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    The Pleistocene to Holocene Transition and Human Economy in Southwest Asia: The Impact ofthe Younger DryasAuthor(s): A. M. T. Moore and G. C. HillmanSource: American Antiquity, Vol. 57, No. 3 (Jul., 1992), pp. 482-494Published by: Society for American ArchaeologyStable URL: http://www.jstor.org/stable/280936.

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  • 7/26/2019 Moore a. M. T. and Hillman C. G. the Ple

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    THE PLEISTOCENE TO HOLOCENE TRANSITION

    AND HUMAN ECONOMY IN SOUTHWEST ASIA:

    THE

    IMPACT

    OF

    THE

    YOUNGER

    DRYAS

    A. M. T. Moore and G. C.

    Hillman

    We

    present

    new evidence

    suggesting

    that

    the

    Late

    Glacial

    worldwide

    episode

    of

    cooling

    known as

    the

    Younger

    Dryas (ca.

    11,000-10,000

    B.P.)

    had a

    significant

    impact

    on

    climate,

    vegetation,

    and human

    economy

    in

    southwest

    Asia. In the Levant

    a

    new

    pollen

    core extracted

    from

    Lake

    Huleh and

    plant

    remains

    from

    the

    early village

    of

    Abu

    Hureyra

    1

    indicate that

    forest gave

    way

    to

    steppe

    in

    response

    to

    the onset

    of

    drier

    climatic

    conditions

    contemporary

    with the

    Younger

    Dryas.

    Similar

    effects

    may

    be seen in

    pollen

    cores

    from

    elsewhere

    in southwest

    Asia.

    This alteration

    in

    climate

    and

    vegetation

    obliged

    the inhabitants

    of

    Abu

    Hureyra

    to

    modify

    their

    plant

    gathering,

    and

    led to

    significant

    disruptions

    in

    culture

    and

    settlement

    over a wide area. We

    argue

    that

    the stresses

    induced

    by

    these

    events

    were a

    contributing

    factor

    in the

    subsequent development

    of

    agriculture

    in southwest

    Asia.

    Presentamos

    nuevas

    evidencias

    que

    indican

    que

    el

    episodio

    de

    enfriamiento

    global

    a

    fines

    del

    periodo

    glacial

    conocido

    como

    Younger

    Dryas (ca.

    11,000-10,000

    A.P.)

    tuvo

    un

    significativo

    impacto

    en el

    clima,

    la

    vegetaci6n

    y

    la

    economia

    humana

    en el suroeste de Asia. En

    el

    Levante,

    una nueva columna

    de

    polen

    extraida del

    Lago

    Huleh

    y

    restos botdnicos

    recuperados

    en la

    aldea

    temprana

    de Abu

    Hureyra

    1 indican

    que

    los

    bosques

    fueron

    reemplazados por estepas

    en

    respuesta

    a condiciones

    climdticas mds secas

    contempordneas

    con

    el

    Younger

    Dryas.

    Consecuencias

    semejantes

    se observan

    en columnas

    de

    polen

    provenientes

    de

    otros

    lugares

    en

    el suroeste

    de Asia.

    Esta

    alteraci6n

    en el clima

    y

    la

    prdctica

    de recolecci6n

    de

    plantas

    produjo significativos

    cambios

    en la cultura

    y

    el asentamiento

    en un

    drea extensa.

    Sostenemos

    que

    las tensiones

    inducidas

    por

    estos hechos

    contribuyeron

    al

    subsiguiente

    desarrollo

    de la

    agricultura

    en

    el suroeste

    de Asia.

    The

    transition

    from

    hunting

    and

    gathering

    to

    farming

    in

    southwest

    Asia

    coincided

    with the

    environmental

    changes

    that

    marked the

    close of the Pleistocene:

    a worldwide increase

    in

    temperature

    that

    melted ice sheets

    and caused

    sea

    levels to

    rise,

    alterations

    in

    atmospheric

    circulation

    systems,

    and

    shifts

    in

    vegetation

    zones.

    The climatic

    amelioration

    was an

    uneven

    process,

    with

    episodes

    of

    increased

    warmth

    alternating

    with

    reversions

    to

    cooler

    conditions.

    Our aim

    in

    this

    paper

    is to

    examine

    new

    evidence

    thatat least

    one

    major

    episode

    of

    cooling,

    the

    Younger Dryas,

    apparently

    had

    a

    profound

    effect on the

    environment of southwest

    Asia,

    and

    contributed

    significantly

    to

    the

    adjustments

    in

    human

    adaptations

    that

    resulted

    in

    the

    development

    of

    agriculture.

