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THAI FOR. BULL. (BOT.) 37: 64–106. 2009.
Towards a stable nomenclature for Thai ferns
STUART LINDSAY1, DAVID J. MIDDLETON1, THAWEESAKDI BOONKERD2 & SOMRAN SUDDEE3
ABSTRACT. There have been many changes to family and genus delimitation in Thai ferns since the publication of volume 3 (Pteridophytes) of the Flora of Thailand. These changes are discussed and current names for all taxa, including those recently described or recorded for Thailand, are presented. A new combination is made in Hymenasplenium.
KEY WORDS: Ferns, Flora of Thailand, nomenclature, Thailand.
INTRODUCTION
There are approximately 670 taxa of ferns in Thailand comprising about 5–7% of the total vascular flora (Middleton, 2003). The ferns were published in volume 3 of the Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989) and 596 taxa were recognised as occurring in Thailand (excluding the lycophytes: Lycopodiaceae, Selaginellaceae and Isoetaceae). A number of papers have since been published in which new taxa have been described or new records have been added to the pteridophyte diversity of Thailand (Mitsuta, 1985; Iwatsuki et al., 1998; Parris, 1998a; Hovenkamp et al., 1998; Nooteboom, 1998; Boonkerd & Nooteboom, 2001; Boonkerd & Pollawatn 2000, 2002a, 2002b, 2006; Lindsay & Middleton, 2004, 2009c; Lindsay et al., 2004, 2008; Suksathan, 2004; Boonkerd et al., 2004; Boonkerd, 2006). Of particular significance amongst these are Boonkerd & Pollawatn (2000), which is a list of all the pteridophytes in Thailand (including an additional 27 taxa) with distribution maps and many photographs, and Boonkerd et al. (2004), in which many more species were added. A total of 71 fern taxa have been newly recorded or newly described for Thailand since the publication of the Flora of Thailand accounts. This rate of addition of new taxa and new records to a recently completed Flora account reflects the fact that pteridophytes had previously often been neglected on traditional collecting expeditions (and were therefore not available to Tagawa & Iwatsuki) and is testament to the relatively poor state of our taxonomic knowledge of them compared to many angiosperm plant groups.
The taxonomy of ferns in Thailand has also been subject to considerable flux due to differences of opinion over the delimitation of families, genera and species. One need only glance at the lists of synonymy to see that very many species have had homotypic combinations made in many different genera (e.g. several species of Selliguea also have combinations in Polypodium, Phymatopsis, Pleopeltis, Phymatodes, Crypsinus
________________________________________________________________________________________________________________________________________________________
1 Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, UK.2 Department of Botany, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand.3 The Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, Chatuchak, Bangkok 10900, Thailand.
65TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
and Phymatopteris - see Lindsay & Middleton, 2009a). This indecision about the best placement of species in genera is also manifest at the family level where the limits of several families have changed considerably over time and between contemporary Flora projects, leading, in some cases, to the abandonment of families altogether in some parts of Flora Malesiana (e.g. Kramer, 1971; Holttum, 1978, 1991) in favour of “groups” of genera.
In recent years phylogenetic techniques, particularly using molecular data, have been used to attempt to solve the many problems in fern familial and generic delimitation (see Smith et al., 2006, 2008 and references cited therein). This has led to a greater consensus on the delimitation of families and genera. We are eventually beginning to see the possibility of stability in the names applied to the ferns of Thailand.
THE TABLES
In this paper we apply the family and genus concepts summarised by Smith et al. (2006, 2008) to the ferns of Thailand and present the data in two tables. For Davalliaceae we follow the more recent generic concepts of Kato & Tsutsumi (2008). In Table 1 the families and genera recognised by Smith et al. (2006, 2008) and Kato & Tsutsumi (2008) are compared to the families and genera recognised in the Flora of Thailand accounts. In Table 2 all fern taxa now known to occur in Thailand are listed using the up-to-date family and genus concepts from Smith et al. (2006, 2008) and Kato & Tsutsumi (2008), with occasional modifications from other recent works (e.g. Parris, 2007). These are compared to the names used for the same taxon from the Flora of Thailand and from Boonkerd & Pollawatn (2000). For species delimitation we follow Tagawa & Iwatsuki (1979, 1985, 1988, 1989), unless other pieces of taxonomic work have superseded these publications (e.g. Hovenkamp et al., 1998), or we have come to our own differing opinion in specific groups. In some cases the authorship of names is not the same as has been previously published. We have checked the problematic ones and only the correct authorships are used without highlighting the discrepancies. Likewise the spelling of specific epithets not in conformity with the International Code of Botanical Nomenclature (McNeill et al., 2006) has been corrected in all columns without comment. We include taxa which have been published as occurring in Thailand. We also include a small number of taxa which we provisionally record for Thailand from our own observations but which still await formal publication. We acknowledge that this list might be incomplete judging by the rate at which taxa have been newly recorded from Thailand in the last 20 years. However, this list has been compiled to facilitate the development of an on-line multi-access key to the ferns of Thailand (Lindsay & Middleton, 2009b) and new taxa can easily be added to this.
The “nine new records” in Mitsuta (1985) were not included in the Flora of Thailand. Some of them are, however, attributable to species already known from Thailand. We have included a genuine additional record (Monachosorum henryi Christ) and as yet unconfirmed new records (Thelypteris laxa (Franch. & Sav.) Ching and Diplazium doerderleinii (Luerss.) Makino) in Table 2. The new record Diplazium okinawaense Tagawa has been synonymised to Diplazium virescens Kunze following Shieh et al. (1994). We have redetermined the other material cited in Mitsuta (1985) to taxa already reported in the Flora of Thailand, thus: Colysis flexiloba (Christ) Ching var. undulato-repanda (C.Chr.) Ching to Leptochilus ellipticus (Thunb.) Noot.; Crypsinus trisectus
66 THaI FOreST BuLLeTIN (BOTaNY) 37
(Baker) Tagawa to Selliguea hirsuta (Baker) S.Linds.; Microlepia trichoclada Ching to Microlepia herbacea Ching & C.Chr. ex Tardieu & C.Chr.; Thelypteris angulariloba Ching to Thelypteris hirsutipes (C.B.Clarke) Ching; and Thelypteris esquirolii (Christ) Ching to Cyclosorus falcilobus (Hook.) Panigrahi.
Major changes in family delimitations
Table 1 summarises the changes in family delimitation between the Flora of Thailand accounts and those currently recognised and lists the genera in each of the families. These changes include losing some familiar families altogether, such as Parkeriaceae (now in Pteridaceae) and Athyriaceae (now mostly in Woodsiaceae), and changing the delimitation of others quite radically, such as Polypodiaceae (which now includes Grammitidaceae) and Dryopteridaceae (which has lost Tectaria and its relatives and gained genera from Lomariopsidaceae, Davalliaceae and Athyriaceae). Pteridaceae has also undergone major changes and is now much larger with five subfamilies which do not correspond well to former families; e.g. Pteridaceae subfamily Pteridoideae includes only Pteris from the previously delimited Pteridaceae, with all of the other genera previously having been placed in Parkeriaceae.
Major changes in generic delimitation
Table 2 lists all the currently used names for Thai ferns and compares them to Pteridophytes in Thailand (Boonkerd & Pollawatn, 2000) and Flora of Thailand (Tagawa & Iwatsuki, 1979, 1985, 1988, 1989). A total of 191 taxa have a name which is now different to the name used in the Flora of Thailand accounts, most of these as a result of changes in generic delimitation. Changes in generic delimitation are particularly extensive in Davalliaceae, Hymenophyllaceae, Polypodiaceae and Thelypteridaceae. Well known genera such as Grammitis, Polypodium, Trichomanes and Vittaria are no longer recognised in Thailand. Table 2 also provides references to the literature on new taxa, new records and in cases where the taxon epithet is different to that used in the Flora of Thailand account (usually due to synonymy). If there is no reference given then the change is made here. Black dots are used to indicate a taxon which is under a different name to that used in the original Flora of Thailand account; circles are used to indicate a taxon which is new to Thailand, either as a newly described taxon or a new record.
ACKNOWLEDGEMENTS
We thank The Leverhulme Trust for financial support for this work; the Curators and Collection Managers of herbaria that have loaned material or welcomed us on visits (notably a, BK, BKF, BM, C, CMu, e, K, SING); the library staff of the royal Botanic Garden edinburgh for their help in locating or obtaining protologues and other references; and Christopher Fraser-Jenkins, John McNeill, Barbara Parris, alan Smith and John Thompson for advice and information.
67TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
Tabl
e 1.
Com
paris
on o
f fam
ily d
elim
itatio
n an
d co
mpo
sitio
n be
twee
n Sm
ith e
t al.
(200
6, 2
008)
and
Flo
ra o
f Tha
iland
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9). N
ote
that
with
in P
terid
acea
e the
subf
amily
nam
e Vitt
ario
idea
e has
prio
rity
over
Adi
anto
idea
e (cf
. Sm
ith et
al.,
2006
, 200
8) an
d th
e sub
fam
ilial
nam
e Cry
ptog
ram
moi
deae
,
w
hich
did
not
exi
st fo
r Sm
ith e
t al.
(200
6, 2
008)
, has
now
bee
n pu
blis
hed
(Lin
dsay
& M
iddl
eton
, 200
9a).
Smith
et a
l. (2
006,
200
8) fa
mily
Gen
era
curr
ently
rec
ogni
zed
in T
haila
nd.
Tho
se in
bo
ld (2
6) w
ere
not r
ecog
nize
d in
Flo
ra o
f Tha
iland
Cor
resp
ondi
ng fa
mily
(or
fam
ilies
) in
Flor
a of
Tha
iland
(T
agaw
a &
Iwat
suki
, 197
9, 1
985,
198
8, 1
989)
ASP
LeN
IaC
eae
Aspl
eniu
m, H
ymen
aspl
eniu
mA
SPLe
NIa
Cea
e
BLe
CH
Na
Cea
eBl
echn
um, B
rain
ea, S
teno
chla
ena,
Woo
dwar
dia
BLe
CH
Na
Cea
e (B
lech
num
, Bra
inea
, Woo
dwar
dia)
, PT
ERID
AC
EAE
(Ste
noch
laen
a)
CIB
OTI
AC
EAE
Cib
otiu
mD
ICK
SON
IaC
eae
CYA
THEA
CEA
EC
yath
eaC
YATH
EAC
EAE
DAV
ALL
IAC
EAE
Ara
iost
egie
lla, D
aval
lia, D
aval
lode
s, H
umat
a, W
ibel
iaD
AVA
LLIA
CEA
E
DeN
NST
aeD
TIa
Cea
eD
enns
taed
tia, H
istio
pter
is, H
ypol
epis
, Mic
role
pia,
Pt
erid
ium
DeN
NST
aeD
TIa
Cea
e
DIP
TER
IDA
CEA
EC
heiro
pleu
ria,
Dip
teri
sD
IPTE
RID
AC
EAE
(Dip
teri
s), C
HEI
RO
PLEU
RIA
CEA
E (C
heiro
pleu
ria)
DRY
OPT
ERID
AC
EAE
Acro
phor
us, A
rach
niod
es, B
olbi
tis, C
teni
tis, C
yrto
miu
m,
Pera
nem
a, D
idym
ochl
aena
, Dry
opte
ris,
Elap
hogl
ossu
m,
Hyp
odem
atiu
m, L
euco
steg
ia, L
omag
ram
ma,
Pol
ystic
hum
, Te
rato
phyl
lum
DRY
OPT
ERID
AC
EAE
(Acr
opho
rus,
Arac
hnio
des,
Cte
nitis
, C
yrto
miu
m,
Did
ymoc
hlae
na, D
ryop
teri
s, Po
lyst
ichu
m),
ATH
YR
IAC
EAE
(Hyp
odem
atiu
m),
DAV
ALL
IAC
EAE
(Leu
cost
egia
), LO
Ma
rIO
PSID
aC
eae
(Bol
bitis
, Ela
phog
loss
um, L
omag
ram
ma,
Te
rato
phyl
lum
)
EQU
ISET
AC
EAE
Equi
setu
mEQ
UIS
ETA
CEA
E
GLe
ICH
eNIa
Cea
eD
icra
nopt
eris
, Dip
lopt
eryg
ium
, Gle
iche
nia,
Stic
heru
sG
LeIC
HeN
IaC
eae
HY
MeN
OPH
YLL
aC
eae
Abro
dict
yum
, Cal
listo
pter
is, C
epha
lom
anes
, Cre
pido
man
es,
Did
ymog
loss
um, H
ymen
ophy
llum
, Van
denb
osch
iaH
YM
eNO
PHY
LLa
Cea
e
LIN
DSa
eaC
eae
Lind
saea
, Sph
enom
eris
, Tap
eini
dium
LIN
DSa
eaC
eae
LOM
ar
IOPS
IDa
Cea
eC
yclo
pelti
s, Lo
mar
iops
is, N
ephr
olep
isLO
Ma
rIO
PSID
aC
eae
(Lom
ario
psis
), D
RYO
PTER
IDA
CEA
E (C
yclo
pelti
s), O
Lea
ND
ra
Cea
e (N
ephr
olep
is)
LYG
OD
IAC
EAE
Lygo
dium
SCH
IZA
EAC
EAE
MA
RAT
TIA
CEA
EAn
giop
teri
s, C
hris
tens
enia
, Ptis
ana
MA
RAT
TIA
CEA
E
68 THaI FOreST BuLLeTIN (BOTaNY) 37
Smith
et a
l. (2
006,
200
8) fa
mily
Gen
era
curr
ently
rec
ogni
zed
in T
haila
nd.
Tho
se in
bol
d (2
6) w
ere
not r
ecog
nize
d in
Flo
ra o
f Tha
iland
Cor
resp
ondi
ng fa
mily
(or
fam
ilies
) in
Flor
a of
Tha
iland
(T
agaw
a &
Iwat
suki
, 197
9, 1
985,
198
8, 1
989)
MA
RSI
LEA
CEA
EM
arsi
lea
MA
RSI
LEA
CEA
E
MaT
ON
IaC
eae
Mat
onia
Abs
ent
OLe
aN
Dr
aC
eae
Ole
andr
aO
Lea
ND
ra
Cea
e
OPH
IOG
LOSS
AC
EAE
Botr
ychi
um, H
elm
inth
osta
chys
, Oph
iogl
ossu
mO
PHIO
GLO
SSA
CEA
E
OSM
uN
Da
Cea
eO
smun
daO
SMu
ND
aC
eae
PLA
GIO
GY
RIA
CEA
EPl
agio
gyri
aPL
AG
IOG
YR
IAC
EAE
POLY
POD
IAC
EAE
Acro
soru
s, Ag
laom
orph
a, A
rthr
omer
is, B
elvi
sia,
C
alym
mod
on, C
hris
tiopt
eris
, Chr
ysog
ram
miti
s, Ct
enop
tere
lla, D
asyg
ram
miti
s, D
ryna
ria, G
onio
phle
bium
, G
ymno
gram
miti
s, Le
cano
pter
is, L
emm
aphy
llum
, Le
piso
rus,
Lept
ochi
lus,
Loxo
gram
me,
Mic
roso
rum
, N
eoch
eiro
pter
is, O
reog
ram
miti
s, Pl
atyc
eriu
m,
Pros
aptia
, Pyr
rosi
a, R
adio
gram
miti
s, Sc
lero
glos
sum
, Se
lligu
ea, T
hem
eliu
m, T
omop
hyllu
m, X
ipho
pter
ella
POLY
POD
IAC
EAE
(Agl
aom
orph
a, A
rthr
omer
is,
Belv
isia
, Chr
istio
pter
is, C
olys
is, D
ryna
ria,
Lem
map
hyllu
m,
Lepi
soru
s, Le
ptoc
hilu
s, Lo
xogr
amm
e, M
icro
soru
m,
Neo
chei
ropt
eris
, Pla
tyce
rium
, Pyr
rosi
a, S
ellig
uea)
, D
AVA
LLIA
CEA
E (G
ymno
gram
miti
s), G
ra
MM
ITID
aC
eae
(Acr
osor
us, C
alym
mod
on, P
rosa
ptia
, Scl
erog
loss
um)
PSIL
OTA
CEA
EPs
ilotu
mPS
ILO
TAC
EAE
PTER
IDA
CEA
E su
bfam
. Che
ilant
hoid
eae
Che
ilant
hes,
Dor
yopt
eris
, Hem
ioni
tis, N
otho
laen
a,
Para
hem
ioni
tis, P
ella
eaPA
RK
ERIA
CEA
E
PTER
IDA
CEA
E su
bfam
. Cry
ptog
ram
moi
deae
Con
iogr
amm
ePA
RK
ERIA
CEA
E
PTER
IDA
CEA
E su
bfam
. Par
kerio
idea
eAc
rost
ichu
m, C
erat
opte
ris
PTER
IDA
CEA
E (A
cros
tichu
m),
PAR
KER
IAC
EAE
(Cer
atop
teri
s)
PTER
IDA
CEA
E su
bfam
. Pte
ridoi
deae
Ony
chiu
m, P
ityro
gram
ma,
Pte
ris,
Syng
ram
ma,
Tae
nitis
PAR
KER
IAC
EAE
(Ony
chiu
m, P
ityro
gram
ma,
Syn
gram
ma,
Ta
eniti
s), P
TER
IDA
CEA
E (P
teri
s)
PTER
IDA
CEA
E su
bfam
. Vitt
ario
idea
eAd
iant
um, A
ntro
phyu
m, H
aplo
pter
is, V
agin
ular
iaPA
RK
ERIA
CEA
E (A
dian
tum
), V
ITTA
RIA
CEA
E (A
ntro
phyu
m, V
agin
ular
ia, V
ittar
ia)
SaLV
INIa
Cea
eAz
olla
, Sal
vini
aSa
LVIN
IaC
eae
(Sal
vini
a), A
ZOLL
AC
EAE
(Azo
lla)
SCH
IZA
EAC
EAE
Schi
zaea
SCH
IZA
EAC
EAE
TEC
TAR
IAC
EAE
Arth
ropt
eris
, Het
erog
oniu
m, P
leoc
nem
ia, P
teri
drys
, Tec
tari
aD
RYO
PTER
IDA
CEA
E (H
eter
ogon
ium
, Ple
ocne
mia
, Pt
erid
rys,
Tect
aria
), O
Lea
ND
ra
Cea
e (A
rthr
opte
ris)
THEL
YPT
ERID
AC
EAE
Cycl
osor
us, M
acro
thel
ypte
ris, P
seud
ophe
gopt
eris,
The
lypt
eris
THEL
YPT
ERID
AC
EAE
WO
OD
SIA
CEA
EAt
hyri
um, C
orno
pter
is, D
epar
ia, D
ipla
zium
ATH
YR
IAC
EAE
69TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
Tabl
e 2.
Com
paris
on o
f the
cur
rent
nam
es fo
r Tha
i fer
n sp
ecie
s w
ith th
e na
mes
use
d fo
r the
sam
e ta
xa in
Pte
ridop
hyte
s in
Tha
iland
(Boo
nker
d &
Pol
law
atn,
200
0) a
nd
Fl
ora
of T
haila
nd (T
agaw
a &
Iwat
suki
, 197
9, 1
985,
198
8, 1
989)
. Tw
o na
mes
for a
sing
le e
ntry
in th
e th
ird c
olum
n lin
ked
by a
‘/’ i
ndic
ate
that
a c
hang
e w
as m
ade
to
the
nam
e in
the
“add
ition
s an
d C
orre
ctio
ns”
sect
ion
in p
art 4
of v
olum
e 3
of th
e Fl
ora
of T
haila
nd a
ccou
nt. T
wo
or m
ore
nam
es fo
r a s
ingl
e en
try li
nked
by
a
‘+’ i
ndic
ate
that
thes
e ta
xa a
re n
ow tr
eate
d as
syn
onym
s. B
lack
dot
s in
dica
te s
peci
es w
hose
nam
es h
ave
chan
ged
sinc
e th
e pu
blic
atio
n of
the
Flor
a of
Tha
iland
or w
ere
chan
ged
in th
e “a
dditi
ons
and
Cor
rect
ions
” se
ctio
n. C
ircle
s in
dica
te s
peci
es th
at h
ave
been
dis
cove
red
or d
escr
ibed
sin
ce th
e pu
blic
atio
n of
the
Flor
a of
Thai
land
. Th
e re
fere
nce
besi
de a
nam
e re
fers
to th
e pl
ace(
s) o
f pub
licat
ion
of th
e ne
w ta
xon
or re
cord
, or t
he so
urce
of i
nfor
mat
ion
abou
t the
cha
nge
of a
nam
e: 1
Boo
nker
d (2
006)
; 2 B
oonk
erd
(in p
ress
); 3
Boo
nker
d &
Noo
tebo
om (2
001)
; 4 B
oonk
erd
& P
olla
wat
n (2
000)
; 5 B
oonk
erd
& P
olla
wat
n (2
002a
); 6
Boo
nker
d &
Polla
wat
n (2
002b
); 7
Boo
nker
d &
Pol
law
atn
(200
4); 8
Boo
nker
d &
Pol
law
atn
(200
6); 9
Boo
nker
d &
Suk
sath
an (
2009
); 10
Boo
nker
d et
al.
(200
4); 1
1 Fr
aser
-
Jenk
ins
(199
7); 1
2 H
olttu
m (
1982
); 13
Hol
ttum
(19
91);
14 H
olttu
m &
Grim
es (
1980
); 15
Hov
enka
mp
(198
6); 1
6 H
oven
kam
p et
al.
(199
8); 1
7 H
oven
kam
p
& M
iyam
oto
(200
5); 1
8 Iw
atsu
ki e
t al.
(199
8); 1
9 K
ato
& T
suts
umi (
2008
); 20
Khu
llar (
1994
); 21
Lin
dsay
& M
iddl
eton
(200
4); 2
2 Li
ndsa
y &
Mid
dlet
on (2
009c
);
23 L
inds
ay e
t al.
(200
4); 2
4 Li
ndsa
y et
al.
(200
8); 2
5 Lu
(199
9); 2
6 M
itsut
a (1
985)
; 27
Mur
dock
(200
8); 2
8 N
akai
ke (1
992)
; 29
Noo
tebo
om (1
997)
; 30
Noo
tebo
om
(1
998)
; 31
Parr
is (
1998
a); 3
2 Pa
rris
(19
98b)
; 33
Parr
is (
2007
); 34
Par
ris &
Lat
iff (
1997
); 35
Pet
chsr
i, B
oonk
erd
& B
aum
(20
09);
36 r
ödl-L
inde
r (1
990)
; 37
Su
ksat
han
(200
4); 3
8 Th
omps
on (2
008)
; 39
Zhan
g (1
999a
); 40
Zha
ng (1
999b
); 41
B. P
arris
, per
s. co
mm
.; 42
J. T
hom
pson
, per
s. co
mm
.; 43
Adi
antu
m p
hilip
pens
e
var.
subj
unon
icum
is re
porte
d fo
r Tha
iland
by
Boo
nker
d &
Pol
law
atn
(200
6) b
ut th
e com
bina
tion
has n
ot b
een
valid
ly p
ublis
hed
and
the s
tatu
s of t
he v
arie
ties o
f thi
s
spec
ies
need
furth
er re
sear
ch; 4
4 Pr
ovis
iona
l new
reco
rds,
curr
ently
und
er in
vest
igat
ion;
45
Hym
enas
plen
ium
exc
isum
(C.P
resl
) S.L
inds
., co
mb.
nov
. Bas
iony
m:
As
plen
ium
exc
isum
C.P
resl
, Epi
mel
. Bot
. 74
(185
1 [‘
1849
’]).
