Ordovician ostracodes from Novaya Zemlya · 2014-01-03 · ter 1993, Pl. 11A, Fig. 1, Pl. 14A, and Pl. 1B, Fig. 2, Pl.2A, Figs 1ñ3, Pl. 3, Fig. 1, Pl. 8A, Pl. 9). In this re-spect,

Journal of the Czech Geological Society 46/3ñ4(2001) 199

Ordovician ostracodes from Novaya Zemlya

OrdoviËtÌ ostrakodi z NovÈ ZemÏ

(4 plates)

ROGER SCHALLREUTER 1 ñ ALEKSANDR V. KANYGIN 2 ñ INGELORE HINZ-SCHALLREUTER 3

1 Geologisch-Pal‰ontologisches Institut und Museum, Bundesstr. 55, D-20416 Hamburg; e-mail: [email protected];Institut f¸r Geologische Wissenschaften, Friedrich-Ludwig-Jahn-Str. 17a, D-17489 Greifswald; [email protected]

2 Russian Academy of Science, Siberian branch, Institute of geology and geophysics, Universitetskyj prosp. 3, RUS-630090 Novosibirsk3 Institut f¸r Geologische Wissenschaften, Friedrich-Ludwig-Jahn-Str. 17a, D-17489 Greifswald; e-mail: [email protected]

From the Ordovician of Novaya Zemlya 31 new species and subspecies of ostracodes are described of 28 (10 new) genera and subgenera.Some of the ostracod genera show relations to Baltica, others to Siberia. This fact is a significant expression for the palaeogeographicallyintermediate position of Novaya Zemlya between these two continents during the Ordovician.

Key words: Ordovician, Ostracodes, Novaya Zemlya, new taxa

E t y m o l o g y : After Cape Petermann, NW coast of Northern NovayaZemlya.

H o l o t y p e : Female (?) left valve from sample 416-33/14 ñ Pl. IV,Fig. 1.

O c c u r r e n c e : sample 416-33/14; sample 416-10/5a (P. cf. peter-manni).

D i a g n o s i s : At least up to 1.22 mm. Large, flat ovalmuscle spot just in front of center of domicilium. Rela-tively narrow velar frill. Reticulation very fine, nearlypunctate.R e m a r k s : P. (R.) temperata temperata Sarv, 1956from the Rakverian of Estonia differs by its larger sizeand relatively broader velar frill.

Genus Cystomatochilina Jaanusson, 1957

Cystomatochilina flotowi sp. nov.Pl. I, Figs 1ñ2

E t y m o l o g y : After Cape Flotow, Eí coast of Ní Novaya Zemlya.H o l o t y p e : Posteriorly incomplete right tecnomorphic valve from

sample 416-30/6 ñ Pl. I, Fig. 1.

D i a g n o s i s : At least up to 0.85 mm. Medium-sizedpreadductorial node in front of center of domicilium. Sul-cal depression behind node (S2) without fissum. No pli-ca. Velar frill relatively narrow. Lateral surface more orless distinctly reticulate and with scattered tubercles (pus-tulae ?).R e m a r k s : The type species of Cystomatochilina,Primitia umbonata (Krause, 1892) from upper Ordovi-cian geschiebes of Ní Germany and Sí Finland and ofNabala to Porkuni stages of Estonia, has a slightly to dis-tinctly larger preadductorial node and a short fissum inthe S2 (Martinsson 1956, Pl. 1, Figs 3ñ6, Jaanusson1957, Fig. 16; Schallreuter 1993, Pl. 61B, Fig. 1, Pl. 62A,Figs 1ñ3; Meidla 1996, Pl. 2, Figs 4ñ5).

Similar to the new species is Gellensia nodoreticula-ta Schallreuter, 1982 from upper Ordovician ÷jlemyrflintgeschiebes of N Germany which is, however, more elon-

Introduction

Presently, Ordovician ostracods from Novaya Zemlya arepoorly known. The first record from the the Llandeilo ofthe neighboured Isle of Vajgach dates back to 1949, whenGlebovskaja described the three new species Primitiajusharensis, Bollia perincerta, and Tetradella variabilis.However, according to a personal comment from A. F.Abushik (1988) the material is apparently lost. The earlyreport by Glebovskaja followed a description of two egor-ovellid species from a collection of Ordovician ostracodsfrom Novaya Zemlya by the present authors in 1999:Debonia micropunctata and Fissebonia retuba. Until now,the family Egorovellidae was known only from Siberia.The same applied for the family Cherskiellidae. Two newspecies of the latter, Nosebiria quadrilobata and Gonia-tor kiesowioides are described in another paper.

The present paper deals with the interposing compo-sition of Novaya Zemlya fauna and the Siberian and Bal-toscandian faunal provinces.

The rock material from which the ostracods were pre-pared by mechanical fracturing is a dark bituminouslimestone. The ostracods usually have their shells pre-served, but some of them appear more or less distorted.

The material will be housed at the Institute of Geolo-gy and Geophysics of the Siberian branch of the RussianAcademy of Sciences.

Systematic paleontology

Family O e p i k e l l i d a e Jaanusson, 1957Subfamily A m p l e t o c h i l i n i n a e Schallreuter, 1975

Genus Platybolbina Henningsmoen, 1953

Platybolbina (Reticulobolbina) Schallreuter, 1969

Platybolbina (Reticulobolbina) temperata petermannisubsp. nov.Pl. IV, Fig. 1

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200 Journal of the Czech Geological Society 46/3ñ4(2001)

gate and reticulate only on the preadductorial node. Thelatter is more closely situated to the dorsal margin thanin the new species. Furthermore, the velar frill seems tobe broader in G. nodoreticulata.

Genus Levisulculus Jaanusson, 1957

Levisulculus posteroventrolobatus sp. nov.Pl. II, Fig. 1

E t y m o l o g y : After the posteroventral lobe.H o l o t y p e : Right female valve from sample 416-30/6 ñ Pl. II, Fig. 1.

