

lum

e) C

.Pre

sl

●C

repi

dom

anes

bip

unct

atum

(Poi

r.) C

opel

. C

repi

dom

anes

bip

unct

atum

(Poi

r.) C

opel

.C

repi

dom

anes

bip

unct

atum

(Poi

r.) C

opel

. +

Crep

idom

anes

bila

biat

um (N

ees &

Blu

me)

Cop

el.

Cre

pido

man

es b

revi

pes (

C.P

resl

) Cop

el.

Cre

pido

man

es b

revi

pes (

C.P

resl

) Cop

el.

Cre

pido

man

es b

revi

pes (

C.P

resl

) Cop

el.

Cre

pido

man

es c

hris

tii (C

opel

.) C

opel

. C

repi

dom

anes

chr

istii

(Cop

el.)

Cop

el.

Cre

pido

man

es c

hris

tii (C

opel

.) C

opel

.

●C

repi

dom

anes

hum

ile (G

.For

st.)

Bos

ch

Cre

pido

man

es h

umile

(G.F

orst

.) B

osch

Reed

iella

hum

ilis (

G.F

orst

.) Pi

c.Se

rm. /

C

repi

dom

anes

hum

ile (G

.For

st.)

Bos

ch

Cre

pido

man

es k

urzi

i (B

edd.

) Tag

awa

& K

.Iwat

s. C

repi

dom

anes

kur

zii (

Bed

d.) T

agaw

a &

K.Iw

ats.

Cre

pido

man

es k

urzi

i (B

edd.

) Tag

awa

& K

.Iwat

s.

Cre

pido

man

es la

teal

atum

(Bos

ch) C

opel

. C

repi

dom

anes

late

alat

um (B

osch

) Cop

el.

Cre

pido

man

es la

teal

atum

(Bos

ch) C

opel

.

Cre

pido

man

es la

tem

argi

nale

(D.C

.Eat

on)

Cop

el.

Cre

pido

man

es la

tem

argi

nale

(D.C

.Eat

on)

Cop

el.

Cre

pido

man

es la

tem

argi

nale

(D.C

.Eat

on)

Cop

el.

Cre

pido

man

es m

egis

tost

omum

(Cop

el.)

Cop

el.

Cre

pido

man

es m

egis

tost

omum

(Cop

el.)

Cop

el.

Cre

pido

man

es m

egis

tost

omum

(Cop

el.)

Cop

el.

●C

repi

dom

anes

min

utum

(Blu

me)

K.Iw

ats.

Cre

pido

man

es m

inut

um (B

lum

e) K

.Iwat

s.G

onoc

orm

us p

rolif

er (B

lum

e) P

rant

l +

Gon

ocor

mus

saxi

frag

oide

s (C

.Pre

sl) B

osch

+

Gon

ocor

mus

siam

ensi

s Tag

awa

& K

.Iwat

s. /

Cre

pido

man

es m

inut

um (B

lum

e) K

.Iwat

s.

●C

repi

dom

anes

par

vifo

lium

(Bak

er) K

.Iwat

s. C

repi

dom

anes

par

vifo

lium

(Bak

er) K

.Iwat

s.M

icro

goni

um p

arvi

foliu

m (B

aker

) Tag

awa

&

K.Iw

ats.

/ Cre

pido

man

es p

arvi

foliu

m (B

aker

) K

.Iwat

s.

●D

idym

oglo

ssum

bim

argi

natu

m (B

osch

) Ebi

hara

&

K.Iw

ats.

Tric

hom

anes

bim

argi

natu

m (B

osch

) Bos

ch

Mic

rogo

nium

bim

argi

natu

m B

osch

/ Tr

icho

man

es

bim

argi

natu

m (B

osch

) Bos

ch

Page 17: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

80 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Did

ymog

loss

um e

xigu

um (B

edd.

) Cop

el.

Tric

hom

anes

exi

guum

(Bed

d.) B

aker

Did

ymog

loss

um e

xigu

um (B

edd.

) Cop

el. /

Tr

icho

man

es e

xigu

um (B

edd.

) Bak

er

●D

idym

oglo

ssum

mot

leyi

(Bos

ch) E

biha

ra &

K.Iw

ats.

Tric

hom

anes

mot

leyi

(Bos

ch) B

osch

Mic

rogo

nium

mot

leyi

Bos

ch /

Tric

hom

anes

m

otle

yi (B

osch

) Bos

ch

●D

idym

oglo

ssum

subl

imba

tum

(Mül

l.Ber

ol.)

ebih

ara

&

K.Iw

ats.

Tric

hom

anes

subl

imba

tum

Mül

l.Ber

ol.

Mic

rogo

nium

subl

imba

tum

(Mül

l.Ber

ol.)

Bos

ch /

Tric

hom

anes

subl

imba

tum

Mül

l.Ber

ol.

●H

ymen

ophy

llum

aca

ntho

ides

(Bos

ch) R

osen

st.

Hym

enop

hyllu

m a

cant

hoid

es (B

osch

) Ros

enst

.M

erin

gium

aca

ntho

ides

(Bos

ch) C

opel

. /

Hym

enop

hyllu

m a

cant

hoid

es (B

osch

) Ros

enst

.

●H

ymen

ophy

llum

bad

ium

Hoo

k. &

Gre

v.

Hym

enop

hyllu

m b

adiu

m H

ook.

& G

rev.

Mec

odiu

m b

adiu

m (H

ook.

& G

rev.

) Cop

el. /

H

ymen

ophy

llum

bad

ium

Hoo

k. &

Gre

v.

Hym

enop

hyllu

m b

arba

tum

(Bos

ch) B

aker

H

ymen

ophy

llum

bar

batu

m (B

osch

) Bak

erH

ymen

ophy

llum

bar

batu

m (B

osch

) Bak

er

●H

ymen

ophy

llum

bla

ndum

Rac

ib.

Hym

enop

hyllu

m b

land

um R

acib

.M

erin

gium

bla

ndum

(Rac

ib.)

Cop

el. /

H

ymen

ophy

llum

bla

ndum

Rac

ib.

●H

ymen

ophy

llum

bon

toce

nse

Cop

el.

Hym

enop

hyllu

m b

onto

cens

e C

opel

.M

erin

gium

bon

toce

nse

(Cop

el.)

Cop

el. /

H

ymen

ophy

llum

bon

toce

nse

Cop

el.

●H

ymen

ophy

llum

den

ticul

atum

Sw.

H

ymen

ophy

llum

den

ticul

atum

Sw.

Mer

ingi

um d

entic

ulat

um (S

w.) C

opel

. /

Hym

enop

hyllu

m d

entic

ulat

um S

w.

●H

ymen

ophy

llum

dig

itatu

m (S

w.) F

osbe

rg

Cre

pido

man

es d

igita

tum

(Sw.

) K.Iw

ats.

Mic

rotr

icho

man

es d

igita

tum

(Sw.

) Cop

el. /

C

repi

dom

anes

dig

itatu

m (S

w.) K

.Iwat

s.

●H

ymen

ophy

llum

exs

ertu

m W

all.

ex H

ook.

H

ymen

ophy

llum

exs

ertu

m W

all.

ex H

ook.

Mec

odiu

m e

xser

tum

(Wal

l. ex

Hoo

k.) C

opel

. /

Hym

enop

hyllu

m e

xser

tum

Wal

l. ex

Hoo

k.

●H

ymen

ophy

llum

hol

ochi

lum

(Bos

ch) C

.Chr

. H

ymen

ophy

llum

hol

ochi

lum

(Bos

ch) C

.Chr

.M

erin

gium

hol

ochi

lum

(Bos

ch) C

opel

. /

Hym

enop

hyllu

m h

oloc

hilu

m (B

osch

) C.C

hr.

●H

ymen

ophy

llum

java

nicu

m S

pren

g.

Hym

enop

hyllu

m ja

vani

cum

Spr

eng.

Mec

odiu

m ja

vani

cum

(Spr

eng.

) Cop

el. /

H

ymen

ophy

llum

java

nicu

m S

pren

g.

●H

ymen

ophy

llum

pal

lidum

(Blu

me)

Ebi

hara

& K

.Iwat

s. C

repi

dom

anes

pal

lidum

(Blu

me)

K.Iw

ats.

Pleu

rom

anes

pal

lidum

(Blu

me)

C.P

resl

. /

Cre

pido

man

es p

allid

um (B

lum

e) K

.Iwat

s.

●H

ymen

ophy

llum

pol

yant

hos (

Sw.)

Sw.

Hym

enop

hyllu

m p

olya

ntho

s (Sw

.) Sw

.M

ecod

ium

pol

yant

hos (

Sw.)

Cop

el. /

H

ymen

ophy

llum

pol

yant

hos (

Sw.)

Sw.

Page 18: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

81TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●H

ymen

ophy

llum

pro

duct

um K

unze

H

ymen

ophy

llum

pro

duct

um K

unze

Mec

odiu

m p

rodu

ctum

(Kun

ze) C

opel

. /

Hym

enop

hyllu

m p

rodu

ctum

Kun

ze

●H

ymen

ophy

llum

riuk

iuen

se C

hris

t H

ymen

ophy

llum

riuk

iuen

se C

hris

tM

ecod

ium

riuk

iuen

se (C

hris

t) C

opel

. /

Hym

enop

hyllu

m ri

ukiu

ense

Chr

ist

●H

ymen

ophy

llum

serr

ulat

um (C

.Pre

sl) C

.Chr

. H

ymen

ophy

llum

serr

ulat

um (C

.Pre

sl) C

.Chr

.M

erin

gium

mey

enia

num

C.P

resl

/ H

ymen

ophy

llum

serr

ulat

um (C

.Pre

sl) C

.Chr

.

●Va

nden

bosc

hia

auri

cula

ta (B

lum

e) C

opel

. C

repi

dom

anes

aur

icul

atum

(Blu

me)

K.Iw

ats.

Tric

hom

anes

aur

icul

atum

Blu

me

/ Cre

pido

man

es

auri

cula

tum

(Blu

me)

K.Iw

ats.

●Va

nden

bosc

hia

birm

anic

a (B

edd.

) Chi

ng

Cre

pido

man

es b

irm

anic

um (B

edd.

) K.Iw

ats.

Tric

hom

anes

bir

man

icum

Bed

d. /

Cre

pido

man

es

birm

anic

um (B

edd.

) K.Iw

ats.

●Va

nden

bosc

hia

max

ima

(Blu

me)

Cop

el.

Cre

pido

man

es m

axim

um (B

lum

e) K

.Iwat

s.Tr

icho

man

es m

axim

um B

lum

e / C

repi

dom

anes

m

axim

um (B

lum

e) K

.Iwat

s.

LIN

DSA

EA

CE

AE

Lind

saea

bou

illod

ii C

hris

t Li

ndsa

ea b

ouill

odii

Chr

ist

Lind

saea

bou

illod

ii C

hris

t

Lind

saea

chi

enii

Chi

ng

Lind

saea

chi

enii

Chi

ngLi

ndsa

ea c

hien

ii C

hing

Lind

saea

cul

trat

a (W

illd.

) Sw.

Li

ndsa

ea c

ultr

ata

(Will

d.) S

w.Li

ndsa

ea c

ultr

ata

(Will

d.) S

w.

○Li

ndsa

ea d

isse

ctifo

rmis

Chi

ng10

Abs

ent

Abs

ent

Lind

saea

div

erge

ns H

ook.

& G

rev.

Lind

saea

div

erge

ns H

ook.

& G

rev.

Lind

saea

div

erge

ns H

ook.

& G

rev.

Lind

saea

dor

ypho

ra K

.U.K

ram

er

Lind

saea

dor

ypho

ra K

.U.K

ram

erLi

ndsa

ea d

oryp

hora

K.U

.Kra

mer

Lind

saea

ens

ifolia

Sw.

Li

ndsa

ea e

nsifo

lia S

w.Li

ndsa

ea e

nsifo

lia S

w.

Lind

saea

het

erop

hylla

Dry

and.

Li

ndsa

ea h

eter

ophy

lla D

ryan

d.Li

ndsa

ea h

eter

ophy

lla D

ryan

d.

Lind

saea

inte

gra

Hol

ttum

Li

ndsa

ea in

tegr

a H

olttu

mLi

ndsa

ea in

tegr

a H

olttu

m

Lind

saea

java

nens

is B

lum

e Li

ndsa

ea ja

vane

nsis

Blu

me

Lind

saea

java

nens

is B

lum

e

Lind

saea

luci

da B

lum

e Li

ndsa

ea lu

cida

Blu

me

Lind

saea

luci

da B

lum

e

Lind

saea

mal

ayen

sis H

olttu

m

Lind

saea

mal

ayen

sis H

olttu

mLi

ndsa

ea m

alay

ensi

s Hol

ttum

Lind

saea

nap

aea

Ald

erw.

Li

ndsa

ea n

apae

a A

lder

w.Li

ndsa

ea n

apae

a A

lder

w.

Page 19: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

82 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Lind

saea

obl

ance

olat

a A

lder

w.

Lind

saea

obl

ance

olat

a A

lder

w.Li

ndsa

ea o

blan

ceol

ata

Ald

erw.

Lind

saea

odo

rata

Rox

b.

Lind

saea

odo

rata

Rox

b.Li

ndsa

ea o

dora

ta R

oxb.

Lind

saea

orb

icul

ata

(Lam

.) M

ett.

ex K

uhn

var.

orbi

cula

ta

Lind

saea

orb

icul

ata

(Lam

.) M

ett.

ex K

uhn

var.

orbi

cula

taLi

ndsa

ea o

rbic

ulat

a (L

am.)

Met

t. ex

Kuh

n va

r. or

bicu

lata

Lind

saea

orb

icul

ata

(Lam

.) M

ett.

ex K

uhn

var.

com

mix

ta (T

agaw

a) K

.U.K

ram

erLi

ndsa

ea o

rbic

ulat

a (L

am.)

Met

t. ex

Kuh

n va

r. co

mm

ixta

(Tag

awa)

K.U

.Kra

mer

Lind

saea

orb

icul

ata

(Lam

.) M

ett.

ex K

uhn

var.

com

mix

ta (T

agaw

a) K

.U.K

ram

er

Lind

saea

par

alle

logr

amm

a A

lder

w.

Lind

saea

par

alle

logr

amm

a A

lder

w.Li

ndsa

ea p

aral

lelo

gram

ma

Ald

erw.

Lind

saea

par

asiti

ca (R

oxb.

ex

Grif

f.) H

iero

n.

Lind

saea

par

asiti

ca (R

oxb.

ex

Grif

f.) H

iero

n.Li

ndsa

ea p

aras

itica

(Rox

b. e

x G

riff.)

Hie

ron.

Lind

saea

repe

ns (B

ory)

Thw

aite

s var

. pec

tinat

a (B

lum

e) M

ett.

ex K

uhn

Lind

saea

repe

ns (B

ory)

Thw

aite

s var

. pec

tinat

a (B

lum

e) M

ett.

ex K

uhn

Lind

saea

repe

ns (B

ory)

Thw

aite

s var

. pec

tinat

a (B

lum

e) M

ett.

ex K

uhn

○Li

ndsa

ea te

nera

Dry

and.