Alth
ough
the
nam
e H
ymen
aspl
eniu
m e
xcis
um h
as b
een
used
in r
ecen
t lite
ratu
re w
e ca
n fin
d no
evid
ence
that
the
com
bina
tion
has p
revi
ousl
y be
en v
alid
ly p
ublis
hed;
46
taxo
nom
ic c
hang
e by
cur
rent
aut
hors
; 47
Ebih
ara
et a
l. (2
006)
.
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
ASP
LE
NIA
CE
AE
Aspl
eniu
m a
ffine
Sw.
Aspl
eniu
m a
ffine
Sw.
Aspl
eniu
m a
ffine
Sw
.
Aspl
eniu
m a
ntro
phyo
ides
Chr
ist
Aspl
eniu
m a
ntro
phyo
ides
Chr
ist
Aspl
eniu
m a
ntro
phyo
ides
Chr
ist
Aspl
eniu
m b
atue
nse A
lder
w.As
plen
ium
bat
uens
e A
lder
w.As
plen
ium
bat
uens
e A
lder
w.
Aspl
eniu
m c
onfu
sum
Tar
dieu
& C
hing
Aspl
eniu
m c
onfu
sum
Tar
dieu
& C
hing
Aspl
eniu
m c
onfu
sum
Tar
dieu
& C
hing
Aspl
eniu
m c
rini
caul
e H
ance
Aspl
eniu
m c
rini
caul
e H
ance
Aspl
eniu
m c
rini
caul
e H
ance
○As
plen
ium
dec
resc
ens K
unze
44A
bsen
tA
bsen
t
Aspl
eniu
m d
elav
ayi (
Fran
ch.)
Cop
el.
Aspl
eniu
m d
elav
ayi (
Fran
ch.)
Cop
el.
Aspl
eniu
m d
elav
ayi (
Fran
ch.)
Cop
el.
Aspl
eniu
m e
nsifo
rme
Wal
l. ex
Hoo
k. &
Gre
v.As
plen
ium
ens
iform
e W
all.
ex H
ook.
& G
rev.
Aspl
eniu
m e
nsifo
rme
Wal
l. ex
Hoo
k. &
Gre
v.
Aspl
eniu
m e
xigu
um B
edd.
Aspl
eniu
m e
xigu
um B
edd.
Aspl
eniu
m e
xigu
um B
edd.
○As
plen
ium
finl
ayso
nian
um W
all.
ex H
ook.
10A
bsen
tA
bsen
t
Aspl
eniu
m g
revi
llei W
all.
ex H
ook.
& G
rev.
Aspl
eniu
m g
revi
llei W
all.
ex H
ook.
& G
rev.
Aspl
eniu
m g
revi
llei W
all.
ex H
ook.
& G
rev.
70 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
○As
plen
ium
gri
ffith
ianu
m H
ook.
44A
bsen
tA
bsen
t
○As
plen
ium
gue
inzi
anum
Met
t. ex
Kuh
n.9
Abs
ent
Abs
ent
Aspl
eniu
m h
umbe
rtii
Tard
ieu
Aspl
eniu
m h
umbe
rtii
Tard
ieu
Aspl
eniu
m h
umbe
rtii
Tard
ieu
○As
plen
ium
inae
quila
tera
le W
illd.
2A
bsen
tA
bsen
t
Aspl
eniu
m in
terj
ectu
m C
hris
tAs
plen
ium
inte
rjec
tum
Chr
ist
Aspl
eniu
m in
terj
ectu
m C
hris
t
○As
plen
ium
kha
sian
um S
ledg
e.44
Abs
ent
Abs
ent
●As
plen
ium
laci
niat
um D
.Don
.20As
plen
ium
var
ians
Wal
l. ex
Hoo
k. &
Gre
v.As
plen
ium
var
ians
Wal
l. ex
Hoo
k. &
Gre
v.
Aspl
eniu
m lo
ngis
sim
um B
lum
eAs
plen
ium
long
issi
mum
Blu
me
Aspl
eniu
m lo
ngis
sim
um B
lum
e
Aspl
eniu
m m
acro
phyl
lum
Sw
.As
plen
ium
mac
roph
yllu
m S
w.As
plen
ium
mac
roph
yllu
m S
w.
Aspl
eniu
m n
idus
L. v
ar. n
idus
As
plen
ium
nid
us L
. var
. nid
usAs
plen
ium
nid
us L
. var
. nid
us
Aspl
eniu
m n
idus
L. v
ar. m
usifo
lium
(Met
t.) C
.Chr
.As
plen
ium
nid
us L
. var
. mus
ifoliu
m (M
ett.)
C.C
hr.
Aspl
eniu
m n
idus
L. v
ar. m
usifo
lium
(Met
t.) C
.Chr
.
Aspl
eniu
m n
itidu
m S
w.
Aspl
eniu
m n
itidu
m S
w.As
plen
ium
niti
dum
Sw.
Aspl
eniu
m n
orm
ale
D.D
on
Aspl
eniu
m n
orm
ale
D.D
onAs
plen
ium
nor
mal
e D
.Don
Aspl
eniu
m p
arad
oxum
Blu
me
Aspl
eniu
m p
arad
oxum
Blu
me
Aspl
eniu
m p
arad
oxum
Blu
me
Aspl
eniu
m p
ellu
cidu
m L
am.
Aspl
eniu
m p
ellu
cidu
m L
am.
Aspl
eniu
m p
ellu
cidu
m L
am.
Aspl
eniu
m p
erak
ense
C.G
.Mat
thew
& C
hris
tAs
plen
ium
per
aken
se C
.G.M
atth
ew &
Chr
ist
Aspl
eniu
m p
erak
ense
C.G
.Mat
thew
& C
hris
t
Aspl
eniu
m p
hylli
tidis
D.D
on. s
ubsp
. phy
llitid
is
Aspl
eniu
m p
hylli
tidis
D.D
on. s
ubsp
. phy
llitid
isAs
plen
ium
phy
llitid
is D
.Don
. sub
sp. p
hylli
tidis
Aspl
eniu
m p
hylli
tidis
D.D
on. s
ubsp
. m
ales
icum
Hol
ttum
As
plen
ium
phy
llitid
is D
.Don
. sub
sp.
mal
esic
um H
olttu
mAs
plen
ium
phy
llitid
is D
.Don
. sub
sp.
mal
esic
um H
olttu
m
●As
plen
ium
pol
yodo
n G
.For
st.34
Aspl
eniu
m fa
lcat
um L
am.
Aspl
eniu
m fa
lcat
um L
am.
Aspl
eniu
m ro
ckii
C.C
hr.
Aspl
eniu
m ro
ckii
C.C
hr.
Aspl
eniu
m ro
ckii
C.C
hr.
Aspl
eniu
m sa
lignu
m B
lum
e As
plen
ium
salig
num
Blu
me
Aspl
eniu
m sa
lignu
m B
lum
e
Aspl
eniu
m sc
orte
chin
ii B
edd.
As
plen
ium
scor
tech
inii
Bed
d.As
plen
ium
scor
tech
inii
Bed
d.
Aspl
eniu
m si
amen
se T
agaw
a &
K.Iw
ats.
Aspl
eniu
m si
amen
se T
agaw
a &
K.Iw
ats.
Aspl
eniu
m si
amen
se T
agaw
a &
K.Iw
ats.
Aspl
eniu
m si
mon
sian
um H
ook.
As
plen
ium
sim
onsi
anum
Hoo
k.As
plen
ium
sim
onsi
anum
Hoo
k.
Aspl
eniu
m te
neru
m G
.For
st.
Aspl
eniu
m te
neru
m G
.For
st.
Aspl
eniu
m te
neru
m G
.For
st.
71TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Aspl
eniu
m te
nuifo
lium
D.D
on
Aspl
eniu
m te
nuifo
lium
D.D
onAs
plen
ium
tenu
ifoliu
m D
.Don
Aspl
eniu
m th
unbe
rgii
Kun
ze
Aspl
eniu
m th
unbe
rgii
Kun
zeAs
plen
ium
thun
berg
ii K
unze
●As
plen
ium
trun
catu
m B
lum
e.46
Aspl
eniu
m c
auda
tum
auc
t. no
n G
.For
st.
Aspl
eniu
m c
auda
tum
auc
t. no
n G
.For
st.
●As
plen
ium
vitt
aefo
rme
Cav
.34As
plen
ium
squa
mul
atum
Blu
me
Aspl
eniu
m sq
uam
ulat
um B
lum
e
Aspl
eniu
m y
oshi
naga
e M
akin
o As
plen
ium
yos
hina
gae
Mak
ino
Aspl
eniu
m y
oshi
naga
e M
akin
o
●H
ymen
aspl
eniu
m a
poga
mum
(N.M
urak
. & H
atan
.) N
akai
keAs
plen
ium
apo
gam
um N
.Mur
ak. &
Hat
an.
Aspl
eniu
m u
nila
tera
le a
uct.
non
Lam
. /
Aspl
eniu
m a
poga
mum
N.M
urak
. & H
atan
.
●H
ymen
aspl
eniu
m c
heilo
soru
m (K
unze
ex
Met
t.) T
agaw
aAs
plen
ium
che
iloso
rum
Kun
ze e
x M
ett.
Aspl
eniu
m c
heilo
soru
m K
unze
ex
Met
t.
Hym
enas
plen
ium
exc
isum
(C.P
resl
) S.L
inds
.45As
plen
ium
exc
isum
C.P
resl
Aspl
eniu
m e
xcis
um C
.Pre
sl
○H
ymen
aspl
eniu
m in
than
onen
se N
.Mur
ak. &
J.Yo
koy.
18H
ymen
aspl
eniu
m in
than
onen
se N
.Mur
ak. &
J.Yo
koy.
Abs
ent
●H
ymen
aspl
eniu
m o
bscu
rum
(Blu
me)
Tag
awa
Aspl
eniu
m o
bscu
rum
Blu
me
Aspl
eniu
m o
bscu
rum
Blu
me
BL
EC
HN
AC
EA
E
Blec
hnum
finl
ayso
nian
um W
all.
ex H
ook.
& G
rev.
Bl
echn
um fi
nlay
soni
anum
Wal
l. ex
Hoo
k. &
Gre
v.Bl
echn
um fi
nlay
soni
anum
Wal
l. ex
Hoo
k. &
Gre
v.
Blec
hnum
indi
cum
Bur
m.f.
Bl
echn
um in
dicu
m B
urm
.f.Bl
echn
um in
dicu
m B
urm
.f.
Blec
hnum
ori
enta
le L
. Bl
echn
um o
rien
tale
L.
Blec
hnum
ori
enta
le L
.
Brai
nea
insi
gnis
(Hoo
k.) J
.Sm
. Br
aine
a in
sign
is (H
ook.
) J.S
m.
Brai
nea
insi
gnis
(Hoo
k.) J
.Sm
.
Sten
ochl
aena
pal
ustr
is (B
urm
.f.) B
edd.
St
enoc
hlae
na p
alus
tris
(Bur
m.f.
) Bed
d.St
enoc
hlae
na p
alus
tris
(Bur
m.f.
) Bed
d.
○W
oodw
ardi
a ha
rlan
dii H
ook.
4,7
Woo
dwar
dia
harl
andi
i Hoo
k.A
bsen
t
●W
oodw
ardi
a ja
poni
ca (L
.f.) S
m.
Woo
dwar
dia
japo
nica
(L.f.
) Sm
.W
oodw
ardi
a co
chin
-chi
nens
is C
hing
CIB
OT
IAC
EA
E
Cib
otiu
m b
arom
etz (
L.) J
.Sm
. C
ibot
ium
bar
omet
z (L.
) J.S
m.
Cib
otiu
m b
arom
etz (
L.) J
.Sm
.
72 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
CYA
TH
EA
CE
AE
Cya
thea
bor
neen
sis C
opel
. C
yath
ea b
orne
ensi
s Cop
el.
Cya
thea
bor
neen
sis C
opel
.
Cya
thea
chi
nens
is C
opel
. C
yath
ea c
hine
nsis
Cop
el.
Cya
thea
chi
nens
is C
opel
.
Cya
thea
con
tam
inan
s (W
all.
ex H
ook.
) Cop
el.
Cya
thea
con
tam
inan
s (W
all.
ex H
ook.
) Cop
el.
Cya
thea
con
tam
inan
s (W
all.
ex H
ook.
) Cop
el.
Cya
thea
gig
ante
a (W
all.
ex H
ook.
) Hol
ttum
C
yath
ea g
igan
tea
(Wal
l. ex
Hoo
k.) H
olttu
mC
yath
ea g
igan
tea
(Wal
l. ex
Hoo
k.) H
olttu
m
○C
yath
ea g
labr
a (B
lum
e) C
opel
.44A
bsen
tA
bsen
t
○C
yath
ea h
ymen
odes
Met
t.44A
bsen
tA
bsen
t
Cya
thea
late
bros
a (W
all.
ex H
ook.
) Cop
el.
Cya
thea
late
bros
a (W
all.
ex H
ook.
) Cop
el.
Cya
thea
late
bros
a (W
all.
ex H
ook.
) Cop
el.
○C
yath
ea m
oluc
cana
R.B
r.10A
bsen
tA
bsen
t
Cya
thea
pod
ophy
lla (H
ook.
) Cop
el.
Cya
thea
pod
ophy
lla (H
ook.
) Cop
el.
Cya
thea
pod
ophy
lla (H
ook.
) Cop
el.
Cya
thea
spin
ulos
a W
all.
ex H
ook.
C
yath
ea sp
inul
osa
Wal
l. ex
Hoo
k.C
yath
ea sp
inul
osa
Wal
l. ex
Hoo
k.
DAV
AL
LIA
CE
AE
●Ar
aios
tegi
ella
fabe
rian
a (C
.Chr
.) M
.Kat
o &
Tsu
tsum
i Ar
aios
tegi
a fa
beri
ana
(C.C
hr.)
Chi
ngAr
aios
tegi
a fa
beri
ana
(C.C
hr.)
Chi
ng
Dav
allia
pet
elot
ii Ta
rdie
u &
C.C
hr.
Dav
allia
pet
elot
ii Ta
rdie
u &
C.C
hr.
Dav
allia
pet
elot
ii Ta
rdie
u &
C.C
hr.
Dav
allia
solid
a (G
.For
st.)
Sw.
Dav
allia
solid
a (G
.For
st.)
Sw.
Dav
allia
solid
a (G
.For
st.)
Sw.
Dav
allia
tric
hom
anoi
des B
lum
e va
r. tr
icho
man
oide
s D
aval
lia tr
icho
man
oide
s Blu
me
var.
tric
hom
anoi
des
Dav
allia
tric
hom
anoi
des B
lum
e va
r. tr
icho
man
oide
s
Dav
allia
tric
hom
anoi
des B
lum
e va
r. lo
rrai
nii
(Han
ce) H
olttu
m
Dav
allia
tric
hom
anoi
des B
lum
e va
r. lo
rrai
nii
(Han
ce) H
olttu
mD
aval
lia tr
icho
man
oide
s Blu
me
var.
lorr
aini
i (H
ance
) Hol
ttum
○D
aval
lode
s bor
neen
se (H
ook.
) Cop
el.30
Abs
ent
Abs
ent
○D
aval
lode
s hym
enop
hyllo
ides
(Blu
me)
M.K
ato
&
Tsut
sum
i30,1
9A
bsen
tA
bsen
t
●D
aval
lode
s im
bric
atum
(Chi
ng) M
.Kat
o &
Tsu
tsum
iAr
aios
tegi
a im
bric
ata
Chi
ngAr
aios
tegi
a im
bric
ata
Chi
ng
Dav
allo
des m
embr
anul
osum
(Hoo
k.) C
opel
. D
aval
lode
s mem
bran
ulos
um (H
ook.
) Cop
el.
Dav
allo
des m
embr
anul
osum
(Hoo
k.) C
opel
.
73TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●D
aval
lode
s pse
udoc
ysto
pter
is (K
unze
) M.K
ato
&
Tsut
sum
i Ar
aios
tegi
a ps
eudo
cyst
opte
ris (
Kun
ze) C
opel
.Ar
aios
tegi
a ps
eudo
cyst
opte
ris (
Kun
ze) C
opel
.
●D
aval
lode
s pul
chra
(D.D
on) M
.Kat
o &
Tsu
tsum
iAr
aios
tegi
a pu
lchr
a (D
.Don
) Cop
el.
Arai
oste
gia
pulc
hra
(D.D
on) C
opel
.
Dav
allo
des v
isci
dulu
m (K
uhn)
Ald
erw
. D
aval
lode
s vis
cidu
lum
(Kuh
n) A
lder
w.D
aval
lode
s vis
cidu
lum
(Kuh
n) A
lder
w.
Hum
ata
angu
stat
a (W
all.
ex H
ook.
& G
rev.
) J.S
m.
Hum
ata
angu
stat
a (W
all.
ex H
ook.
& G
rev.
) J.S
m.
Hum
ata
angu
stat
a (W
all.
ex H
ook.
& G
rev.
) J.S
m.
/ Pac
hypl
euri
a an
gust
ata
(Wal
l. ex
Hoo
k. &
G
rev.
) C.P
resl
●H
umat
a co
rnic
ulat
a (T
.Moo
re) M
.Kat
o &
Tsu
tsum
iD
aval
lia c
orni
cula
ta T
.Moo
reD
aval
lia c
orni
cula
ta T
.Moo
re
Hum
ata
hete
roph
ylla
(Sm
.) D
esv.
H
umat
a he
tero
phyl
la (S
m.)
Des
v.H
umat
a he
tero
phyl
la (S
m.)
Des
v. /
Dav
allia
het
erop
hylla
Sm
.
Hum
ata
pect
inat
a (S
m.)
Des
v.
Hum
ata
pect
inat
a (S
m.)
Des
v.H
umat
a pe
ctin
ata
(Sm
.) D
esv.
/ Pa
chyp
leur
ia p
ectin
ata
(Sm
.) C
.Pre
sl
Hum
ata
repe
ns (L
.f.) D
iels
Hum
ata
repe
ns (L
.f.) J
.Sm
ex
Die
lsH
umat
a re
pens
(L.f.
) J.S
m e
x D
iels
/ Pa
chyp
leur
ia re
pens
(L.f.
) M.K
ato
Hum
ata
vest
ita (B
lum
e) T
.Moo
reH
umat
a ve
stita
(Blu
me)
T.M
oore
Hum
ata
vest
ita (B
lum
e) T
.Moo
re /
Pach
yple
uria
ves
tita
(Blu
me)
C.P
resl
●W
ibel
ia d
entic
ulat
a ( B
urm
.f.) M
.Kat
o &
Tsu
tsum
iD
aval
lia d
entic
ulat
a (B
urm
.f.) M
ett.
ex K
uhn
Dav
allia
den
ticul
ata
(Bur
m.f.
) Met
t. ex
Kuh
n
●W
ibel
ia d
ivar
icat
a (B
lum
e) M
.Kat
o &
Tsu
tsum
iD
aval
lia d
ivar
icat
a B
lum
eD
aval
lia d
ivar
icat
a B
lum
e
○W
ibel
ia e
mbo
lost
egia
(Cop
el.)
M.K
ato
& T
suts
umi10
,19
Abs
ent
Abs
ent
DE
NN
STA
ED
TIA
CE
AE
Den
nsta
edtia
scab
ra (W
all.
ex H
ook.
) T.M
oore
D
enns
taed
tia sc
abra
(Wal
l. ex
Hoo
k.) T
.Moo
reD
enns
taed
tia sc
abra
(Wal
l. ex
Hoo
k.) T
.Moo
re
His
tiopt
eris
inci
sa (T
hunb
.) J.S
m.
His
tiopt
eris
inci
sa (T
hunb
.) J.S
m.
His
tiopt
eris
inci
sa (T
hunb
.) J.S
m.
Hyp
olep
is p
unct
ata
(Thu
nb.)
Met
t. ex
Kuh
n H
ypol
epis
pun
ctat
a (T
hunb
.) M
ett.
ex K
uhn
Hyp
olep
is p
unct
ata
(Thu
nb.)
Met
t. ex
Kuh
n
Mic
role
pia
calv
esce
ns (W
all.
ex H
ook.
) C.P
resl
M
icro
lepi
a ca
lves
cens
(Wal
l. ex
Hoo
k.) C
.Pre
slM
icro
lepi
a ca
lves
cens
(Wal
l. ex
Hoo
k.) C
.Pre
sl
Mic
role
pia
firm
a M
ett.
ex K
uhn
Mic
role
pia
firm
a M
ett.
ex K
uhn
Mic
role
pia
firm
a M
ett.
ex K
uhn
Mic
role
pia
herb
acea
Chi
ng &
C.C
hr. e
x Ta
rdie
u &
C.C
hr.
Mic
role
pia
herb
acea
Chi
ng &
C.C
hr. e
x Ta
rdie
u &
C.C
hr.
Mic
role
pia
herb
acea
Chi
ng &
C.C
hr. e
x Ta
rdie
u &
C.C
hr.
74 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Mic
role
pia
hook
eria
na (W
all.
ex H
ook.
) C
.Pre
sl
Mic
role
pia
hook
eria
na (W
all.
ex H
ook.
) C
.Pre
slM
icro
lepi
a ho
oker
iana
(Wal
l. ex
Hoo
k.)
C.P
resl
Mic
role
pia
kurz
ii (C
.B.C
lark
e) B
edd.
M
icro
lepi
a ku
rzii
(C.B
.Cla
rke)
Bed
d.M
icro
lepi
a ku
rzii
(C.B
.Cla
rke)
Bed
d.
Mic
role
pia
plat
yphy
lla (D
.Don
) J.S
m.
Mic
role
pia
plat
yphy
lla (D
.Don
) J.S
m.
Mic
role
pia
plat
yphy
lla (D
.Don
) J.S
m.
Mic
role
pia
pube
rula
Ald
erw.
M
icro
lepi
a pu
beru
la A
lder
w.M
icro
lepi
a pu
beru
la A
lder
w.
Mic
role
pia
ridl
eyi C
opel
. M
icro
lepi
a ri
dley
i Cop
el.
Mic
role
pia
ridl
eyi C
opel
.
Mic
role
pia
spel
unca
e (L
.) T.
Moo
re
Mic
role
pia
spel
unca
e (L
.) T.
Moo
reM
icro
lepi
a sp
elun
cae
(L.)
T.M
oore
Mic
role
pia
stri
gosa
(Thu
nb.)
C.P
resl
M
icro
lepi
a st
rigo
sa (T
hunb
.) C
.Pre
slM
icro
lepi
a st
rigo
sa (T
hunb
.) C
.Pre
sl
Mic
role
pia
taiw
ania
na T
agaw
a M
icro
lepi
a ta
iwan
iana
Tag
awa
Mic
role
pia
taiw
ania
na T
agaw
a
Mic
role
pia
trap
ezifo
rmis
(Rox
b.) K
uhn
Mic
role
pia
trap
ezifo
rmis
(Rox
b.) K
uhn
Mic
role
pia
trap
ezifo
rmis
(Rox
b.) K
uhn
○M
onac
hoso
rum
hen
ryi C
hris
t26
Abs
ent
Abs
ent
○Pt
erid
ium
aqu
ilinu
m (L
.) K
uhn
subs
p. ja
poni
cum
(N
akai
) Á.L
öve
& D
.Löv
e4,6,
38,4
2Pt
erid
ium
aqu
ilinu
m (L
.) K
uhn
var.
latiu
scul
um
(Des
v.) U
nder
w. e
x A
.Hel
ler
Abs
ent
●Pt
erid
ium
aqu
ilinu
m (L
.) K
uhn
subs
p. w
ight
ianu
m
(J.a
gard
h) W
.C.S
hieh
38,4
2Pt
erid
ium
aqu
ilinu
m (L
.) K
uhn
var.
wig
htia
num
(J
.aga
rdh)
r.M
.Try
onPt
erid
ium
aqu
ilinu
m (L
.) K
uhn
subs
p.
aqui
linum
var
. wig
htia
num
(J.a
gard
h)
r.M
.Try
on
●Pt
erid
ium
sem
ihas
tatu
m (W
all.
ex J.
aga
rdh)
S.