D i a g n o s i s : At least up to > 0.63 mm. Sulcal depres-sion with large muscle scar behind distinct preadducto-rial node. Distinct lobe with cone-like top situated pos-teroventrally to muscle scar. Females with long antrumfrom anterocentral to posteroventral region. Lateral sur-face reticulate.R e m a r k s : The species is characterized by the poster-oventral lobe with its cone-like top.

Genus Bulbosulculus Ivanova, 1979

Bulbosulculus dactylus sp. nov.Pl. II, Fig. 7

E t y m o l o g y : dactylo Ö, according to the fingerprint-like ornamen-tation.

H o l o t y p e : Posteriorly incomplete right female valve from sam-ple 416-30/6 ñ Pl. II, Fig. 7.

D i a g n o s i s : At least up to > 0.62 mm. Sulcal depres-sion (S2) weak, most distinct in front of and dorsal toflat and indistinct posteroventral inflation which is locat-ed in the ventral half of the domicilium. Dolon broad andstrongly convex, extending from anterocentral to at leastposteroventral region. Lateral surface striated parallel toanterodorsal corner. Dolon weaker striated than lateralsurface, running perpendicular to lateral surface withwhich it forms a flat but distinct laterovelar furrow.R e m a r k s : The new species differs from B. fragilis(Kanygin 1971) by its broader dolon and less distinct S2(Kanygin 1971, Pl. 4, Figs 6ñ10; Ivanova 1979, Pl. 8,Figs 3ñ4).

Genus Chilobolbina Ulrich ñ Bassler, 1923

Chilobolbina pseudola sp. nov.Pl. III, Fig. 6

E t y m o l o g y : According to the similarity with C. ola Schallreuter,1993.

H o l o t y p e : Right valve from sample 416-10/5a ñ Pl. III, Fig. 6.

D i a g n o s i s : At least up to 1.05 mm. Well-developedplica. Lateral surface not distinctly sculptured.R e m a r k s : The new species is very similar to C. ola.In the holotype of C. ola the dorsal region is still ob-

scured by rock material, but the presence of a distinctplica is observable in another specimen which was re-cently discovered from a Rogˆ sandstone geschiebe ofN Niedersachsen. The only distinct difference is the sur-face ornamentation which is reticulate (or spinose ?) inC. ola (Schallreuter 1993, Pl. 43A, Fig. 1).

Family T v a e r e n e l l i d a e Jaanusson, 1957Subfamily N o d a m b i c h i l i n i n a e Schallreuter, 1967

Genus Havlicekites gen. nov.

E t y m o l o g y : In honour of V. HavlÌËek.I n c l u d e d s p e c i e s :

Apatochilina ? falacata Sarv, 1962 (type species)Macronotella porkunica Neckaja in Abushik ñ Neckaja et al., 1958Havlicekites fissuratus sp. nov.

O c c u r r e n c e : Upper Ordovician of the Baltic region, Ordovicianof Novaya Zemlya.

D i a g n o s i s : Medium-sized. Outline subamplete toslightly preplete, rounded-triangular. Nearly nonsulcatewith the S2 only as a faint sulcal depression. Dorsal lobesvery weak, and converging, forming a broad umbo thatprojects beyond the hinge-line. Small velar ridge in tec-nomorphs, females with a long and broad anteroventralantrum. Surface smooth or reticulate.R e m a r k s : Havlicekites differs from the North Ameri-can Nodambichilina by its less distinct sulcus and the dor-sal node-like lobes which converge to a broad umbo. Sarv1962 (with ?) and Meidla 1996 placed H. falacatus in Ap-atochilina. Apatochilina (= Colacchilina Kesling ñ Hall ñMelik, 1962) differs from Havlicekites by the umbo presentonly in posterior half of valve, a more rectangular outlineand the frill-like velum (Ulrich 1890, Pl. 9, Fig. 13).

Havlicekites fissuratus sp. nov.Pl. I, Fig. 4

E t y m o l o g y : After the fissum-like furrow in the ventrocentral re-gion.

H o l o t y p e : Left tecnomorphic valve from sample 416-30/6 ñ Pl. I,Fig. 4.

D i a g n o s i s : At least up to 1.85 mm. Outline round-ed-triangular. Ventrocentral region with a short fissum-like furrow. Surface smooth.R e m a r k s : H. porkunica (= H. falacatus) is finely re-ticulate and has a less triangular outline. The laterovelarfurrow is less distinct in H. fissuratus and a fissum-likefurrow is not known in H. porkunica (Abushik ñ Necka-ja et al. 1958, Pl. 2, Fig. 1; Sarv 1962, Pl. 3, Figs 1ñ5).

Family D o l b o r e l l i d a e Melnikova, 1976

Genus Parajonesites V. A. Ivanova in E. A. Ivanovaal., 1955

Parajonesites sawina sp. nov.Pl. I, Fig. 9.

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E t y m o l o g y : After the Sawina Bay, eastern coast of Sí NowajaZemlya.

H o l o t y p e : Right tecnomorphic valve from sample 416-30/6 ñ Pl. I,Fig. 9.

D i a g n o s i s : At least up to 0.78 mm. Two verticalridges in front of the S2, another ridge behind the S2.Anterior ridges rather long, dorsally almost reaching theplica. The converge and join ventrally just before reach-ing the velum. Posterior ridge extending from dorsal plicato a line just below the S2.R e m a r k s : In P. notabilis V. A. Ivanova in E. A. Ivano-va et al., 1955 the ridges are more distinctly developedin the ventral half of the valve with only one prominentvertical ridge in front of the S2 (V. A. Ivanova in E. A.Ivanova et al., 1955, Pl. 23, Figs 4ñ5).

Family C t e n o n o t e l l i d a e Schmidt, 1941Subfamily T a l l i n n e l l i n a e Schallreuter, 1976

Genus Tetrada Neckaja in Abushik et al., 1958

Tetrada ? indistincta sp. nov.Pl. I, Fig. 8

E t y m o l o g y : According to the flat lobation.H o l o t y p e : Left valve from sample 416-30/6 ñ Pl. I, Fig. 8.