4Li

ndsa

ea te

nera

Dry

and.

Abs

ent

Sphe

nom

eris

chi

nens

is (L

.) M

axon

va

r. ch

inen

sis

Sphe

nom

eris

chi

nens

is (L

.) M

axon

va

r. ch

inen

sis

Sphe

nom

eris

chi

nens

is (L

.) M

axon

va

r. ch

inen

sis

Sphe

nom

eris

chi

nens

is (L

.) M

axon

var.

diva

rica

ta (C

hris

t) K

.U.K

ram

er

Sphe

nom

eris

chi

nens

is (L

.) M

axon

va

r. di

vari

cata

(Chr

ist)

K.U

.Kra

mer

Sphe

nom

eris

chin

ensis

(L.)

Max

on

var.

diva

ricat

a (C

hris

t) K

.U.K

ram

er

Tape

inid

ium

luzo

nicu

m (H

ook.

) K.U

.Kra

mer

Ta

pein

idiu

m lu

zoni

cum

(Hoo

k.) K

.U.K

ram

erTa

pein

idiu

m lu

zoni

cum

(Hoo

k.) K

.U.K

ram

er

Tape

inid

ium

pin

natu

m (C

av.)

C.C

hr.

Tape

inid

ium

pin

natu

m (C

av.)

C.C

hr.

Tape

inid

ium

pin

natu

m (C

av.)

C.C

hr.

LO

MA

RIO

PSID

AC

EA

E

Cyc

lope

ltis c

rena

ta (F

ée) C

.Chr

. C

yclo

pelti

s cre

nata

(Fée

) C.C

hr.

Cyc

lope

ltis c

rena

ta (F

ée) C

.Chr

.

Lom

ario

psis

line

ata

(C.P

resl

) Hol

ttum

Lo

mar

iops

is li

neat

a (C

.Pre

sl) H

olttu

mLo

mar

iops

is li

neat

a (C

.Pre

sl) H

olttu

m

Nep

hrol

epis

acu

tifol

ia (D

esv.

) Chr

ist

Nep

hrol

epis

acu

tifol

ia (D

esv.

) Chr

ist

Nep

hrol

epis

acu

tifol

ia (D

esv.

) Chr

ist

Nep

hrol

epis

bis

erra

ta (S

w.)

Scho

tt N

ephr

olep

is b

iser

rata

(Sw.

) Sch

ott

Nep

hrol

epis

bis

erra

ta (S

w.) S

chot

t

Nep

hrol

epis

cor

difo

lia (L

.) C

.Pre

sl

Nep

hrol

epis

cor

difo

lia (L

.) C

.Pre

slN

ephr

olep

is c

ordi

folia

(L.)

C.P

resl

Nep

hrol

epis

dav

allio

ides

(Sw.

) Kun

ze

Nep

hrol

epis

dav

allio

ides

(Sw.

) Kun

zeN

ephr

olep

is d

aval

lioid

es (S

w.) K

unze

●N

ephr

olep

is fa

lcifo

rmis

J.Sm

.16N

ephr

olep

is fa

lcat

a au

ct. n

on (C

av.)

C.C

hr.

Nep

hrol

epis

falc

ata

auct

. non

(Cav

.) C

.Chr

.

Nep

hrol

epis

hir

sutu

la (G

.For

st.)

C.P

resl

N

ephr

olep

is h

irsu

tula

(G.F

orst

.) C

.Pre

slN

ephr

olep

is h

irsu

tula

(G.F

orst

.) C

.Pre

sl

Nep

hrol

epis

radi

cans

(Bur

m.f.

) Kuh

n N

ephr

olep

is ra

dica

ns (B

urm

.f.) K

uhn

Nep

hrol

epis

radi

cans

(Bur

m.f.

) Kuh

n

Page 20: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

83TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●N

ephr

olep

is u

ndul

ata

(afz

el.)

J.Sm

.16N

ephr

olep

is d

elic

atul

a (D

ecne

.) Pi

c.Se

rm.

Nep

hrol

epis

del

icat

ula

(Dec

ne.)

Pic.

Serm

.

LYG

OD

IAC

EA

E

Lygo

dium

circ

inat

um (B

urm

.f.) S

w.

Lygo

dium

circ

inat

um (B

urm

.f.) S

w.Ly

godi

um c

ircin

atum

(Bur

m.f.

) Sw.

Lygo

dium

flex

uosu

m (L

.) Sw

. Ly

godi

um fl

exuo

sum

(L.)

Sw.

Lygo

dium

flex

uosu

m (L

.) Sw

.

Lygo

dium

gig

ante

um T

agaw

a &

K.Iw

ats.

Lygo

dium

gig

ante

um T

agaw

a &

K.Iw

ats.

Lygo

dium

gig

ante

um T

agaw

a &

K.Iw

ats.

Lygo

dium

japo

nicu

m (T

hunb

.) Sw

. Ly

godi

um ja

poni

cum

(Thu

nb.)

Sw.

Lygo

dium

japo

nicu

m (T

hunb

.) Sw

.

Lygo

dium

mic

roph

yllu

m (C

av.)

R.B

r. Ly

godi

um m

icro

phyl

lum

(Cav

.) R

.Br.

Lygo

dium

mic

roph

yllu

m (C

av.)

R.B

r.

Lygo

dium

pol

ysta

chyu

m W

all.

ex T

.Moo

re

Lygo

dium

pol

ysta

chyu

m W

all.

ex T

.Moo

reLy

godi

um p

olys

tach

yum

Wal

l. ex

T.M

oore

Lygo

dium

salic

ifoliu

m C

.Pre

sl

Lygo

dium

salic

ifoliu

m C

.Pre

slLy

godi

um sa

licifo

lium

C.P

resl

MA

RAT

TIA

CE

AE

○An

giop

teri

s ang

ustif

olia

C.P

resl

10A

bsen

tA

bsen

t

Angi

opte

ris e

vect

a (G

.For

st.)

Hof

fm.

Angi

opte

ris e

vect

a (G

.For

st.)

Hof

fm.

Angi

opte

ris e

vect

a (G

.For

st.)

Hof

fm.

Chr

iste

nsen

ia a

escu

lifol

ia (B

lum

e) M

axon

C

hris

tens

enia

aes

culif

olia

(Blu

me)

Max

onC

hris

tens

enia

aes

culif

olia

(Blu

me)

Max

on

●Pt

isan

a sa

mbu

cina

(Blu

me)

Mur

dock

27M

arat

tia sa

mbu

cina

Blu

me

Mar

attia

sam

buci

na B

lum

e

MA

RSI

LE

AC

EA

E

Mar

sile

a cr

enat

a C

.Pre

sl

Mar

sile

a cr

enat

a C

.Pre

slM

arsi

lea

cren

ata

C.P

resl

○M

arsi

lea

scal

arip

es D

.M.Jo

hnso

n44A

bsen

tA

bsen

t

MAT

ON

IAC

EA

E

○M

aton

ia p

ectin

ata

R.B

r.23A

bsen

tA

bsen

t

OL

EA

ND

RA

CE

AE

Ole

andr

a m

usifo

lia (B

lum

e) C

.Pre

sl

Ole

andr

a m

usifo

lia (B

lum

e) C

.Pre

slO

lean

dra

mus

ifolia

(Blu

me)

C.P

resl

Page 21: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

84 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Ole

andr

a pi

still

aris

(Sw.

) C.C

hr.

Ole

andr

a pi

still

aris

(Sw.

) C.C

hr.

Ole

andr

a pi

still

aris

(Sw.

) C.C

hr.

Ole

andr

a un

dula

ta (W

illd.

) Chi

ng

Ole

andr

a un

dula

ta (W

illd.

) Chi

ngO

lean

dra

undu

lata

(Will

d.) C

hing

Ole

andr

a w

allic

hii (

Hoo

k.) C

.Pre

sl

Ole

andr

a w

allic

hii (

Hoo

k.) C

.Pre

slO

lean

dra

wal

lichi

i (H

ook.

) C.P

resl

OPH

IOG

LO

SSA

CE

AE

Botr

ychi

um la

nugi

nosu

m W

all.

ex H

ook.

& G

rev.

Bo

tryc

hium

lanu

gino

sum

Wal

l. ex

Hoo

k. &

Gre

v.Bo

tryc

hium

lanu

gino

sum

Wal

l. ex

Hoo

k. &

Gre

v./ J

apan

obot

rych

ium

lanu

gino

sum

(W

all.

ex H

ook.

& G

rev.

) Nish

ida

ex T

agaw

a

Hel

min

thos

tach

ys ze

ylan

ica

(L.)

Hoo

k.

Hel

min

thos

tach

ys ze

ylan

ica

(L.)

Hoo

k.H

elm

inth

osta

chys

zeyl

anic

a (L

.) H

ook.

Oph

iogl

ossu

m c

osta

tum

R.B

r. O

phio

glos

sum

cos

tatu

m R

.Br.

Oph

iogl

ossu

m c

osta

tum

R.B

r.

Oph

iogl

ossu

m g

ram

ineu

m W

illd.

O

phio

glos

sum

gra

min

eum

Will

d.O

phio

glos

sum

gra

min

eum

Will

d.

Oph

iogl

ossu

m p

endu

lum

L.

Oph

iogl

ossu

m p

endu

lum

L.

Oph

iogl

ossu

m p

endu

lum

L.

Oph

iogl

ossu

m p

etio

latu

m H

ook.

O

phio

glos

sum

pet

iola

tum

Hoo

k.O

phio

glos

sum

pet

iola

tum

Hoo

k.

OSM

UN

DA

CE

AE

Osm

unda

ang

ustif

olia

Chi

ng

Osm

unda

ang

ustif

olia

Chi

ngO

smun

da a

ngus

tifol

ia C

hing

Osm

unda

cin

nam

omea

L.

Osm

unda

cin

nam

omea

L.

Osm

unda

cin

nam

omea

L.

○O

smun

da ja

vani

ca (C

.Pre

sl) B

lum

e4,10

Osm

unda

java

nica

(C.P

resl

) Blu

me

Abs

ent

Osm

unda

vac

helli

i Hoo

k.

Osm

unda

vac

helli

i Hoo

k.O

smun

da v

ache

llii H

ook.

PLA

GIO

GY

RIA

CE

AE

Plag

iogy

ria

adna

ta (B

lum

e) B

edd.

Pl

agio

gyri

a ad

nata

(Blu

me)

Bed

d.Pl

agio

gyri

a ad

nata

(Blu

me)

Bed

d.

Plag

iogy

ria

com

mun

is C

hing

Pl

agio

gyri

a co

mm

unis

Chi

ngPl

agio

gyri

a co

mm

unis

Chi

ng

POLY

POD

IAC

EA

E

●Ac

roso

rus f

ride

rici

-et-p

auli

(Chr

ist)

Cop

el.31

Ac

roso

rus t

rian

gula

ris (

Scor

t. ex

Bed

d.) C

opel

.Ac

roso

rus t

rian

gula

ris (

Scor

t. ex

Bed

d.) C

opel

.

Page 22: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

85TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

○Ac

roso

rus s

trept

ophy

llus (

Bak

er) C

opel

.31A

bsen

tA

bsen

t

●Ag

laom

orph

a ac

umin

ata

(Will

d.) H

oven

kam

p Ph

otin

opte

ris s

peci

osa

(Blu

me)

C.P

resl

Phot

inop

teri

s acu

min

ata

(Will

d.) C

.V.M

orto

n

Agla

omor

pha

coro

nans

(Wal

l. ex

Met

t.) C

opel

. Ag

laom

orph

a co

rona

ns (W

all.

ex M

ett.)

Cop

el.

Agla

omor

pha

coro

nans

(Wal

l. ex

Met

t.) C

opel

.

○Ag

laom

orph

a dr

ynar

ioid

es (H

ook.

) Roo

s10A

bsen

tA

bsen

t

○Ag

laom

orph

a he

racl

ea (K

unze

) Cop

el.4

Agla

omor

pha

hera

clea

(Kun

ze) C

opel

.A

bsen

t

Arth

rom

eris

am

plex

ifolia

(Chr

ist)

Chi

ng

Arth

rom

eris

am

plex

ifolia

(Chr

ist)

Chi

ngAr

thro

mer

is a

mpl

exifo

lia (C

hris

t) C

hing

Arth

rom

eris

lehm

anni

(Met

t.) C

hing

Ar

thro

mer

is le

hman

ni (M

ett.)

Chi

ngAr

thro

mer

is le

hman

ni (M

ett.)

Chi

ng

Arth

rom

eris

phu

luan

gens

is T

agaw

a &

K.Iw

ats.

Arth

rom

eris

phu

luan

gens

is T

agaw

a &

K.Iw

ats.

Arth

rom

eris

phu

luan

gens

is T

agaw

a &

K.Iw

ats.

○Ar

thro

mer

is p

rote

us (C

opel

.) Ta

gaw

a16A

bsen

tA

bsen

t

Arth

rom

eris

tats

iene

nsis

(Fra

nch.

& B

urea

u ex

Chr

ist)

Chi

ng

Arth

rom

eris

tats

iene

nsis

(Fra

nch.

& B

urea

u ex

Chr

ist)

Chi

ngAr

thro

mer

is ta

tsie

nens

is (F

ranc

h. &

Bur

eau

ex C

hris

t) C

hing

Belv

isia

ann

amen

sis (

C.C

hr.)

Taga

wa

Belv

isia

ann

amen

sis (

C.C

hr.)

Taga

wa

Belv

isia

ann

amen

sis (

C.C

hr.)

Taga

wa

Belv

isia

hen

ryi (

Hie

ron.

ex

C.C

hr.)

Taga

wa

Belv

isia

hen

ryi (

Hie

ron.

ex

C.C

hr.)

Taga

wa

Belv

isia

hen

ryi (

Hie

ron.

ex

C.C

hr.)

Taga

wa

Belv

isia

muc

rona

ta (F

ée) C

opel

. Be

lvis

ia m

ucro

nata

(Fée

) Cop

el.

Belv

isia

muc

rona

ta (F

ée) C

opel

.

●Be

lvis

ia sp

icat

a (L

.f.) M

irbel

ex

Cop

el.16

Belv

isia

revo

luta

(Blu

me)

Cop

el.

Belv

isia

revo

luta

(Blu

me)

Cop

el.

Cal

ymm

odon

asi

atic

us C

opel

. C

alym

mod

on a

siat

icus

Cop

el.

Cal

ymm

odon

asi

atic

us C

opel

.

●C

alym

mod

on c

urtu

s Par

ris41

Cal

ymm

odon

cuc

ulla

tus a

uct.

non

(Nee

s & B

lum

e) C

.Pre

sl

Cal

ymm

odon

cuc

ulla

tus a

uct.

non

(Nee

s & B

lum

e) C

.Pre

sl

Chr

istio

pter

is tr

icus

pis (

Hoo

k.) C

hris

t C

hris

tiopt

eris

tric

uspi

s (H

ook.