B.A
ndre
ws42
Pter
idiu
m a
quili
num
(L.)
Kuh
n va
r. ya
rrab
ense
D
omin
Pter
idiu
m a
quili
num
(L.)
Kuh
n su
bsp.
ca
udat
um v
ar. y
arra
bens
e D
omin
DIP
TE
RID
AC
EA
E
Che
iropl
euri
a bi
cusp
is (B
lum
e) C
.Pre
sl
Che
iropl
euri
a bi
cusp
is (B
lum
e) C
.Pre
slC
heiro
pleu
ria
bicu
spis
(Blu
me)
C.P
resl
Dip
teri
s con
juga
ta R
einw
. D
ipte
ris c
onju
gata
Rei
nw.
Dip
teri
s con
juga
ta R
einw
.
DR
YO
PTE
RID
AC
EA
E
●Ac
roph
orus
nod
osus
C.P
resl
Acro
phor
us st
ipel
latu
s T.M
oore
, nom
. nud
.Ac
roph
orus
stip
ella
tus T
.Moo
re, n
om. n
ud.
Arac
hnio
des a
ssam
ica
(Kuh
n) O
hwi
Arac
hnio
des a
ssam
ica
(Kuh
n) O
hwi
Arac
hnio
des a
ssam
ica
(Kuh
n) O
hwi
75TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Arac
hnio
des c
aval
erii
(Chr
ist)
Ohw
i Ar
achn
iode
s cav
aler
ii (C
hris
t) O
hwi
Arac
hnio
des c
aval
erii
(Chr
ist)
Ohw
i
Arac
hnio
des c
hine
nsis
(Ros
enst
.) C
hing
Ar
achn
iode
s chi
nens
is (R
osen
st.)
Chi
ngAr
achn
iode
s chi
nens
is (R
osen
st.)
Chi
ng
○Ar
achn
iode
s con
iifol
ia (T
.Moo
re) C
hing
8,10
Abs
ent
Abs
ent
Arac
hnio
des h
enry
i (C
hris
t) C
hing
Ar
achn
iode
s hen
ryi (
Chr
ist)
Chi
ngAr
achn
iode
s hen
ryi (
Chr
ist)
Chi
ng
Arac
hnio
des s
peci
osa
(D.D
on) C
hing
Ar
achn
iode
s spe
cios
a (D
.Don
) Chi
ngAr
achn
iode
s spe
cios
a (D
.Don
) Chi
ng
Arac
hnio
des s
pect
abili
s (C
hing
) Chi
ng
Arac
hnio
des s
pect
abili
s (C
hing
) Chi
ngAr
achn
iode
s spe
ctab
ilis (
Chi
ng) C
hing
○Ar
achn
iode
s sub
refle
xipi
nna
(Oga
ta) C
hing
4Ar
achn
iode
s sub
refle
xipi
nna
(Oga
ta) C
hing
A
bsen
t
Bolb
itis a
ngus
tipin
na (H
ayat
a) H
.Ito
Bolb
itis a
ngus
tipin
na (H
ayat
a) H
.Ito
Bolb
itis a
ngus
tipin
na (H
ayat
a) H
.Ito
Bolb
itis a
ppen
dicu
lata
(Will
d.) K
.Iwat
s. su
bsp.
ap
pend
icul
ata
Bolb
itis a
ppen
dicu
lata
(Will
d.) K
.Iwat
s. su
bsp.
ap
pend
icul
ata
Bolb
itis a
ppen
dicu
lata
(Will
d.) K
.Iwat
s. su
bsp.
ap
pend
icul
ata
Bolb
itis c
opel
andi
i Chi
ng e
x C
.Chr
. & T
ardi
eu
Bolb
itis c
opel
andi
i Chi
ng e
x C
.Chr
. & T
ardi
euBo
lbiti
s cop
elan
dii C
hing
ex
C.C
hr. &
Tar
dieu
Bolb
itis c
osta
ta (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis c
osta
ta (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis c
osta
ta (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis d
eltig
era
(Bed
d.) C
.Chr
. Bo
lbiti
s del
tiger
a (B
edd.
) C.C
hr.
Bolb
itis d
eltig
era
(Bed
d.) C
.Chr
.
Bolb
itis h
eter
oclit
a (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis h
eter
oclit
a (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis h
eter
oclit
a (C
.Pre
sl) C
hing
ex
C.C
hr.
Bolb
itis h
ooke
rian
a K
.Iwat
s. Bo
lbiti
s hoo
keri
ana
K.Iw
ats.
+ Bo
lbiti
s app
endi
cula
ta
(Will
d.) K
.Iwat
s. su
bsp.
viv
ipar
a (B
uch.
-Ham
. ex
Hoo
k.)
Hen
nipm
an
Bolb
itis h
ooke
rian
a K
.Iwat
s.
Bolb
itis s
calp
tura
ta (F
ée) C
hing
Bo
lbiti
s sca
lptu
rata
(Fée
) Chi
ngBo
lbiti
s sca
lptu
rata
(Fée
) Chi
ng
Bolb
itis s
inen
sis (
Bak
er) K
.Iwat
s. va
r. si
nens
is
Bolb
itis s
inen
sis (
Bak
er) K
.Iwat
s. va
r. si
nens
isBo
lbiti
s sin
ensi
s (B
aker
) K.Iw
ats.
var.
sine
nsis
Bolb
itis s
inen
sis (
Bak
er) K
.Iwat
s. va
r. co
stul
ata
(Hoo
k.) T
agaw
a &
K.Iw
ats.
Bolb
itis s
inen
sis (
Bak
er) K
.Iwat
s. va
r. co
stul
ata
(Hoo
k.) T
agaw
a &
K.Iw
ats.
Bolb
itis s
inen
sis (
Bak
er) K
.Iwat
s. va
r. co
stul
ata
(Hoo
k.) T
agaw
a &
K.Iw
ats
Bolb
itis s
inua
ta (C
.Pre
sl) H
enni
pman
Bo
lbiti
s sin
uata
(C.P
resl
) Hen
nipm
anBo
lbiti
s sin
uata
(C.P
resl
) Hen
nipm
an
Bolb
itis t
onki
nens
is (C
.Chr
. ex
Chi
ng) K
.Iwat
s.Bo
lbiti
s ton
kine
nsis
(C.C
hr. e
x C
hing
) K.Iw
ats.
Bolb
itis t
onki
nens
is (C
.Chr
. ex
Chi
ng) K
.Iwat
s
Bolb
itis v
irens
(Wal
l. ex
Hoo
k. &
Gre
v.) S
chot
t va
r. vi
rens
Bolb
itis v
irens
(Wal
l. ex
Hoo
k. &
Gre
v.) S
chot
t va
r. vi
rens
Bolb
itis v
irens
(Wal
l. ex
Hoo
k. &
Gre
v.) S
chot
t va
r. vi
rens
Bolb
itis v
irens
(Wal
l. ex
Hoo
k. &
Gre
v.) S
chot
t va
r. co
mpa
cta
Hen
nipm
anBo
lbiti
s vire
ns (W
all.
ex H
ook.
& G
rev.
) Sch
ott
var.
com
pact
a H
enni
pman
Bolb
itis v
irens
(Wal
l. ex
Hoo
k. &
Gre
v.) S
chot
t va
r. co
mpa
cta
Hen
nipm
an
76 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Cte
nitis
dum
rong
ii Ta
gaw
a &
K.Iw
ats.
Cte
nitis
dum
rong
ii Ta
gaw
a &
K.Iw
ats.
Cte
nitis
dum
rong
ii Ta
gaw
a &
K.Iw
ats.
●C
teni
tis su
bobs
cura
(Chr
ist)
Hol
ttum
C
teni
tis su
bobs
cura
(Chr
ist)
Hol
ttum
Cte
nitis
man
nii a
uct.
non
(C.H
ope)
Chi
ng /
Cte
nitis
subo
bscu
ra (C
hris
t) H
olttu
m
Cte
nitis
vili
s (K
unze
) Chi
ng
Cte
nitis
vili
s (K
unze
) Chi
ngC
teni
tis v
ilis (
Kun
ze) C
hing
Cyr
tom
ium
fort
unei
J.Sm
. C
yrto
miu
m fo
rtun
ei J.
Sm.
Cyr
tom
ium
fort
unei
J.Sm
.
Did
ymoc
hlae
na tr
unca
tula
(Sw.
) J.S
m.
Did
ymoc
hlae
na tr
unca
tula
(Sw.
) J.S
m.
Did
ymoc
hlae
na tr
unca
tula
(Sw.
) J.S
m.
Dry
opte
ris c
ochl
eata
(D.D
on) C
.Chr
. D
ryop
teri
s coc
hlea
ta (D
.Don
) C.C
hr.
Dry
opte
ris c
ochl
eata
(D.D
on) C
.Chr
.
○D
ryop
teri
s diff
ract
a (B
aker
) C.C
hr.10
Abs
ent
Abs
ent
Dry
opte
ris g
ymno
phyl
la (B
aker
) C.C
hr.
Dry
opte
ris g
ymno
phyl
la (B
aker
) C.C
hr.
Dry
opte
ris g
ymno
phyl
la (B
aker
) C.C
hr.
●D
ryop
teri
s has
selti
i (B
lum
e) C
.Chr
. Ar
achn
iode
s has
selti
i (B
lum
e) C
hing
Arac
hnio
des h
asse
ltii (
Blu
me)
Chi
ng
Dry
opte
ris h
ende
rson
ii (B
edd.
) C.C
hr.
Dry
opte
ris h
ende
rson
ii (B
edd.
) C.C
hr.
Dry
opte
ris h
ende
rson
ii (B
edd.
) C.C
hr.
Dry
opte
ris h
irtip
es (B
lum
e) K
untz
e D
ryop
teri
s hir
tipes
(Blu
me)
Kun
tze
Dry
opte
ris h
irtip
es (B
lum
e) K
untz
e
Dry
opte
ris i
nteg
rilo
ba C
.Chr
. D
ryop
teri
s int
egri
loba
C.C
hr.
Dry
opte
ris i
nteg
rilo
ba C
.Chr
.
Dry
opte
ris n
eoas
sam
ensi
s Chi
ng
Dry
opte
ris n
eoas
sam
ensi
s Chi
ngD
ryop
teri
s neo
assa
men
sis C
hing
Dry
opte
ris n
eoch
ryso
com
a C
hing
D
ryop
teri
s neo
chry
soco
ma
Chi
ngD
ryop
teri
s neo
chry
soco
ma
Chi
ng
Dry
opte
ris p
olita
Ros
enst
. D
ryop
teri
s pol
ita R
osen
st.
Dry
opte
ris p
olita
Ros
enst
.
Dry
opte
ris p
oros
a C
hing
D
ryop
teri
s por
osa
Chi
ngD
ryop
teri
s por
osa
Chi
ng
Dry
opte
ris p
seud
ospa
rsa
Chi
ng
Dry
opte
ris p
seud
ospa
rsa
Chi
ngD
ryop
teri
s pse
udos
pars
a C
hing
Dry
opte
ris r
heop
hila
Mits
uta
ex D
arna
edi,
M.K
ato
& K
.Iwat
s. D
ryop
teri
s rhe
ophi
la M
itsut
a ex
Dar
naed
i, M
.Kat
o &
K.Iw
ats.
Dry
opte
ris r
heop
hila
Mits
uta
ex D
arna
edi,
M.K
ato
& K
.Iwat
s.
Dry
opte
ris s
cotti
i (B
edd.
) Chi
ng
Dry
opte
ris s
cotti
i (B
edd.
) Chi
ngD
ryop
teri
s sco
ttii (
Bed
d.) C
hing
Dry
opte
ris s
pars
a (D
.Don
) Kun
tze
Dry
opte
ris s
pars
a (D
.Don
) Kun
tze
Dry
opte
ris s
pars
a (D
.Don
) Kun
tze
Dry
opte
ris s
ubtr
iang
ular
is (C
.Hop
e) C
.Chr
. D
ryop
teri
s sub
tria
ngul
aris
(C.H
ope)
C.C
hr.
Dry
opte
ris s
ubtr
iang
ular
is (C
.Hop
e) C
.Chr
.
Elap
hogl
ossu
m a
ngul
atum
(Blu
me)
T.M
oore
El
apho
glos
sum
ang
ulat
um (B
lum
e) T
.Moo
reEl
apho
glos
sum
ang
ulat
um (B
lum
e) T
.Moo
re
Elap
hogl
ossu
m d
umro
ngii
Taga
wa
& K
.Iwat
s. El
apho
glos
sum
dum
rong
ii Ta
gaw
a &
K.Iw
ats.
Elap
hogl
ossu
m d
umro
ngii
Taga
wa
& K
.Iwat
s.
Elap
hogl
ossu
m m
alay
ense
Hol
ttum
El
apho
glos
sum
mal
ayen
se H
olttu
mEl
apho
glos
sum
mal
ayen
se H
olttu
m
77TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Elap
hogl
ossu
m m
argi
natu
m (F
ée) T
.Moo
re
Elap
hogl
ossu
m m
argi
natu
m (F
ée) T
.Moo
reEl
apho
glos
sum
mar
gina
tum
(Fée
) T.M
oore
Elap
hogl
ossu
m m
elan
ostic
tum
(Blu
me)
T.M
oore
El
apho
glos
sum
mel
anos
tictu
m (B
lum
e) T
.Moo
reEl
apho
glos
sum
mel
anos
tictu
m (B
lum
e) T
.Moo
re
Elap
hogl
ossu
m st
ellig
erum
(Wal
l. ex
Bak
er)
T.M
oore
ex
als
ton
& B
onne
rEl
apho
glos
sum
stel
liger
um (W
all.
ex B
aker
) T.
Moo
re e
x a
lsto
n &
Bon
ner
Elap
hogl
ossu
m st
ellig
erum
(Wal
l. ex
Bak
er)
T.M
oore
ex
als
ton
& B
onne
r
Elap
hogl
ossu
m su
belli
ptic
um R
osen
st.
Elap
hogl
ossu
m su
belli
ptic
um R
osen
st.
Elap
hogl
ossu
m su
belli
ptic
um R
osen
st.
Elap
hogl
ossu
m y
oshi
naga
e (Y
atab
e) M
akin
o El
apho
glos
sum
yos
hina
gae
(Yat
abe)
Mak
ino
Elap
hogl
ossu
m y
oshi
naga
e (Y
atab
e) M
akin
o
Hyp
odem
atiu
m c
rena
tum
(For
ssk.
) Kuh
n H
ypod
emat
ium
cre
natu
m (F
orss
k.) K
uhn
Hyp
odem
atiu
m c
rena
tum
(For
ssk.
) Kuh
n
Hyp
odem
atiu
m g
land
ulos
o-pi
losu
m (T
agaw
a)
Ohw
i H
ypod
emat
ium
gla
ndul
oso-
pilo
sum
(Tag
awa)
O
hwi
Hyp
odem
atiu
m g
land
ulos
o-pi
losu
m (T
agaw
a)
Ohw
i
Leuc
oste
gia
imm
ersa
(Wal
l. ex
Hoo
k.) C
.Pre
sl
Leuc
oste
gia
imm
ersa
(Wal
l. ex
Hoo
k.) C
.Pre
slLe
ucos
tegi
a im
mer
sa (W
all.
ex H
ook.
) C.P
resl
Lom
agra
mm
a gr
osso
serr
ata
Hol
ttum
Lo
mag
ram
ma
gros
sose
rrat
a H
olttu
mLo
mag
ram
ma
gros
sose
rrat
a H
olttu
m
●Pe
rane
ma
aspi
dioi
des (
Blu
me)
Met
t. D
iaca
lpe
aspi
dioi
des B
lum
eD
iaca
lpe
aspi
dioi
des B
lum
e
Poly
stic
hum
atte
nuat
um T
agaw
a &
K.Iw
ats.
Poly
stic
hum
atte
nuat
um T
agaw
a &
K.Iw
ats.
Poly
stic
hum
atte
nuat
um T
agaw
a &
K.Iw
ats.
Poly
stic
hum
bia
rist
atum
(Blu
me)
T.M
oore
Po
lyst
ichu
m b
iari
stat
um (B
lum
e) T
.Moo
rePo
lyst
ichu
m b
iari
stat
um (B
lum
e) T
.Moo
re
Poly
stic
hum
exi
miu
m (K
uhn)
C.C
hr.
Poly
stic
hum
exi
miu
m (K
uhn)
C.C
hr.
Poly
stic
hum
exi
miu
m (K
uhn)
C.C
hr.
Poly
stic
hum
lind
saei
foliu
m S
cort.
ex
Rid
l. Po
lyst
ichu
m li
ndsa
eifo
lium
Sco
rt. e
x. R
idl.
Poly
stic
hum
lind
saei
foliu
m S
cort.
ex
Rid
l.
Poly
stic
hum
pro
lifica
ns A
lder
w.
Poly
stic
hum
pro
lifica
ns A
lder
w.Po
lyst
ichu
m p
rolifi
cans
Ald
erw.
○Po
lyst
ichu
m p
seud
otsu
s-si
men
se C
hing
8A
bsen
tA
bsen
t
○Po
lyst
ichu
m sc
ario
sum
(rox
b.) C
.V.M
orto
n8,10
Abs
ent
Abs
ent
Poly
stic
hum
sem
ifert
ile (C
.B.C
lark
e) C
hing
Po
lyst
ichu
m se
mife
rtile
(C.B
.Cla
rke)
Chi
ngPo
lyst
ichu
m se
mife
rtile
(C.B
.Cla
rke)
Chi
ng
Poly
stic
hum
teng
gere
nse
Ros
enst
. Po
lyst
ichu
m te
ngge
rens
e R
osen
st.
Poly
stic
hum
teng
gere
nse
Ros
enst
.
Tera
toph
yllu
m a
cule
atum
(Blu
me)
Met
t. ex
Kuh
n Te
rato
phyl
lum
acu
leat
um (B
lum
e) M
ett.
ex K
uhn
Tera
toph
yllu
m a
cule
atum
(Blu
me)
Met
t. ex
Kuh
n
Tera
toph
yllu
m lu
dens
(Fée
) Hol
ttum
Te
rato
phyl
lum
lude
ns (F
ée) H
olttu
mTe
rato
phyl
lum
lude
ns (F
ée) H
olttu
m
EQ
UIS
ETA
CE
AE
○Eq
uise
tum
diff
usum
D.D
on4
Equi
setu
m d
iffus
um D
.Don
Abs
ent
78 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●Eq
uise
tum
ram
osis
sim
um D
esf.
subs
p. d
ebile
(Rox
b. e
x Va
uche
r) H
auke
Eq
uise
tum
deb
ile R
oxb.
ex
Vauc
her
Equi
setu
m d
ebile
Rox
b. e
x Va
uche
r
GL
EIC
HE
NIA
CE
AE
Dic
rano
pter
is c
urra
nii C
opel
. D
icra
nopt
eris
cur
rani
i Cop
el.
Dic
rano
pter
is c
urra
nii C
opel
.
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
linea
ris
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
linea
ris
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
linea
ris
○D
icra
nopt
eris
line
aris
(Bur
m.f)
Und
erw.
va
r. m
onta
na H
olttu
m4
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
mon
tana
Hol
ttum
Abs
ent
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
subp
ectin
ata
(Chr
ist)
Hol
ttum
D
icra
nopt
eris
line
aris
(Bur
m.f.
) Und
erw.
va
r. su
bpec
tinat
a (C
hris
t) H
olttu
mD
icra
nopt
eris
line
aris
(Bur
m.f.
) Und
erw.
va
r. su
bpec
tinat
a (C
hris
t) H
olttu
m
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
tetr
aphy
lla (r
osen
st.)
Nak
ai
Dic
rano
pter
is li
near
is (B
urm
.f.) U
nder
w.
var.
tetr
aphy
lla (r
osen
st.)
Nak
aiD
icra
nopt
eris
line
aris
(Bur
m.f.
) Und
erw.
va
r. te
trap
hylla
(ros
enst
.) N
akai
Dic
rano
pter
is sp
ecio
sa (C
.Pre
sl) H
olttu
m
Dic
rano
pter
is sp
ecio
sa (C
.Pre
sl) H
olttu
mD
icra
nopt
eris
spec
iosa
(C.P
resl
) Hol
ttum
Dic
rano
pter
is sp
lend
ida
(Han
d.-M
azz.
) Tag
awa
Dic
rano
pter
is sp
lend
ida
(Han
d.-M
azz.
) Tag
awa
Dic
rano
pter
is sp
lend
ida
(Han
d.-M
azz.
) Tag
awa
●D
iplo
pter
ygiu
m b
lotia
num
(C.C
hr.)
Nak
aiG
leic
heni
a bl
otia
na C
.Chr
.G
leic
heni
a bl
otia
na C
.Chr
.
●D
iplo
pter
ygiu
m lo
ngis
sim
um (B
lum
e) N
akai
Gle
iche
nia
long
issi
ma
Blu
me
Gle
iche
nia
long
issi
ma
Blu
me
●D
iplo
pter
ygiu
m n
orri
sii (
Met
t.) N
akai
Gle
iche
nia
norr
isii
Met
t.G
leic
heni
a no
rris
ii M
ett.
Gle
iche
nia
mic
roph
ylla
R.B
r. G
leic
heni
a m
icro
phyl
la R
.Br.
Gle
iche
nia
mic
roph
ylla
R.B
r.
○St
iche
rus h
irtu
s (B
lum
e) C
hing
10A
bsen
tA
bsen
t
●St
iche
rus t
runc
atus
(Will
d.) N
akai
Gle
iche
nia
trun
cata
(Will
d.) S
pren
g.G
leic
heni
a tr
unca
ta (W
illd.
) Spr
eng.
HY
ME
NO
PHY
LL
AC
EA
E
●Ab
rodi
ctyu
m id
oneu
m (C
.V.M
orto
n)
Ebih
ara
& K
.Iwat
s.47
Cep
halo
man
es g
emm
atum
(J.S
m. e
x B
aker
) K
.Iwat
s. M
acro
glen
a ge
mm
ata
(J.S
m. e
x B
aker
) Cop
el.
/ Cep
halo
man
es g
emm
atum
(J.S
m. e
x B
aker
) K
.Iwat
s.
●Ab
rodi
ctyu
m o
bscu
rum
(Blu
me)
Ebi
hara
& K
.Iwat
s. va
r. ob
scur
um
Cep
halo
man
es o
bscu
rum
(Blu
me)
K.Iw
ats.
var.
obsc
urum
Sele
node
smiu
m o
bscu
rum
(Blu
me)
Cop
el.
79TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●Ab
rodi
ctyu
m o
bscu
rum
(Blu
me)
Ebi
hara
& K
.Iwat
s. va
r. si
amen
se (C
hris
t) K
.Iwat
s.C
epha
lom
anes
obs
curu
m (B
lum
e) K
.Iwat
s. va
r. si
amen
se (C
hris
t) K
.Iwat
s.Se
leno
desm
ium
obs
curu
m (B
lum
e) C
opel
., pr
o pa
rte /
Cep
halo
man
es o
bscu
rum
(Blu
me)
K
.Iwat
s. va
r. si
amen
se (C
hris
t) K
.Iwat
s.
●Ab
rodi
ctyu
m p
lum
a (H
ook.
) Ebi
hara
& K
.Iwat
s.47
Cep
halo
man
es m
eifo
lium
(Bor
y ex
Will
d.) K
.Iwat
s.M
acro
glen
a m
eifo
lia (B
ory
ex W
illd.
) Cop
el.
/ Cep
halo
man
es m
eifo
lium
(Bor
y ex
Will
d.)
K.Iw
ats.
Cal
listo
pter
is a
piifo
lia (C
.Pre
sl) C
opel
. C
epha
lom
anes
api
ifoliu
m (C
.Pre
sl) K
.Iwat
s.C
allis
topt
eris
api
ifolia
(C.P
resl
) Cop
el. /
C
epha
lom
anes
api
ifoliu
m (C
.Pre
sl) K
.Iwat
s.
Cep
halo
man
es ja
vani
cum
(Blu
me)
C.P
resl
C
epha
lom
anes
java
nicu
m (B
lum
e) C
.Pre
slC
epha
lom
anes
java
nicu
m (B
lum
e) C
.Pre
sl
●C
repi
dom
anes
bip
unct
atum
(Poi
r.) C
opel
. C
repi
dom
anes
bip
unct
atum
(Poi
r.) C
opel
.C
repi
dom
anes
bip
unct
atum
(Poi
r.) C
opel
. +
Crep
idom
anes
bila
biat
um (N
ees &
Blu
me)
Cop
el.
Cre
pido
man
es b
revi
pes (
C.P
resl
) Cop
el.
Cre
pido
man
es b
revi
pes (
C.P
resl
) Cop
el.
Cre
pido
man
es b
revi
pes (
C.P
resl
) Cop
el.
Cre
pido
man
es c
hris
tii (C
opel
.) C
opel
. C
repi
dom
anes
chr
istii
(Cop
el.)
Cop
el.
Cre
pido
man
es c
hris
tii (C
opel
.) C
opel
.
●C
repi
dom
anes
hum
ile (G
.For
st.)
Bos
ch
Cre
pido
man
es h
umile
(G.F
orst
.) B
osch
Reed
iella
hum
ilis (
G.F
orst
.) Pi
c.Se
rm. /
C
repi
dom
anes
hum
ile (G
.For
st.)
Bos
ch
Cre
pido
man
es k
urzi
i (B
edd.
) Tag
awa
& K
.Iwat
s. C
repi
dom
anes
kur
zii (
Bed
d.) T
agaw
a &
K.Iw
ats.
Cre
pido
man
es k
urzi
i (B
edd.
) Tag
awa
& K
.Iwat
s.
Cre
pido
man
es la
teal
atum
(Bos
ch) C
opel
. C
repi
dom
anes
late
alat
um (B
osch
) Cop
el.
Cre
pido
man
es la
teal
atum
(Bos
ch) C
opel
.
Cre
pido
man
es la
tem
argi
nale
(D.C
.Eat
on)
Cop
el.
Cre
pido
man
es la
tem
argi
nale
(D.C
.Eat
on)
Cop
el.
Cre
pido
man
es la
tem
argi
nale
(D.C
.Eat
on)
Cop
el.
Cre
pido
man
es m
egis
tost
omum
(Cop
el.)
Cop
el.
Cre
pido
man
es m
egis
tost
omum
(Cop
el.)
Cop
el.
Cre
pido
man
es m
egis
tost
omum
(Cop
el.)
Cop
el.
●C
repi
dom
anes
min
utum
(Blu
me)
K.Iw
ats.
Cre
pido
man
es m
inut
um (B
lum
e) K
.Iwat
s.G
onoc
orm
us p
rolif
er (B
lum
e) P
rant
l +
Gon
ocor
mus
saxi
frag
oide
s (C
.Pre
sl) B
osch
+
Gon
ocor
mus
siam
ensi
s Tag
awa
& K
.Iwat
s. /
Cre
pido
man
es m
inut
um (B
lum
e) K
.Iwat
s.
●C
repi
dom
anes
par
vifo
lium
(Bak
er) K
.Iwat
s. C
repi
dom
anes
par
vifo
lium
(Bak
er) K
.Iwat
s.M
icro
goni
um p
arvi
foliu
m (B
aker
) Tag
awa
&
K.Iw
ats.
/ Cre
pido
man
es p
arvi
foliu
m (B
aker
) K
.Iwat
s.
●D
idym
oglo
ssum
bim
argi
natu
m (B
osch
) Ebi
hara
&
K.Iw
ats.
Tric
hom
anes
bim
argi
natu
m (B
osch
) Bos
ch
Mic
rogo
nium
bim
argi
natu
m B
osch
/ Tr
icho
man
es
bim
argi
natu
m (B
osch
) Bos
ch
80 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Did
ymog
loss
um e
xigu
um (B
edd.
) Cop
el.
Tric
hom
anes
exi
guum
(Bed
d.) B
aker
Did
ymog
loss
um e
xigu
um (B
edd.
) Cop
el. /
Tr
icho
man
es e
xigu
um (B
edd.
) Bak
er
●D
idym
oglo
ssum
mot
leyi
(Bos
ch) E
biha
ra &
K.Iw
ats.
Tric
hom
anes
mot
leyi
(Bos
ch) B
osch
Mic
rogo
nium
mot
leyi
Bos
ch /
Tric
hom
anes
m
otle
yi (B
osch
) Bos
ch
●D
idym
oglo
ssum
subl
imba
tum
(Mül
l.Ber
ol.)
ebih
ara
&
K.Iw
ats.
Tric
hom
anes
subl
imba
tum
Mül
l.Ber
ol.
Mic
rogo
nium
subl
imba
tum
(Mül
l.Ber
ol.)
Bos
ch /
Tric
hom
anes
subl
imba
tum
Mül
l.Ber
ol.
●H
ymen
ophy
llum
aca
ntho
ides
(Bos
ch) R
osen
st.
Hym
enop
hyllu
m a
cant
hoid
es (B
osch
) Ros
enst
.M
erin
gium
aca
ntho
ides
(Bos
ch) C
opel
. /
Hym
enop
hyllu
m a
cant
hoid
es (B
osch
) Ros
enst
.
●H
ymen
ophy
llum
bad
ium
Hoo
k. &
Gre
v.
Hym
enop
hyllu
m b
adiu
m H
ook.
& G
rev.
Mec
odiu
m b
adiu
m (H
ook.
& G
rev.
) Cop
el. /
H
ymen
ophy
llum
bad
ium
Hoo
k. &
Gre
v.
Hym
enop
hyllu
m b
arba
tum
(Bos
ch) B
aker
H
ymen
ophy
llum
bar
batu
m (B
osch
) Bak
erH
ymen
ophy
llum
bar
batu
m (B
osch
) Bak
er
●H
ymen
ophy
llum
bla
ndum
Rac
ib.
Hym
enop
hyllu
m b
land
um R
acib
.M
erin
gium
bla
ndum
(Rac
ib.)
Cop
el. /
H
ymen
ophy
llum
bla
ndum
Rac
ib.
●H
ymen
ophy
llum
bon
toce
nse
Cop
el.
Hym
enop
hyllu
m b
onto
cens
e C
opel
.M
erin
gium
bon
toce
nse
(Cop
el.)
Cop
el. /
H
ymen
ophy
llum
bon
toce
nse
Cop
el.
●H
ymen
ophy
llum
den
ticul
atum
Sw.
H
ymen
ophy
llum
den
ticul
atum
Sw.
Mer
ingi
um d
entic
ulat
um (S
w.) C
opel
. /
Hym
enop
hyllu
m d
entic
ulat
um S
w.
●H
ymen
ophy
llum
dig
itatu
m (S
w.) F
osbe
rg
Cre
pido
man
es d
igita
tum
(Sw.
) K.Iw
ats.
Mic
rotr
icho
man
es d
igita
tum
(Sw.
) Cop
el. /
C
repi
dom
anes
dig
itatu
m (S
w.) K
.Iwat
s.
●H
ymen
ophy
llum
exs
ertu
m W
all.
ex H
ook.
H
ymen
ophy
llum
exs
ertu
m W
all.
ex H
ook.
Mec
odiu
m e
xser
tum
(Wal
l. ex
Hoo
k.) C
opel
. /
Hym
enop
hyllu
m e
xser
tum
Wal
l. ex
Hoo
k.
●H
ymen
ophy
llum
hol
ochi
lum
(Bos
ch) C
.Chr
. H
ymen
ophy
llum
hol
ochi
lum
(Bos
ch) C
.Chr
.M
erin
gium
hol
ochi
lum
(Bos
ch) C
opel
. /
Hym
enop
hyllu
m h
oloc
hilu
m (B
osch
) C.C
hr.
●H
ymen
ophy
llum
java
nicu
m S
pren
g.
Hym
enop
hyllu
m ja
vani
cum
Spr
eng.
Mec
odiu
m ja
vani
cum
(Spr
eng.
) Cop
el. /
H
ymen
ophy
llum
java
nicu
m S
pren
g.
●H
ymen
ophy
llum
pal
lidum
(Blu
me)
Ebi
hara
& K
.Iwat
s. C
repi
dom
anes
pal
lidum
(Blu
me)
K.Iw
ats.
Pleu
rom
anes
pal
lidum
(Blu
me)
C.P
resl
. /
Cre
pido
man
es p
allid
um (B
lum
e) K
.Iwat
s.
●H
ymen
ophy
llum
pol
yant
hos (
Sw.)
Sw.
Hym
enop
hyllu
m p
olya
ntho
s (Sw
.) Sw
.M
ecod
ium
pol
yant
hos (
Sw.)
Cop
el. /
H
ymen
ophy
llum
pol
yant
hos (
Sw.)
Sw.
81TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●H
ymen
ophy
llum
pro
duct
um K
unze
H
ymen
ophy
llum
pro
duct
um K
unze
Mec
odiu
m p
rodu
ctum
(Kun
ze) C
opel
. /
Hym
enop
hyllu
m p
rodu
ctum
Kun
ze
●H
ymen
ophy
llum
riuk
iuen
se C
hris
t H
ymen
ophy
llum
riuk
iuen
se C
hris
tM
ecod
ium
riuk
iuen
se (C
hris
t) C
opel
. /
Hym
enop
hyllu
m ri
ukiu
ense
Chr
ist
●H
ymen
ophy
llum
serr
ulat
um (C
.Pre
sl) C
.Chr
. H
ymen
ophy
llum
serr
ulat
um (C
.Pre
sl) C
.Chr
.M
erin
gium
mey
enia
num
C.P
resl
/ H
ymen
ophy
llum
serr
ulat
um (C
.Pre
sl) C
.Chr
.
●Va
nden
bosc
hia
auri
cula
ta (B
lum
e) C
opel
. C
repi
dom
anes
aur
icul
atum
(Blu
me)
K.Iw
ats.
Tric
hom
anes
aur
icul
atum
Blu
me
/ Cre
pido
man
es
auri
cula
tum
(Blu
me)
K.Iw
ats.
●Va
nden
bosc
hia
birm
anic
a (B
edd.
) Chi
ng
Cre
pido
man
es b
irm
anic
um (B
edd.
) K.Iw
ats.
Tric
hom
anes
bir
man
icum
Bed
d. /
Cre
pido
man
es
birm
anic
um (B
edd.
) K.Iw
ats.
●Va
nden
bosc
hia
max
ima
(Blu
me)
Cop
el.
Cre
pido
man
es m
axim
um (B
lum
e) K
.Iwat
s.Tr
icho
man
es m
axim
um B
lum
e / C
repi
dom
anes
m
axim
um (B
lum
e) K
.Iwat
s.
LIN
DSA
EA
CE
AE
Lind
saea
bou
illod
ii C
hris
t Li
ndsa
ea b
ouill
odii
Chr
ist
Lind
saea
bou
illod
ii C
hris
t
Lind
saea
chi
enii
Chi
ng
Lind
saea
chi
enii
Chi
ngLi
ndsa
ea c
hien
ii C
hing
Lind
saea
cul
trat
a (W
illd.
) Sw.
Li
ndsa
ea c
ultr
ata
(Will
d.) S
w.Li
ndsa
ea c
ultr
ata
(Will
d.) S
w.
○Li
ndsa
ea d
isse
ctifo
rmis
Chi
ng10
Abs
ent
Abs
ent
Lind
saea
div
erge
ns H
ook.
& G
rev.
Lind
saea
div
erge
ns H
ook.
& G
rev.
Lind
saea
div
erge
ns H
ook.
& G
rev.
Lind
saea
dor
ypho
ra K
.U.K
ram
er
Lind
saea
dor
ypho
ra K
.U.K
ram
erLi
ndsa
ea d
oryp
hora
K.U
.Kra
mer
Lind
saea
ens
ifolia
Sw.
Li
ndsa
ea e
nsifo
lia S
w.Li
ndsa
ea e
nsifo
lia S
w.
Lind
saea
het
erop
hylla
Dry
and.
Li
ndsa
ea h
eter
ophy
lla D
ryan
d.Li
ndsa
ea h
eter
ophy
lla D
ryan
d.
Lind
saea
inte
gra
Hol
ttum
Li
ndsa
ea in
tegr
a H
olttu
mLi
ndsa
ea in
tegr
a H
olttu
m
Lind
saea
java
nens
is B
lum
e Li
ndsa
ea ja
vane
nsis
Blu
me
Lind
saea
java
nens
is B
lum
e
Lind
saea
luci
da B
lum
e Li
ndsa
ea lu
cida
Blu
me
Lind
saea
luci
da B
lum
e
Lind
saea
mal
ayen
sis H
olttu
m
Lind
saea
mal
ayen
sis H
olttu
mLi
ndsa
ea m
alay
ensi
s Hol
ttum
Lind
saea
nap
aea
Ald
erw.
Li
ndsa
ea n
apae
a A
lder
w.Li
ndsa
ea n
apae
a A
lder
w.
82 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Lind
saea
obl
ance
olat
a A
lder
w.
Lind
saea
obl
ance
olat
a A
lder
w.Li
ndsa
ea o
blan
ceol
ata
Ald
erw.
Lind
saea
odo
rata
Rox
b.
Lind
saea
odo
rata
Rox
b.Li
ndsa
ea o
dora
ta R
oxb.
Lind
saea
orb
icul
ata
(Lam
.) M
ett.
ex K
uhn
var.
orbi
cula
ta
Lind
saea
orb
icul
ata
(Lam
.) M
ett.
ex K
uhn
var.
orbi
cula
taLi
ndsa
ea o
rbic
ulat
a (L
am.)
Met
t. ex
Kuh
n va
r. or
bicu
lata
Lind
saea
orb
icul
ata
(Lam
.) M
ett.
ex K
uhn
var.
com
mix
ta (T
agaw
a) K
.U.K
ram
erLi
ndsa
ea o
rbic
ulat
a (L
am.)
Met
t. ex
Kuh
n va
r. co
mm
ixta
(Tag
awa)
K.U
.Kra
mer
Lind
saea
orb
icul
ata
(Lam
.) M
ett.
ex K
uhn
var.
com
mix
ta (T
agaw
a) K
.U.K
ram
er
Lind
saea
par
alle
logr
amm
a A
lder
w.
Lind
saea
par
alle
logr
amm
a A
lder
w.Li
ndsa
ea p
aral
lelo
gram
ma
Ald
erw.
Lind
saea
par
asiti
ca (R
oxb.
ex
Grif
f.) H
iero
n.
Lind
saea
par
asiti
ca (R
oxb.
ex
Grif
f.) H
iero
n.Li
ndsa
ea p
aras
itica
(Rox
b. e
x G
riff.)
Hie
ron.
Lind
saea
repe
ns (B
ory)
Thw
aite
s var
. pec
tinat
a (B
lum
e) M
ett.
ex K
uhn
Lind
saea
repe
ns (B
ory)
Thw
aite
s var
. pec
tinat
a (B
lum
e) M
ett.
ex K
uhn
Lind
saea
repe
ns (B
ory)
Thw
aite
s var
. pec
tinat
a (B
lum
e) M
ett.
ex K
uhn
○Li
ndsa
ea te
nera
Dry
and.
4Li
ndsa
ea te
nera
Dry
and.
Abs
ent
Sphe
nom
eris
chi
nens
is (L
.) M
axon
va
r. ch
inen
sis
Sphe
nom
eris
chi
nens
is (L
.) M
axon
va
r. ch
inen
sis
Sphe
nom
eris
chi
nens
is (L
.) M
axon
va
r. ch
inen
sis
Sphe
nom
eris
chi
nens
is (L
.) M
axon
var.
diva
rica
ta (C
hris
t) K
.U.K
ram
er
Sphe
nom
eris
chi
nens
is (L
.) M
axon
va
r. di
vari
cata
(Chr
ist)
K.U
.Kra
mer
Sphe
nom
eris
chin
ensis
(L.)
Max
on
var.
diva
ricat
a (C
hris
t) K
.U.K
ram
er
Tape
inid
ium
luzo
nicu
m (H
ook.
) K.U
.Kra
mer
Ta
pein
idiu
m lu
zoni
cum
(Hoo
k.) K
.U.K
ram
erTa
pein
idiu
m lu
zoni
cum
(Hoo
k.) K
.U.K
ram
er
Tape
inid
ium
pin
natu
m (C
av.)
C.C
hr.
Tape
inid
ium
pin
natu
m (C
av.)
C.C
hr.
Tape
inid
ium
pin
natu
m (C
av.)
C.C
hr.
LO
MA
RIO
PSID
AC
EA
E
Cyc
lope
ltis c
rena
ta (F
ée) C
.Chr
. C
yclo
pelti
s cre
nata
(Fée
) C.C
hr.
Cyc
lope
ltis c
rena
ta (F
ée) C
.Chr
.
Lom
ario
psis
line
ata
(C.P
resl
) Hol
ttum
Lo
mar
iops
is li
neat
a (C
.Pre
sl) H
olttu
mLo
mar
iops
is li
neat
a (C
.Pre
sl) H
olttu
m
Nep
hrol
epis
acu
tifol
ia (D
esv.
) Chr
ist
Nep
hrol
epis
acu
tifol
ia (D
esv.
) Chr
ist
Nep
hrol
epis
acu
tifol
ia (D
esv.
) Chr
ist
Nep
hrol
epis
bis
erra
ta (S
w.)
Scho
tt N
ephr
olep
is b
iser
rata
(Sw.
) Sch
ott
Nep
hrol
epis
bis
erra
ta (S
w.) S
chot
t
Nep
hrol
epis
cor
difo
lia (L
.) C
.Pre
sl
Nep
hrol
epis
cor
difo
lia (L
.) C
.Pre
slN
ephr
olep
is c
ordi
folia
(L.)
C.P
resl
Nep
hrol
epis
dav
allio
ides
(Sw.
) Kun
ze
Nep
hrol
epis
dav
allio
ides
(Sw.
) Kun
zeN
ephr
olep
is d
aval
lioid
es (S
w.) K
unze
●N
ephr
olep
is fa
lcifo
rmis
J.Sm
.16N
ephr
olep
is fa
lcat
a au
ct. n
on (C
av.)
C.C
hr.
Nep
hrol
epis
falc
ata
auct
. non
(Cav
.) C
.Chr
.
Nep
hrol
epis
hir
sutu
la (G
.For
st.)
C.P
resl
N
ephr
olep
is h
irsu
tula
(G.F
orst
.) C
.Pre
slN
ephr
olep
is h
irsu
tula
(G.F
orst
.) C
.Pre
sl
Nep
hrol
epis
radi
cans
(Bur
m.f.
) Kuh
n N
ephr
olep
is ra
dica
ns (B
urm
.f.) K
uhn
Nep
hrol
epis
radi
cans
(Bur
m.f.
) Kuh
n
83TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●N
ephr
olep
is u
ndul
ata
(afz
el.)
J.Sm
.16N
ephr
olep
is d
elic
atul
a (D
ecne
.) Pi
c.Se
rm.
Nep
hrol
epis
del
icat
ula
(Dec
ne.)
Pic.
Serm
.
LYG
OD
IAC
EA
E
Lygo
dium
circ
inat
um (B
urm
.f.) S
w.
Lygo
dium
circ
inat
um (B
urm
.f.) S
w.Ly
godi
um c
ircin
atum
(Bur
m.f.
) Sw.
Lygo
dium
flex
uosu
m (L
.) Sw
. Ly
godi
um fl
exuo
sum
(L.)
Sw.
Lygo
dium
flex
uosu
m (L
.) Sw
.
Lygo
dium
gig
ante
um T
agaw
a &
K.Iw
ats.
Lygo
dium
gig
ante
um T
agaw
a &
K.Iw
ats.
Lygo
dium
gig
ante
um T
agaw
a &
K.Iw
ats.
Lygo
dium
japo
nicu
m (T
hunb
.) Sw
. Ly
godi
um ja
poni
cum
(Thu
nb.)
Sw.
Lygo
dium
japo
nicu
m (T
hunb
.) Sw
.
Lygo
dium
mic
roph
yllu
m (C
av.)
R.B
r. Ly
godi
um m
icro
phyl
lum
(Cav
.) R
.Br.
Lygo
dium
mic
roph
yllu
m (C
av.)
R.B
r.
Lygo
dium
pol
ysta
chyu
m W
all.
ex T
.Moo
re
Lygo
dium
pol
ysta
chyu
m W
all.
ex T
.Moo
reLy
godi
um p
olys
tach
yum
Wal
l. ex
T.M
oore
Lygo
dium
salic
ifoliu
m C
.Pre
sl
Lygo
dium
salic
ifoliu
m C
.Pre
slLy
godi
um sa
licifo
lium
C.P
resl
MA
RAT
TIA
CE
AE
○An
giop
teri
s ang
ustif
olia
C.P
resl
10A
bsen
tA
bsen
t
Angi
opte
ris e
vect
a (G
.For
st.)
Hof
fm.
Angi
opte
ris e
vect
a (G
.For
st.)
Hof
fm.
Angi
opte
ris e
vect
a (G
.For
st.)
Hof
fm.
Chr
iste
nsen
ia a
escu
lifol
ia (B
lum
e) M
axon
C
hris
tens
enia
aes
culif
olia
(Blu
me)
Max
onC
hris
tens
enia
aes
culif
olia
(Blu
me)
Max
on
●Pt
isan
a sa
mbu
cina
(Blu
me)
Mur
dock
27M
arat
tia sa
mbu
cina
Blu
me
Mar
attia
sam
buci
na B
lum
e
MA
RSI
LE
AC
EA
E
Mar
sile
a cr
enat
a C
.Pre
sl
Mar
sile
a cr
enat
a C
.Pre
slM
arsi
lea
cren
ata
C.P
resl
○M
arsi
lea
scal
arip
es D
.M.Jo
hnso
n44A
bsen
tA
bsen
t
MAT
ON
IAC
EA
E
○M
aton
ia p
ectin
ata
R.B
r.23A
bsen
tA
bsen
t
OL
EA
ND
RA
CE
AE
Ole
andr
a m
usifo
lia (B
lum
e) C
.Pre
sl
Ole
andr
a m
usifo
lia (B
lum
e) C
.Pre
slO
lean
dra
mus
ifolia
(Blu
me)
C.P
resl
84 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Ole
andr
a pi
still
aris
(Sw.