D i a g n o s i s : Length at least up to 0.54 mm. Quadri-lobate, lobes flat. S2 is the broadest sulcus and situatedin the centrodorsal part; other sulci weak, narrow and fis-sum-like. Parallel to free margin extends a more or lessbroad, bulge-like lateroventral ridge. It starts anteriorlyin front of the L1 and incorporating the posterior L4. Be-tween this ridge and the free margin there is anotherbulge-like marginal ridge. Lobes being connected ven-trally with the lateroventral ridge.R e m a r k s : The congeneric T. ? fragilis (Kanygin, 1971)from the Middle Ordovician of Sette-Daban is of about thesame size (0.60 mm), but differs in the development of thelobes. Except for L4, they are not connected with the lat-eroventral ridge (Kanygin 1971, Pl. 4, Figs 11ñ12). Be-cause of the unkown velar dimorphism the assignment ofboth species to the genus Tetrada is questionable.

Subfamily C t e n e n t o m i n a e Schmidt, 1941

Genus Octocristatia gen. nov.

E t y m o l o g y : According to the eight vertical ridges (adventral ridg-es included).

T y p e s p e c i e s : Octocristatia ocris sp. nov.

D i a g n o s i s : Small. Quadrilobate with lobes reach-ing dorsal border except for L2. Lobes cristate, ventrallyconnected by a ridge-like lobe. An entire histium-likeridge is present and parallels L1 and L4 together withthe connecting ridge as well as the completely devel-

oped ridge-like velum. Velar dimorphism unknown.Surface reticulate.R e m a r k s : The genus resembles Tallinnellina andRigidella but differs by the histium-like ridge (Schallreu-ter 1993, Pl. 11A, Fig. 1, Pl. 14A, and Pl. 1B, Fig. 2,Pl. 2A, Figs 1ñ3, Pl. 3, Fig. 1, Pl. 8A, Pl. 9). In this re-spect, the genus resembles the phylogenetically more ad-vanced Pseudostrepula which differs by the noncristateL2, the posteriorly dividing cristae which do not reachthe dorsal margin and the restricted velum [Schallreuter1976, Pl. 39(6), Fig. 4; 1983, Pl. 25(11), Figs 4ñ5].

Octocristatia octo sp. nov.Pl. II, Fig. 2.

E t y m o l o g y : Artificially formed.H o l o t y p e : Tecnomorphic left valve from sample 416-10/5a ñ Pl. II,

Fig. 2.

D i a g n o s i s : As for genus.

Genus Pseudostrepula ÷pik, 1937

Pseudostrepula ? severa sp. nov.Pl. IV, Fig. 2

E t y m o l o g y : From sever (Russian) North.H o l o t y p e : Posteroventrally incomplete tecnomorphic left valve

from sample 416-30/4 ñ Pl. IV, Fig. 2.

D i a g n o s i s : Length at leat up to 0.69 mm. Lateralsurface with three ridges parallel to free margin. Dorsalridge branching anteriorly with the posterior branchcrossing preadductorial node. Surface reticulated.R e m a r k s : The type species, P. kuckersiana (Bonne-ma, 1909), is characterized by two posteriorly branch-ing ridges and a preadductorial node without ridge(Schallreuter 1966, Pl. 4, Figs 1ñ2; 1976, Pl. 6, Fig. 4;1983, Pl. 11, Figs 4ñ5).

Genus Strephithis gen. nov.

E t y m o l o g y : Formed from the genera names Strepula + HithisT y p e s p e c i e s : Strephithis multicostatus sp. nov.

D i a g n o s i s : Medium-sized. Unisulcate with moder-ately long S2. Distinct preadductorial node (L2), a moreor less distinct bulb-like large node (L3) located poster-odorsally to S2 at the dorsal border. Distinct plica. Lat-eral surface between N2/S2 and velum marked by threesubparallel ridges. Median ridge completely developedor only with its ventral part; innermost ridge commenc-es at plica and splits in two or three branches posterily.A short ridge connects the innermost ridge with the ven-tral end of the N2 or a ridge on the N2, respectively. Di-morphism unknown.R e m a r k s : Pseudostrepula possesses two parallel ridg-es which both divides in the posterior half.

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Strephithis multicostatus sp. nov.Pl. II, Fig. 3; Pl. III, Fig. 2

E t y m o l o g y : after the several ridges of the lateral surface.H o l o t y p e : Posterodorsally incomplete right tecnomorphic valve

from sample 416-33/14 ñ Pl. III, Fig. 2.

D i a g n o s i s : As for genus which is presently mono-typic.R e m a r k s : The posterodorsal node is missing in thesmaller valve (Pl. II, Fig. 3); in the holotype it is distinctbut not as strong as in S. multicostatus binodus. The me-dian ridge is present mainly in the ventral half of thevalve and might be the strongest of all ridges.

Strephithis multicostatus binodus subsp. nov.Pl. II, Fig. 8

E t y m o l o g y : after the several ridges of the lateral surface.H o l o t y p e : Right tecnomorphic valve from sample 416-30/6 ñ

Pl. II, Fig. 8.

D i a g n o s i s : Small. Strong posterodorsal bulb-likenode. Median ridge completely developed.R e m a r k s : The subspecies differs from S. multicosta-tus multicostatus in its smaller size, the stronger poster-odorsal node and the longer median ridge.

Genus Cuphithis gen. nov.

E t y m o l o g y : cupa, lat. ñ tub; according to the tub-like crista.T y p e s p e c i e s : Lennukella hendricksi Schallreuter, 1985.

D i a g n o s i s : Medium-sized. Unisulcate with distinct,oblique, funnel-like sulcus and pronounced preadducto-rial node. Large indistinctly delimited posteroventral lobetopped by a longitudinal crista which continues anteri-orly. Surface reticulate or granulose and tuberculate.R e m a r k s : Similar to Henningsmoenia Schallreuter,1964 with less funnel-like sulcus and less distinct node,a posteroventral lobe is not developed. In both the typespecies and H. billingensis the crista is only short andwith spine-like posterior ends. H. ? wenningstedtensis isvery similar to the type species but has a much longercrista (Schallreuter 1984, Figs 1.4ñ5). This species isconsidered to be transitional to Pseudostrepula ? acuta(Bonnema, 1909) in which the crista branches off pos-teriorly (Schallreuter 1976, Pl. 6, Figs 9ñ10).