) Chr

ist

Chr

istio

pter

is tr

icus

pis (

Hoo

k.) C

hris

t

○C

hrys

ogra

mm

itis m

usgr

avia

na (B

aker

) Par

ris32

Abs

ent

Abs

ent

●C

teno

pter

ella

ble

chno

ides

(Gre

v.) P

arris

33C

teno

pter

is m

oulto

nii (

Cop

el.)

C.C

hr. &

Tar

dieu

Cte

nopt

eris

mou

ltoni

i (C

opel

.) C

.Chr

. & T

ardi

eu

●C

teno

pter

ella

kha

olua

ngen

sis (

Taga

wa

& K

.Iwat

s.)

Parr

is 33

Xiph

opte

ris k

haol

uang

ensi

s Tag

awa

& K

.Iwat

s.Xi

phop

teri

s kha

olua

ngen

sis T

agaw

a &

K.Iw

ats.

○D

asyg

ram

miti

s bre

vive

nosa

(Ald

erw.

) Par

ris41

,44

Abs

ent

Abs

ent

●D

asyg

ram

miti

s mol

licom

a (N

ees &

Blu

me)

Par

ris33

Cte

nopt

eris

mol

licom

a (N

ees.

& B

lum

e) K

unze

Cte

nopt

eris

mol

licom

a (N

ees.

& B

lum

e) K

unze

Dry

nari

a bo

nii C

hris

t D

ryna

ria

boni

i Chr

ist

Dry

nari

a bo

nii C

hris

t

Page 23: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

86 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●D

ryna

ria

roos

ii N

akai

ke28

Dry

nari

a fo

rtun

ei (K

unze

ex

Met

t.) J.

Sm.

Dry

nari

a fo

rtun

ei (K

unze

ex

Met

t.) J.

Sm.

Dry

nari

a pa

rish

ii (B

edd.

) Bed

d.

Dry

nari

a pa

rish

ii (B

edd.

) Bed

d.D

ryna

ria

pari

shii

(Bed

d.) B

edd.

Dry

nari

a pr

opin

qua

(Wal

l. ex

Met

t.) J.

Sm.

ex B

edd.

D

ryna

ria

prop

inqu

a (W

all.

ex M

ett.)

J.Sm

. ex

Bed

d.D

ryna

ria

prop

inqu

a (W

all.

ex M

ett.)

J.Sm

. ex

Bed

d.

Dry

nari

a qu

erci

folia

(L.)

J.Sm

. D

ryna

ria

quer

cifo

lia (L

.) J.S

m.

Dry

nari

a qu

erci

folia

(L.)

J.Sm

.

Dry

nari

a ri

gidu

la (S

w.) B

edd.

D

ryna

ria

rigi

dula

(Sw.

) Bed

d.D

ryna

ria

rigi

dula

(Sw.

) Bed

d.

Dry

nari

a sp

arsi

sora

(Des

v.) T

.Moo

re

Dry

nari

a sp

arsi

sora

(Des

v.) T

.Moo

reD

ryna

ria

spar

siso

ra (D

esv.

) T.M

oore

●G

onio

phle

bium

am

oenu

m (

Wal

l. ex

Met

t.) B

edd.

G

onio

phle

bium

am

oenu

m (W

all.

ex M

ett.)

Bed

d.Po

lypo

dium

am

oenu

m W

all.

ex M

ett.

●G

onio

phle

bium

lach

nopu

s J.S

m.36

Poly

podi

um g

arre

ttii C

.H.W

right

Poly

podi

um g

arre

ttii C

.H.W

right

●G

onio

phle

bium

man

mei

ense

(Chr

ist)

Röd

l-Lin

der

Poly

podi

um m

anm

eien

se C

hris

tPo

lypo

dium

man

mei

ense

Chr

ist

●G

onio

phle

bium

men

gtze

ense

(Chr

ist)

Röd

l-Lin

der7

Gon

ioph

lebi

um a

rgut

um J.

Sm. e

x H

ook.

& G

rev.

Poly

podi

um a

rgut

um (J

.Sm

. ex

Hoo

k. &

Gre

v.)

Hoo

k.

●G

onio

phle

bium

mic

rorh

izom

a (C

.B.C

lark

e ex

Bak

er)

Bed

d.G

onio

phle

bium

mic

rorh

izom

a (C

.B.C

lark

e ex

Bak

er)

Bed

d.Po

lypo

dium

mic

rorh

izom

a C

.B.C

lark

e ex

Bak

er

●G

onio

phle

bium

per

cuss

um (C

av.)

Wag

ner &

Gre

ther

16G

onio

phle

bium

ver

ruco

sum

(Hoo

k.) J

.Sm

.Po

lypo

dium

ver

ruco

sum

(Hoo

k.) M

ett.

●G

onio

phle

bium

per

sici

foliu

m (D

esv.

) Bed

d.

Gon

ioph

lebi

um p

ersi

cifo

lium

(Des

v.) B

edd.

Poly

podi

um p

ersi

cifo

lium

Des

v.

●G

onio

phle

bium

suba

uric

ulat

um (B

lum

e) C

.Pre

sl

Gon

ioph

lebi

um su

baur

icul

atum

(Blu

me)

C.P

resl

+

Gon

ioph

lebi

um m

olle

Bed

d.Po

lypo

dium

suba

uric

ulat

um B

lum

e +

Poly

podi

um b

eddo

mei

Bak

er

●G

ymno

gram

miti

s dar

eifo

rmis

(Hoo

k.) C

hing

ex

Tard

ieu

& C

.Chr

.G

ymno

gram

miti

s dar

eifo

rmis

(Hoo

k.) C

hing

ex

Tard

ieu

& C

.Chr

.Ar

aios

tegi

a da

reifo

rmis

(Hoo

k.) C

opel

. /

Gym

nogr

amm

itis d

arei

form

is (H

ook.

) C

hing

ex

Tard

ieu

& C

.Chr

.

●Le

cano

pter

is c

rust

acea

Cop

el.

Leca

nopt

eris

cru

stac

ea C

opel

.M

yrm

ecop

hila

cru

stac

ea (C

opel

.) Ta

gaw

a

○Le

cano

pter

is p

umila

Blu

me22

Abs

ent

Abs

ent

●Le

cano

pter

is si

nuos

a (W

all.

ex H

ook.

) Cop

el.

Leca

nopt

eris

sinu

osa

(Wal

l. ex

Hoo

k.) C

opel

.M

yrm

ecop

hila

sinu

osa

(Wal

l. ex

Hoo

k.) N

akai

ex

H.It

o

Lem

map

hyllu

m a

cced

ens (

Blu

me)

Don

k Le

mm

aphy

llum

acc

eden

s (B

lum

e) D

onk

Lem

map

hyllu

m a

cced

ens (

Blu

me)

Don

k

Page 24: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

87TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Lem

map

hyllu

m c

arno

sum

(Hoo

k.) C

.Pre

sl

Lem

map

hyllu

m c

arno

sum

(Hoo

k.) C

.Pre

slLe

mm

aphy

llum

car

nosu

m (H

ook.

) C.P

resl

Lepi

soru

s bic

olor

Chi

ng

Lepi

soru

s bic

olor

Chi

ngLe

piso

rus b

icol

or C

hing

Lepi

soru

s con

tort

us (C

hris

t) C

hing

Le

piso

rus c

onto

rtus

(Chr

ist)

Chi

ngLe

piso

rus c

onto

rtus

(Chr

ist)

Chi

ng

Lepi

soru

s het

erol

epis

(Ros

enst

.) C

hing

Le

piso

rus h

eter

olep

is (R

osen

st.)

Chi

ngLe

piso

rus h

eter

olep

is (R

osen

st.)

Chi

ng

Lepi

soru

s lon

gifo

lius (

Blu

me)

Hol

ttum

Le

piso

rus l

ongi

foliu

s (B

lum

e) H

olttu

mLe

piso

rus l

ongi

foliu

s (B

lum

e) H

olttu

m

Lepi

soru

s nud

us (H

ook.

) Chi

ng

Lepi

soru

s nud

us (H

ook.

) Chi

ngLe

piso

rus n

udus

(Hoo

k.) C

hing

Lepi

soru

s oos

phae

rus (

C.C

hr.)

Chi

ng

Lepi

soru

s oos

phae

rus (

C.C

hr.)

Chi

ngLe

piso

rus o

osph

aeru

s (C

.Chr

.) C

hing

Lepi

soru

s sco

lope

ndri

um (C

hing

) M

ehra

& B

irLe

piso

rus s

colo

pend

rium

(Chi

ng)

Meh

ra &

Bir

Lepi

soru

s sco

lope

ndri

um (C

hing

) M

ehra

& B

ir

Lepi

soru

s sin

ensi

s (C

hris

t) C

hing

Le

piso

rus s

inen

sis (

Chr

ist)

Chi

ngLe

piso

rus s

inen

sis (

Chr

ist)

Chi

ng

Lepi

soru

s sub

confl

uens

Chi

ng

Lepi

soru

s sub

confl

uens

Chi

ngLe

piso

rus s

ubco

nflue

ns C

hing

Lepi

soru

s sub

linea

ris (

Bak

er e

x Ta

keda

) Chi

ng

Lepi

soru

s sub

linea

ris (

Bak

er e

x Ta

keda

) Chi

ngLe

piso

rus s

ublin

eari

s (B

aker

ex

Take

da) C

hing

Lepi

soru

s sub

olig

olep

idus

Chi

ng

Lepi

soru

s sub

olig

olep

idus

Chi

ngLe

piso

rus s

ubol

igol

epid

us C

hing

Lept

ochi

lus a

xilla

ris (

Cav

.) K

aulf.

Le

ptoc

hilu

s axi

llari

s (C

av.)

Kau

lf.Le

ptoc

hilu

s axi

llari

s (C

av.)

Kau

lf.

Lept

ochi

lus d

ecur

rens

Blu

me

Lept

ochi

lus d

ecur

rens

Blu

me

Lept

ochi

lus d

ecur

rens

Blu

me

●Le

ptoc

hilu

s elli

ptic

us (T

hunb

.) N

oot.29

Col

ysis

pen

taph

ylla

(Bak

er) C

hing

+ C

olys

is p

othi

folia

(D

.Don

) C.P

resl

Col

ysis

pen

taph

ylla

(Bak

er) C

hing

+

Col

ysis

pot

hifo

lia (D

.Don

) C.P

resl

●Le

ptoc

hilu

s hem

ioni

tideu

s (C

.Pre

sl) N

oot.

Col

ysis

hem

ioni

tidea

C.P

resl

Col

ysis

hem

ioni

tidea

C.P

resl

●Le

ptoc

hilu

s mac

roph

yllu

s (B

lum

e) N

oot.

C

olys

is m

acro

phyl

la (B

lum

e) C

.Pre

slC

olys

is m

acro

phyl

la (B

lum

e) C

.Pre

sl

○Le

ptoc

hilu

s min

or F

ée5

Abs

ent

Abs

ent

●Le

ptoc

hilu

s ped

uncu

latu

s (H

ook.

& G

rev.

) Fra

ser-J

enk.

C

olys

is p

edun

cula

ta (H

ook.

& G

rev.

) Chi

ng +

C

olys

is w

ui (C

.Chr

.) C

hing

Col

ysis

ped

uncu

lata

(Hoo

k. &

Gre

v.) C

hing

+

Col

ysis

wui

(C.C

hr.)

Chi

ng

○Lo

xogr

amm

e ac

rosc

opa

(Chr

ist)

C.C

hr.4

Loxo

gram

me

acro

scop

a (C

hris

t) C

.Chr

.A

bsen

t

○Lo

xogr

amm

e as

sim

ilis C

hing

4Lo

xogr

amm

e as

sim

ilis C

hing

Abs

ent

Page 25: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

88 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Loxo

gram

me

aven

ia (B

lum

e) C

.Pre

sl

Loxo

gram

me

aven

ia (B

lum

e) C

.Pre

slLo

xogr

amm

e av

enia

(Blu

me)

C.P

resl

○Lo

xogr

amm

e ce

ntic

ola

M.G

.Pric

e4,8

Loxo

gram

me

cent

icol

a M

.G.P

rice

Abs

ent

Loxo

gram

me

chin

ensi

s Chi

ng

Loxo

gram

me

chin

ensi

s Chi

ngLo

xogr

amm

e ch

inen

sis C

hing

○Lo

xogr

amm

e cu

spid

ata

(Zen

ker)

M.G

.Pric

e4Lo

xogr

amm

e cu

spid

ata

(Zen

ker)

M.G

.Pric

e A

bsen

t

Loxo

gram

me

ducl

ouxi

i Chr

ist

Loxo

gram

me

ducl

ouxi

i Chr

ist

Loxo

gram

me

ducl

ouxi

i Chr

ist

Loxo

gram

me

invo

luta

(D.D

on) C

.Pre

sl

Loxo

gram

me

invo

luta

(D.D

on) C

.Pre

slLo

xogr

amm

e in

volu

ta (D

.Don

) C.P

resl

Loxo

gram

me

lank

okie

nsis

(Ros

enst

.) C

.Chr

. Lo

xogr

amm

e la

nkok

iens

is (R

osen

st.)

C.C

hr.

Loxo

gram

me

lank

okie

nsis

(Ros

enst

.) C

.Chr

.

○Lo

xogr

amm

e po

rcat

a M

.G.P

rice4

Loxo

gram

me

porc

ata

M.G

.Pric

eA

bsen

t

●Lo

xogr

amm

e sc

olop

endr

ioid

es (G

audi

ch.)

C.V

.Mor

ton34

Loxo

gram

me

scol

open

drin

a (B

ory)

C

.Pre

slLo

xogr

amm

e sc

olop

endr

ina

(Bor

y)

C.P

resl

Loxo

gram

me

sube

cost

ata

(Hoo

k.) C

.Chr

. Lo

xogr

amm

e su

beco

stat

a (H

ook.

) C.C

hr.

Loxo

gram

me

sube

cost

ata

(Hoo

k.) C

.Chr

.

Mic

roso

rum

het

eroc

arpu

m (B

lum

e) C

hing

M

icro

soru

m h

eter

ocar

pum

(Blu

me)

Chi

ngM

icro

soru

m h

eter

ocar

pum

(Blu

me)

Chi

ng

●M

icro

soru

m in

sign

e (B

lum

e) C

opel

.29M

icro

soru

m d

ilata

tum

(Bed

d.) S

ledg

eM

icro

soru

m d

ilata

tum

(Bed

d.) S

ledg

e

Mic

roso

rum

mem

bran

aceu

m (D

.Don

) C

hing

M

icro

soru

m m

embr

anac

eum

(D.D

on)

Chi

ngM

icro

soru

m m

embr

anac

eum

(D.D

on)

Chi

ng

○M

icro

soru

m m

usifo

lium

Cop

el.35

Abs

ent

Abs

ent

Mic

roso

rum

pte

ropu

s (B

lum

e) C

opel

. M

icro

soru

m p

tero

pus (

Blu

me)

Cop

el.