) C.C
hr.
Ole
andr
a pi
still
aris
(Sw.
) C.C
hr.
Ole
andr
a pi
still
aris
(Sw.
) C.C
hr.
Ole
andr
a un
dula
ta (W
illd.
) Chi
ng
Ole
andr
a un
dula
ta (W
illd.
) Chi
ngO
lean
dra
undu
lata
(Will
d.) C
hing
Ole
andr
a w
allic
hii (
Hoo
k.) C
.Pre
sl
Ole
andr
a w
allic
hii (
Hoo
k.) C
.Pre
slO
lean
dra
wal
lichi
i (H
ook.
) C.P
resl
OPH
IOG
LO
SSA
CE
AE
Botr
ychi
um la
nugi
nosu
m W
all.
ex H
ook.
& G
rev.
Bo
tryc
hium
lanu
gino
sum
Wal
l. ex
Hoo
k. &
Gre
v.Bo
tryc
hium
lanu
gino
sum
Wal
l. ex
Hoo
k. &
Gre
v./ J
apan
obot
rych
ium
lanu
gino
sum
(W
all.
ex H
ook.
& G
rev.
) Nish
ida
ex T
agaw
a
Hel
min
thos
tach
ys ze
ylan
ica
(L.)
Hoo
k.
Hel
min
thos
tach
ys ze
ylan
ica
(L.)
Hoo
k.H
elm
inth
osta
chys
zeyl
anic
a (L
.) H
ook.
Oph
iogl
ossu
m c
osta
tum
R.B
r. O
phio
glos
sum
cos
tatu
m R
.Br.
Oph
iogl
ossu
m c
osta
tum
R.B
r.
Oph
iogl
ossu
m g
ram
ineu
m W
illd.
O
phio
glos
sum
gra
min
eum
Will
d.O
phio
glos
sum
gra
min
eum
Will
d.
Oph
iogl
ossu
m p
endu
lum
L.
Oph
iogl
ossu
m p
endu
lum
L.
Oph
iogl
ossu
m p
endu
lum
L.
Oph
iogl
ossu
m p
etio
latu
m H
ook.
O
phio
glos
sum
pet
iola
tum
Hoo
k.O
phio
glos
sum
pet
iola
tum
Hoo
k.
OSM
UN
DA
CE
AE
Osm
unda
ang
ustif
olia
Chi
ng
Osm
unda
ang
ustif
olia
Chi
ngO
smun
da a
ngus
tifol
ia C
hing
Osm
unda
cin
nam
omea
L.
Osm
unda
cin
nam
omea
L.
Osm
unda
cin
nam
omea
L.
○O
smun
da ja
vani
ca (C
.Pre
sl) B
lum
e4,10
Osm
unda
java
nica
(C.P
resl
) Blu
me
Abs
ent
Osm
unda
vac
helli
i Hoo
k.
Osm
unda
vac
helli
i Hoo
k.O
smun
da v
ache
llii H
ook.
PLA
GIO
GY
RIA
CE
AE
Plag
iogy
ria
adna
ta (B
lum
e) B
edd.
Pl
agio
gyri
a ad
nata
(Blu
me)
Bed
d.Pl
agio
gyri
a ad
nata
(Blu
me)
Bed
d.
Plag
iogy
ria
com
mun
is C
hing
Pl
agio
gyri
a co
mm
unis
Chi
ngPl
agio
gyri
a co
mm
unis
Chi
ng
POLY
POD
IAC
EA
E
●Ac
roso
rus f
ride
rici
-et-p
auli
(Chr
ist)
Cop
el.31
Ac
roso
rus t
rian
gula
ris (
Scor
t. ex
Bed
d.) C
opel
.Ac
roso
rus t
rian
gula
ris (
Scor
t. ex
Bed
d.) C
opel
.
85TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
○Ac
roso
rus s
trept
ophy
llus (
Bak
er) C
opel
.31A
bsen
tA
bsen
t
●Ag
laom
orph
a ac
umin
ata
(Will
d.) H
oven
kam
p Ph
otin
opte
ris s
peci
osa
(Blu
me)
C.P
resl
Phot
inop
teri
s acu
min
ata
(Will
d.) C
.V.M
orto
n
Agla
omor
pha
coro
nans
(Wal
l. ex
Met
t.) C
opel
. Ag
laom
orph
a co
rona
ns (W
all.
ex M
ett.)
Cop
el.
Agla
omor
pha
coro
nans
(Wal
l. ex
Met
t.) C
opel
.
○Ag
laom
orph
a dr
ynar
ioid
es (H
ook.
) Roo
s10A
bsen
tA
bsen
t
○Ag
laom
orph
a he
racl
ea (K
unze
) Cop
el.4
Agla
omor
pha
hera
clea
(Kun
ze) C
opel
.A
bsen
t
Arth
rom
eris
am
plex
ifolia
(Chr
ist)
Chi
ng
Arth
rom
eris
am
plex
ifolia
(Chr
ist)
Chi
ngAr
thro
mer
is a
mpl
exifo
lia (C
hris
t) C
hing
Arth
rom
eris
lehm
anni
(Met
t.) C
hing
Ar
thro
mer
is le
hman
ni (M
ett.)
Chi
ngAr
thro
mer
is le
hman
ni (M
ett.)
Chi
ng
Arth
rom
eris
phu
luan
gens
is T
agaw
a &
K.Iw
ats.
Arth
rom
eris
phu
luan
gens
is T
agaw
a &
K.Iw
ats.
Arth
rom
eris
phu
luan
gens
is T
agaw
a &
K.Iw
ats.
○Ar
thro
mer
is p
rote
us (C
opel
.) Ta
gaw
a16A
bsen
tA
bsen
t
Arth
rom
eris
tats
iene
nsis
(Fra
nch.
& B
urea
u ex
Chr
ist)
Chi
ng
Arth
rom
eris
tats
iene
nsis
(Fra
nch.
& B
urea
u ex
Chr
ist)
Chi
ngAr
thro
mer
is ta
tsie
nens
is (F
ranc
h. &
Bur
eau
ex C
hris
t) C
hing
Belv
isia
ann
amen
sis (
C.C
hr.)
Taga
wa
Belv
isia
ann
amen
sis (
C.C
hr.)
Taga
wa
Belv
isia
ann
amen
sis (
C.C
hr.)
Taga
wa
Belv
isia
hen
ryi (
Hie
ron.
ex
C.C
hr.)
Taga
wa
Belv
isia
hen
ryi (
Hie
ron.
ex
C.C
hr.)
Taga
wa
Belv
isia
hen
ryi (
Hie
ron.
ex
C.C
hr.)
Taga
wa
Belv
isia
muc
rona
ta (F
ée) C
opel
. Be
lvis
ia m
ucro
nata
(Fée
) Cop
el.
Belv
isia
muc
rona
ta (F
ée) C
opel
.
●Be
lvis
ia sp
icat
a (L
.f.) M
irbel
ex
Cop
el.16
Belv
isia
revo
luta
(Blu
me)
Cop
el.
Belv
isia
revo
luta
(Blu
me)
Cop
el.
Cal
ymm
odon
asi
atic
us C
opel
. C
alym
mod
on a
siat
icus
Cop
el.
Cal
ymm
odon
asi
atic
us C
opel
.
●C
alym
mod
on c
urtu
s Par
ris41
Cal
ymm
odon
cuc
ulla
tus a
uct.
non
(Nee
s & B
lum
e) C
.Pre
sl
Cal
ymm
odon
cuc
ulla
tus a
uct.
non
(Nee
s & B
lum
e) C
.Pre
sl
Chr
istio
pter
is tr
icus
pis (
Hoo
k.) C
hris
t C
hris
tiopt
eris
tric
uspi
s (H
ook.
) Chr
ist
Chr
istio
pter
is tr
icus
pis (
Hoo
k.) C
hris
t
○C
hrys
ogra
mm
itis m
usgr
avia
na (B
aker
) Par
ris32
Abs
ent
Abs
ent
●C
teno
pter
ella
ble
chno
ides
(Gre
v.) P
arris
33C
teno
pter
is m
oulto
nii (
Cop
el.)
C.C
hr. &
Tar
dieu
Cte
nopt
eris
mou
ltoni
i (C
opel
.) C
.Chr
. & T
ardi
eu
●C
teno
pter
ella
kha
olua
ngen
sis (
Taga
wa
& K
.Iwat
s.)
Parr
is 33
Xiph
opte
ris k
haol
uang
ensi
s Tag
awa
& K
.Iwat
s.Xi
phop
teri
s kha
olua
ngen
sis T
agaw
a &
K.Iw
ats.
○D
asyg
ram
miti
s bre
vive
nosa
(Ald
erw.
) Par
ris41
,44
Abs
ent
Abs
ent
●D
asyg
ram
miti
s mol
licom
a (N
ees &
Blu
me)
Par
ris33
Cte
nopt
eris
mol
licom
a (N
ees.
& B
lum
e) K
unze
Cte
nopt
eris
mol
licom
a (N
ees.
& B
lum
e) K
unze
Dry
nari
a bo
nii C
hris
t D
ryna
ria
boni
i Chr
ist
Dry
nari
a bo
nii C
hris
t
86 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●D
ryna
ria
roos
ii N
akai
ke28
Dry
nari
a fo
rtun
ei (K
unze
ex
Met
t.) J.
Sm.
Dry
nari
a fo
rtun
ei (K
unze
ex
Met
t.) J.
Sm.
Dry
nari
a pa
rish
ii (B
edd.
) Bed
d.
Dry
nari
a pa
rish
ii (B
edd.
) Bed
d.D
ryna
ria
pari
shii
(Bed
d.) B
edd.
Dry
nari
a pr
opin
qua
(Wal
l. ex
Met
t.) J.
Sm.
ex B
edd.
D
ryna
ria
prop
inqu
a (W
all.
ex M
ett.)
J.Sm
. ex
Bed
d.D
ryna
ria
prop
inqu
a (W
all.
ex M
ett.)
J.Sm
. ex
Bed
d.
Dry
nari
a qu
erci
folia
(L.)
J.Sm
. D
ryna
ria
quer
cifo
lia (L
.) J.S
m.
Dry
nari
a qu
erci
folia
(L.)
J.Sm
.
Dry
nari
a ri
gidu
la (S
w.) B
edd.
D
ryna
ria
rigi
dula
(Sw.
) Bed
d.D
ryna
ria
rigi
dula
(Sw.
) Bed
d.
Dry
nari
a sp
arsi
sora
(Des
v.) T
.Moo
re
Dry
nari
a sp
arsi
sora
(Des
v.) T
.Moo
reD
ryna
ria
spar
siso
ra (D
esv.
) T.M
oore
●G
onio
phle
bium
am
oenu
m (
Wal
l. ex
Met
t.) B
edd.
G
onio
phle
bium
am
oenu
m (W
all.
ex M
ett.)
Bed
d.Po
lypo
dium
am
oenu
m W
all.
ex M
ett.
●G
onio
phle
bium
lach
nopu
s J.S
m.36
Poly
podi
um g
arre
ttii C
.H.W
right
Poly
podi
um g
arre
ttii C
.H.W
right
●G
onio
phle
bium
man
mei
ense
(Chr
ist)
Röd
l-Lin
der
Poly
podi
um m
anm
eien
se C
hris
tPo
lypo
dium
man
mei
ense
Chr
ist
●G
onio
phle
bium
men
gtze
ense
(Chr
ist)
Röd
l-Lin
der7
Gon
ioph
lebi
um a
rgut
um J.
Sm. e
x H
ook.
& G
rev.
Poly
podi
um a
rgut
um (J
.Sm
. ex
Hoo
k. &
Gre
v.)
Hoo
k.
●G
onio
phle
bium
mic
rorh
izom
a (C
.B.C
lark
e ex
Bak
er)
Bed
d.G
onio
phle
bium
mic
rorh
izom
a (C
.B.C
lark
e ex
Bak
er)
Bed
d.Po
lypo
dium
mic
rorh
izom
a C
.B.C
lark
e ex
Bak
er
●G
onio
phle
bium
per
cuss
um (C
av.)
Wag
ner &
Gre
ther
16G
onio
phle
bium
ver
ruco
sum
(Hoo
k.) J
.Sm
.Po
lypo
dium
ver
ruco
sum
(Hoo
k.) M
ett.
●G
onio
phle
bium
per
sici
foliu
m (D
esv.
) Bed
d.
Gon
ioph
lebi
um p
ersi
cifo
lium
(Des
v.) B
edd.
Poly
podi
um p
ersi
cifo
lium
Des
v.
●G
onio
phle
bium
suba
uric
ulat
um (B
lum
e) C
.Pre
sl
Gon
ioph
lebi
um su
baur
icul
atum
(Blu
me)
C.P
resl
+
Gon
ioph
lebi
um m
olle
Bed
d.Po
lypo
dium
suba
uric
ulat
um B
lum
e +
Poly
podi
um b
eddo
mei
Bak
er
●G
ymno
gram
miti
s dar
eifo
rmis
(Hoo
k.) C
hing
ex
Tard
ieu
& C
.Chr
.G
ymno
gram
miti
s dar
eifo
rmis
(Hoo
k.) C
hing
ex
Tard
ieu
& C
.Chr
.Ar
aios
tegi
a da
reifo
rmis
(Hoo
k.) C
opel
. /
Gym
nogr
amm
itis d
arei
form
is (H
ook.
) C
hing
ex
Tard
ieu
& C
.Chr
.
●Le
cano
pter
is c
rust
acea
Cop
el.
Leca
nopt
eris
cru
stac
ea C
opel
.M
yrm
ecop
hila
cru
stac
ea (C
opel
.) Ta
gaw
a
○Le
cano
pter
is p
umila
Blu
me22
Abs
ent
Abs
ent
●Le
cano
pter
is si
nuos
a (W
all.
ex H
ook.
) Cop
el.
Leca
nopt
eris
sinu
osa
(Wal
l. ex
Hoo
k.) C
opel
.M
yrm
ecop
hila
sinu
osa
(Wal
l. ex
Hoo
k.) N
akai
ex
H.It
o
Lem
map
hyllu
m a
cced
ens (
Blu
me)
Don
k Le
mm
aphy
llum
acc
eden
s (B
lum
e) D
onk
Lem
map
hyllu
m a
cced
ens (
Blu
me)
Don
k
87TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Lem
map
hyllu
m c
arno
sum
(Hoo
k.) C
.Pre
sl
Lem
map
hyllu
m c
arno
sum
(Hoo
k.) C
.Pre
slLe
mm
aphy
llum
car
nosu
m (H
ook.
) C.P
resl
Lepi
soru
s bic
olor
Chi
ng
Lepi
soru
s bic
olor
Chi
ngLe
piso
rus b
icol
or C
hing
Lepi
soru
s con
tort
us (C
hris
t) C
hing
Le
piso
rus c
onto
rtus
(Chr
ist)
Chi
ngLe
piso
rus c
onto
rtus
(Chr
ist)
Chi
ng
Lepi
soru
s het
erol
epis
(Ros
enst
.) C
hing
Le
piso
rus h
eter
olep
is (R
osen
st.)
Chi
ngLe
piso
rus h
eter
olep
is (R
osen
st.)
Chi
ng
Lepi
soru
s lon
gifo
lius (
Blu
me)
Hol
ttum
Le
piso
rus l
ongi
foliu
s (B
lum
e) H
olttu
mLe
piso
rus l
ongi
foliu
s (B
lum
e) H
olttu
m
Lepi
soru
s nud
us (H
ook.
) Chi
ng
Lepi
soru
s nud
us (H
ook.
) Chi
ngLe
piso
rus n
udus
(Hoo
k.) C
hing
Lepi
soru
s oos
phae
rus (
C.C
hr.)
Chi
ng
Lepi
soru
s oos
phae
rus (
C.C
hr.)
Chi
ngLe
piso
rus o
osph
aeru
s (C
.Chr
.) C
hing
Lepi
soru
s sco
lope
ndri
um (C
hing
) M
ehra
& B
irLe
piso
rus s
colo
pend
rium
(Chi
ng)
Meh
ra &
Bir
Lepi
soru
s sco
lope
ndri
um (C
hing
) M
ehra
& B
ir
Lepi
soru
s sin
ensi
s (C
hris
t) C
hing
Le
piso
rus s
inen
sis (
Chr
ist)
Chi
ngLe
piso
rus s
inen
sis (
Chr
ist)
Chi
ng
Lepi
soru
s sub
confl
uens
Chi
ng
Lepi
soru
s sub
confl
uens
Chi
ngLe
piso
rus s
ubco
nflue
ns C
hing
Lepi
soru
s sub
linea
ris (
Bak
er e
x Ta
keda
) Chi
ng
Lepi
soru
s sub
linea
ris (
Bak
er e
x Ta
keda
) Chi
ngLe
piso
rus s
ublin
eari
s (B
aker
ex
Take
da) C
hing
Lepi
soru
s sub
olig
olep
idus
Chi
ng
Lepi
soru
s sub
olig
olep
idus
Chi
ngLe
piso
rus s
ubol
igol
epid
us C
hing
Lept
ochi
lus a
xilla
ris (
Cav
.) K
aulf.
Le
ptoc
hilu
s axi
llari
s (C
av.)
Kau
lf.Le
ptoc
hilu
s axi
llari
s (C
av.)
Kau
lf.
Lept
ochi
lus d
ecur
rens
Blu
me
Lept
ochi
lus d
ecur
rens
Blu
me
Lept
ochi
lus d
ecur
rens
Blu
me
●Le
ptoc
hilu
s elli
ptic
us (T
hunb
.) N
oot.29
Col
ysis
pen
taph
ylla
(Bak
er) C
hing
+ C
olys
is p
othi
folia
(D
.Don
) C.P
resl
Col
ysis
pen
taph
ylla
(Bak
er) C
hing
+
Col
ysis
pot
hifo
lia (D
.Don
) C.P
resl
●Le
ptoc
hilu
s hem
ioni
tideu
s (C
.Pre
sl) N
oot.
Col
ysis
hem
ioni
tidea
C.P
resl
Col
ysis
hem
ioni
tidea
C.P
resl
●Le
ptoc
hilu
s mac
roph
yllu
s (B
lum
e) N
oot.
C
olys
is m
acro
phyl
la (B
lum
e) C
.Pre
slC
olys
is m
acro
phyl
la (B
lum
e) C
.Pre
sl
○Le
ptoc
hilu
s min
or F
ée5
Abs
ent
Abs
ent
●Le
ptoc
hilu
s ped
uncu
latu
s (H
ook.
& G
rev.
) Fra
ser-J
enk.
C
olys
is p
edun
cula
ta (H
ook.
& G
rev.
) Chi
ng +
C
olys
is w
ui (C
.Chr
.) C
hing
Col
ysis
ped
uncu
lata
(Hoo
k. &
Gre
v.) C
hing
+
Col
ysis
wui
(C.C
hr.)
Chi
ng
○Lo
xogr
amm
e ac
rosc
opa
(Chr
ist)
C.C
hr.4
Loxo
gram
me
acro
scop
a (C
hris
t) C
.Chr
.A
bsen
t
○Lo
xogr
amm
e as
sim
ilis C
hing
4Lo
xogr
amm
e as
sim
ilis C
hing
Abs
ent
88 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Loxo
gram
me
aven
ia (B
lum
e) C
.Pre
sl
Loxo
gram
me
aven
ia (B
lum
e) C
.Pre
slLo
xogr
amm
e av
enia
(Blu
me)
C.P
resl
○Lo
xogr
amm
e ce
ntic
ola
M.G
.Pric
e4,8
Loxo
gram
me
cent
icol
a M
.G.P
rice
Abs
ent
Loxo
gram
me
chin
ensi
s Chi
ng
Loxo
gram
me
chin
ensi
s Chi
ngLo
xogr
amm
e ch
inen
sis C
hing
○Lo
xogr
amm
e cu
spid
ata
(Zen
ker)
M.G
.Pric
e4Lo
xogr
amm
e cu
spid
ata
(Zen
ker)
M.G
.Pric
e A
bsen
t
Loxo
gram
me
ducl
ouxi
i Chr
ist
Loxo
gram
me
ducl
ouxi
i Chr
ist
Loxo
gram
me
ducl
ouxi
i Chr
ist
Loxo
gram
me
invo
luta
(D.D
on) C
.Pre
sl
Loxo
gram
me
invo
luta
(D.D
on) C
.Pre
slLo
xogr
amm
e in
volu
ta (D
.Don
) C.P
resl
Loxo
gram
me
lank
okie
nsis
(Ros
enst
.) C
.Chr
. Lo
xogr
amm
e la
nkok
iens
is (R
osen
st.)
C.C
hr.
Loxo
gram
me
lank
okie
nsis
(Ros
enst
.) C
.Chr
.
○Lo
xogr
amm
e po
rcat
a M
.G.P
rice4
Loxo
gram
me
porc
ata
M.G
.Pric
eA
bsen
t
●Lo
xogr
amm
e sc
olop
endr
ioid
es (G
audi
ch.)
C.V
.Mor
ton34
Loxo
gram
me
scol
open
drin
a (B
ory)
C
.Pre
slLo
xogr
amm
e sc
olop
endr
ina
(Bor
y)
C.P
resl
Loxo
gram
me
sube
cost
ata
(Hoo
k.) C
.Chr
. Lo
xogr
amm
e su
beco
stat
a (H
ook.
) C.C
hr.
Loxo
gram
me
sube
cost
ata
(Hoo
k.) C
.Chr
.
Mic
roso
rum
het
eroc
arpu
m (B
lum
e) C
hing
M
icro
soru
m h
eter
ocar
pum
(Blu
me)
Chi
ngM
icro
soru
m h
eter
ocar
pum
(Blu
me)
Chi
ng
●M
icro
soru
m in
sign
e (B
lum
e) C
opel
.29M
icro
soru
m d
ilata
tum
(Bed
d.) S
ledg
eM
icro
soru
m d
ilata
tum
(Bed
d.) S
ledg
e
Mic
roso
rum
mem
bran
aceu
m (D
.Don
) C
hing
M
icro
soru
m m
embr
anac
eum
(D.D
on)
Chi
ngM
icro
soru
m m
embr
anac
eum
(D.D
on)
Chi
ng
○M
icro
soru
m m
usifo
lium
Cop
el.35
Abs
ent
Abs
ent
Mic
roso
rum
pte
ropu
s (B
lum
e) C
opel
. M
icro
soru
m p
tero
pus (
Blu
me)
Cop
el.
Mic
roso
rum
pte
ropu
s (B
lum
e) C
opel
.
Mic
roso
rum
pun
ctat
um (L
.) C
opel
. M
icro
soru
m p
unct
atum
(L.)
Cop
el.
Mic
roso
rum
pun
ctat
um (L
.) C
opel
.
○M
icro
soru
m si
amen
se B
oonk
erd1
Abs
ent
Abs
ent
Mic
roso
rum
supe
rfici
ale
(Blu
me)
Chi
ng
Mic
roso
rum
supe
rfici
ale
(Blu
me)
Chi
ngM
icro
soru
m su
perfi
cial
e (B
lum
e) C
hing
○M
icro
soru
m th
aila
ndic
um B
oonk
erd
& N
oot.3
Mic
roso
rum
sp.