Cuphithis liobqua sp. nov.Pl. II, Figs 9ñ10

E t y m o l o g y : Artificially formed from obliqua, lat. ñ oblique.H o l o t y p e : Posteriorly incomplete right tecnomorphic valve from

sample 416-10/5a ñ Pl. II, Fig. 10.

D i a g n o s i s : Up to 0.73 mm. Crista gently converg-ing towards dorsal margin in posterior direction. Surfacereticulate and tuberculate.

R e m a r k s : C. hendricksi from a Rogˆ sandstone ge-schiebe (upper Lower Ordovician) of Westphalia differsfrom the new subspecies mainly by its crista which runsmore parallel to the dorsal margin (Schallreuter 1985,Pl. 3, Figs 1ñ2).

Genus Telegraphia Olempska, 1994

Telegraphia novasemljaensis sp. nov.Pl. IV, Fig. 8

E t y m o l o g y : After Novaya Zemlya.Holotype. Right tecnomorphic valve from sample 416-5/2 ñ Pl. IV,

Fig. 8.

D i a g n o s i s : Up to 0.74 mm. Outline distinctly pre-plete. In the ventral half of the valve and posteroventrallyto the S2 two ridges converge but do not meet in ventraldirection.R e m a r k s : The type species, Telegraphia prima Ole-mpska, 1994 from the Early Ordovician of the HolyCross Mountains, is characterized by three ridges behindthe S2; the two posterior ridges are paralleling each oth-er and are ventrally connected. Telegraphia neglecta(Sarv, 1959) from the Late Ordovician of the Baltic areais more similar to T. novasemljaensis, but differs by itsdistinctly preplete outline and by the ridges that almostparallel each other (Sarv 1959, Pl. 15, Figs 5ñ8; Meidla1996, Pl. 7, Fig. 1). The shorter ridges may be a juve-nile feature.

Subfamily We h r l i i n a e Schallreuter, 1965

Genus Platybolbinoides gen. nov.

E t y m o l o g y : After the similarity with Platybolbina.T y p e s p e c i e s : Platybolbinoides guttasulcata sp. nov.

D i a g n o s i s : Medium-sized. Preplete. Unisulcate.Muscle scar located in a weak sulcal depression withinthe ventral half of the valve. Scar pit-like, ventrally slight-ly curved in anterior direction. Lateral surface reticulateexcept for the two cardinal corner fields.R e m a r k s : The species seems to belong to the Wehrli-inae which are characterized by a velum with a periph-erical row of spines. The latter forms a rake-like antralsculpture in females. The new genus differs from all otherknown genera of the subfamily by its weak sulcus andthe lacking lobale features. The genus is homeomorphicwith Platybolbina Henningsmoen, 1953 but it is distinctby the marginal surface with a comparatively high posi-tion of the velum and the spinose border of the latter.

Platybolbinoides guttasulcata sp. nov.Pl. III, Fig. 1

E t y m o l o g y : After the drop-like sulcus.H o l o t y p e : Anterodorsally incomplete right valve from sample 416-

10/5a ñ Pl. III, Fig. 1.

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D i a g n o s i s : As for genus which is currently mono-typic.

Family T e t r a d e l l i d a e Swartz, 1936Subfamily P e r s p i c i l l i n a e Schallreuter, 1966

Genus Sigmobolbina Henningsmoen, 1953

Sigmobolbina w-formis sp. nov.Pl. IV, Fig. 9añb

E t y m o l o g y : After the lobes and the histiovelum constituting a w.H o l o t y p e : Right female valve from sample 416-30/6, IGiG ñ Pl. IV,

Fig. 9añb.

D i a g n o s i s : Up to 0.78 mm. Long, flat and almost ver-tical preadductorial lobe (L2) situated in front of a longsigmoidal sulcus (S2). Anterior to the L2, an indistinct,narrow, ridge-like, vertical L1 is separated from both theL2 and the histiovelum by weak, narrow furrows. Poster-oventral lobe (L3) flat, obliquely extending towardsstraight dorsal margin. It is separated from the posteroven-tral part of the histium by a flat semisulcus (S3). Velumforms anterocentrally an admarginal loculus and ventral-ly the inner antral fence of an admarginal histial antrumwhich diverges ventrally from the free margin in posteri-or direction but the velum does not form a rudimentaryantrum. Lobate area antero- and posteroventrally corrugatereticulate. Ventral side of velar keel weakly reticulate.R e m a r k s : The species is characterized by the distinct-ly separated preadductorial and posteroventral lobes; to-gether with the histium they serve for the quadrilobateappearance of the valve.

Genus Podolibolbina Abushik and Sarv, 1983

Podolibolbina luetkei sp. nov.Pl. III, Fig. 3

E t y m o l o g y : After the L¸tke-Land on Ní Nowaya Zemlya.H o l o t y p e : Right female valve from sample 416-33/14 ñ Pl. III,

Fig. 3.

D i a g n o s i s : Up to 1.00 mm. Distinct laterovelar fur-row in posterior half of the female valve. Lateral surfacecoarsely tuberculate.R e m a r k s : In the holotype the three anteroventral lo-culi typical for Podolibolbina seem to be obscured byrock material. The type species of the genus, P. podoli-ca Abushik ñ Sarv, 1983, reaches nearly the same size(0.95 mm), but differs from the new species mainly bythe weak or missing laterovelar furrow in the females.Furthermore, the lateral surface is not tuberculate(Abushik ñ Sarv 1983, Pl. 2, Figs 9, 11). By the weakor missing laterovelar furrow P. podolica resembles Fo-ramenella and might even be its predecessor. With thedistinct laterovelar furrow and the coarse-tuberculate lat-eral surface the new species is also similar to the type

species of Deefgella, D. dajsiveteri Schallreuter, 1981,but the latter differs markedly by having developed sev-eral conical nodes (Schallreuter 1981, Fig. 7).