Mic

roso

rum

pte

ropu

s (B

lum

e) C

opel

.

Mic

roso

rum

pun

ctat

um (L

.) C

opel

. M

icro

soru

m p

unct

atum

(L.)

Cop

el.

Mic

roso

rum

pun

ctat

um (L

.) C

opel

.

○M

icro

soru

m si

amen

se B

oonk

erd1

Abs

ent

Abs

ent

Mic

roso

rum

supe

rfici

ale

(Blu

me)

Chi

ng

Mic

roso

rum

supe

rfici

ale

(Blu

me)

Chi

ngM

icro

soru

m su

perfi

cial

e (B

lum

e) C

hing

○M

icro

soru

m th

aila

ndic

um B

oonk

erd

& N

oot.3

Mic

roso

rum

sp.

Abs

ent

Mic

roso

rum

zipp

elii

(Blu

me)

Chi

ng

Mic

roso

rum

zipp

elii

(Blu

me)

Chi

ngM

icro

soru

m zi

ppel

ii (B

lum

e) C

hing

Neo

chei

ropt

eris

nor

mal

is (D

.Don

) Tag

awa

Neo

chei

ropt

eris

nor

mal

is (D

.Don

) Tag

awa

Neo

chei

ropt

eris

nor

mal

is (D

.Don

) Tag

awa

●O

reog

ram

miti

s ads

pers

a (B

lum

e) P

arris

33G

ram

miti

s ads

pers

a B

lum

eG

ram

miti

s ads

pers

a B

lum

e

●O

reog

ram

miti

s con

gene

r (B

lum

e) P

arris

41,3

3G

ram

miti

s set

osa

auct

. non

Blu

me

Gra

mm

itis s

etos

a au

ct. n

on B

lum

e

Page 26: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

89TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●O

reog

ram

miti

s dor

sipi

la (C

hris

t) Pa

rris

33G

ram

miti

s dor

sipi

la (C

hris

t) C

.Chr

. &

Tard

ieu

Gra

mm

itis d

orsi

pila

(Chr

ist)

C.C

hr. &

Tard

ieu

●O

reog

ram

miti

s rei

nwar

dtii

(Blu

me)

Par

ris33

Gra

mm

itis b

ongo

ensi

s (C

opel

.) C

opel

. G

ram

miti

s bon

goen

sis (

Cop

el.)

Cop

el.

●Ph

ymat

osor

us c

uspi

datu

s (D

.Don

) Pic

.Ser

m.

Phym

atos

orus

cus

pida

tus (

D.D

on) P

ic.S

erm

.M

icro

soru

m c

uspi

datu

m (D

.Don

) Tag

awa

●Ph

ymat

osor

us lo

ngis

sim

us (B

lum

e) P

ic.S

erm

.Ph

ymat

osor

us lo

ngis

sim

us (B

lum

e) P

ic.S

erm

.M

icro

soru

m ru

bidu

m (K

unze

) Cop

el.

●Ph

ymat

osor

us m

embr

anifo

lius (

R.B

r.) S

.G.L

u25Ph

ymat

osor

us n

igre

scen

s (B

lum

e) P

ic.S

erm

.M

icro

soru

m n

igre

scen

s (B

lum

e) C

opel

.

●Ph

ymat

osor

us sc

olop

endr

ia (B

urm

.f.) P

ic.S

erm

.Ph

ymat

osor

us sc

olop

endr

ia (B

urm

.f.) P

ic.S

erm

.M

icro

soru

m sc

olop

endr

ia (B

urm

.f.) C

opel

.

Plat

ycer

ium

cor

onar

ium

(J.K

oeni

g) D

esv.

Pl

atyc

eriu

m c

oron

ariu

m (J

.Koe

nig)

Des

v.Pl

atyc

eriu

m c

oron

ariu

m (J

.Koe

nig)

Des

v.

Plat

ycer

ium

hol

ttum

ii de

Jonc

h. &

Hen

nipm

an

Plat

ycer

ium

hol

ttum

ii de

Jonc

h. &

Hen

nipm

anPl

atyc

eriu

m h

olttu

mii

de Jo

nch.

& H

enni

pman

○Pl

atyc

eriu

m ri

dley

i Chr

ist7

Plat

ycer

ium

ridl

eyi C

hris

tA

bsen

t

Plat

ycer

ium

wal

lichi

i Hoo

k.

Plat

ycer

ium

wal

lichi

i Hoo

k.Pl

atyc

eriu

m w

allic

hii H

ook.

Pros

aptia

ala

ta (B

lum

e) C

hris

t Pr

osap

tia a

lata

(Blu

me)

Chr

ist

Pros

aptia

ala

ta (B

lum

e) C

hris

t

●Pr

osap

tia b

arat

hrop

hylla

(Bak

er) M

.G.P

rice41

Pr

osap

tia k

hasy

ana

auct

. non

(Hoo

k.) C

.Chr

. &

Tard

ieu

Pros

aptia

kha

syan

a au

ct. n

on (H

ook.

) C.C

hr. &

Ta

rdie

u

Pros

aptia

cel

ebic

a (B

lum

e) T

agaw

a &

K

.Iwat

s. Pr

osap

tia c

eleb

ica

(Blu

me)

Tag

awa

&

K.Iw

ats.

Pros

aptia

cel

ebic

a (B

lum

e) T

agaw

a &

K

.Iwat

s.

Pros

aptia

con

tigua

(G.F

orst

.) C

.Pre

sl

Pros

aptia

con

tigua

(G.F

orst

.) C

.Pre

slPr

osap

tia c

ontig

ua (G

.For

st.)

C.P

resl

Pros

aptia

obl

iqua

ta (B

lum

e) M

ett.

Pros

aptia

obl

iqua

ta (B

lum

e) M

ett.

Pros

aptia

obl

iqua

ta (B

lum

e) M

ett.

●Pr

osap

tia p

ectin

ata

T.M

oore

41Pr

osap

tia le

ysii

(Bak

er) C

hing

Pros

aptia

leys

ii (B

aker

) Chi

ng

●Py

rros

ia a

lbic

ans (

Blu

me)

Chi

ng15

Pyrr

osia

floc

cige

ra (B

lum

e) C

hing

Pyrr

osia

floc

cige

ra (B

lum

e) C

hing

Pyrr

osia

ang

usta

ta (S

w.) C

hing

Py

rros

ia a

ngus

tata

(Sw.

) Chi

ngPy

rros

ia a

ngus

tata

(Sw.

) Chi

ng

Pyrr

osia

ang

ustis

sim

a (G

iese

nh. e

x D

iels

) Ta

gaw

a &

K.Iw

ats.

Pyrr

osia

ang

ustis

sim

a (G

iese

nh. e

x D

iels

) Ta

gaw

a &

K.Iw

ats.

Pyrr

osia

ang

ustis

sim

a (G

iese

nh. e

x D

iels

) Ta

gaw

a &

K.Iw

ats.

Pyrr

osia

cos

tata

(C.P

resl

ex

Bed

d.)

Taga

wa

& K

.Iwat

s. Py

rros

ia c

osta

ta (C

.Pre

sl e

x B

edd.

) Ta

gaw

a &

K.Iw

ats.

Pyrr

osia

cos

tata

(C.P

resl

ex

Bed

d.)

Taga

wa

& K

.Iwat

s.

Pyrr

osia

floc

culo

sa (D

.Don

) Chi

ng

Pyrr

osia

floc

culo

sa (D

.Don

) Chi

ngPy

rros

ia fl

occu

losa

(D.D

on) C

hing

Page 27: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

90 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●Py

rros

ia la

nceo

lata

(L.)

Farw

.15Py

rros

ia la

nceo

lata

(L.)

Farw

. +Py

rros

ia a

dnas

cens

(Sw.

) Chi

ng +

Py

rros

ia n

uda

(Gie

senh

.) C

hing

+

Pyrr

osia

var

ia (K

aulf.

) Far

w.

Pyrr

osia

lanc

eola

ta (L

.) Fa

rw. +

Pyrr

osia

adn

asce

ns (S

w.) C

hing

+

Pyrr

osia

nud

a (G

iese

nh.)

Chi

ng +

Py

rros

ia v

aria

(Kau

lf.) F

arw.

○Py

rros

ia li

ngua

(Thu

nb.)

Farw

. var

. lin

gua4

Pyrr

osia

ling

ua (T

hunb

.) Fa

rw. v

ar. l

ingu

aA

bsen

t

●Py

rros

ia li

ngua

(Thu

nb.)

Farw

. var

. het

erac

tis (M

ett.

ex

Kuh

n) H

oven

kam

pPy

rros

ia li

ngua

(Thu

nb.)

Farw

. var

. het

erac

tis (M

ett.

ex

Kuh

n) H

oven

kam

pPy

rros

ia e

berh

ardt

ii (C

hris

t) C

hing

+

Pyrr

osia

het

erac

tis (M

ett.

ex K

uhn)

Chi

ng

var.

hete

ract

is +

Pyr

rosi

a he

tera

ctis

(Met

t. ex

K

uhn)

Chi

ng v

ar. m

inor

(C.C

hr.)

Chi

ng

Pyrr

osia

long

ifolia

(Bur

m.f.

) C.V

.Mor

ton

Pyrr

osia

long

ifolia

(Bur

m.f.

) C.V

.Mor

ton

Pyrr

osia

long

ifolia

(Bur

m.f.

) C.V

.Mor

ton

Pyrr

osia

man

nii (

Gie

senh

.) C

hing

Py

rros

ia m

anni

i (G

iese

nh.)

Chi

ngPy

rros

ia m

anni

i (G

iese

nh.)

Chi

ng

Pyrr

osia

num

mul

arifo

lia (S

w.)

Chi

ng

Pyrr

osia

num

mul

arifo

lia (S

w.) C

hing

Pyrr

osia

num

mul

arifo

lia (S

w.) C

hing

●Py

rros

ia p

enan

gian

a (H

ook.

) Hol

ttum

Py

rros

ia p

enan

gian

a (H

ook.

) Hol

ttum

+

Pyrr

osia

mol

lis (K

unze

) Chi

ngPy

rros

ia p

enan

gian

a (H

ook.

) Hol

ttum

+

Pyrr

osia

mol

lis (K

unze

) Chi

ng

●Py

rros

ia p

ilose

lloid

es (L

.) M

.G.P

rice

Pyrr

osia

pilo

sello

ides

(L.)

M.G

.Pric

eD

rym

oglo

ssum

pilo

sello

ides

(L.)

C.P

resl

○Py

rros

ia p

oros

a (C

.Pre

sl) H

oven

kam

p va

r. po

rosa

23A

bsen

tA

bsen

t

●Py

rros

ia p

oros

a (C

.Pre

sl) H

oven

kam

p va

r. to

nkin

ensi

s (G

iese

nh.)

Chi

ng

Pyrr

osia

tonk

inen

sis (

Gie

senh

.) C

hing

Pyrr

osia

tonk

inen

sis (

Gie

senh

.) C

hing

○Py

rros

ia ra

sam

alae

(Rac

ib.)

K.H

. Shi

ng10

,15

Abs

ent

Abs

ent

Pyrr

osia

stig

mos

a (S

w.) C

hing

Py

rros

ia st

igm

osa

(Sw.

) Chi

ngPy

rros

ia st

igm

osa

(Sw.

) Chi

ng

●Ra

diog

ram

miti

s jag

oria

na (M

ett.

ex K

uhn)

Par

ris33

Gra

mm

itis j

agor

iana

(Met

t.) T

agaw

aG

ram

miti

s jag

oria

na (M

ett.)

Tag

awa

●Ra

diog

ram

miti

s mul

tifol

ia (C

opel

.) Pa

rris

41,3

3G

ram

miti

s hir

tella

auc

t. no

n (B

lum

e)

Tuya

ma

Gra

mm

itis h

irte

lla a

uct.

non

(Blu

me)

Tu

yam

a

●Sc

lero

glos

sum

pus

illum

(Blu

me)

Ald

erw.

41Sc

lero

glos

sum

min

us a

uct.

non

(Fée

) C.C

hr.

Scle

rogl

ossu

m m

inus

auc

t. no

n (F

ée) C

.Chr

.

●Sc

lero

glos

sum

sulc

atum

(Kuh

n) A

lder

w.41

Sc

lero

glos

sum

pus

illum

auc

t. no

n (B

lum

e)

Ald

erw.

Scle

rogl

ossu

m p

usill

um a

uct.

non

(Blu

me)

A

lder

w.

●Se

lligu

ea c

ruci

form

is (C

hing

) Fra

ser-J

enk.

Cry

psin

us c

ruci

form

is (C

hing

) Tag

awa

Cry

psin

us c

ruci

form

is (C

hing

) Tag

awa

Page 28: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

91TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●Se

lligu

ea e

beni

pes (

Hoo

k.) S

.Lin

ds.

Cry

psin

us e

beni

pes (

Hoo

k.) C

opel

.C

ryps

inus

ebe

nipe

s (H

ook.

) Cop

el.

●Se

lligu

ea e

nerv

is (C

av.)

Chi

ng

Cry

psin

us e

nerv

is (C

av.)

Cop

el.

Cry

psin

us e

nerv

is (C

av.)

Cop

el.

●Se

lligu

ea g

riffi

thia

na (H

ook.

) Fra

ser-J

enk.

Cry

psin

us g

riffi

thia

nus (

Hoo

k.) C

opel

. C

ryps

inus

gri

ffith

ianu

s (H

ook.

) Cop

el.

Selli

guea

het

eroc

arpa

Blu

me

Selli

guea

het

eroc

arpa

Blu

me

Selli

guea

het

eroc

arpa

Blu

me

●Se

lligu

ea h

irsu

ta (T

agaw

a &

K.Iw

ats.)

S.L

inds

. C

ryps

inus

hir

sutu

s Tag

awa

& K

.Iwat

s. C

ryps

inus

hir

sutu

s Tag

awa

& K

.Iwat

s.

●Se

lligu

ea la

cini

ata

(C.P

resl

) Par

risC

ryps

inus

laci

niat

us (C

.Pre

sl) H

olttu

mC

ryps

inus

laci

niat

us (C

.Pre

sl) H

olttu

m

○Se

lligu

ea la

teri

tia (B

aker

) Hov

enka

mp16

Abs

ent

Abs

ent

●Se

lligu

ea o

xylo

ba (W

all.

ex K

unze

) Fra

ser-J

enk.

Cry

psin

us o

xylo

bus (

Wal

l. ex

Kun

ze)

Sled

geC

ryps

inus

oxy

lobu

s (W

all.

ex K

unze

) Sl

edge

●Se

lligu

ea rh

ynch

ophy

lla (H

ook.

) Fra

ser-J

enk.

Cry

psin

us rh

ynch

ophy

llus (

Hoo

k.) C

opel

.C

ryps

inus

rhyn

chop

hyllu

s (H

ook.

) Cop

el.