Abs
ent
Mic
roso
rum
zipp
elii
(Blu
me)
Chi
ng
Mic
roso
rum
zipp
elii
(Blu
me)
Chi
ngM
icro
soru
m zi
ppel
ii (B
lum
e) C
hing
Neo
chei
ropt
eris
nor
mal
is (D
.Don
) Tag
awa
Neo
chei
ropt
eris
nor
mal
is (D
.Don
) Tag
awa
Neo
chei
ropt
eris
nor
mal
is (D
.Don
) Tag
awa
●O
reog
ram
miti
s ads
pers
a (B
lum
e) P
arris
33G
ram
miti
s ads
pers
a B
lum
eG
ram
miti
s ads
pers
a B
lum
e
●O
reog
ram
miti
s con
gene
r (B
lum
e) P
arris
41,3
3G
ram
miti
s set
osa
auct
. non
Blu
me
Gra
mm
itis s
etos
a au
ct. n
on B
lum
e
89TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●O
reog
ram
miti
s dor
sipi
la (C
hris
t) Pa
rris
33G
ram
miti
s dor
sipi
la (C
hris
t) C
.Chr
. &
Tard
ieu
Gra
mm
itis d
orsi
pila
(Chr
ist)
C.C
hr. &
Tard
ieu
●O
reog
ram
miti
s rei
nwar
dtii
(Blu
me)
Par
ris33
Gra
mm
itis b
ongo
ensi
s (C
opel
.) C
opel
. G
ram
miti
s bon
goen
sis (
Cop
el.)
Cop
el.
●Ph
ymat
osor
us c
uspi
datu
s (D
.Don
) Pic
.Ser
m.
Phym
atos
orus
cus
pida
tus (
D.D
on) P
ic.S
erm
.M
icro
soru
m c
uspi
datu
m (D
.Don
) Tag
awa
●Ph
ymat
osor
us lo
ngis
sim
us (B
lum
e) P
ic.S
erm
.Ph
ymat
osor
us lo
ngis
sim
us (B
lum
e) P
ic.S
erm
.M
icro
soru
m ru
bidu
m (K
unze
) Cop
el.
●Ph
ymat
osor
us m
embr
anifo
lius (
R.B
r.) S
.G.L
u25Ph
ymat
osor
us n
igre
scen
s (B
lum
e) P
ic.S
erm
.M
icro
soru
m n
igre
scen
s (B
lum
e) C
opel
.
●Ph
ymat
osor
us sc
olop
endr
ia (B
urm
.f.) P
ic.S
erm
.Ph
ymat
osor
us sc
olop
endr
ia (B
urm
.f.) P
ic.S
erm
.M
icro
soru
m sc
olop
endr
ia (B
urm
.f.) C
opel
.
Plat
ycer
ium
cor
onar
ium
(J.K
oeni
g) D
esv.
Pl
atyc
eriu
m c
oron
ariu
m (J
.Koe
nig)
Des
v.Pl
atyc
eriu
m c
oron
ariu
m (J
.Koe
nig)
Des
v.
Plat
ycer
ium
hol
ttum
ii de
Jonc
h. &
Hen
nipm
an
Plat
ycer
ium
hol
ttum
ii de
Jonc
h. &
Hen
nipm
anPl
atyc
eriu
m h
olttu
mii
de Jo
nch.
& H
enni
pman
○Pl
atyc
eriu
m ri
dley
i Chr
ist7
Plat
ycer
ium
ridl
eyi C
hris
tA
bsen
t
Plat
ycer
ium
wal
lichi
i Hoo
k.
Plat
ycer
ium
wal
lichi
i Hoo
k.Pl
atyc
eriu
m w
allic
hii H
ook.
Pros
aptia
ala
ta (B
lum
e) C
hris
t Pr
osap
tia a
lata
(Blu
me)
Chr
ist
Pros
aptia
ala
ta (B
lum
e) C
hris
t
●Pr
osap
tia b
arat
hrop
hylla
(Bak
er) M
.G.P
rice41
Pr
osap
tia k
hasy
ana
auct
. non
(Hoo
k.) C
.Chr
. &
Tard
ieu
Pros
aptia
kha
syan
a au
ct. n
on (H
ook.
) C.C
hr. &
Ta
rdie
u
Pros
aptia
cel
ebic
a (B
lum
e) T
agaw
a &
K
.Iwat
s. Pr
osap
tia c
eleb
ica
(Blu
me)
Tag
awa
&
K.Iw
ats.
Pros
aptia
cel
ebic
a (B
lum
e) T
agaw
a &
K
.Iwat
s.
Pros
aptia
con
tigua
(G.F
orst
.) C
.Pre
sl
Pros
aptia
con
tigua
(G.F
orst
.) C
.Pre
slPr
osap
tia c
ontig
ua (G
.For
st.)
C.P
resl
Pros
aptia
obl
iqua
ta (B
lum
e) M
ett.
Pros
aptia
obl
iqua
ta (B
lum
e) M
ett.
Pros
aptia
obl
iqua
ta (B
lum
e) M
ett.
●Pr
osap
tia p
ectin
ata
T.M
oore
41Pr
osap
tia le
ysii
(Bak
er) C
hing
Pros
aptia
leys
ii (B
aker
) Chi
ng
●Py
rros
ia a
lbic
ans (
Blu
me)
Chi
ng15
Pyrr
osia
floc
cige
ra (B
lum
e) C
hing
Pyrr
osia
floc
cige
ra (B
lum
e) C
hing
Pyrr
osia
ang
usta
ta (S
w.) C
hing
Py
rros
ia a
ngus
tata
(Sw.
) Chi
ngPy
rros
ia a
ngus
tata
(Sw.
) Chi
ng
Pyrr
osia
ang
ustis
sim
a (G
iese
nh. e
x D
iels
) Ta
gaw
a &
K.Iw
ats.
Pyrr
osia
ang
ustis
sim
a (G
iese
nh. e
x D
iels
) Ta
gaw
a &
K.Iw
ats.
Pyrr
osia
ang
ustis
sim
a (G
iese
nh. e
x D
iels
) Ta
gaw
a &
K.Iw
ats.
Pyrr
osia
cos
tata
(C.P
resl
ex
Bed
d.)
Taga
wa
& K
.Iwat
s. Py
rros
ia c
osta
ta (C
.Pre
sl e
x B
edd.
) Ta
gaw
a &
K.Iw
ats.
Pyrr
osia
cos
tata
(C.P
resl
ex
Bed
d.)
Taga
wa
& K
.Iwat
s.
Pyrr
osia
floc
culo
sa (D
.Don
) Chi
ng
Pyrr
osia
floc
culo
sa (D
.Don
) Chi
ngPy
rros
ia fl
occu
losa
(D.D
on) C
hing
90 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●Py
rros
ia la
nceo
lata
(L.)
Farw
.15Py
rros
ia la
nceo
lata
(L.)
Farw
. +Py
rros
ia a
dnas
cens
(Sw.
) Chi
ng +
Py
rros
ia n
uda
(Gie
senh
.) C
hing
+
Pyrr
osia
var
ia (K
aulf.
) Far
w.
Pyrr
osia
lanc
eola
ta (L
.) Fa
rw. +
Pyrr
osia
adn
asce
ns (S
w.) C
hing
+
Pyrr
osia
nud
a (G
iese
nh.)
Chi
ng +
Py
rros
ia v
aria
(Kau
lf.) F
arw.
○Py
rros
ia li
ngua
(Thu
nb.)
Farw
. var
. lin
gua4
Pyrr
osia
ling
ua (T
hunb
.) Fa
rw. v
ar. l
ingu
aA
bsen
t
●Py
rros
ia li
ngua
(Thu
nb.)
Farw
. var
. het
erac
tis (M
ett.
ex
Kuh
n) H
oven
kam
pPy
rros
ia li
ngua
(Thu
nb.)
Farw
. var
. het
erac
tis (M
ett.
ex
Kuh
n) H
oven
kam
pPy
rros
ia e
berh
ardt
ii (C
hris
t) C
hing
+
Pyrr
osia
het
erac
tis (M
ett.
ex K
uhn)
Chi
ng
var.
hete
ract
is +
Pyr
rosi
a he
tera
ctis
(Met
t. ex
K
uhn)
Chi
ng v
ar. m
inor
(C.C
hr.)
Chi
ng
Pyrr
osia
long
ifolia
(Bur
m.f.
) C.V
.Mor
ton
Pyrr
osia
long
ifolia
(Bur
m.f.
) C.V
.Mor
ton
Pyrr
osia
long
ifolia
(Bur
m.f.
) C.V
.Mor
ton
Pyrr
osia
man
nii (
Gie
senh
.) C
hing
Py
rros
ia m
anni
i (G
iese
nh.)
Chi
ngPy
rros
ia m
anni
i (G
iese
nh.)
Chi
ng
Pyrr
osia
num
mul
arifo
lia (S
w.)
Chi
ng
Pyrr
osia
num
mul
arifo
lia (S
w.) C
hing
Pyrr
osia
num
mul
arifo
lia (S
w.) C
hing
●Py
rros
ia p
enan
gian
a (H
ook.
) Hol
ttum
Py
rros
ia p
enan
gian
a (H
ook.
) Hol
ttum
+
Pyrr
osia
mol
lis (K
unze
) Chi
ngPy
rros
ia p
enan
gian
a (H
ook.
) Hol
ttum
+
Pyrr
osia
mol
lis (K
unze
) Chi
ng
●Py
rros
ia p
ilose
lloid
es (L
.) M
.G.P
rice
Pyrr
osia
pilo
sello
ides
(L.)
M.G
.Pric
eD
rym
oglo
ssum
pilo
sello
ides
(L.)
C.P
resl
○Py
rros
ia p
oros
a (C
.Pre
sl) H
oven
kam
p va
r. po
rosa
23A
bsen
tA
bsen
t
●Py
rros
ia p
oros
a (C
.Pre
sl) H
oven
kam
p va
r. to
nkin
ensi
s (G
iese
nh.)
Chi
ng
Pyrr
osia
tonk
inen
sis (
Gie
senh
.) C
hing
Pyrr
osia
tonk
inen
sis (
Gie
senh
.) C
hing
○Py
rros
ia ra
sam
alae
(Rac
ib.)
K.H
. Shi
ng10
,15
Abs
ent
Abs
ent
Pyrr
osia
stig
mos
a (S
w.) C
hing
Py
rros
ia st
igm
osa
(Sw.
) Chi
ngPy
rros
ia st
igm
osa
(Sw.
) Chi
ng
●Ra
diog
ram
miti
s jag
oria
na (M
ett.
ex K
uhn)
Par
ris33
Gra
mm
itis j
agor
iana
(Met
t.) T
agaw
aG
ram
miti
s jag
oria
na (M
ett.)
Tag
awa
●Ra
diog
ram
miti
s mul
tifol
ia (C
opel
.) Pa
rris
41,3
3G
ram
miti
s hir
tella
auc
t. no
n (B
lum
e)
Tuya
ma
Gra
mm
itis h
irte
lla a
uct.
non
(Blu
me)
Tu
yam
a
●Sc
lero
glos
sum
pus
illum
(Blu
me)
Ald
erw.
41Sc
lero
glos
sum
min
us a
uct.
non
(Fée
) C.C
hr.
Scle
rogl
ossu
m m
inus
auc
t. no
n (F
ée) C
.Chr
.
●Sc
lero
glos
sum
sulc
atum
(Kuh
n) A
lder
w.41
Sc
lero
glos
sum
pus
illum
auc
t. no
n (B
lum
e)
Ald
erw.
Scle
rogl
ossu
m p
usill
um a
uct.
non
(Blu
me)
A
lder
w.
●Se
lligu
ea c
ruci
form
is (C
hing
) Fra
ser-J
enk.
Cry
psin
us c
ruci
form
is (C
hing
) Tag
awa
Cry
psin
us c
ruci
form
is (C
hing
) Tag
awa
91TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●Se
lligu
ea e
beni
pes (
Hoo
k.) S
.Lin
ds.
Cry
psin
us e
beni
pes (
Hoo
k.) C
opel
.C
ryps
inus
ebe
nipe
s (H
ook.
) Cop
el.
●Se
lligu
ea e
nerv
is (C
av.)
Chi
ng
Cry
psin
us e
nerv
is (C
av.)
Cop
el.
Cry
psin
us e
nerv
is (C
av.)
Cop
el.
●Se
lligu
ea g
riffi
thia
na (H
ook.
) Fra
ser-J
enk.
Cry
psin
us g
riffi
thia
nus (
Hoo
k.) C
opel
. C
ryps
inus
gri
ffith
ianu
s (H
ook.
) Cop
el.
Selli
guea
het
eroc
arpa
Blu
me
Selli
guea
het
eroc
arpa
Blu
me
Selli
guea
het
eroc
arpa
Blu
me
●Se
lligu
ea h
irsu
ta (T
agaw
a &
K.Iw
ats.)
S.L
inds
. C
ryps
inus
hir
sutu
s Tag
awa
& K
.Iwat
s. C
ryps
inus
hir
sutu
s Tag
awa
& K
.Iwat
s.
●Se
lligu
ea la
cini
ata
(C.P
resl
) Par
risC
ryps
inus
laci
niat
us (C
.Pre
sl) H
olttu
mC
ryps
inus
laci
niat
us (C
.Pre
sl) H
olttu
m
○Se
lligu
ea la
teri
tia (B
aker
) Hov
enka
mp16
Abs
ent
Abs
ent
●Se
lligu
ea o
xylo
ba (W
all.
ex K
unze
) Fra
ser-J
enk.
Cry
psin
us o
xylo
bus (
Wal
l. ex
Kun
ze)
Sled
geC
ryps
inus
oxy
lobu
s (W
all.
ex K
unze
) Sl
edge
●Se
lligu
ea rh
ynch
ophy
lla (H
ook.
) Fra
ser-J
enk.
Cry
psin
us rh
ynch
ophy
llus (
Hoo
k.) C
opel
.C
ryps
inus
rhyn
chop
hyllu
s (H
ook.
) Cop
el.
●Se
lligu
ea st
enop
hylla
(Blu
me)
Par
risC
ryps
inus
sten
ophy
llus (
Blu
me)
Hol
ttum
C
ryps
inus
sten
ophy
llus (
Blu
me)
Hol
ttum
●Se
lligu
ea tr
iloba
(Hou
tt.) M
.G.P
rice
Cry
psin
us tr
ilobu
s (H
outt.
) Cop
el.
Cry
psin
us tr
ilobu
s (H
outt.
) Cop
el.
●Th
emel
ium
tenu
isec
tum
(Blu
me)
Par
ris
Cte
nopt
eris
tenu
isec
ta (B
lum
e) J.
Sm.
Cte
nopt
eris
tenu
isec
ta (B
lum
e) J.
Sm.
●To
mop
hyllu
m su
bfal
catu
m (B
lum
e) P
arris
33C
teno
pter
is su
bfal
cata
(Blu
me)
Kun
zeC
teno
pter
is su
bfal
cata
(Blu
me)
Kun
ze
●Xi
phop
tere
lla h
iero
nym
usii
(C.C
hr.)
Parr
is33
Xiph
opte
ris h
iero
nym
usii
(C.C
hr.)
Hol
ttum
Xi
phop
teri
s hie
rony
mus
ii (C
.Chr
.) H
olttu
m
PSIL
OTA
CE
AE
Psilo
tum
com
plan
atum
Sw.
Ps
ilotu
m c
ompl
anat
um S
w.Ps
ilotu
m c
ompl
anat
um S
w.
Psilo
tum
nud
um (L
.) B
eauv
. Ps
ilotu
m n
udum
(L.)
Bea
uv.
Psilo
tum
nud
um (L
.) B
eauv
.
PTE
RID
AC
EA
E su
bfam
. Che
ilant
hoid
eae
○C
heila
nthe
s arg
ente
a (S
.G.G
mel
) Kun
ze4
Che
ilant
hes a
rgen
tea
(S.G
.Gm
el) K
unze
Abs
ent
Che
ilant
hes b
elan
geri
(Bor
y) C
.Chr
. C
heila
nthe
s bel
ange
ri (B
ory)
C.C
hr.
Che
ilant
hes b
elan
geri
(Bor
y) C
.Chr
.
Che
ilant
hes d
elic
atul
a Ta
gaw
a &
K.Iw
ats.
Che
ilant
hes d
elic
atul
a Ta
gaw
a &
K.Iw
ats.
Che
ilant
hes d
elic
atul
a Ta
gaw
a &
K.Iw
ats.
92 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●C
heila
nthe
s for
mos
ana
Hay
ata
Che
ilant
hes f
orm
osan
a H
ayat
aC
heila
nthe
s far
inos
a au
ct. n
on (F
orss
k.) K
aulf.
, pr
o pa
rte /
Che
ilant
hes f
orm
osan
a H
ayat
a
Che
ilant
hes f
ragi
lis H
ook.
C
heila
nthe
s fra
gilis
Hoo
k.C
heila
nthe
s fra
gilis
Hoo
k.
●C
heila
nthe
s kra
mer
i Fra
nch.
& S
av.
Che
ilant
hes k
ram
eri F
ranc
h. &
Sav
.C
heila
nthe
s far
inos
a au
ct. n
on (F
orss
k.) K
aulf.
, pr
o pa
rte /
Che
ilant
hes k
ram
eri F
ranc
h. &
Sav
.
●C
heila
nthe
s pse
udoa
rgen
tea
(S.K
.Wu)
K.Iw
ats.
Che
ilant
hes p
seud
oarg
ente
a (S
.K.W
u) K
.Iwat
s.C
heila
nthe
s far
inos
a au
ct. n
on (F
orss
k.)
Kau
lf., p
ro p
arte
/ C
heila
nthe
s pse
udoa
rgen
tea
(S.K
.Wu)
K.Iw
ats.
●C
heila
nthe
s pse
udof
arin
osa
(Chi
ng &
S.K
.Wu)
K.Iw
ats.
Che
ilant
hes p
seud
ofar
inos
a (C
hing
& S
.K.W
u) K
.Iwat
s.C
heila
nthe
s far
inos
a au
ct. n
on (F
orss
k.) K
aulf.
, pr
o pa
rte /
Che
ilant
hes p
seud
ofar
inos
a (C
hing
&
S.K
.Wu)
K.Iw
ats.
●C
heila
nthe
s ruf
a D
.Don
C
heila
nthe
s ruf
a D
.Don
Che
ilant
hes r
ufa
D.D
on +
Che
ilant
hes s
ubru
fa
auct
. non
Bak
er
●C
heila
nthe
s sia
men
sis (
S.K
.Wu)
K.Iw
ats.
Che
ilant
hes s
iam
ensi
s (S.
K.W
u) K
.Iwat
s.C
heila
nthe
s far
inos
a au
ct. n
on (F
orss
k.) K
aulf.
, pr
o pa
rte /
Che
ilant
hes s
iam
ensi
s (S.
K.W
u)
K.Iw
ats.
Che
ilant
hes t
enui
folia
(Bur
m.f.
) Sw.
C
heila
nthe
s ten
uifo
lia (B
urm
.f.) S
w.C
heila
nthe
s ten
uifo
lia (B
urm
.f.) S
w.
○D
oryo
pter
is a
lleni
ae r
.M.T
ryon
10A
bsen
tA
bsen
t
Dor
yopt
eris
lude
ns (W
all.
ex H
ook.
) J.S
m.
Dor
yopt
eris
lude
ns (W
all.
ex H
ook.
) J.S
m.
Dor
yopt
eris
lude
ns (W
all.
ex H
ook.
) J.S
m.
●H
emio
nitis
ves
tita
(Wal
l. ex
Hoo
k.) J
.Sm
. G
ymno
pter
is v
estit
a (W
all.
ex H
ook.
) Und
erw.
Gym
nopt
eris
ves
tita
(Wal
l. ex
Hoo
k.) U
nder
w.
Not
hola
ena
velu
tina
Tard
ieu
& C
.Chr
. N
otho
laen
a ve
lutin
a Ta
rdie
u &
C.C
hr.
Not
hola
ena
velu
tina
Tard
ieu
& C
.Chr
.
●Pa
rahe
mio
nitis
cor
data
(Rox
b. e
x H
ook.
& G
rev.
) Fr
aser
-Jen
k.11
Hem
ioni
tis a
rifo
lia (B
urm
.f.) T
.Moo
reH
emio
nitis
ari
folia
(Bur
m.f.
) T.M
oore
○Pe
llaea
tim
oren
sis A
lder
w.4
Pella
ea ti
mor
ensi
s Ald
erw.
Abs
ent
PTE
RID
AC
EA
E su
bfam
. Cry
ptog
ram
moi
deae
Con
iogr
amm
e fr
axin
ea (D
.Don
) Die
ls
var.
serr
ulat
a (B
lum
e) H
iero
n.
Con
iogr
amm
e fr
axin
ea (D
.Don
) Die
ls
var.
serr
ulat
a (B
lum
e) H
iero
n.C
onio
gram
me
frax
inea
(D.D
on) D
iels
va
r. se
rrul
ata
(Blu
me)
Hie
ron.
93TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Con
iogr
amm
e pe
telo
tii T
ardi
eu
Con
iogr
amm
e pe
telo
tii T
ardi
euC
onio
gram
me
pete
lotii
Tar
dieu
Con
iogr
amm
e pr
ocer
a Fé
e C
onio
gram
me
proc
era
Fée
Con
iogr
amm
e pr
ocer
a Fé
e
PTE
RID
AC
EA
E su
bfam
. Par
keri
oide
ae
Acro
stic
hum
aur
eum
L.
Acro
stic
hum
aur
eum
L.
Acro
stic
hum
aur
eum
L.
Acro
stic
hum
spec
iosu
m W
illd.
Ac
rost
ichu
m sp
ecio
sum
Will
d.Ac
rost
ichu
m sp
ecio
sum
Will
d.
Cer
atop
teri
s tha
lictro
ides
(L.)
Bro
ngn.
C
erat
opte
ris t
halic
troid
es (L
.) B
rong
n.C
erat
opte
ris t
halic
troid
es (L
.) B
rong
n.
PTE
RID
AC
EA
E su
bfam
. Pte
rido
idea
e
Ony
chiu
m c
ontig
uum
C.H
ope
Ony
chiu
m c
ontig
uum
C.H
ope
Ony
chiu
m c
ontig
uum
C.H
ope
Ony
chiu
m si
licul
osum
(Des
v.) C
.Chr
. O
nych
ium
silic
ulos
um (D
esv.
) C.C
hr.
Ony
chiu
m si
licul
osum
(Des
v.) C
.Chr
.
Pity
rogr
amm
a ca
lom
elan
os (L
.) Li
nk
Pity
rogr
amm
a ca
lom
elan
os (L
.) Li
nkPi
tyro
gram
ma
calo
mel
anos
(L.)
Link
Pter
is a
sper
icau
lis W
all.
ex J.
aga
rdh
Pter
is a
sper
icau
lis W
all.
ex J.
aga
rdh
Pter
is a
sper
icau
lis W
all.
ex J.
aga
rdh
Pter
is a
sper
ula
J.Sm
. ex
Hie
ron.
Pt
eris
asp
erul
a J.S
m. e
x H
iero
n.Pt
eris
asp
erul
a J.S
m. e
x H
iero
n.
Pter
is b
ella
Tag
awa
Pter
is b
ella
Tag
awa
Pter
is b
ella
Tag
awa
Pter
is b
iaur
ita L
. Pt
eris
bia
urita
L.
Pter
is b
iaur
ita L
.
Pter
is b
lum
eana
J.a
gard
h Pt
eris
blu
mea
na J.
aga
rdh
Pter
is b
lum
eana
J.a
gard
h
Pter
is c
retic
a L.
Pt
eris
cre
tica
L.Pt
eris
cre
tica
L.
Pter
is d
alho
usia
e H
ook.