Subfamily G l o s s o m o r p h i t i n a e Hessland, 1954Tribus G l o s s o m o r p h i t i n i Hessland, 1954

Genus Ctenomorphites gen. nov.

E t y m o l o g y : Formed by a combination of the generic namesCtenobolbina and Glossomorphites.

Ty p e s p e c i e s : Ctenomorphites bila sp. nov.

D i a g n o s i s : Medium-sized. Distinctly preplete. Un-isulcate. S2 long and only slightly sigmoidal. Long pre-adductorial lobe (L2). Posteroventral lobe (L3v) high andsimilar to L2 extending almost perpendicular to straightdorsal margin. Lobes forming the maximum width of thevalve. Histial ridge from anterodorsal region to postero-central region. Thick velar bulge from anterodorsal re-gion to posteroventral region.R e m a r k s : The monotypic Lingulibolbina Melnikova,1986 from the Ordovician of Kazakhstan is very similarin its lobation, but contrary to Ctenomorphites it has ashort drop-like preadductorial node which is distinctly setoff from a flat L1 (Melnikova 1986, Pl. 8, Figs 4ñ5).

Concerning shape, outline, strong development of theL3v, and especially the bulgy velar sculpture, Ctenom-orphites is similar to Lenatella which has been assignedto the Egorovellidae (Schallreuter ñ Kanygin ñ Hinz-Schallreuter 1999, 278, Fig. 4). However, relationshipsto that genus and/or family could not be excluded.

Ctenomorphites brevis sp. nov.Pl. I, Fig. 6.

E t y m o l o g y : Brevis, short ñ according to the relatively high L:Hratio.

H o l o t y p e : Right valve from sample 416-30/6 ñ Pl. I, Fig. 6.

D i a g n o s i s : As for genus which is currently mono-typic.

Genus Vittella Schallreuter, 1964

Vittella noze sp. nov.Pl. III, Fig. 10; Pl. IV, Fig. 3

E t y m o l o g y : Artificially formed from Novaya Zemlya.H o l o t y p e : Right female valve from sample 416-10/5a ñ Pl. III,

Fig. 10.

D i a g n o s i s : Up to 0.86 mm. Deep and narrow sul-cus distinct. Posteroventral lobe narrow and withoutspine. Dolon restricted to anteroventral region. Surfacereticulate.R e m a r k s : Most similar Among all Vittella speciesV. gullhoegensis Schallreuter, 1984 from the Middle Or-dovician of Sweden ist most similar to the new species.

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V. gullhoegensis is only slightly larger (0.98 mm) and hasalso a short dolon which is restricted to the anteroven-tral region. The sulcus, however, is less distinct.

Tribus H i p p u l i n i Schallreuter, 1983

Genus Hippula Tromelin and Lebesconte, 1876

Subgenus Hippula (Interruptula) subgen. nov.

E t y m o l o g y : after the interrupted histiovelum.T y p e s p e c i e s : Hippula (Interruptula) interrupta sp. nov.

D i a g n o s i s : L1 developed bulb-like at dorsal marginand as an elongate lobe in its ventral half. L2 bulb-likedorsally (just above mid-length), its narrow ventral endconnected with L1. Posteroventral lobe (L3) distinct. His-tiovelum interrupted anteroventrally.

Hippula (Interruptula) interrupta sp. nov.Pl. I, Fig. 7; Pl. III, Fig. 9

E t y m o l o g y : after the interrupted histiovelum.H o l o t y p e : Left tecnomorphic valve from sample 416-10/5a ñ Pl. I,

Fig. 7.

D i a g n o s i s : At least up to 0.88 mm.

Subgenus Hippula (Cetona) Schallreuter, 1964

Hippula (Cetona) brevisulcata sp. nov.Pl. IV, Fig. 7

E t y m o l o g y : According to the short sulcus.H o l o t y p e : Right tecnomorphic valve from sample 416-30/6 ñ

Pl. IV, Fig. 7.

D i a g n o s i s : At least up to 1.01 mm. Sulcus short andrestricted to dorsal half. Both preadductorial node andposteroventral lobe indistinct. Velar flange parallelingventral margin. Surface indistinctly reticulate.R e m a r k s : The species is characterized by its shortsulcus and the indistinct lobes. The similar H. (C.) serraSchallreuter, 1984 from a Backsteinkalk geschiebe (Mid-dle Ordovician) of Northern Germany differs by the ve-lar ridge that diverges posteriorly from the free margin(Schallreuter 1984, Fig. 4.1).

Hippula (Cetona ?) facies sp. nov.Pl. I, Fig. 10; Pl. II, Fig. 6; Pl. IV, Fig. 5

E t y m o l o g y : According to the two lobes, the sulcus and the ve-lum consituting a sculpture similar to a face.

H o l o t y p e : Left tecnomorphic valve from sample 416-10/5a ñ Pl. I,Fig. 10.

D i a g n o s i s : At least up to 1.11 mm. Shape elongate.Sulcus not indistinctly delimited. Preadductorial node andposteroventral lobe similarly developed as nodes, poster-

oventral lobe something higher and with short ridge-liketop. Velar flange equally developed from anterocentral tocentroventral region. Surface weakly reticulate.R e m a r k s : The species is characterized by the distinctpreadductorial node and posteroventral lobe. However,because of the broken velar ridge the subgeneric assign-ment is uncertain.

Genus Ctenyginia gen. nov.

E t y m o l o g y : Formed by combination of the generic names Cten-onotella and Kanyginia.

T y p e s p e c i e s : Ctenyginia crassivelata sp. nov.

D i a g n o s i s : Small. Unisulcate. S2 long and onlyslightly sigmoidal. Long preadductorial lobe of aboutthe same length as the posteroventral lobe. Strong,broad histiovelum that reaches the same height as thelobes.R e m a r k s : Concerning lobation, Ctenyginia is verysimilar to Ctenomorphites gen. nov. The main differencelies in the development of the stout histiovelum inCtenyginia which resembles that of Kanyginia Schallreu-ter ñ Kanygin, 1992.