●Se

lligu

ea st

enop

hylla

(Blu

me)

Par

risC

ryps

inus

sten

ophy

llus (

Blu

me)

Hol

ttum

C

ryps

inus

sten

ophy

llus (

Blu

me)

Hol

ttum

●Se

lligu

ea tr

iloba

(Hou

tt.) M

.G.P

rice

Cry

psin

us tr

ilobu

s (H

outt.

) Cop

el.

Cry

psin

us tr

ilobu

s (H

outt.

) Cop

el.

●Th

emel

ium

tenu

isec

tum

(Blu

me)

Par

ris

Cte

nopt

eris

tenu

isec

ta (B

lum

e) J.

Sm.

Cte

nopt

eris

tenu

isec

ta (B

lum

e) J.

Sm.

●To

mop

hyllu

m su

bfal

catu

m (B

lum

e) P

arris

33C

teno

pter

is su

bfal

cata

(Blu

me)

Kun

zeC

teno

pter

is su

bfal

cata

(Blu

me)

Kun

ze

●Xi

phop

tere

lla h

iero

nym

usii

(C.C

hr.)

Parr

is33

Xiph

opte

ris h

iero

nym

usii

(C.C

hr.)

Hol

ttum

Xi

phop

teri

s hie

rony

mus

ii (C

.Chr

.) H

olttu

m

PSIL

OTA

CE

AE

Psilo

tum

com

plan

atum

Sw.

Ps

ilotu

m c

ompl

anat

um S

w.Ps

ilotu

m c

ompl

anat

um S

w.

Psilo

tum

nud

um (L

.) B

eauv

. Ps

ilotu

m n

udum

(L.)

Bea

uv.

Psilo

tum

nud

um (L

.) B

eauv

.

PTE

RID

AC

EA

E su

bfam

. Che

ilant

hoid

eae

○C

heila

nthe

s arg

ente

a (S

.G.G

mel

) Kun

ze4

Che

ilant

hes a

rgen

tea

(S.G

.Gm

el) K

unze

Abs

ent

Che

ilant

hes b

elan

geri

(Bor

y) C

.Chr

. C

heila

nthe

s bel

ange

ri (B

ory)

C.C

hr.

Che

ilant

hes b

elan

geri

(Bor

y) C

.Chr

.

Che

ilant

hes d

elic

atul

a Ta

gaw

a &

K.Iw

ats.

Che

ilant

hes d

elic

atul

a Ta

gaw

a &

K.Iw

ats.

Che

ilant

hes d

elic

atul

a Ta

gaw

a &

K.Iw

ats.

Page 29: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

92 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●C

heila

nthe

s for

mos

ana

Hay

ata

Che

ilant

hes f

orm

osan

a H

ayat

aC

heila

nthe

s far

inos

a au

ct. n

on (F

orss

k.) K

aulf.

, pr

o pa

rte /

Che

ilant

hes f

orm

osan

a H

ayat

a

Che

ilant

hes f

ragi

lis H

ook.

C

heila

nthe

s fra

gilis

Hoo

k.C

heila

nthe

s fra

gilis

Hoo

k.

●C

heila

nthe

s kra

mer

i Fra

nch.

& S

av.

Che

ilant

hes k

ram

eri F

ranc

h. &

Sav

.C

heila

nthe

s far

inos

a au

ct. n

on (F

orss

k.) K

aulf.

, pr

o pa

rte /

Che

ilant

hes k

ram

eri F

ranc

h. &

Sav

.

●C

heila

nthe

s pse

udoa

rgen

tea

(S.K

.Wu)

K.Iw

ats.

Che

ilant

hes p

seud

oarg

ente

a (S

.K.W

u) K

.Iwat

s.C

heila

nthe

s far

inos

a au

ct. n

on (F

orss

k.)

Kau

lf., p

ro p

arte

/ C

heila

nthe

s pse

udoa

rgen

tea

(S.K

.Wu)

K.Iw

ats.

●C

heila

nthe

s pse

udof

arin

osa

(Chi

ng &

S.K

.Wu)

K.Iw

ats.

Che

ilant

hes p

seud

ofar

inos

a (C

hing

& S

.K.W

u) K

.Iwat

s.C

heila

nthe

s far

inos

a au

ct. n

on (F

orss

k.) K

aulf.

, pr

o pa

rte /

Che

ilant

hes p

seud

ofar

inos

a (C

hing

&

S.K

.Wu)

K.Iw

ats.

●C

heila

nthe

s ruf

a D

.Don

C

heila

nthe

s ruf

a D

.Don

Che

ilant

hes r

ufa

D.D

on +

Che

ilant

hes s

ubru

fa

auct

. non

Bak

er

●C

heila

nthe

s sia

men

sis (

S.K

.Wu)

K.Iw

ats.

Che

ilant

hes s

iam

ensi

s (S.

K.W

u) K

.Iwat

s.C

heila

nthe

s far

inos

a au

ct. n

on (F

orss

k.) K

aulf.

, pr

o pa

rte /

Che

ilant

hes s

iam

ensi

s (S.

K.W

u)

K.Iw

ats.

Che

ilant

hes t

enui

folia

(Bur

m.f.

) Sw.

C

heila

nthe

s ten

uifo

lia (B

urm

.f.) S

w.C

heila

nthe

s ten

uifo

lia (B

urm

.f.) S

w.

○D

oryo

pter

is a

lleni

ae r

.M.T

ryon

10A

bsen

tA

bsen

t

Dor

yopt

eris

lude

ns (W

all.

ex H

ook.

) J.S

m.

Dor

yopt

eris

lude

ns (W

all.

ex H

ook.

) J.S

m.

Dor

yopt

eris

lude

ns (W

all.

ex H

ook.

) J.S

m.

●H

emio

nitis

ves

tita

(Wal

l. ex

Hoo

k.) J

.Sm

. G

ymno

pter

is v

estit

a (W

all.

ex H

ook.

) Und

erw.

Gym

nopt

eris

ves

tita

(Wal

l. ex

Hoo

k.) U

nder

w.

Not

hola

ena

velu

tina

Tard

ieu

& C

.Chr

. N

otho

laen

a ve

lutin

a Ta

rdie

u &

C.C

hr.

Not

hola

ena

velu

tina

Tard

ieu

& C

.Chr

.

●Pa

rahe

mio

nitis

cor

data

(Rox

b. e

x H

ook.

& G

rev.

) Fr

aser

-Jen

k.11

Hem

ioni

tis a

rifo

lia (B

urm

.f.) T

.Moo

reH

emio

nitis

ari

folia

(Bur

m.f.

) T.M

oore

○Pe

llaea

tim

oren

sis A

lder

w.4

Pella

ea ti

mor

ensi

s Ald

erw.

Abs

ent

PTE

RID

AC

EA

E su

bfam

. Cry

ptog

ram

moi

deae

Con

iogr

amm

e fr

axin

ea (D

.Don

) Die

ls

var.

serr

ulat

a (B

lum

e) H

iero

n.

Con

iogr

amm

e fr

axin

ea (D

.Don

) Die

ls

var.

serr

ulat

a (B

lum

e) H

iero

n.C

onio

gram

me

frax

inea

(D.D

on) D

iels

va

r. se

rrul

ata

(Blu

me)

Hie

ron.

Page 30: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

93TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Con

iogr

amm

e pe

telo

tii T

ardi

eu

Con

iogr

amm

e pe

telo

tii T

ardi

euC

onio

gram

me

pete

lotii

Tar

dieu

Con

iogr

amm

e pr

ocer

a Fé

e C

onio

gram

me

proc

era

Fée

Con

iogr

amm

e pr

ocer

a Fé

e

PTE

RID

AC

EA

E su

bfam

. Par

keri

oide

ae

Acro

stic

hum

aur

eum

L.

Acro

stic

hum

aur

eum

L.

Acro

stic

hum

aur

eum

L.

Acro

stic

hum

spec

iosu

m W

illd.

Ac

rost

ichu

m sp

ecio

sum

Will

d.Ac

rost

ichu

m sp

ecio

sum

Will

d.

Cer

atop

teri

s tha

lictro

ides

(L.)

Bro

ngn.

C

erat

opte

ris t

halic

troid

es (L

.) B

rong

n.C

erat

opte

ris t

halic

troid

es (L

.) B

rong

n.

PTE

RID

AC

EA

E su

bfam

. Pte

rido

idea

e

Ony

chiu

m c

ontig

uum

C.H

ope

Ony

chiu

m c

ontig

uum

C.H

ope

Ony

chiu

m c

ontig

uum

C.H

ope

Ony

chiu

m si

licul

osum

(Des

v.) C

.Chr

. O

nych

ium

silic

ulos

um (D

esv.

) C.C

hr.

Ony

chiu

m si

licul

osum

(Des

v.) C

.Chr

.

Pity

rogr

amm

a ca

lom

elan

os (L

.) Li

nk

Pity

rogr

amm

a ca

lom

elan

os (L

.) Li

nkPi

tyro

gram

ma

calo

mel

anos

(L.)

Link

Pter

is a

sper

icau

lis W

all.

ex J.

aga

rdh

Pter

is a

sper

icau

lis W

all.

ex J.

aga

rdh

Pter

is a

sper

icau

lis W

all.

ex J.

aga

rdh

Pter

is a

sper

ula

J.Sm

. ex

Hie

ron.

Pt

eris

asp

erul

a J.S

m. e

x H

iero

n.Pt

eris

asp

erul

a J.S

m. e

x H

iero

n.

Pter

is b

ella

Tag

awa

Pter

is b

ella

Tag

awa

Pter

is b

ella

Tag

awa

Pter

is b

iaur

ita L

. Pt

eris

bia

urita

L.

Pter

is b

iaur

ita L

.

Pter

is b

lum

eana

J.a

gard

h Pt

eris

blu

mea

na J.

aga

rdh

Pter

is b

lum

eana

J.a

gard

h

Pter

is c

retic

a L.

Pt

eris

cre

tica

L.Pt

eris

cre

tica

L.

Pter

is d

alho

usia

e H

ook.

Pt

eris

dal

hous

iae

Hoo

k.Pt

eris

dal

hous

iae

Hoo

k.

Pter

is d

ecre

scen

s Chr

ist

Pter

is d

ecre

scen

s Chr

ist

Pter

is d

ecre

scen

s Chr

ist

Pter

is e

nsifo

rmis

Bur

m.f.

Pt

eris

ens

iform

is B

urm

.f.Pt

eris

ens

iform

is B

urm

.f.

Pter

is g

revi

llean

a W

all.

ex J.

aga

rdh

Pter

is g

revi

llean

a W

all.

ex J.

aga

rdh

Pter

is g

revi

llean

a W

all.

ex J.

aga

rdh

Pter

is h

eter

omor

pha

Fée

Pter

is h

eter

omor

pha

Fée

Pter

is h

eter

omor

pha

Fée

Pter

is li

near

is P

oir.

Pter

is li

near

is P

oir.

Pter

is li

near

is P

oir.

Page 31: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

94 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Pter

is lo

ngip

es D

.Don

Pt

eris

long

ipes

D.D

onPt

eris

long

ipes

D.D

on

Pter

is lo

ngip

innu

la W

all.

ex J.

aga

rdh

Pter

is lo

ngip

innu

la W

all.

ex J.

aga

rdh

Pter

is lo

ngip

innu

la W

all.

ex J.

aga

rdh

Pter

is m

erte

nsio

ides

Will

d.

Pter

is m

erte

nsio

ides

Will

d.Pt

eris

mer

tens

ioid

es W

illd.

Pter

is m

ultifi

da P

oir.

Pter

is m

ultifi

da P

oir.

Pter

is m

ultifi

da P

oir.

Pter

is n

epal

ensi

s H.It

o Pt

eris

nep

alen

sis H

.Ito

Pter

is n

epal

ensi

s H.It

o

Pter

is p

hulu

ange

nsis

Tag

awa

& K

.Iwat

s. Pt

eris

phu

luan

gens

is T

agaw

a &

K.Iw

ats.

Pter

is p

hulu

ange

nsis

Tag

awa

& K

.Iwat

s.

Pter

is p

lum

bea

Chr

ist

Pter

is p

lum

bea

Chr

ist

Pter

is p

lum

bea

Chr

ist

Pter

is sc

abri

pes W

all.

ex J.

aga

rdh

Pter

is sc

abri

pes W

all.

ex J.

aga

rdh

Pter

is sc

abri

pes W

all.

ex J.

aga

rdh

Pter

is se

mip

inna

ta L

. Pt

eris

sem

ipin

nata

L.

Pter

is se

mip

inna

ta L

.

Pter

is st

enop

hylla

Wal

l. ex

Hoo

k. &

Gre

v.

Pter

is st

enop

hylla

Wal

l. ex

Hoo

k. &

Gre

v.Pt

eris

sten

ophy

lla W

all.

ex H

ook.

& G

rev.

Pter

is su

bqui

nata

Wal

l. ex

J.a

gard

h Pt

eris

subq

uina

ta W

all.

ex J.

aga

rdh

Pter

is su

bqui

nata

Wal

l. ex

J.a

gard

h

Pter

is to

kioi

Mas

am.

Pter

is to

kioi

Mas

am.

Pter

is to

kioi

Mas

am.

Pter

is tr

ipar

tita

Sw.

Pter

is tr

ipar

tita

Sw.

Pter

is tr

ipar

tita

Sw.

Pter

is v

enus

ta K

unze

Pt

eris

ven

usta

Kun

zePt

eris

ven

usta

Kun

ze

Pter

is v

ittat

a L.

Pt

eris

vitt

ata

L.Pt

eris

vitt

ata

L.

Pter

is w

allic

hian

a J.a

gard

h Pt

eris

wal

lichi

ana

J.aga

rdh

Pter

is w

allic

hian

a J.a

gard

h

Syng

ram

ma

alis

mifo

lia (C

.Pre

sl) J

.Sm

. Sy

ngra

mm

a al

ism

ifolia

(C.P

resl

) J.S

m.

Syng

ram

ma

alis

mifo

lia (C

.Pre

sl) J

.Sm

.

Taen

itis b

lech

noid

es (W

illd.

) Sw

. Ta

eniti

s ble

chno

ides

(Will

d.) S

w.Ta

eniti

s ble

chno

ides

(Will

d.) S

w.

○Ta

eniti

s int

erru

pta

Hoo

k. &

Gre

v.10

Abs

ent

Abs

ent

PTE

RID

AC

EA

E su

bfam

. Vitt

ario

idea

e

Adia

ntum

cap

illus

-ven

eris

L.

Adia

ntum

cap

illus

-ven

eris

L.

Adia

ntum

cap

illus

-ven

eris

L.

Adia

ntum

cau

datu

m L

. Ad

iant

um c

auda

tum

L.

Adia

ntum

cau

datu

m L

.

Adia

ntum

edg

ewor

thii

Hoo

k.

Adia

ntum

edg

ewor

thii

Hoo

k.Ad

iant

um e

dgew

orth

ii H

ook.