Pt
eris
dal
hous
iae
Hoo
k.Pt
eris
dal
hous
iae
Hoo
k.
Pter
is d
ecre
scen
s Chr
ist
Pter
is d
ecre
scen
s Chr
ist
Pter
is d
ecre
scen
s Chr
ist
Pter
is e
nsifo
rmis
Bur
m.f.
Pt
eris
ens
iform
is B
urm
.f.Pt
eris
ens
iform
is B
urm
.f.
Pter
is g
revi
llean
a W
all.
ex J.
aga
rdh
Pter
is g
revi
llean
a W
all.
ex J.
aga
rdh
Pter
is g
revi
llean
a W
all.
ex J.
aga
rdh
Pter
is h
eter
omor
pha
Fée
Pter
is h
eter
omor
pha
Fée
Pter
is h
eter
omor
pha
Fée
Pter
is li
near
is P
oir.
Pter
is li
near
is P
oir.
Pter
is li
near
is P
oir.
94 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Pter
is lo
ngip
es D
.Don
Pt
eris
long
ipes
D.D
onPt
eris
long
ipes
D.D
on
Pter
is lo
ngip
innu
la W
all.
ex J.
aga
rdh
Pter
is lo
ngip
innu
la W
all.
ex J.
aga
rdh
Pter
is lo
ngip
innu
la W
all.
ex J.
aga
rdh
Pter
is m
erte
nsio
ides
Will
d.
Pter
is m
erte
nsio
ides
Will
d.Pt
eris
mer
tens
ioid
es W
illd.
Pter
is m
ultifi
da P
oir.
Pter
is m
ultifi
da P
oir.
Pter
is m
ultifi
da P
oir.
Pter
is n
epal
ensi
s H.It
o Pt
eris
nep
alen
sis H
.Ito
Pter
is n
epal
ensi
s H.It
o
Pter
is p
hulu
ange
nsis
Tag
awa
& K
.Iwat
s. Pt
eris
phu
luan
gens
is T
agaw
a &
K.Iw
ats.
Pter
is p
hulu
ange
nsis
Tag
awa
& K
.Iwat
s.
Pter
is p
lum
bea
Chr
ist
Pter
is p
lum
bea
Chr
ist
Pter
is p
lum
bea
Chr
ist
Pter
is sc
abri
pes W
all.
ex J.
aga
rdh
Pter
is sc
abri
pes W
all.
ex J.
aga
rdh
Pter
is sc
abri
pes W
all.
ex J.
aga
rdh
Pter
is se
mip
inna
ta L
. Pt
eris
sem
ipin
nata
L.
Pter
is se
mip
inna
ta L
.
Pter
is st
enop
hylla
Wal
l. ex
Hoo
k. &
Gre
v.
Pter
is st
enop
hylla
Wal
l. ex
Hoo
k. &
Gre
v.Pt
eris
sten
ophy
lla W
all.
ex H
ook.
& G
rev.
Pter
is su
bqui
nata
Wal
l. ex
J.a
gard
h Pt
eris
subq
uina
ta W
all.
ex J.
aga
rdh
Pter
is su
bqui
nata
Wal
l. ex
J.a
gard
h
Pter
is to
kioi
Mas
am.
Pter
is to
kioi
Mas
am.
Pter
is to
kioi
Mas
am.
Pter
is tr
ipar
tita
Sw.
Pter
is tr
ipar
tita
Sw.
Pter
is tr
ipar
tita
Sw.
Pter
is v
enus
ta K
unze
Pt
eris
ven
usta
Kun
zePt
eris
ven
usta
Kun
ze
Pter
is v
ittat
a L.
Pt
eris
vitt
ata
L.Pt
eris
vitt
ata
L.
Pter
is w
allic
hian
a J.a
gard
h Pt
eris
wal
lichi
ana
J.aga
rdh
Pter
is w
allic
hian
a J.a
gard
h
Syng
ram
ma
alis
mifo
lia (C
.Pre
sl) J
.Sm
. Sy
ngra
mm
a al
ism
ifolia
(C.P
resl
) J.S
m.
Syng
ram
ma
alis
mifo
lia (C
.Pre
sl) J
.Sm
.
Taen
itis b
lech
noid
es (W
illd.
) Sw
. Ta
eniti
s ble
chno
ides
(Will
d.) S
w.Ta
eniti
s ble
chno
ides
(Will
d.) S
w.
○Ta
eniti
s int
erru
pta
Hoo
k. &
Gre
v.10
Abs
ent
Abs
ent
PTE
RID
AC
EA
E su
bfam
. Vitt
ario
idea
e
Adia
ntum
cap
illus
-ven
eris
L.
Adia
ntum
cap
illus
-ven
eris
L.
Adia
ntum
cap
illus
-ven
eris
L.
Adia
ntum
cau
datu
m L
. Ad
iant
um c
auda
tum
L.
Adia
ntum
cau
datu
m L
.
Adia
ntum
edg
ewor
thii
Hoo
k.
Adia
ntum
edg
ewor
thii
Hoo
k.Ad
iant
um e
dgew
orth
ii H
ook.
95TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Adia
ntum
ery
lliae
C.C
hr. &
Tar
dieu
Ad
iant
um e
rylli
ae C
.Chr
. & T
ardi
euAd
iant
um e
rylli
ae C
.Chr
. & T
ardi
eu
Adia
ntum
flab
ellu
latu
m L
. Ad
iant
um fl
abel
lula
tum
L.
Adia
ntum
flab
ellu
latu
m L
.
●Ad
iant
um fr
agili
form
e C
.Chr
.46Ad
iant
um st
enoc
hlam
ys B
aker
, pro
par
teAd
iant
um st
enoc
hlam
ys B
aker
, pro
par
te
○Ad
iant
um la
tifol
ium
Lam
.10A
bsen
tA
bsen
t
○Ad
iant
um p
hano
men
sis S
.Lin
ds. &
D.J.
Mid
dlet
on21
Abs
ent
Abs
ent
Adia
ntum
phi
lippe
nse
L.43
Adia
ntum
phi
lippe
nse
L.Ad
iant
um p
hilip
pens
e L.
Adia
ntum
siam
ense
Tag
awa
& K
.Iwat
s. Ad
iant
um si
amen
se T
agaw
a &
K.Iw
ats.
Adia
ntum
siam
ense
Tag
awa
& K
.Iwat
s.
Adia
ntum
sobo
lifer
um W
all.
ex H
ook.
Ad
iant
um so
bolif
erum
Wal
l. ex
Hoo
k.Ad
iant
um so
bolif
erum
Wal
l. ex
Hoo
k.
Adia
ntum
sten
ochl
amys
Bak
er
Adia
ntum
sten
ochl
amys
Bak
er, p
ro p
arte
Adia
ntum
sten
ochl
amys
Bak
er, p
ro p
arte
○Ad
iant
um th
ongt
ham
ii Su
ksat
han37
Abs
ent
Abs
ent
Adia
ntum
zolli
nger
i Met
t. ex
Kuh
n Ad
iant
um zo
lling
eri M
ett.
ex K
uhn
Adia
ntum
zolli
nger
i Met
t. ex
Kuh
n
Antro
phyu
m c
allif
oliu
m B
lum
e An
troph
yum
cal
lifol
ium
Blu
me
Antro
phyu
m c
allif
oliu
m B
lum
e
Antro
phyu
m o
bova
tum
Bak
er
Antro
phyu
m o
bova
tum
Bak
erAn
troph
yum
obo
vatu
m B
aker
Antro
phyu
m p
arvu
lum
Blu
me
Antro
phyu
m p
arvu
lum
Blu
me
Antro
phyu
m p
arvu
lum
Blu
me
●An
troph
yum
vitt
ario
ides
Bak
er39
Antro
phyu
m st
enop
hyllu
m B
aker
Antro
phyu
m st
enop
hyllu
m B
aker
●H
aplo
pter
is a
mbo
inen
sis (
Fée)
X.C
.Zha
ng40
Vitta
ria
ambo
inen
sis F
ée +
Vitt
aria
fo
rres
tiana
auc
t. no
n C
hing
Vitta
ria
ambo
inen
sis F
ée +
Vitt
aria
fo
rres
tiana
auc
t. no
n C
hing
●H
aplo
pter
is a
ngus
tifol
ia (B
lum
e) E
.H.C
rane
Vi
ttari
a an
gust
ifolia
Blu
me
Vitta
ria
angu
stifo
lia B
lum
e
●H
aplo
pter
is e
long
ata
(Sw
.) E.
H.C
rane
Vi
ttari
a el
onga
ta S
w.Vi
ttari
a el
onga
ta S
w.
●H
aplo
pter
is e
nsifo
rmis
(Sw.
) E.H
.Cra
ne
Vitta
ria
ensi
form
is S
w.Vi
ttari
a en
sifo
rmis
Sw.
●H
aplo
pter
is fl
exuo
sa (F
ée) E
.H.C
rane
Vi
ttari
a fle
xuos
a Fé
eVi
ttari
a fle
xuos
a Fé
e
●H
aplo
pter
is sc
olop
endr
ina
C.P
resl
Vi
ttari
a sc
olop
endr
ina
(Bor
y) T
hwai
tes &
H
ook.
Vitta
ria
scol
open
drin
a (B
ory)
Thw
aite
s &
Hoo
k.
●H
aplo
pter
is si
kkim
ensi
s (K
uhn)
E.H
.Cra
ne
Vitta
ria
sikk
imen
sis K
uhn
Vitta
ria
sikk
imen
sis K
uhn
●H
aplo
pter
is ta
enio
phyl
la (C
opel
.) E.
H.C
rane
Vi
ttari
a ta
enio
phyl
la C
opel
.Vi
ttari
a ta
enio
phyl
la C
opel
.
96 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●H
aplo
pter
is w
initi
i (Ta
gaw
a &
K.Iw
ats.)
S.L
inds
. An
troph
yum
win
itii T
agaw
a &
K.Iw
ats.
Antro
phyu
m w
initi
i Tag
awa
& K
.Iwat
s.
Vagi
nula
ria
para
doxa
(Fée
) Met
t. Va
ginu
lari
a pa
rado
xa (F
ée) M
ett.
Vagi
nula
ria
para
doxa
(Fée
) Met
t.
Vagi
nula
ria
tric
hoid
ea F
ée
Vagi
nula
ria
tric
hoid
ea F
éeVa
ginu
lari
a tr
icho
idea
Fée
SALV
INIA
CE
AE
Azol
la c
arol
inia
na W
illd.
Az
olla
car
olin
iana
Will
d.Az
olla
car
olin
iana
Will
d.
Azol
la p
inna
ta R
.Br.
Azol
la p
inna
ta R
.Br.
Azol
la p
inna
ta R
.Br.
Salv
inia
cuc
ulla
ta R
oxb.
ex
Bor
y Sa
lvin
ia c
ucul
lata
Rox
b. e
x B
ory
Salv
inia
cuc
ulla
ta R
oxb.
ex
Bor
y
Salv
inia
nat
ans (
L.) A
ll.
Salv
inia
nat
ans (
L.) A
ll.Sa
lvin
ia n
atan
s (L.
) All.
SCH
IZA
EA
CE
AE
Schi
zaea
dic
hoto
ma
(L.)
Sm.
Schi
zaea
dic
hoto
ma
(L.)
Sm.
Schi
zaea
dic
hoto
ma
(L.)
Sm.
Schi
zaea
dig
itata
(L.)
Sw.
Schi
zaea
dig
itata
(L.)
Sw.
Schi
zaea
dig
itata
(L.)
Sw.
TE
CTA
RIA
CE
AE
Arth
ropt
eris
pal
isot
ii (D
esv.
) Als
ton
Arth
ropt
eris
pal
isot
ii (D
esv.
) Als
ton
Arth
ropt
eris
pal
isot
ii (D
esv.
) Als
ton
Het
erog
oniu
m a
lder
wer
eltii
Hol
ttum
H
eter
ogon
ium
ald
erw
erel
tii H
olttu
mH
eter
ogon
ium
ald
erw
erel
tii H
olttu
m
Het
erog
oniu
m g
urup
ahen
se (C
.Chr
.) H
olttu
m
Het
erog
oniu
m g
urup
ahen
se (C
.Chr
.) H
olttu
mH
eter
ogon
ium
gur
upah
ense
(C.C
hr.)
Hol
ttum
Het
erog
oniu
m h
enni
pman
ii Ta
gaw
a &
K.Iw
ats.
Het
erog
oniu
m h
enni
pman
ii Ta
gaw
a &
K.Iw
ats.
Het
erog
oniu
m h
enni
pman
ii Ta
gaw
a &
K.Iw
ats.
Het
erog
oniu
m p
inna
tum
(Cop
el.)
Hol
ttum
H
eter
ogon
ium
pin
natu
m (C
opel
.) H
olttu
mH
eter
ogon
ium
pin
natu
m (C
opel
.) H
olttu
m
Het
erog
oniu
m sa
geni
oide
s (M
ett.)
Hol
ttum
H
eter
ogon
ium
sage
nioi
des (
Met
t.) H
olttu
mH
eter
ogon
ium
sage
nioi
des (
Met
t.) H
olttu
m
Pleo
cnem
ia h
emite
liifo
rmis
(Rac
ib.)
Hol
ttum
Pl
eocn
emia
hem
itelii
form
is (R
acib
.) H
olttu
mPl
eocn
emia
hem
itelii
form
is (R
acib
.) H
olttu
m
Pleo
cnem
ia ir
regu
lari
s (C
.Pre
sl) H
olttu
m
Pleo
cnem
ia ir
regu
lari
s (C
.Pre
sl) H
olttu
mPl
eocn
emia
irre
gula
ris (
C.P
resl
) Hol
ttum
Pleo
cnem
ia su
bmem
bran
acea
(Hay
ata)
Tag
awa
&
K.Iw
ats.
Pleo
cnem
ia su
bmem
bran
acea
(Hay
ata)
Tag
awa
&
K.Iw
ats.
Pleo
cnem
ia su
bmem
bran
acea
(Hay
ata)
Tag
awa
&
K.Iw
ats.
97TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Pter
idry
s aus
tral
is C
hing
Pt
erid
rys a
ustr
alis
Chi
ngPt
erid
rys a
ustr
alis
Chi
ng
Pter
idry
s cne
mid
aria
(Chr
ist)
C.C
hr. &
C
hing
Pt
erid
rys c
nem
idar
ia (C
hris
t) C
.Chr
. &
Chi
ngPt
erid
rys c
nem
idar
ia (C
hrist
) C.C
hr. &
Chi
ng
Pter
idry
s syr
mat
ica
(Will
d.) C
.Chr
. & C
hing
Pt
erid
rys s
yrm
atic
a (W
illd.
) C.C
hr. &
Chi
ngPt
erid
rys s
yrm
atic
a (W
illd.
) C.C
hr. &
Chi
ng
Tect
aria
am
plifo
lia (A
lder
w.) C
.Chr
. Te
ctar
ia a
mpl
ifolia
(Ald
erw.
) C.C
hr.
Tect
aria
am
plifo
lia (A
lder
w.) C
.Chr
.
Tect
aria
ang
ulat
a (W
illd.
) C.C
hr.
Tect
aria
ang
ulat
a (W
illd.
) C.C
hr.
Tect
aria
ang
ulat
a (W
illd.
) C.C
hr.
Tect
aria
bar
beri
(Hoo
k.) C
opel
. Te
ctar
ia b
arbe
ri (H
ook.
) Cop
el.
Tect
aria
bar
beri
(Hoo
k.) C
opel
.
○Te
ctar
ia b
rach
iata
(Zol
l. &
Mor
itzi)
C.V
.Mor
ton4
Tect
aria
bra
chia
ta (Z
oll.
& M
oritz
i) C
.V.M
orto
nA
bsen
t
●Te
ctar
ia c
hris
tii C
opel
.13Te
ctar
ia c
hris
tii C
opel
.Te
ctar
ia c
oadu
nata
(J.S
m.)
C.Ch
r., p
ro p
arte
Tect
aria
coa
duna
ta (J
.Sm
.) C
.Chr
.ab
sent
Tect
aria
coa
duna
ta (J
.Sm
.) C.
Chr.,
pro
par
te
Tect
aria
cre
nata
Cav
. Te
ctar
ia c
rena
ta C
av.
Tect
aria
cre
nata
Cav
.
Tect
aria
dec
urre
ns (C
.Pre
sl) C
opel
. Te
ctar
ia d
ecur
rens
(C.P
resl
) Cop
el.
Tect
aria
dec
urre
ns (C
.Pre
sl) C
opel
.
Tect
aria
dev
exa
(Kun
ze) C
opel
. Te
ctar
ia d
evex
a (K
unze
) Cop
el.
Tect
aria
dev
exa
(Kun
ze) C
opel
.
Tect
aria
faur
iei T
agaw
a Te
ctar
ia fa
urie
i Tag
awa
Tect
aria
faur
iei T
agaw
a
●Te
ctar
ia fi
ssa
(Kun
ze) H
olttu
m13
Tect
aria
rum
icifo
lia (R
idl.)
C.C
hr.
Tect
aria
rum
icifo
lia (R
idl.)
C.C
hr.
Tect
aria
fusc
ipes
(Wal
l. ex
Bed
d.) C
.Chr
. Te
ctar
ia fu
scip
es (W
all.
ex B
edd.
) C.C
hr.
Tect
aria
fusc
ipes
(Wal
l. ex
Bed
d.) C
.Chr
.
Tect
aria
gri
ffith
ii (B
aker
) C.C
hr.
Tect
aria
gri
ffith
ii (B
aker
) C.C
hr.
Tect
aria
gri
ffith
ii (B
aker
) C.C
hr.
Tect
aria
gym
noso
ra H
olttu
m
Tect
aria
gym
noso
ra H
olttu
mTe
ctar
ia g
ymno
sora
Hol
ttum
Tect
aria
her
peto
caul
os H
olttu
m
Tect
aria
her
peto
caul
os H
olttu
mTe
ctar
ia h
erpe
toca
ulos
Hol
ttum
●Te
ctar
ia im
pres
sa (F
ée) H
olttu
m
Tect
aria
impr
essa
(Fée
) Hol
ttum
Tect
aria
var
iolo
sa (W
all.
ex H
ook.
) C.C
hr.
○Te
ctar
ia k
ecki
i (Lu
erss
.) C
.Chr
.4Te
ctar
ia k
ecki
i (Lu
erss
.) C
.Chr
.A
bsen
t
Tect
aria
laot
ica
Tard
ieu
& C
.Chr
. Te
ctar
ia la
otic
a Ta
rdie
u &
C.C
hr.
Tect
aria
laot
ica
Tard
ieu
& C
.Chr
.
●Te
ctar
ia m
anile
nsis
(C.P
resl
) Hol
ttum
Te
ctar
ia m
anile
nsis
(C.P
resl
) Hol
ttum
va
r. m
anile
nsis
+ T
ecta
ria
man
ilens
is (C
.Pre
sl) H
olttu
m
var.
chup
enge
nsis
(Rid
l.) H
olttu
m
Cte
nitis
man
ilens
is (C
.Pre
sl) H
olttu
m
98 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Tect
aria
mel
anoc
aulis
(Blu
me)
Cop
el.
Tect
aria
mel
anoc
aulis
(Blu
me)
Cop
el.
Tect
aria
mel
anoc
aulis
(Blu
me)
Cop
el.
Tect
aria
pha
eoca
ulis
(Ros
enst
.) C
.Chr
. Te
ctar
ia p
haeo
caul
is (R
osen
st.)
C.C
hr.
Tect
aria
pha
eoca
ulis
(Ros
enst
.) C
.Chr
.
○Te
ctar
ia p
hano
men
sis S
.Lin
ds.24
Abs
ent
Abs
ent
○Te
ctar
ia p
ilosa
(Fée
) r.C
.Mor
an4
Tect
aria
pilo
sa (F
ée) r
.C.M
oran
Abs
ent
Tect
aria
pol
ymor
pha
(Wal
l. ex
Hoo
k.)
Cop
el.
Tect
aria
pol
ymor
pha
(Wal
l. ex
Hoo
k.)
Cop
el.
Tect
aria
pol
ymor
pha
(Wal
l. ex
Hoo
k.)
Cop
el.
Tect
aria
rock
ii C
.Chr
. Te
ctar
ia ro
ckii
C.C
hr.
Tect
aria
rock
ii C
.Chr
.
●Te
ctar
ia se
mip
inna
ta (r
oxb.
) C.V
.Mor
ton
Tect
aria
sem
ipin
nata
(rox
b.) C
.V.M
orto
nTe
ctar
ia m
aing
ayi (
Bak
er) C
.Chr
.
○Te
ctar
ia sh
ahid
ania
na R
usea
44A
bsen
tA
bsen
t
○Te
ctar
ia si
ifolia
(Will
d.) C
opel
.4Te
ctar
ia si
ifolia
(Will
d.) C
opel
.A
bsen
t
Tect
aria
sim
onsi
i (B
aker
) Chi
ng
Tect
aria
sim
onsi
i (B
aker
) Chi
ngTe
ctar
ia si
mon
sii (
Bak
er) C
hing
Tect
aria
sing
apor
eana
(Wal
l. ex
Hoo
k. &
G
rev.
) Chi
ng
Tect
aria
sing
apor
eana
(Wal
l. ex
Hoo
k. &
G
rev.
) Chi
ngTe
ctar
ia si
ngap
orea
na (W
all.
ex H
ook.
&
Gre
v.) C
hing
Tect
aria
tene
rifro
ns (H
ook.
) Chi
ng
Tect
aria
tene
rifro
ns (H
ook.
) Chi
ngTe
ctar
ia te
neri
frons
(Hoo
k.) C
hing
Tect
aria
tern
ifolia
(Ald
erw.
) C.C
hr.
Tect
aria
tern
ifolia
(Ald
erw.
) C.C
hr.
Tect
aria
tern
ifolia
(Ald
erw.
) C.C
hr.
Tect
aria
vas
ta (B
lum
e) C
opel
. Te
ctar
ia v
asta
(Blu
me)
Cop
el.
Tect
aria
vas
ta (B
lum
e) C
opel
.
●Te
ctar
ia ze
ilani
ca (H
outt.
) Sle
dge
Que
rcifi
lix ze
ilani
ca (H
outt.
) Cop
el.
Que
rcifi
lix ze
ilani
ca (H
outt.
) Cop
el.
TH
ELY
PTE
RID
AC
EA
E
●C
yclo
soru
s ari
dus (
D.D
on) C
hing
C
hris
tella
ari
da (D
.Don
) Hol
ttum
Thel
ypte
ris a
rida
(D.D
on) C
.V.M
orto
n
●C
yclo
soru
s art
icul
atus
(Hou
lsto
n &
T.M
oore
) Pa
nigr
ahi
Pron
ephr
ium
art
icul
atum
(Hou
lsto
n &
T.M
oore
) H
olttu
mTh
elyp
teri
s art
icul
ata
(Hou
lsto
n &
T.M
oore
) Ta
gaw
a &
K.Iw
ats.
●C
yclo
soru
s asp
erus
(C.P
resl
) B.K
.Nay
ar &
Kau
r Pr
onep
hriu
m a
sper
um (C
.Pre
sl) H
olttu
mTh
elyp
teri
s asp
era
(C.P
resl
) K.Iw
ats.
●C
yclo
soru
s can
us (B
aker
) S.L
inds
.14Th
elyp
teri
s rep
ens (
C.H
ope)
Chi
ngTh
elyp
teri
s rep
ens (
C.H
ope)
Chi
ng
●C
yclo
soru
s cili
atus
(Wal
l. ex
Ben
th.)