Ctenyginia crassivelata sp. nov.Pl. III, Fig. 8

E t y m o l o g y : According to the stout histiovelum.H o l o t y p e : Posteriorly incomplete right valve from sample 416-

10/5a ñ Pl. III, Fig. 8.

D i a g n o s i s : As for genus which is currently mono-typic. Length of holotype (without posterior end)0.64 mm.

Genus Spinohippula Vannier, Kr˘ta and Marek, 1977

Spinohippula ? biserrata sp. nov.Pl. I, Fig. 5

E t y m o l o g y : Serra, lat. ñ saw, according to the saw-like dentitionof the adventral sculpture.

H o l o t y p e : Left tecnomorphic valve from sample 416-10/5a ñ Pl. I,Fig. 5.

D i a g n o s i s : Length at least up to 0.54 mm. Unisul-cate with short oblique sulcus (S2) in dorsal half. Twoflat indistinct lobes (L2, L3) on both sides of the sulcusthat fuse ventrally. Thick velar bend from anterocentralto posterocentral region, with two rows of triangular den-ticles in the plane of the lateral surface; tips of denticlesvis-à-vis and alternating. Surface indistinctly reticulate.R e m a r k s : The type species, Spinohippula esurialisfrom the Bohemian Ordovician, the only yet known spe-cies, is nonsulcate and differs from the new species byhaving only the outer row of triangular spines developed(Vannier ñ Kr˘ta ñ Marek 1977, Pls 50, 52, 54, 56).

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Genus Quaca gen. nov.

E t y m o l o g y : Formed artificically from quadricarinata.T y p e s p e c i e s : Quaca muca sp. nov.

D i a g n o s i s : Unisulcate. S2 long and sigmoidal, be-coming weak ventrally. Both preadductorial node andposteroventral lobe indistinctly flat. Anterocentral to cen-troventral regions between keel-like histial ridge and freemargin with three perimarginal ridges. A single short per-imarginal ridge occurs in ventral half of posterior part.Marginal sculpture developed as a row of spines that be-come flange-like posteroventrally. Lateral surface tuber-culate and reticulate, except for a flat and smooth, indis-tinct, node-like swelling in the anterodorsal region.R e m a r k s : The genus resembles Hippula but lacks thecharacteristical histiovelum with the special compart-ments.

Quaca muca sp. nov.Pl. I, Fig. 3

E t y m o l o g y : Formed artificically from multicarinata.H o l o t y p e : Left tecnomorphic valve from sample 416-30/6 ñ Pl. I,

Fig. 3.

D i a g n o s i s : Length at leat up to 0.91 mm. Preadduc-torial bulb and posteroventral lobe weakly developed.Ventral surface above fine admarginal ridge characterizedby three ridges with the uppermost ridge forming a his-tial edge. Surface reticulate and with scattered tubercles.

Quaca pura sp. nov.Pl. III, Fig. 7; Pl. IV, Fig. 6

E t y m o l o g y : purus, Latin ñ poor; according to the fewer admar-ginal ridges compared with the type species.

H o l o t y p e : Left tecnomorphic valve from sample 416-10/5a ñPl. IV, Fig. 6.

D i a g n o s i s : Length at leat up to 0.76 mm. No distinctpreadductorial node and posteroventral lobe. Ventral sur-face with two ridges, but no histial ridge. Surface wrin-kled-reticulate.R e m a r k s : Q. pura differs from Q. muca by lack ofthe distinct preaductorial bulb, histial ridge, and tuber-culation; instead its surface is wrinkled reticulate.

Genus Nowehrlina gen. nov.

E t y m o l o g y : Combined from Nowaya Zemlya and the genus nameWehrlina.

T y p e s p e c i e s : Nowehrlina sperata sp. nov.

D i a g n o s i s : Small with relatively weak, long, strong-ly sigmoidal sulcus. Preadductoral node and posteroven-tral lobe distinct. L1 developed as a strong sperum. Do-lon fairly strongly convex in anterocentral, anteroventraland centroventral regions, forming a long botulate an-trum. Distinct laterovelar furrow.

R e m a r k s : The new genus differs from WehrlinaSchallreuter, 1964 by the distinct preadductorial node, theposteroventral lobe, the development of a sperum, thedistinct laterovelar furrow, and the strongly convex do-lon. The assignment to the Wehrliinae is uncertain.

Nowehrlina sperata sp. nov.Pl. II, Fig. 11

E t y m o l o g y : According to the anterodorsal sperum.H o l o t y p e : Posteriorly incomplete left female valve from sample

416-30/6 ñ Pl. II, Fig. 11.

D i a g n o s i s : As for genus which is presently mono-typic.

Family Monotiopleuridae Guber and Jaanusson, 1964

Genus Tior Schallreuter, 1998

Tior verticalis sp. nov.Pl. IV, Fig. 4

E t y m o l o g y : After the vertical sulcus.H o l o t y p e : Left tecnomorphic valve (posterior part lost after scan-

ning) from sample 416-30/6 ñ Pl. IV, Fig. 4.O c c u r r e n c e : Sample 416-30/6.

D i a g n o s i s : Up to 0.73 mm. Sulcus (S2) extendingperpendicular to dorsal margin and in some distance fromthe latter. Dorsally it is trumpet-like opened, but rathernarrow ventrally. Surface weakly and irregularly reticu-late.R e m a r k s : The type species T. altior (Schallreuter,1993) from an upper Viruan geschiebe of Ní Germanyis very similar to the new species concerning outline aswell as in the construction and position of the S2 withits trumpet-like dorsal opening. T. altior is, however, larg-er (0.81 mm) and more distinctly reticulate. The sulcusin T. verticalis is ventrally narrower than in T. altior.

T. fastidiosus (Sarv, 1959) from the Kundan of Esto-nia possesses a broader sulcus and a less convex poste-rior end (Sarv 1959, Pl. 31, Figs 1ñ4; Schallreuter 1999,Pl. 3, Fig. 5).