Page 32: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

95TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Adia

ntum

ery

lliae

C.C

hr. &

Tar

dieu

Ad

iant

um e

rylli

ae C

.Chr

. & T

ardi

euAd

iant

um e

rylli

ae C

.Chr

. & T

ardi

eu

Adia

ntum

flab

ellu

latu

m L

. Ad

iant

um fl

abel

lula

tum

L.

Adia

ntum

flab

ellu

latu

m L

.

●Ad

iant

um fr

agili

form

e C

.Chr

.46Ad

iant

um st

enoc

hlam

ys B

aker

, pro

par

teAd

iant

um st

enoc

hlam

ys B

aker

, pro

par

te

○Ad

iant

um la

tifol

ium

Lam

.10A

bsen

tA

bsen

t

○Ad

iant

um p

hano

men

sis S

.Lin

ds. &

D.J.

Mid

dlet

on21

Abs

ent

Abs

ent

Adia

ntum

phi

lippe

nse

L.43

Adia

ntum

phi

lippe

nse

L.Ad

iant

um p

hilip

pens

e L.

Adia

ntum

siam

ense

Tag

awa

& K

.Iwat

s. Ad

iant

um si

amen

se T

agaw

a &

K.Iw

ats.

Adia

ntum

siam

ense

Tag

awa

& K

.Iwat

s.

Adia

ntum

sobo

lifer

um W

all.

ex H

ook.

Ad

iant

um so

bolif

erum

Wal

l. ex

Hoo

k.Ad

iant

um so

bolif

erum

Wal

l. ex

Hoo

k.

Adia

ntum

sten

ochl

amys

Bak

er

Adia

ntum

sten

ochl

amys

Bak

er, p

ro p

arte

Adia

ntum

sten

ochl

amys

Bak

er, p

ro p

arte

○Ad

iant

um th

ongt

ham

ii Su

ksat

han37

Abs

ent

Abs

ent

Adia

ntum

zolli

nger

i Met

t. ex

Kuh

n Ad

iant

um zo

lling

eri M

ett.

ex K

uhn

Adia

ntum

zolli

nger

i Met

t. ex

Kuh

n

Antro

phyu

m c

allif

oliu

m B

lum

e An

troph

yum

cal

lifol

ium

Blu

me

Antro

phyu

m c

allif

oliu

m B

lum

e

Antro

phyu

m o

bova

tum

Bak

er

Antro

phyu

m o

bova

tum

Bak

erAn

troph

yum

obo

vatu

m B

aker

Antro

phyu

m p

arvu

lum

Blu

me

Antro

phyu

m p

arvu

lum

Blu

me

Antro

phyu

m p

arvu

lum

Blu

me

●An

troph

yum

vitt

ario

ides

Bak

er39

Antro

phyu

m st

enop

hyllu

m B

aker

Antro

phyu

m st

enop

hyllu

m B

aker

●H

aplo

pter

is a

mbo

inen

sis (

Fée)

X.C

.Zha

ng40

Vitta

ria

ambo

inen

sis F

ée +

Vitt

aria

fo

rres

tiana

auc

t. no

n C

hing

Vitta

ria

ambo

inen

sis F

ée +

Vitt

aria

fo

rres

tiana

auc

t. no

n C

hing

●H

aplo

pter

is a

ngus

tifol

ia (B

lum

e) E

.H.C

rane

Vi

ttari

a an

gust

ifolia

Blu

me

Vitta

ria

angu

stifo

lia B

lum

e

●H

aplo

pter

is e

long

ata

(Sw

.) E.

H.C

rane

Vi

ttari

a el

onga

ta S

w.Vi

ttari

a el

onga

ta S

w.

●H

aplo

pter

is e

nsifo

rmis

(Sw.

) E.H

.Cra

ne

Vitta

ria

ensi

form

is S

w.Vi

ttari

a en

sifo

rmis

Sw.

●H

aplo

pter

is fl

exuo

sa (F

ée) E

.H.C

rane

Vi

ttari

a fle

xuos

a Fé

eVi

ttari

a fle

xuos

a Fé

e

●H

aplo

pter

is sc

olop

endr

ina

C.P

resl

Vi

ttari

a sc

olop

endr

ina

(Bor

y) T

hwai

tes &

H

ook.

Vitta

ria

scol

open

drin

a (B

ory)

Thw

aite

s &

Hoo

k.

●H

aplo

pter

is si

kkim

ensi

s (K

uhn)

E.H

.Cra

ne

Vitta

ria

sikk

imen

sis K

uhn

Vitta

ria

sikk

imen

sis K

uhn

●H

aplo

pter

is ta

enio

phyl

la (C

opel

.) E.

H.C

rane

Vi

ttari

a ta

enio

phyl

la C

opel

.Vi

ttari

a ta

enio

phyl

la C

opel

.

Page 33: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

96 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●H

aplo

pter

is w

initi

i (Ta

gaw

a &

K.Iw

ats.)

S.L

inds

. An

troph

yum

win

itii T

agaw

a &

K.Iw

ats.

Antro

phyu

m w

initi

i Tag

awa

& K

.Iwat

s.

Vagi

nula

ria

para

doxa

(Fée

) Met

t. Va

ginu

lari

a pa

rado

xa (F

ée) M

ett.

Vagi

nula

ria

para

doxa

(Fée

) Met

t.

Vagi

nula

ria

tric

hoid

ea F

ée

Vagi

nula

ria

tric

hoid

ea F

éeVa

ginu

lari

a tr

icho

idea

Fée

SALV

INIA

CE

AE

Azol

la c

arol

inia

na W

illd.

Az

olla

car

olin

iana

Will

d.Az

olla

car

olin

iana

Will

d.

Azol

la p

inna

ta R

.Br.

Azol

la p

inna

ta R

.Br.

Azol

la p

inna

ta R

.Br.

Salv

inia

cuc

ulla

ta R

oxb.

ex

Bor

y Sa

lvin

ia c

ucul

lata

Rox

b. e

x B

ory

Salv

inia

cuc

ulla

ta R

oxb.

ex

Bor

y

Salv

inia

nat

ans (

L.) A

ll.

Salv

inia

nat

ans (

L.) A

ll.Sa

lvin

ia n

atan

s (L.

) All.

SCH

IZA

EA

CE

AE

Schi

zaea

dic

hoto

ma

(L.)

Sm.

Schi

zaea

dic

hoto

ma

(L.)

Sm.

Schi

zaea

dic

hoto

ma

(L.)

Sm.

Schi

zaea

dig

itata

(L.)

Sw.

Schi

zaea

dig

itata

(L.)

Sw.

Schi

zaea

dig

itata

(L.)

Sw.

TE

CTA

RIA

CE

AE

Arth

ropt

eris

pal

isot

ii (D

esv.

) Als

ton

Arth

ropt

eris

pal

isot

ii (D

esv.

) Als

ton

Arth

ropt

eris

pal

isot

ii (D

esv.

) Als

ton

Het

erog

oniu

m a

lder

wer

eltii

Hol

ttum

H

eter

ogon

ium

ald

erw

erel

tii H

olttu

mH

eter

ogon

ium

ald

erw

erel

tii H

olttu

m

Het

erog

oniu

m g

urup

ahen

se (C

.Chr

.) H

olttu

m

Het

erog

oniu

m g

urup

ahen

se (C

.Chr

.) H

olttu

mH

eter

ogon

ium

gur

upah

ense

(C.C

hr.)

Hol

ttum

Het

erog

oniu

m h

enni

pman

ii Ta

gaw

a &

K.Iw

ats.

Het

erog

oniu

m h

enni

pman

ii Ta

gaw

a &

K.Iw

ats.

Het

erog

oniu

m h

enni

pman

ii Ta

gaw

a &

K.Iw

ats.

Het

erog

oniu

m p

inna

tum

(Cop

el.)

Hol

ttum

H

eter

ogon

ium

pin

natu

m (C

opel

.) H

olttu

mH

eter

ogon

ium

pin

natu

m (C

opel

.) H

olttu

m

Het

erog

oniu

m sa

geni

oide

s (M

ett.)

Hol

ttum

H

eter

ogon

ium

sage

nioi

des (

Met

t.) H

olttu

mH

eter

ogon

ium

sage

nioi

des (

Met

t.) H

olttu

m

Pleo

cnem

ia h

emite

liifo

rmis

(Rac

ib.)

Hol

ttum

Pl

eocn

emia

hem

itelii

form

is (R

acib

.) H

olttu

mPl

eocn

emia

hem

itelii

form

is (R

acib

.) H

olttu

m

Pleo

cnem

ia ir

regu

lari

s (C

.Pre

sl) H

olttu

m

Pleo

cnem

ia ir

regu

lari

s (C

.Pre

sl) H

olttu

mPl

eocn

emia

irre

gula

ris (

C.P

resl

) Hol

ttum

Pleo

cnem

ia su

bmem

bran

acea

(Hay

ata)

Tag

awa

&

K.Iw

ats.

Pleo

cnem

ia su

bmem

bran

acea

(Hay

ata)

Tag

awa

&

K.Iw

ats.

Pleo

cnem

ia su

bmem

bran

acea

(Hay

ata)

Tag

awa

&

K.Iw

ats.

Page 34: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

97TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Pter

idry

s aus

tral

is C

hing

Pt

erid

rys a

ustr

alis

Chi

ngPt

erid

rys a

ustr

alis

Chi

ng

Pter

idry

s cne

mid

aria

(Chr

ist)

C.C

hr. &

C

hing

Pt

erid

rys c

nem

idar

ia (C

hris

t) C

.Chr

. &

Chi

ngPt

erid

rys c

nem

idar

ia (C

hrist

) C.C

hr. &

Chi

ng

Pter

idry

s syr

mat

ica

(Will

d.) C

.Chr

. & C

hing

Pt

erid

rys s

yrm

atic

a (W

illd.

) C.C

hr. &

Chi

ngPt

erid

rys s

yrm

atic

a (W

illd.

) C.C

hr. &

Chi

ng

Tect

aria

am

plifo

lia (A

lder

w.) C

.Chr

. Te

ctar

ia a

mpl

ifolia

(Ald

erw.

) C.C

hr.

Tect

aria

am

plifo

lia (A

lder

w.) C

.Chr

.

Tect

aria

ang

ulat

a (W

illd.

) C.C

hr.

Tect

aria

ang

ulat

a (W

illd.

) C.C

hr.

Tect

aria

ang

ulat

a (W

illd.

) C.C

hr.

Tect

aria

bar

beri

(Hoo

k.) C

opel

. Te

ctar

ia b

arbe

ri (H

ook.

) Cop

el.

Tect

aria

bar

beri

(Hoo

k.) C

opel

.

○Te

ctar

ia b

rach

iata

(Zol

l. &

Mor

itzi)

C.V

.Mor

ton4

Tect

aria

bra

chia

ta (Z

oll.

& M

oritz

i) C

.V.M

orto

nA

bsen

t

●Te

ctar

ia c

hris

tii C

opel

.13Te

ctar

ia c

hris

tii C

opel

.Te

ctar

ia c

oadu

nata

(J.S

m.)

C.Ch

r., p

ro p

arte

Tect

aria

coa

duna

ta (J

.Sm

.) C

.Chr

.ab

sent

Tect

aria

coa

duna

ta (J

.Sm

.) C.

Chr.,

pro

par

te

Tect

aria

cre

nata

Cav

. Te

ctar

ia c

rena

ta C

av.

Tect

aria

cre

nata

Cav

.

Tect

aria

dec

urre

ns (C

.Pre

sl) C

opel

. Te

ctar

ia d

ecur

rens

(C.P

resl

) Cop

el.

Tect

aria

dec

urre

ns (C

.Pre

sl) C

opel

.

Tect

aria

dev

exa

(Kun

ze) C

opel

. Te

ctar

ia d

evex

a (K

unze

) Cop

el.

Tect

aria

dev

exa

(Kun

ze) C

opel

.

Tect

aria

faur

iei T

agaw

a Te

ctar

ia fa

urie

i Tag

awa

Tect

aria

faur

iei T

agaw

a

●Te

ctar

ia fi

ssa

(Kun

ze) H

olttu

m13

Tect

aria

rum

icifo

lia (R

idl.)

C.C

hr.

Tect

aria

rum

icifo

lia (R

idl.)

C.C

hr.

Tect

aria

fusc

ipes

(Wal

l. ex

Bed

d.) C

.Chr

. Te

ctar

ia fu

scip

es (W

all.

ex B

edd.

) C.C

hr.

Tect

aria

fusc

ipes

(Wal

l. ex

Bed

d.) C

.Chr

.

Tect

aria

gri

ffith

ii (B

aker

) C.C

hr.

Tect

aria

gri

ffith

ii (B

aker

) C.C

hr.

Tect

aria

gri

ffith

ii (B

aker

) C.C

hr.

Tect

aria

gym

noso

ra H

olttu

m

Tect

aria

gym

noso

ra H

olttu

mTe

ctar

ia g

ymno

sora

Hol

ttum

Tect

aria

her

peto

caul

os H

olttu

m

Tect

aria

her

peto

caul

os H

olttu

mTe

ctar

ia h

erpe

toca

ulos

Hol

ttum

●Te

ctar

ia im

pres

sa (F

ée) H

olttu

m

Tect

aria

impr

essa

(Fée

) Hol

ttum

Tect

aria

var

iolo

sa (W

all.

ex H

ook.

) C.C

hr.

○Te

ctar

ia k

ecki

i (Lu

erss

.) C

.Chr

.4Te

ctar

ia k

ecki

i (Lu

erss

.) C

.Chr

.A

bsen

t

Tect

aria

laot

ica

Tard

ieu

& C

.Chr

. Te

ctar

ia la

otic

a Ta

rdie

u &

C.C

hr.

Tect

aria

laot

ica

Tard

ieu

& C

.Chr

.

●Te

ctar

ia m

anile

nsis

(C.P

resl

) Hol

ttum

Te

ctar

ia m

anile

nsis

(C.P

resl

) Hol

ttum

va

r. m

anile

nsis

+ T

ecta

ria

man

ilens

is (C

.Pre

sl) H

olttu

m

var.

chup

enge

nsis

(Rid

l.) H

olttu

m

Cte

nitis

man

ilens

is (C

.Pre

sl) H

olttu

m

Page 35: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

98 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

Tect

aria

mel

anoc

aulis

(Blu

me)

Cop

el.

Tect

aria

mel

anoc

aulis

(Blu

me)

Cop

el.

Tect

aria

mel

anoc

aulis

(Blu

me)

Cop

el.

Tect

aria

pha

eoca

ulis

(Ros

enst

.) C

.Chr

. Te

ctar

ia p

haeo

caul

is (R

osen

st.)

C.C

hr.

Tect

aria

pha

eoca

ulis

(Ros

enst

.) C

.Chr

.

○Te

ctar

ia p

hano

men

sis S

.Lin

ds.24

Abs

ent

Abs

ent

○Te

ctar

ia p

ilosa

(Fée

) r.C

.Mor

an4

Tect

aria

pilo

sa (F

ée) r

.C.M

oran

Abs

ent

Tect

aria

pol

ymor

pha

(Wal

l. ex

Hoo

k.)