Pani
grah
i Tr
igon
ospo
ra c
iliat
a (W
all.
ex B
enth
.) H
olttu
mTh
elyp
teri
s cili
ata
(Wal
l. ex
Ben
th.)
Chi
ng
99TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●C
yclo
soru
s cra
ssifo
lius (
Blu
me)
S.L
inds
. M
esop
hleb
ion
cras
sifo
lium
(Blu
me)
Hol
ttum
Thel
ypte
ris c
rass
ifolia
(Blu
me)
Chi
ng
●C
yclo
soru
s cri
nipe
s (H
ook.
) Chi
ng
Chr
iste
lla c
rini
pes (
Hoo
k.) H
olttu
mTh
elyp
teri
s cri
nipe
s (H
ook.
) K.Iw
ats.
●C
yclo
soru
s cla
rkei
(Bed
d.) C
hing
11C
yclo
soru
s cyl
indr
othr
ix (R
osen
st.)
Chi
ngTh
elyp
teri
s cyl
indr
othr
ix (R
osen
st.)
K.Iw
ats.
●C
yclo
soru
s den
tatu
s (Fo
rssk
.) C
hing
C
hris
tella
den
tata
(For
ssk.
) Bro
wns
ey &
Jerm
yTh
elyp
teri
s den
tata
(For
ssk.
) e.P
.St.J
ohn
●C
yclo
soru
s evo
lutu
s (C
.B.C
lark
e &
Bak
er)
Chi
ng
Chr
iste
lla e
volu
ta (C
.B.C
lark
e &
Bak
er)
Hol
ttum
Thel
ypte
ris e
volu
ta (C
.B.C
lark
e &
Bak
er)
Taga
wa
& K
.Iwat
s
●C
yclo
soru
s exs
culp
tus (
Bak
er) S
.Lin
ds.
Pron
ephr
ium
exs
culp
tum
(Bak
er) H
olttu
mTh
elyp
teri
s exs
culp
ta (B
aker
) K.Iw
ats.
●C
yclo
soru
s fal
cilo
bus (
Hoo
k.) P
anig
rahi
Th
elyp
teri
s fal
cilo
ba (H
ook.
) Chi
ngTh
elyp
teri
s fal
cilo
ba (H
ook.
) Chi
ng
●C
yclo
soru
s fer
ox (B
lum
e) C
hing
C
hing
ia fe
rox
(Blu
me)
Hol
ttum
Thel
ypte
ris f
erox
(Blu
me)
Tag
awa
&
K.Iw
ats.
●C
yclo
soru
s gla
ndul
osus
(Blu
me)
Chi
ng
Pron
ephr
ium
gla
ndul
osum
(Blu
me)
Hol
ttum
Thel
ypte
ris g
land
ulos
a (B
lum
e) T
agaw
a &
K
.Iwat
s., n
om. i
lleg.
●C
yclo
soru
s het
eroc
arpu
s (B
lum
e)
Chi
ng
Spha
eros
teph
anos
het
eroc
arpu
s (B
lum
e)
Hol
ttum
Thel
ypte
ris h
eter
ocar
pa (B
lum
e)
C.V
.Mor
ton
●C
yclo
soru
s im
mer
sus (
Blu
me)
S.L
inds
. Am
phin
euro
n im
mer
sum
(Blu
me)
Hol
ttum
Thel
ypte
ris i
mm
ersa
(Blu
me)
Chi
ng
●C
yclo
soru
s int
erru
ptus
(Will
d.) H
.Ito
Cyc
loso
rus i
nter
rupt
us (W
illd.
) H.It
oTh
elyp
teri
s int
erru
pta
(Will
d.) K
.Iwat
s.
●C
yclo
soru
s lak
him
pure
nse
(ros
enst
.) B
.K.N
ayar
& K
aur
Pron
ephr
ium
lakh
impu
rens
e (R
osen
st.)
Hol
ttum
Thel
ypte
ris l
akhi
mpu
rens
is (R
osen
st.)
K.Iw
ats.
●C
yclo
soru
s lar
uten
sis (
Bed
d.) C
hing
Sp
haer
oste
phan
os la
rute
nsis
(Bed
d.) C
.Chr
.Th
elyp
teri
s lar
uten
sis (
Bed
d.) T
agaw
a &
K
.Iwat
s.
●C
yclo
soru
s leb
eufii
(Bak
er) W
.M.C
hu
Chr
iste
lla le
beufi
i (B
aker
) Hol
ttum
Thel
ypte
ris l
ebeu
fii (B
aker
) Pan
igra
hi
●C
yclo
soru
s meg
aphy
llus (
Met
t.) C
hing
Spha
eros
teph
anos
pen
nige
r Hol
ttum
, no
m. n
ud.
Thel
ypte
ris m
egap
hylla
(Met
t.) K
.Iwat
s.
●C
yclo
soru
s men
isci
icar
pus (
Blu
me)
H
olttu
m
Pron
ephr
ium
men
isci
icar
pon
(Blu
me)
H
olttu
mTh
elyp
teri
s men
isci
icar
pa (B
lum
e)
K.Iw
ats.
100 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●C
yclo
soru
s mol
liusc
ulus
(Kuh
n) C
hing
C
hris
tella
app
endi
cula
ta (C
.Pre
sl) H
olttu
mTh
elyp
teri
s mol
liusc
ula
(Kuh
n) K
.Iwat
s.
●C
yclo
soru
s nud
atus
(rox
b.) B
.K.N
ayar
& K
aur
Pron
ephr
ium
nud
atum
(Rox
b.) H
olttu
mTh
elyp
teri
s nud
ata
(rox
b.) C
.V.M
orto
n
●C
yclo
soru
s opu
lent
us (K
aulf.
) Nak
aike
Am
phin
euro
n op
ulen
tum
(Kau
lf.) H
olttu
mTh
elyp
teri
s opu
lent
a (K
aulf.
) Fos
berg
●C
yclo
soru
s pap
ilio
(C.H
ope)
Chi
ng
Chr
iste
lla p
apili
o (C
.Hop
e) H
olttu
mTh
elyp
teri
s pap
ilio
(C.H
ope)
K.Iw
ats.
●C
yclo
soru
s par
asiti
cus (
L.) F
arw
. C
hris
tella
par
asiti
ca (L
.) H
.Lév
.Th
elyp
teri
s par
asiti
ca (L
.) Ta
rdie
u
●C
yclo
soru
s pen
angi
anus
(Hoo
k.) C
opel
. Pr
onep
hriu
m p
enan
gian
um (H
ook.
) Hol
ttum
Thel
ypte
ris p
enan
gian
a (H
ook.
) C.F
.Ree
d
●C
yclo
soru
s pol
ycar
pus (
Blu
me)
Hol
ttum
Sp
haer
oste
phan
os p
olyc
arpa
(Blu
me)
Cop
el.
Thel
ypte
ris p
olyc
arpa
(Blu
me)
K.Iw
ats.
●C
yclo
soru
s pro
lifer
(Ret
z.) T
ardi
eu e
x Ta
rdie
u &
C.C
hr.
Men
isci
um p
rolif
erum
(Ret
z.) S
w.
Men
isci
um p
rolif
erum
(Ret
z.) S
w.
●C
yclo
soru
s rep
andu
s (Fé
e) B
.K.N
ayar
& K
aur
Pron
ephr
ium
repa
ndum
(Fée
) Hol
ttum
Thel
ypte
ris r
epan
da (F
ée) C
.V.M
orto
n
○C
yclo
soru
s rub
icun
dus (
Ald
erw
.) S.
Lind
s.4Pr
onep
hriu
m ru
bicu
ndum
(Ald
erw.
) Hol
ttum
Abs
ent
○C
yclo
soru
s sal
icifo
lius (
Wal
l. ex
Hoo
k.) C
opel
.4Pr
onep
hriu
m sa
licifo
lium
(Wal
l. ex
Hoo
k.) H
olttu
mA
bsen
t
●C
yclo
soru
s sia
men
sis (
Taga
wa
& K
.Iwat
s.) P
anig
rahi
C
hris
tella
siam
ensi
s (Ta
gaw
a &
K.Iw
ats.)
Hol
ttum
Thel
ypte
ris s
iam
ensi
s Tag
awa
& K
.Iwat
s.
●C
yclo
soru
s sub
elat
us (B
aker
) Chi
ng
Chr
iste
lla su
bela
ta (B
aker
) Hol
ttum
Thel
ypte
ris s
ubel
ata
(Bak
er) K
.Iwat
s.
●C
yclo
soru
s sub
pube
scen
s (B
lum
e) C
hing
12C
hris
tella
subp
ubes
cens
(Blu
me)
Hol
ttum
+
Chr
iste
lla la
tipin
na (B
enth
.) H
.Lév
.Th
elyp
teri
s sub
pube
scen
s (B
lum
e) K
.Iwat
s. +
Thel
ypte
ris l
atip
inna
(Hoo
k.) K
.Iwat
s. +
Thel
ypte
ris s
umat
rana
(Ald
erw.
) Tag
awa
&
K.Iw
ats.
●C
yclo
soru
s ter
min
ans (
J.Sm
. ex
Hoo
k.)
Pani
grah
i Am
phin
euro
n te
rmin
ans (
J.Sm
. ex
Hoo
k.)
Hol
ttum
Th
elyp
teri
s ter
min
ans (
J.Sm
. ex
Hoo
k.)
Taga
wa
& K
.Iwat
s.
○C
yclo
soru
s tha
iland
icus
S.L
inds
.24A
bsen
tA
bsen
t
●C
yclo
soru
s tri
phyl
lus (
Sw.)
Tard
ieu
ex T
ardi
eu &
C.C
hr.
var.
trip
hyllu
sPr
onep
hriu
m tr
iphy
llum
(Sw.
) Hol
ttum
va
r. tr
iphy
llum
Thel
ypte
ris t
riph
ylla
(Sw.
) K.Iw
ats.
var.
trip
hylla
●C
yclo
soru
s tri
phyl
lus (
Sw.)
Tard
ieu
ex T
ardi
eu &
C.C
hr.
var.
pari
shii
(Bed
d.) S
.Lin
ds.
Pron
ephr
ium
trip
hyllu
m (S
w.) H
olttu
m
var.
pari
shii
(Bed
d.) N
akai
keTh
elyp
teri
s tri
phyl
la (S
w.) K
.Iwat
s. va
r. pa
rish
ii (B
edd.
) K.Iw
ats.
●C
yclo
soru
s tru
ncat
us (P
oir.)
Far
w.
Pneu
mat
opte
ris t
runc
ata
(Poi
r.) H
olttu
mTh
elyp
teri
s tru
ncat
a (P
oir.)
K.Iw
ats.
●C
yclo
soru
s tub
ercu
lifer
(C.C
hr.)
Pani
grah
i Th
elyp
teri
s tub
ercu
lifer
a (C
.Chr
.) C
hing
Thel
ypte
ris t
uber
culif
era
(C.C
hr.)
Chi
ng
101TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●C
yclo
soru
s tyl
odes
(Kun
ze) P
anig
rahi
14Th
elyp
teri
s xyl
odes
(Kun
ze) C
hing
Thel
ypte
ris x
ylod
es (K
unze
) Chi
ng
●C
yclo
soru
s uni
tus (
L.) C
hing
Sp
haer
oste
phan
os u
nitu
s (L.
) Hol
ttum
Thel
ypte
ris u
nita
(L.)
C.V
.Mor
ton
●C
yclo
soru
s val
idus
(Chr
ist)
Chi
ng
Cyc
loso
rus h
irtis
orus
(C.C
hr.)
Chi
ngTh
elyp
teri
s hir
tisor
a (C
.Chr
.) K
.Iwat
s. +
Thel
ypte
ris v
alid
a (C
hris
t) Ta
gaw
a &
K
.Iwat
s.
●M
acro
thel
ypte
ris o
rnat
a (W
all.
ex B
edd.
) Chi
ng
Mac
roth
elyp
teri
s orn
ata
(Wal
l. ex
Bed
d.) C
hing
Thel
ypte
ris o
rnat
a (W
all.
ex B
edd.
) Chi
ng
●M
acro
thel
ypte
ris t
orre
sian
a (G
audi
ch.)
Chi
ng
Mac
roth
elyp
teri
s tor
resi
ana
(Gau
dich
.) C
hing
Thel
ypte
ris t
orre
sian
a (G
audi
ch.)
Als
ton
●Ps
eudo
pheg
opte
ris s
umat
rana
Hol
ttum
Ps
eudo
pheg
opte
ris s
umat
rana
Hol
ttum
Abs
ent
Thel
ypte
ris c
onflu
ens (
Thun
b.) C
.V.M
orto
n Th
elyp
teri
s con
fluen
s (Th
unb.
) C.V
.Mor
ton
Thel
ypte
ris c
onflu
ens (
Thun
b.) C
.V.M
orto
n
Thel
ypte
ris fl
acci
da (B
lum
e) C
hing
M
etat
hely
pter
is fl
acci
da (B
lum
e) C
hing
Thel
ypte
ris fl
acci
da (B
lum
e) C
hing
●Th
elyp
teri
s hir
sutip
es (C
lark
e) C
hing
C
oryp
hopt
eris
hir
sutip
es (C
.B.C
lark
e) H
olttu
mTh
elyp
teri
s hir
sutip
es (C
lark
e) C
hing
○Th
elyp
teri
s lax
a (F
ranc
h. &
Sav
.) C
hing
26
Abs
ent
Abs
ent
●Th
elyp
teri
s sin
gala
nens
is (B
aker
) Chi
ng
Met
athe
lypt
eris
sing
alan
ensi
s (B
aker
) Chi
ngTh
elyp
teri
s sin
gala
nens
is (B
aker
) Chi
ng
Thel
ypte
ris v
isco
sa (B
aker
) Chi
ng
Cor
ypho
pter
is v
isco
sa (B
aker
) Hol
ttum
Thel
ypte
ris v
isco
sa (B
aker
) Chi
ng
WO
OD
SIA
CE
AE
Athy
rium
ani
sopt
erum
Chr
ist
Athy
rium
ani
sopt
erum
Chr
ist
Athy
rium
ani
sopt
erum
Chr
ist
●At
hyri
um c
umin
gian
um (C
.Pre
sl) M
ilde
Anis
ocam
pium
cum
ingi
anum
C.P
resl
Anis
ocam
pium
cum
ingi
anum
C.P
resl
●At
hyri
um c
uspi
datu
m (B
edd.
) M.K
ato
Kun
iwat
suki
a cu
spid
ata
(Bed
d.) P
ichi
-Ser
m.
Kun
iwat
suki
a cu
spid
ata
(Bed
d.) P
ic.S
erm
.
Athy
rium
dis
sitif
oliu
m (B
aker
) C.C
hr.
Athy
rium
dis
sitif
oliu
m (B
aker
) C.C
hr.
Athy
rium
dis
sitif
oliu
m (B
aker
) C.C
hr.
Athy
rium
mac
kinn
onii
(C.H
ope)
C.C
hr.
Athy
rium
mac
kinn
onii
(C.H
ope)
C.C
hr.
Athy
rium
mac
kinn
onii
(C.H
ope)
C.C
hr.
Athy
rium
setif
erum
C.C
hr.
Athy
rium
setif
erum
C.C
hr.
Athy
rium
setif
erum
C.C
hr.
Cor
nopt
eris
opa
ca (D
.Don
) Tag
awa
Cor
nopt
eris
opa
ca (D
.Don
) Tag
awa
Cor
nopt
eris
opa
ca (D
.Don
) Tag
awa
●D
epar
ia p
eter
seni
i (K
unze
) M.K
ato
Athy
rium
japo
nicu
m a
uct.
non
(Thu
nb.)
Cop
el.
Dep
aria
japo
nica
auc
t. no
n (T
hunb
.) M
.Kat
o / D
epar
ia p
eter
seni
i (K
unze
) M.K
ato
102 THaI FOreST BuLLeTIN (BOTaNY) 37
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
●D
epar
ia su
bfluv
ialis
(Hay
ata)
M.K
ato
Athy
rium
bor
yanu
m a
uct.
non
(Will
d.) T
agaw
aD
epar
ia b
orya
na a
uct.
non
(Will
d.) M
.Kat
o /
Dep
aria
subfl
uvia
lis (H
ayat
a) M
.Kat
o
Dip
lazi
um a
cced
ens B
lum
e D
ipla
zium
acc
eden
s Blu
me
Dip
lazi
um a
cced
ens B
lum
e
Dip
lazi
um b
anta
men
se B
lum
e D
ipla
zium
ban
tam
ense
Blu
me
Dip
lazi
um b
anta
men
se B
lum
e
Dip
lazi
um c
onte
rmin
um C
hris
t D
ipla
zium
con
term
inum
Chr
ist
Dip
lazi
um c
onte
rmin
um C
hris
t
Dip
lazi
um c
ordi
foliu
m B
lum
e D
ipla
zium
cor
difo
lium
Blu
me
Dip
lazi
um c
ordi
foliu
m B
lum
e
Dip
lazi
um c
rena
to-s
erra
tum
(Blu
me)
T.
Moo
re
Dip
lazi
um c
rena
to-s
erra
tum
(Blu
me)
T.
Moo
reD
ipla
zium
cre
nato
-ser
ratu
m (B
lum
e)
T.M
oore
Dip
lazi
um d
ilata
tum
Blu
me
Dip
lazi
um d
ilata
tum
Blu
me
Dip
lazi
um d
ilata
tum
Blu
me
○D
ipla
zium
doe
derl
eini
i (Lu
erss
.) M
akin
o26A
bsen
tA
bsen
t
Dip
lazi
um d
onia
num
(Met
t.) T
ardi
eu
Dip
lazi
um d
onia
num
(Met
t.) T
ardi
euD
ipla
zium
don
ianu
m (M
ett.)
Tar
dieu
Dip
lazi
um e
scul
entu
m (R
etz.
) Sw.
D
ipla
zium
esc
ulen
tum
(Ret
z.) S
w.D
ipla
zium
esc
ulen
tum
(Ret
z.) S
w.
Dip
lazi
um h
eter
ophl
ebiu
m (M
ett.
ex B
aker
) D
iels
D
ipla
zium
het
erop
hleb
ium
(Met
t. ex
Bak
er)
Die
lsD
ipla
zium
het
erop
hleb
ium
(Met
t. ex
Bak
er)
Die
ls
Dip
lazi
um le
ptop
hyllu
m C
hris
t D
ipla
zium
lept
ophy
llum
Chr
ist
Dip
lazi
um le
ptop
hyllu
m C
hris
t
Dip
lazi
um m
alac
cens
e C
.Pre
sl
Dip
lazi
um m
alac
cens
e C
.Pre
slD
ipla
zium
mal
acce
nse
C.P
resl
Dip
lazi
um m
egap
hyllu
m (B
aker
) Chr
ist
Dip
lazi
um m
egap
hyllu
m (B
aker
) Chr
ist
Dip
lazi
um m
egap
hyllu
m (B
aker
) Chr
ist
Dip
lazi
um m
ette
nian
um (M
iq.)
C.C
hr.
Dip
lazi
um m
ette
nian
um (M
iq.)
C.C
hr.
Dip
lazi
um m
ette
nian
um (M
iq.)
C.C
hr.
Dip
lazi
um m
uric
atum
(Met
t.) a
lder
w.
Dip
lazi
um m
uric
atum
(Met
t.) a
lder
w.D
ipla
zium
mur
icat
um (M
ett.)
ald
erw.
Dip
lazi
um p
etel
otii
Tard
ieu
Dip
lazi
um p
etel
otii
Tard
ieu
Dip
lazi
um p
etel
otii
Tard
ieu
Dip
lazi
um p
etri
Tar
dieu
D
ipla
zium
pet
ri T
ardi
euD
ipla
zium
pet
ri T
ardi
eu
Dip
lazi
um p
olyp
odio
ides
Blu
me
Dip
lazi
um p
olyp
odio
ides
Blu
me
Dip
lazi
um p
olyp
odio
ides
Blu
me
Dip
lazi
um p
resc
ottia
num
(Wal
l. ex
Hoo
k.)
T.M
oore
D
ipla
zium
pre
scot
tianu
m (W
all.
ex H
ook.
) T.
Moo
reD
ipla
zium
pre
scot
tianu
m (W
all.
ex H
ook.
) T.
Moo
re
○D
ipla
zium
pro
cum
bens
Hol
ttum
10A
bsen
tA
bsen
t
103TOWarDS a STaBLe NOMeNCLaTure FOr THaI FerNS (S. LINDSaY, D.J. MIDDLeTON, T. BOONKerD & S. SuDDee)
CU
RR
EN
T N
AM
E
NA
ME
IN P
TE
RID
OPH
YT
ES
IN T
HA
ILA
ND
(B
oonk
erd
& P
olla
wat
n, 2
000)
NA
ME
IN F
LO
RA
OF
TH
AIL
AN
D
(Tag
awa
& Iw
atsu
ki, 1
979,
198
5, 1
988,
198
9)
Dip
lazi
um ri
pari
um H
olttu
m
Dip
lazi
um ri
pari
um H
olttu
mD
ipla
zium
ripa
rium
Hol
ttum
Dip
lazi
um si
amen
se C
.Chr
. D
ipla
zium
siam
ense
C.C
hr.
Dip
lazi
um si
amen
se C
.Chr
.
Dip
lazi
um si
lvat
icum
(Bor
y) S
w.
Dip
lazi
um si
lvat
icum
(Bor
y) S
w.D
ipla
zium
silv
atic
um (B
ory)
Sw.
Dip
lazi
um si
mpl
iciv
eniu
m H
olttu
m
Dip
lazi
um si
mpl
iciv
eniu
m H
olttu
mD
ipla
zium
sim
plic
iven
ium
Hol
ttum
Dip
lazi
um so
rzog
onen
se (C
.Pre
sl) C
.Pre
sl
Dip
lazi
um so
rzog
onen
se (C
.Pre
sl) C
.Pre
slD
ipla
zium
sorz
ogon
ense
(C.P
resl
) C.P
resl
Dip
lazi
um su
bint
egru
m H
olttu
m
Dip
lazi
um su
bint
egru
m H
olttu
mD
ipla
zium
subi
nteg
rum
Hol
ttum
Dip
lazi
um su
bser
ratu
m B
lum
e D
ipla
zium
subs
erra
tum
Blu
me
Dip
lazi
um su
bser
ratu
m B
lum
e
Dip
lazi
um su
bsin
uatu
m (W
all.
ex H
ook.
& G
rev.
) Ta
gaw
a D
ipla
zium
subs
inua
tum
(Wal
l. ex
Hoo
k. &
Gre
v.)
Taga
wa
Dip
lazi
um su
bsin
uatu
m (W
all.
ex H
ook.
& G
rev.
) Ta
gaw
a
Dip
lazi
um ta
iwan
ense
Tag
awa
Dip
lazi
um ta
iwan
ense
Tag
awa
Dip
lazi
um ta
iwan
ense
Tag
awa
Dip
lazi
um to
men
tosu
m B
lum
e D
ipla
zium
tom
ento
sum
Blu
me
Dip
lazi
um to
men
tosu
m B
lum
e
○D
ipla
zium
vire
scen
s Kun
ze26
A
bsen
tA
bsen
t
Dip
lazi
um x
ipho
phyl
lum
(Bak
er) C
.Chr
. D
ipla
zium
xip
hoph
yllu
m (B
aker
) C.C
hr.
Dip
lazi
um x
ipho
phyl
lum
(Bak
er) C
.Chr
.
104 THaI FOreST BuLLeTIN (BOTaNY) 37
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