? Genus Disparigonya Schallreuter, 1985

R e m a r k s : The type species, Disparigonya voigtiSchallreuter, 1985 from an late Viruan (D

2) geschiebe of

Northern Germany, seems to be a junior synonym ofHaploprimitia kogermani ÷pik, 1937 from the Kukrusebeds (C

2) of Estonia where it is a very rare faunal ele-

ment (÷pik 1937, Pl. 1, Fig. 2).

Disparigonya ? umbona sp. nov.Pl. III, Fig. 5; Pl. IV, Fig. 10

E t y m o l o g y : After the umbo.H o l o t y p e : Right valve from sample 416-30/4 ñ Pl. IV, Fig. 10.O c c u r r e n c e : Samples 416-30/4 and 416-30/6.

F13_x199_212_Schali_10_3P.p65 18.10.2001, 0:49205


D i a g n o s i s : Up to 1.42 mm. Strongly postplete.Weak, broad sulcal depression (S2) in the center of dor-sal half. Posterior half of valve forming a broad infla-tion which is wider than the anterior half. A broad (long.)triangular umbo located posterodorsally to the S2 givesthe dorsal margin its characteristic kink. Its anterior partis separated by a very weak depression within the weak-ly bulb-like inflation of the whole lobal element.R e m a r k s : The type species differs mainly by its small-er size, the amplete outline, the weak but distinct sulcusand the missing umbo (÷pik 1937, Pl. 1, Figs 2ñ3; Pl. 13,Fig. 30; Sarv 1959, Pl. 31, Figs 9ñ11). These differenc-es are so fundamental that the new species could be re-ferred to the genus with great reservations only.

Disparigonya ? umbona sp. nov. also resembles Me-dianella Neckaja, 1966 but without seeing its inner sidethe relationship is speculative only.

Family unknown

Genus Hallatina Ivanova in Chugaeva, Rozman andIvanova, 1964

Hallatina ? bulbata sp. nov.Pl. II, Fig. 5

E t y m o l o g y : According to the dorsal bulb in the anterior half.H o l o t y p e : Left valve from sample 416-33/14 ñ Pl. II, Fig. 5.

D i a g n o s i s : Up to 0.81 mm. Outline slightly preplete.S2 developed only as a sulcal depression in the anteriorcentrodorsal region. Distinct velar ridge near free mar-gin. Outer surface reticulate.R e m a r k s : The other known species of the genus,H. chanae Ivanova in Chugaeva ñ Rozman ñ Ivanova,1964, H. orlovi Ivanova in Chugaeva ñ Rozman ñ Ivano-va, 1964 and H. dentata Kanygin, 1967 from the Ordov-ician of NEí Siberia becomes much larger (1.98 ñ 2.48mm) and has a deeper S2 (Chugaeva ñ Rozman ñ Ivano-va 1964, Pl. 24, Figs 8ñ9; Kanygin 1967, Pl. 6, Figs 2ñ11).

Genus Estonaceratella Schallreuter, 1984

Estonaceratella aspinata sp. nov.Pl. III, Fig. 4

E t y m o l o g y : After the missing ventrolateral spine.H o l o t y p e : Left valve from sample 416-30/6 ñ Pl. III, Fig. 4.

D i a g n o s i s : Up to 0.70 mm. Dorsal area with distinctbroad (long.) umbo that forms an epicline dorsum. Lat-eral surface without any spine.R e m a r k s : The type species E. estona (Sarv, 1962) ischaracterized by a spine in the posteroventral region, buthas a less distinct umbo (Schallreuter 1986, Pl. 6, Fig. 4).

Discussion

The Ordovician ostracod faunas of Baltica and Siberiaare quite different and have only few genera in common.Before that background the faunas from Novaya Zem-lya with their interposing role become extremely signif-icant. The yet described Ordovician ostracods fromNovaya Zemlya represent about half of the entire collec-tion. All species are new, but the genera show relationsto both the Baltic region and Siberia. Typical Baltic gen-era are Platybolbina, Cystomatochilina, Chilobolbina,Levisulculus, Havlicekites, Telegraphia, Sigmobolbina,Tior, and Estonaceratella. Typical Siberian taxa are theegorovellids, the cherskiellids, and the genera Bulbosul-culus, Parajonesites, and Hallatina. Some genera showrelations to Laurentia, e.g., Levisulculus, and Platybol-bina (Kesling 1960a, b).

In the latest palaeogeographic reconstructions Balti-ca and Siberia take positions on both sides of the Iape-tus Ocean, Siberia at the equator, Baltica at about 30∞ S(e.g., Lehnert ñ Hinz-Schallreuter ñ Krueger 1999,Fig. 4). Until now Novaya Zemlya has been consideredto be part of Baltica. The ostracod faunas, however, castsome doubt on this assumption and may suggest thatNovaya Zemlya was a separate microcontinent, posi-tioned somewhere between Baltica and Siberia, but closerto Baltica.

Submitted December 13, 2000

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Explanation of plates

P l a t e I1 ñ Cystomatochilina flotowi sp. nov. Holotype, posteriorly incomplete tecnomorphic right valve, length 0.71 mm. Sample 416-30/6.2 ñ Cystomatochilina flotowi sp. nov. Paratype, posteriorly incomplete tecnomorphic right valve, length 0.85 mm. Sample 416-30/6.3 ñ Quaca muca gen. et sp. nov. Holotype, tecnomorphic left valve, length 0.91 mm. Sample 416-30/6.4 ñ Havlicekites fissuratus gen. et sp. nov. Holotype, tecnomorphic left valve, length 1.85 mm. Sample 416-30/6.5 ñ Spinohippula biserrata sp. nov. Holotype, tecnomorphic left valve, length 0.54 mm. Sample 416-10/5a.6 ñ Ctenomorphites brevis gen. et sp. nov. Holotype, tecnomorphic right valve, length 1.13 mm. Sample 416-30/6.7 ñ Hippula (Interruptula) interrupta subgen. et sp. nov. Holotype, tecnomorphic left valve, length 0.88 mm. Sample 416-10/5a.8 ñ Tetrada ? indistincta sp. nov. Holotype, tecnomorphic left valve, length 0.54 mm. Sample 4126-30/6.9 ñ Parajonesites sawina sp. nov. Holotype, right valve, length 0.78 mm. Sample 416-30/6.10 ñ Hippula (Cetona) ? facies sp. nov. Holotype, tecnomorphic left valve, length 1.11 mm. Sample 416-10/5a.