Cop

el.

Tect

aria

pol

ymor

pha

(Wal

l. ex

Hoo

k.)

Cop

el.

Tect

aria

pol

ymor

pha

(Wal

l. ex

Hoo

k.)

Cop

el.

Tect

aria

rock

ii C

.Chr

. Te

ctar

ia ro

ckii

C.C

hr.

Tect

aria

rock

ii C

.Chr

.

●Te

ctar

ia se

mip

inna

ta (r

oxb.

) C.V

.Mor

ton

Tect

aria

sem

ipin

nata

(rox

b.) C

.V.M

orto

nTe

ctar

ia m

aing

ayi (

Bak

er) C

.Chr

.

○Te

ctar

ia sh

ahid

ania

na R

usea

44A

bsen

tA

bsen

t

○Te

ctar

ia si

ifolia

(Will

d.) C

opel

.4Te

ctar

ia si

ifolia

(Will

d.) C

opel

.A

bsen

t

Tect

aria

sim

onsi

i (B

aker

) Chi

ng

Tect

aria

sim

onsi

i (B

aker

) Chi

ngTe

ctar

ia si

mon

sii (

Bak

er) C

hing

Tect

aria

sing

apor

eana

(Wal

l. ex

Hoo

k. &

G

rev.

) Chi

ng

Tect

aria

sing

apor

eana

(Wal

l. ex

Hoo

k. &

G

rev.

) Chi

ngTe

ctar

ia si

ngap

orea

na (W

all.

ex H

ook.

&

Gre

v.) C

hing

Tect

aria

tene

rifro

ns (H

ook.

) Chi

ng

Tect

aria

tene

rifro

ns (H

ook.

) Chi

ngTe

ctar

ia te

neri

frons

(Hoo

k.) C

hing

Tect

aria

tern

ifolia

(Ald

erw.

) C.C

hr.

Tect

aria

tern

ifolia

(Ald

erw.

) C.C

hr.

Tect

aria

tern

ifolia

(Ald

erw.

) C.C

hr.

Tect

aria

vas

ta (B

lum

e) C

opel

. Te

ctar

ia v

asta

(Blu

me)

Cop

el.

Tect

aria

vas

ta (B

lum

e) C

opel

.

●Te

ctar

ia ze

ilani

ca (H

outt.

) Sle

dge

Que

rcifi

lix ze

ilani

ca (H

outt.

) Cop

el.

Que

rcifi

lix ze

ilani

ca (H

outt.

) Cop

el.

TH

ELY

PTE

RID

AC

EA

E

●C

yclo

soru

s ari

dus (

D.D

on) C

hing

C

hris

tella

ari

da (D

.Don

) Hol

ttum

Thel

ypte

ris a

rida

(D.D

on) C

.V.M

orto

n

●C

yclo

soru

s art

icul

atus

(Hou

lsto

n &

T.M

oore

) Pa

nigr

ahi

Pron

ephr

ium

art

icul

atum

(Hou

lsto

n &

T.M

oore

) H

olttu

mTh

elyp

teri

s art

icul

ata

(Hou

lsto

n &

T.M

oore

) Ta

gaw

a &

K.Iw

ats.

●C

yclo

soru

s asp

erus

(C.P

resl

) B.K

.Nay

ar &

Kau

r Pr

onep

hriu

m a

sper

um (C

.Pre

sl) H

olttu

mTh

elyp

teri

s asp

era

(C.P

resl

) K.Iw

ats.

●C

yclo

soru

s can

us (B

aker

) S.L

inds

.14Th

elyp

teri

s rep

ens (

C.H

ope)

Chi

ngTh

elyp

teri

s rep

ens (

C.H

ope)

Chi

ng

●C

yclo

soru

s cili

atus

(Wal

l. ex

Ben

th.)

Pani

grah

i Tr

igon

ospo

ra c

iliat

a (W

all.

ex B

enth

.) H

olttu

mTh

elyp

teri

s cili

ata

(Wal

l. ex

Ben

th.)

Chi

ng

Page 36: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

99TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●C

yclo

soru

s cra

ssifo

lius (

Blu

me)

S.L

inds

. M

esop

hleb

ion

cras

sifo

lium

(Blu

me)

Hol

ttum

Thel

ypte

ris c

rass

ifolia

(Blu

me)

Chi

ng

●C

yclo

soru

s cri

nipe

s (H

ook.

) Chi

ng

Chr

iste

lla c

rini

pes (

Hoo

k.) H

olttu

mTh

elyp

teri

s cri

nipe

s (H

ook.

) K.Iw

ats.

●C

yclo

soru

s cla

rkei

(Bed

d.) C

hing

11C

yclo

soru

s cyl

indr

othr

ix (R

osen

st.)

Chi

ngTh

elyp

teri

s cyl

indr

othr

ix (R

osen

st.)

K.Iw

ats.

●C

yclo

soru

s den

tatu

s (Fo

rssk

.) C

hing

C

hris

tella

den

tata

(For

ssk.

) Bro

wns

ey &

Jerm

yTh

elyp

teri

s den

tata

(For

ssk.

) e.P

.St.J

ohn

●C

yclo

soru

s evo

lutu

s (C

.B.C

lark

e &

Bak

er)

Chi

ng

Chr

iste

lla e

volu

ta (C

.B.C

lark

e &

Bak

er)

Hol

ttum

Thel

ypte

ris e

volu

ta (C

.B.C

lark

e &

Bak

er)

Taga

wa

& K

.Iwat

s

●C

yclo

soru

s exs

culp

tus (

Bak

er) S

.Lin

ds.

Pron

ephr

ium

exs

culp

tum

(Bak

er) H

olttu

mTh

elyp

teri

s exs

culp

ta (B

aker

) K.Iw

ats.

●C

yclo

soru

s fal

cilo

bus (

Hoo

k.) P

anig

rahi

Th

elyp

teri

s fal

cilo

ba (H

ook.

) Chi

ngTh

elyp

teri

s fal

cilo

ba (H

ook.

) Chi

ng

●C

yclo

soru

s fer

ox (B

lum

e) C

hing

C

hing

ia fe

rox

(Blu

me)

Hol

ttum

Thel

ypte

ris f

erox

(Blu

me)

Tag

awa

&

K.Iw

ats.

●C

yclo

soru

s gla

ndul

osus

(Blu

me)

Chi

ng

Pron

ephr

ium

gla

ndul

osum

(Blu

me)

Hol

ttum

Thel

ypte

ris g

land

ulos

a (B

lum

e) T

agaw

a &

K

.Iwat

s., n

om. i

lleg.

●C

yclo

soru

s het

eroc

arpu

s (B

lum

e)

Chi

ng

Spha

eros

teph

anos

het

eroc

arpu

s (B

lum

e)

Hol

ttum

Thel

ypte

ris h

eter

ocar

pa (B

lum

e)

C.V

.Mor

ton

●C

yclo

soru

s im

mer

sus (

Blu

me)

S.L

inds

. Am

phin

euro

n im

mer

sum

(Blu

me)

Hol

ttum

Thel

ypte

ris i

mm

ersa

(Blu

me)

Chi

ng

●C

yclo

soru

s int

erru

ptus

(Will

d.) H

.Ito

Cyc

loso

rus i

nter

rupt

us (W

illd.

) H.It

oTh

elyp

teri

s int

erru

pta

(Will

d.) K

.Iwat

s.

●C

yclo

soru

s lak

him

pure

nse

(ros

enst

.) B

.K.N

ayar

& K

aur

Pron

ephr

ium

lakh

impu

rens

e (R

osen

st.)

Hol

ttum

Thel

ypte

ris l

akhi

mpu

rens

is (R

osen

st.)

K.Iw

ats.

●C

yclo

soru

s lar

uten

sis (

Bed

d.) C

hing

Sp

haer

oste

phan

os la

rute

nsis

(Bed

d.) C

.Chr

.Th

elyp

teri

s lar

uten

sis (

Bed

d.) T

agaw

a &

K

.Iwat

s.

●C

yclo

soru

s leb

eufii

(Bak

er) W

.M.C

hu

Chr

iste

lla le

beufi

i (B

aker

) Hol

ttum

Thel

ypte

ris l

ebeu

fii (B

aker

) Pan

igra

hi

●C

yclo

soru

s meg

aphy

llus (

Met

t.) C

hing

Spha

eros

teph

anos

pen

nige

r Hol

ttum

, no

m. n

ud.

Thel

ypte

ris m

egap

hylla

(Met

t.) K

.Iwat

s.

●C

yclo

soru

s men

isci

icar

pus (

Blu

me)

H

olttu

m

Pron

ephr

ium

men

isci

icar

pon

(Blu

me)

H

olttu

mTh

elyp

teri

s men

isci

icar

pa (B

lum

e)

K.Iw

ats.

Page 37: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

100 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●C

yclo

soru

s mol

liusc

ulus

(Kuh

n) C

hing

C

hris

tella

app

endi

cula

ta (C

.Pre

sl) H

olttu

mTh

elyp

teri

s mol

liusc

ula

(Kuh

n) K

.Iwat

s.

●C

yclo

soru

s nud

atus

(rox

b.) B

.K.N

ayar

& K

aur

Pron

ephr

ium

nud

atum

(Rox

b.) H

olttu

mTh

elyp

teri

s nud

ata

(rox

b.) C

.V.M

orto

n

●C

yclo

soru

s opu

lent

us (K

aulf.

) Nak

aike

Am

phin

euro

n op

ulen

tum

(Kau

lf.) H

olttu

mTh

elyp

teri

s opu

lent

a (K

aulf.

) Fos

berg

●C

yclo

soru

s pap

ilio

(C.H

ope)

Chi

ng

Chr

iste

lla p

apili

o (C

.Hop

e) H

olttu

mTh

elyp

teri

s pap

ilio

(C.H

ope)

K.Iw

ats.

●C

yclo

soru

s par

asiti

cus (

L.) F

arw

. C

hris

tella

par

asiti

ca (L

.) H

.Lév

.Th

elyp

teri

s par

asiti

ca (L

.) Ta

rdie

u

●C

yclo

soru

s pen

angi

anus

(Hoo

k.) C

opel

. Pr

onep

hriu

m p

enan

gian

um (H

ook.

) Hol

ttum

Thel

ypte

ris p

enan

gian

a (H

ook.

) C.F

.Ree

d

●C

yclo

soru

s pol

ycar

pus (

Blu

me)

Hol

ttum

Sp

haer

oste

phan

os p

olyc

arpa

(Blu

me)

Cop

el.

Thel

ypte

ris p

olyc

arpa

(Blu

me)

K.Iw

ats.

●C

yclo

soru

s pro

lifer

(Ret

z.) T

ardi

eu e

x Ta

rdie

u &

C.C

hr.

Men

isci

um p

rolif

erum

(Ret

z.) S

w.

Men

isci

um p

rolif

erum

(Ret

z.) S

w.

●C

yclo

soru

s rep

andu

s (Fé

e) B

.K.N

ayar

& K

aur

Pron

ephr

ium

repa

ndum

(Fée

) Hol

ttum

Thel

ypte

ris r

epan

da (F

ée) C

.V.M

orto

n

○C

yclo

soru

s rub

icun

dus (

Ald

erw

.) S.

Lind

s.4Pr

onep

hriu

m ru

bicu

ndum

(Ald

erw.

) Hol

ttum

Abs

ent

○C

yclo

soru

s sal

icifo

lius (

Wal

l. ex

Hoo

k.) C

opel

.4Pr

onep

hriu

m sa

licifo

lium

(Wal

l. ex

Hoo

k.) H

olttu

mA

bsen

t

●C

yclo

soru

s sia

men

sis (

Taga

wa

& K

.Iwat

s.) P

anig

rahi

C

hris

tella

siam

ensi

s (Ta

gaw

a &

K.Iw

ats.)

Hol

ttum

Thel

ypte

ris s

iam

ensi

s Tag

awa

& K

.Iwat

s.

●C

yclo

soru

s sub

elat

us (B

aker

) Chi

ng

Chr

iste

lla su

bela

ta (B

aker

) Hol

ttum

Thel

ypte

ris s

ubel

ata

(Bak

er) K

.Iwat

s.

●C

yclo

soru

s sub

pube

scen

s (B

lum

e) C

hing

12C

hris

tella

subp

ubes

cens

(Blu

me)

Hol

ttum

+

Chr

iste

lla la

tipin

na (B

enth

.) H

.Lév

.Th

elyp

teri

s sub

pube

scen

s (B

lum

e) K

.Iwat

s. +

Thel

ypte

ris l

atip

inna

(Hoo

k.) K

.Iwat

s. +

Thel

ypte

ris s

umat

rana

(Ald

erw.

) Tag

awa

&

K.Iw

ats.

●C

yclo

soru

s ter

min

ans (

J.Sm

. ex

Hoo

k.)

Pani

grah

i Am

phin

euro

n te

rmin

ans (

J.Sm

. ex

Hoo

k.)

Hol

ttum

Th

elyp

teri

s ter

min

ans (

J.Sm

. ex

Hoo

k.)

Taga

wa

& K

.Iwat

s.

○C

yclo

soru

s tha

iland

icus

S.L

inds

.24A

bsen

tA

bsen

t

●C

yclo

soru

s tri

phyl

lus (

Sw.)

Tard

ieu

ex T

ardi

eu &

C.C

hr.

var.

trip

hyllu

sPr

onep

hriu

m tr

iphy

llum

(Sw.

) Hol

ttum

va

r. tr

iphy

llum

Thel

ypte

ris t

riph

ylla

(Sw.

) K.Iw

ats.

var.

trip

hylla

●C

yclo

soru

s tri

phyl

lus (

Sw.)

Tard

ieu

ex T

ardi

eu &

C.C

hr.

var.

pari

shii

(Bed

d.) S

.Lin

ds.

Pron

ephr

ium

trip

hyllu

m (S

w.) H

olttu

m

var.

pari

shii

(Bed

d.) N

akai

keTh

elyp

teri

s tri

phyl

la (S

w.) K

.Iwat

s. va

r. pa

rish

ii (B

edd.

) K.Iw

ats.

●C

yclo

soru

s tru

ncat

us (P

oir.)

Far

w.

Pneu

mat

opte

ris t

runc

ata

(Poi

r.) H

olttu

mTh

elyp

teri

s tru

ncat

a (P

oir.)

K.Iw

ats.

●C

yclo

soru

s tub

ercu

lifer

(C.C

hr.)

Pani

grah

i Th

elyp

teri

s tub

ercu

lifer

a (C

.Chr

.) C

hing

Thel

ypte

ris t

uber

culif

era

(C.C

hr.)

Chi

ng

Page 38: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

101TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●C

yclo

soru

s tyl

odes

(Kun

ze) P

anig

rahi

14Th

elyp

teri

s xyl

odes

(Kun

ze) C

hing

Thel

ypte

ris x

ylod

es (K

unze

) Chi

ng

●C

yclo

soru

s uni

tus (

L.) C

hing

Sp

haer

oste

phan

os u

nitu

s (L.

) Hol

ttum

Thel

ypte

ris u

nita

(L.)

C.V

.Mor

ton

●C

yclo

soru

s val

idus

(Chr

ist)

Chi

ng

Cyc

loso

rus h

irtis

orus

(C.C

hr.)

Chi

ngTh

elyp

teri

s hir

tisor

a (C

.Chr

.) K

.Iwat

s. +

Thel

ypte

ris v

alid

a (C

hris

t) Ta

gaw

a &

K

.Iwat

s.

●M

acro

thel

ypte

ris o

rnat

a (W

all.

ex B

edd.

) Chi

ng

Mac

roth

elyp

teri

s orn

ata

(Wal

l. ex

Bed

d.) C

hing

Thel

ypte

ris o

rnat

a (W

all.

ex B

edd.

) Chi

ng

●M

acro

thel

ypte

ris t

orre

sian

a (G

audi

ch.)

Chi

ng

Mac

roth

elyp

teri

s tor

resi

ana

(Gau

dich

.) C

hing

Thel

ypte

ris t

orre

sian

a (G

audi

ch.)

Als

ton

●Ps

eudo

pheg

opte

ris s

umat

rana

Hol

ttum

Ps

eudo

pheg

opte

ris s

umat

rana

Hol

ttum

Abs

ent

Thel

ypte

ris c

onflu

ens (

Thun

b.) C

.V.M

orto

n Th

elyp

teri

s con

fluen

s (Th

unb.

) C.V

.Mor

ton

Thel

ypte

ris c

onflu

ens (

Thun

b.) C

.V.M

orto

n

Thel

ypte

ris fl

acci

da (B

lum

e) C

hing

M

etat

hely

pter

is fl

acci

da (B

lum

e) C

hing

Thel

ypte

ris fl

acci

da (B

lum

e) C

hing

●Th

elyp

teri

s hir

sutip

es (C

lark

e) C

hing

C

oryp

hopt

eris

hir

sutip

es (C

.B.C

lark

e) H

olttu

mTh

elyp

teri

s hir

sutip

es (C

lark

e) C

hing

○Th

elyp

teri

s lax

a (F

ranc

h. &

Sav

.) C

hing

26

Abs

ent

Abs

ent

●Th

elyp

teri

s sin

gala

nens

is (B

aker

) Chi

ng

Met

athe

lypt

eris

sing

alan

ensi

s (B

aker

) Chi

ngTh

elyp

teri

s sin

gala

nens

is (B

aker

) Chi

ng

Thel

ypte

ris v

isco

sa (B

aker

) Chi

ng

Cor

ypho

pter

is v

isco

sa (B

aker

) Hol

ttum

Thel

ypte

ris v

isco

sa (B

aker

) Chi

ng

WO

OD

SIA

CE

AE

Athy

rium

ani

sopt

erum

Chr

ist

Athy

rium

ani

sopt

erum

Chr

ist

Athy

rium

ani

sopt

erum

Chr

ist

●At

hyri

um c

umin

gian

um (C

.Pre

sl) M

ilde

Anis

ocam

pium

cum

ingi

anum

C.P

resl

Anis

ocam

pium

cum

ingi

anum

C.P

resl

●At

hyri

um c

uspi

datu

m (B

edd.

) M.K

ato

Kun

iwat

suki

a cu

spid

ata

(Bed

d.) P

ichi

-Ser

m.

Kun

iwat

suki

a cu

spid

ata

(Bed

d.) P

ic.S

erm

.

Athy

rium

dis

sitif

oliu

m (B

aker

) C.C

hr.

Athy

rium

dis

sitif

oliu

m (B

aker

) C.C

hr.

Athy

rium

dis

sitif

oliu

m (B

aker

) C.C

hr.

Athy

rium

mac

kinn

onii

(C.H

ope)

C.C

hr.

Athy

rium

mac

kinn

onii

(C.H

ope)

C.C

hr.

Athy

rium

mac

kinn

onii

(C.H

ope)

C.C

hr.

Athy

rium

setif

erum

C.C

hr.

Athy

rium

setif

erum

C.C

hr.

Athy

rium

setif

erum

C.C

hr.

Cor

nopt

eris

opa

ca (D

.Don

) Tag

awa

Cor

nopt

eris

opa

ca (D

.Don

) Tag

awa

Cor

nopt

eris

opa

ca (D

.Don

) Tag

awa

●D

epar

ia p

eter

seni

i (K

unze

) M.K

ato

Athy

rium

japo

nicu

m a

uct.

non

(Thu

nb.)

Cop

el.

Dep

aria

japo

nica

auc

t. no

n (T

hunb

.) M

.Kat

o / D

epar

ia p

eter

seni

i (K

unze

) M.K

ato

Page 39: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

102 THaI FOreST BuLLeTIN (BOTaNY) 37

CU

RR

EN

T N

AM

E

NA

ME

IN P

TE

RID

OPH

YT

ES

IN T

HA

ILA

ND

(B

oonk

erd

& P

olla

wat

n, 2

000)

NA

ME

IN F

LO

RA

OF

TH

AIL

AN

D

(Tag

awa

& Iw

atsu

ki, 1

979,

198

5, 1

988,

198

9)

●D

epar

ia su

bfluv

ialis

(Hay

ata)

M.K

ato

Athy

rium

bor

yanu

m a

uct.

non

(Will

d.) T

agaw

aD

epar

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Page 40: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

103TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

CU

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104 THaI FOreST BuLLeTIN (BOTaNY) 37

REFERENCES

Boonkerd, T. (2006). A new species of Microsorum (Polypodiaceae) from Thailand. Blumea 51: 143–145.

________. Asplenium inaequilaterale Willd. (Aspleniaceae) a new record for Thailand. ScienceAsia, in press.

Boonkerd, T. & Nooteboom, H.P. (2001). a new species of Microsorum (Polypodiaceae) from Thailand. Blumea 46: 581–583.

Boonkerd, T. & Pollawatn, r. (2000). Pteridophytes in Thailand. Office of environmental Policy and Planning, Bangkok.

________. (2002a). Leptochilus minor Fée (Polypodiaceae), a new record for Thailand. The Natural History Journal of Chulalongkorn university 2: 1–3.

________. (2002b). The first record of Pteridium aquilinum (L.) Kuhn var. latiusculum (Desv.) Underw. ex A.Heller (Dennstaedtiaceae) in Thailand. Thai Forest Bulletin (Botany) 30: 72–74.

________. (2004). A revised taxonomic account of the fern genus Woodwardia (Blechnaceae) in Thailand. Thai Forest Bulletin (Botany) 32: 1–5.

________. (2006). Pteridophyte flora of Thong Pha Phum National Park, Kanchanaburi Province, Thailand. The Natural History Journal of Chulalongkorn university 6: 17–30.

Boonkerd, T. & Suksathan, P. (2009). Asplenium gueinzianum Mett. ex Kuhn (aspleniaceae), newly discovered in Thailand. The Natural History Journal of Chulalongkorn University 9(1): 95–98.

Boonkerd, T., Lindsay, S., Middleton, D.J. & Suddee, S. (2004). additions to the Pteridophyte flora of Thailand. Thai Forest Bulletin (Botany) 32: 6–11.

ebihara, a., Dubuisson, J.-Y., Iwatsuki, K., Hennequin, S. & Ito, M. (2006). a taxonomic revision of Hymenophyllaceae. Blumea 51: 221–280.

Fraser-Jenkins, C.r. (1997). New Species Syndrome in Indian Pteridology and the Ferns of Nepal. Dehra Dun, India.

Holttum, R.E. (1978). Lomariopsis Group. Flora Malesiana, ser. II, Ferns and Fern allies, 2: 255–330. Rijksherbarium/Hortus Botanicus, Leiden.

________. (1982 [‘1981’]). Thelypteridaceae. Flora Malesiana ser. II, Ferns and Fern Allies, 1: 331–560. Rijksherbarium/Hortus Botanicus, Leiden.

________. (1991). Tectaria Group. Flora Malesiana, ser. II, Ferns and Fern allies, 2: 1–132. Rijksherbarium/Hortus Botanicus, Leiden.

Holttum, r.e. & Grimes, J.W. (1980). The genus Pseudocyclosorus Ching (Thelypteridaceae). Kew Bulletin 34: 499–516.

Hovenkamp, P.H. (1986). a Monograph of the Fern Genus Pyrrosia. Leiden University Press, Leiden., 280 pages.

Page 42: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

105TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)

Hovenkamp, P.H., Bosman, M.T.M., Hennipman, e., Nooteboom, H.P., rödl-Linder, G. & roos, M.C. (1998). Polypodiaceae. In: Kalkman, C. et al. (eds), Flora Malesiana, ser. II, Ferns and Fern allies, 3: 1–234. rijksherbarium/Hortus Botanicus, Leiden.

Hovenkamp, P.H. & Miyamoto, F. (2005). a conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279–322.

Iwatsuki, K., Yokoyama, J. & Murakami, N. (1998). Hymenasplenium inthanonense (Aspleniaceae), a new fern species from Doi Inthanon, and its phylogenetic status. Thai Forest Bulletin (Botany) 26: 4–52.

Khullar, S.P. 1994. An Illustrated Fern Flora of the West Himalaya. International Book Distributors.

Kato, M. & Tsutsumi, C. (2008). Generic classification of Davalliaceae. acta Phytotaxonomica et Geobotanica 59: 1–14.

Kramer, K.U. (1971). Lindsaea-Group. Flora Malesiana, ser. II, Ferns and Fern allies, 1: 177–254. Rijksherbarium/Hortus Botanicus, Leiden.

Lindsay, S. & Middleton, D.J. (2004). Adiantum phanomensis (Adiantaceae), a new fern species from Peninsular Thailand. Harvard Papers in Botany 8: 137–140.

________. (2009a). New combinations in the ferns of Thailand. edinburgh Journal of Botany 66: 355–361.

________. (2009b). Development of a multi-access key to the ferns of Thailand. Thai Forest Bulletin (Botany), Special Issue: 134–137.

________. (2009c). Lecanopteris pumila Blume (Polypodiaceae), a new record for Thailand. Thai Forest Bulletin (Botany) 37: 59–63.

Lindsay, S., Suddee, S., Middleton, D.J. & Pooma, r. (2004 [‘2003’]). Matoniaceae (Pteridophyta) – a new family record for Thailand. Thai Forest Bulletin (Botany) 31: 47–52.

Lindsay, S., Middleton, D.J. & Suddee, S. (2008). Two new species of ferns from Thailand. Thai Forest Bulletin (Botany) 36: 46–51.

Lu, S.G. (1999). Three new names for Chinese ferns. Guihaia 19: 27–28.McNeill, J., Barrie, F.r., Burdet, H.M., Demoulin, V., Hawksworth, D.L., Marhold,

K., Nicolson, D.H., Prado, J., Silva, P.C., Skog, J.e., Wiersema, J.H. & Turland, N.J. (eds). (2006). International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005. regnum Vegetabile 146. ruggell: a.r.G. Gantner Verlag KG.

Middleton, D.J. (2003). Progress on the Flora of Thailand. Telopea 10: 33–42. Mitsuta, S. (1985). Nine new records of ferns to Thailand flora. acta Phytotaxonomica et

Geobotanica 36: 148. Murdock, a.G. (2008). a taxonomic revision of the eusporangiate fern family Marattiaceae,

with description of a new genus Ptisana. Taxon 57(3): 737–755.

Page 43: rbg-web2.rbge.org.uk · THAI FOR. BULL. (BOT.) 37: 64–106. 2009. Towards a stable nomenclature for Thai ferns STUART LINDSAY 1, DAVID J. MIDDLETON , THAWEESAKDI BOONKERD2 & SOMRAN

106 THaI FOreST BuLLeTIN (BOTaNY) 37

Nakaike, T. (1992). New Flora of Japan. Pteridophytes. Shibundo Co., Tokyo.Nooteboom, H. (1997). The microsoroid ferns (Polypodiaceae). Blumea 42: 261–395. ________. (1998). Davalliaceae. In: Kalkman, C. et al. (eds), Flora Malesiana, ser. II,

Ferns and Fern Allies, 3: 235–276. Rijksherbarium/Hortus Botanicus, Leiden.Parris, B.S. (1998a). The addition of Acrosorus streptophyllus (Baker) Copel.

(Grammitidaceae: Pteridophyta) to the flora of Thailand. Fern Gazette 15: 215–216.________. (1998b). Chrysogrammitis, a new genus of Grammitidaceae (Filicales). Kew

Bulletin 53: 909–918.________. (2007). Five new genera and three new species of Grammitidaceae (Filicales)

and the re-establishment of Oreogrammitis. Gardens’ Bulletin Singapore 58: 233–273.

Parris, B.S. & Latiff, a. (1997). Towards a pteridophyte flora of Malaysia: a provisional checklist of taxa. Malayan Nature Journal 50: 235–280.

Petchsri, S., Boonkerd, T. & Baum, B.r. (2009). a first record of Microsorum musifolium Copel. (Polypodiaceae) from Thailand. The Natural History Journal of Chulalongkorn University 9(1): 99–104.

rödl-Linder, G. (1990). Monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277–423.

Shieh, W.-C., Devol, C.e. & Kuo, C.-M. (1994). athyriaceae. In: Huang, T.-C. (ed.), Flora of Taiwan, 2nd ed., vol. 1: 414–448.

Smith, a.r., Pryer, K.M., Schuettpelz, e., Korall, P., Schneider, H. & Wolf, P.G. (2006). a classification of extant ferns. Taxon 55: 705–731.

________. (2008). Fern Classification. Pp. 417–467 In: ranker, T.a. & Haufler, C.H. (eds), Biology and Evolution of Ferns and Lycophytes, CUP, Cambridge.

Suksathan, P. (2004). A new species of Adiantum (Pteridaceae) from Thailand. American Fern Journal 94: 77–80.

Tagawa, M. & Iwatsuki, K. (1979). In: Smitinand, T. & Larsen, K. (eds), Flora of Thailand, Vol. 3, part 1. Royal Forest Department, Bangkok.

________. (1985). In: Smitinand, T. & Larsen, K. (eds), Flora of Thailand, Vol. 3, part 2. Royal Forest Department, Bangkok.

________. (1988). In: Smitinand, T. & Larsen, K. (eds), Flora of Thailand, Vol. 3, part 3. Royal Forest Department, Bangkok.

________. (1989). In: Smitinand, T. & Larsen, K. (eds), Flora of Thailand, Vol. 3, part 4. Royal Forest Department, Bangkok.

Thompson, J.a. (2008). Morphotype and conflicting taxonomies in Pteridium (Dennstaedtiaceae; Pteridophyta). Fern Gazette 18: 101–109.

Zhang, X.C. (1999a). Antrophyaceae. Flora Reipublicae Popularis Sinicae 3(2): 1–12 (in Chinese).

________. (1999b). Vittariaceae. Flora Reipublicae Popularis Sinicae 3(2): 12–31 (in Chinese).