P l a t e I I1 ñ Levisulculus posteroventrolobatus sp. nov. Holotype, female right valve, length 0.63 mm. Sample 416-30/6.2 ñ Octocristatia octo gen. et sp. nov. Holotype, tecnomorphic left valve, length 0.73 mm. Sample 416-10/5a.3 ñ Strephites multicostatus gen. et sp. nov. Paratype, anterodorsally incomplete tecnomorphic left valve, length 0.97 mm. Sample 416-33/14.4 ñ Octocristatia octo gen. et sp. nov. Paratype, anterodorsally incomplete tecnomorphic right valve, length 0.64 mm. Sample 416-10/5a.5 ñ Hallatia ? bulbata sp. nov. Holotype, left valve, length 0.81 mm. Sample 416-33/14.6 ñ Hippula (Cetona) ? facies sp. nov. Paratype, tecnomorphic left valve, length 1.05 mm. Sample 416-10/5a.7 ñ Bulbosulculus daytylus sp. nov. Holotype, posteriorly incomplete female right valve, length 0.62 mm. Sample 416-30/6.8 ñ Strephites multicostatus binodus subsp. nov. Holotype, tecnomorphic right valve, length 0.56 mm. Sample 416-30/6.9 ñ Cuphithis liobqua gen. et sp. nov. Paratype, anteroventrally incomplete tecnomorphic right valve, length 0.59 mm. Sample 416-10/5a.10 ñ Cuphithis liobqua gen. et sp. nov. Holotype, posteriorly incomplete tecnomorphic right valve, length 0.73 mm. Sample 416-10/5a.11 ñ Nowehrlina sperata gen. et sp. nov. Holotype, posteriorly incomplete female left valve, length 0.68 mm. Sample 416-30/6.

P l a t e I I I1 ñ Platybolbinoides guttasulcata sp. nov. Holotype, female (?) right valve, length 0.98 mm. Sample 416-10/5a.2 ñ Strephites multicostatus gen. et sp. nov. Holotype, tecnomorphic right valve, length 1.37 mm. Sample 416-33/14.3 ñ Podolibolbina luetkei sp. nov. Holotype, female right valve, length 1.00 mm. Sample 416-33/14.4 ñ Estonaceratella aspinata sp. nov. Holotype, left valve, length 0.66 mm. Sample 416-30/6.5 ñ Disparigonya ? umbona sp. nov. Paratype, right valve, length 1.09 mm. Sample 416-30/6.6 ñ Chilobolbina pseudola sp. nov. Holotype, right tecnomorphic valve, length 1.05 mm. Sample 416-10/5a.7 ñ Quaca pura sp. nov. Paratype, posteriorly incomplete tecnomorphic left valve, height 0.44 mm. Sample 416-10/5a.8 ñ Ctenyginia crassivelata gen. et sp. nov. Holotype, posterodorsally incomplete tecnomorphic right valve, length >0.64 mm. Sample 416-10/5a.9 ñ Hippula (Interruptula) interrupta subgen. et sp. nov. Paratype, posteriorly incomplete tecnomorphic left valve, height 0.58 mm. Sample 416-10/5a.10 ñ Vittella noze sp. nov. Holotype, female rigtht valve, length 0.86 mm. Sample 416-10/5a.

P l a t e I V1 ñ Platybolbina (R.) temperata petermanni subsp. nov. Holotype, posteroventrally incomplete female (?) left valve, length 1.22 mm. Sample 416-33/14.2 ñ Pseudostrepula ? severa sp. nov. Holotype, posteroventrally incomplete tecnomorophic left valve, length 0.69 mm. Sample 416-30/4.3 ñ Vittella noze sp. nov. Paratype, slightly distorted female right valve, length 0.78 mm. Sample 416-10/5a.4 ñ Tior verticalis sp. nov. Holotype, left valve, length 0.73 mm. Sample 416-30/16.5 ñ Hippula (Cetona) ? facies sp. nov. Paratype, tecnomorphic left valve, length 0.80 mm. Sample 416-10/5a.6 ñ Quaca pura sp. nov. Holotype, tecnomorphic left valve, height 0.73 mm. Sample 416-10/5a.7 ñ Hippula (Cetona) brevisulcata sp. nov. Holotype, tecnomorphic right valve, length 1.01 mm. Sample 416-30/6.8 ñ Telegraphia novasemljaensis sp. nov. Holotype, tecnomorphic right valve, length 0.71 mm. Sample 416-5/2.9 ñ Sigmobolbina w-formis sp. nov. Holotype, female right valve in lateral (a) and ventral views (b), length 0.78 mm. Sample 416-30/6.10 ñ Disparigonya ? umbona sp. nov. Holotype, right valve, length 1.42 mm. Sample 416-30/4.

All figures stereo-pairs.

OrdoviËtÌ ostrakodi z NovÈ ZemÏ

Z ordoviku NovÈ ZemÏ je pops·no celkem 31 nov˝ch druh˘ a poddruh˘ ostrakod˘ pat¯ÌcÌch do celkem 28 rod˘ nebo podrod˘ (z toho 10 nov˝ch).NÏkte¯Ì ostrakodi jevÌ vztahy k paleokontinentu Baltika, jinÈ druhy k Siberii. Tyto ˙daje silnÏ podporujÌ paleogeograficky mezilehlou pozici NovÈZemÏ mezi tÏmito dvÏma paleokontinenty v ordovickÈm obdobÌ.

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Ordovician ostracodes from Novaya Zemlya · 2014-01-03 · ter 1993, Pl. 11A, Fig. 1, Pl. 14A, and Pl. 1B, Fig. 2, Pl.2A, Figs 1ñ3, Pl. 3, Fig. 1, Pl. 8A, Pl. 9). In this re-spect,