    The

    region

    within

    southwest Asia

    with the

    most

    substantial

    record of

    environmental

    and

    cultural

    change

    is the Levant.

    The

    Younger

    Dryas

    climatic

    episode

    occurred there

    during

    the second

    stage

    of

    the

    Epipaleolithic,

    Epipaleolithic

    2

    (ca.

    12,500-10,000

    B.P.),

    one constituent

    culture

    of

    which

    was

    the Natufian

    in Palestine. This

    stage

    is

    important

    because

    it

    was the

    last

    period

    of

    hunting

    and

    gathering

    before

    the

    dvent of

    agriculture;

    it

    was

    also

    during

    the

    Epipaleolithic

    2

    that

    the inhabitants

    of some

    sites

    on

    the

    middle

    Euphrates

    River and

    in

    Palestine

    adopted

    a more

    sedentary

    mode

    of

    life. The

    changes

    in

    economy

    and settlement that

    took

    place

    then

    are

    obviously

    of

    crucial

    importance

    for

    understanding

    the circumstances

    in

    which

    agriculture

    developed.

    THE

    YOUNGER

    DRYAS

    The

    Younger

    Dryas

    was first

    recognized

    in

    the

    pollen

    record

    of

    northern

    Europe.

    During

    the Late

    Glacial,

    as the

    temperature

    rose and the

    glaciers began

    to

    retreat,

    the tundra

    vegetation

    was

    replaced

    by

    birch and

    pine

    woodland;

    these

    trends

    began

    during

    the

    Bo6lling

    nd Allerod

    pollen

    phases

    (Iversen

    A. M.

    T.

    Moore,

    The Graduate

    School,

    Yale

    University,

    1504A

    Yale

    Station,

    New

    Haven,

    CT 06520-7425

    G.

    C.

    Hillman,

    Department

    of

    Human

    Environment,

    Institute

    ofArchaeology,

    University

    College

    London,

    31-

    34

    Gordon

    Square,

    London,

    WC1H

    OPY,

    England

    AmericanAntiquity, 57(3), 1992,pp. 482-494.

    Copyright

    ?)

    1992

    by

    the

    Society

    for

    American

    Archaeology

    482

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    REPORTS

    1954).

    Then

    in

    the

    ensuing Younger

    Dryas (pollen

    zone

    III)

    the

    temperature

    fell

    sharply,

    the

    glaciers

    advanced

    once

    more,

    and the woodlands

    retreated

    southward,

    giving way

    to

    open

    tundra

    charac-

    terized

    by Dryas

    octopetala

    from which the

    name of the

    episode

    is

    derived. This

    phase

    lasted

    approximately

    1,000

    radiocarbon

    years,

    from

    11,000

    to

    10,000

    B.P.

    (Berger 1990).1

    Radiocarbon

    dates for the end of the Younger Dryas are subject to uncertainty because they fall in a period during

    which the calibration

    curve levels

    off

    for at least three

    centuries;

    the

    age

    of this

    event

    has,

    however,

    been established

    at

    11,100 dendroyears

    B.P.

    (Becker

    and Kromer

    1986;

    Becker et

    al.

    1991).

    The

    pollen

    evidence

    suggests

    that

    the

    climate

    turned

    so cold

    during

    the

    Younger

    Dryas

    that

    it

    approached

    the conditions

    of the full

    glacial.

    This has been confirmed

    by

    studies of fossil

    coleoptera

    in

    Britain

    (Coope

    1977:330).

    In

    the

    succeeding

    Preboreal and Boreal

    phases

    the

    temperature

    rose once

    more

    and the forest

    advanced

    rapidly

    northward.

    The

    precise

    timing

    and effects of the rises and falls

    in

    temperature

    are still

    subjects

    of

    discussion;

    the evidence

    of

    fossil

    coleoptera,

    for

    example,

    while

    confirming

    the

    oscillatory

    nature

    of

    late

    glacial

    environmental

    change, suggests

    that the

    fluctuations

    in

    temperature may

    have

    preceded

    by

    several centuries

    the

    advances and retreats of the forest

    zones

    (Coope

    1975:167).

    The Younger Dryas period of cooling was originally defined in northern Europe, but parallel

    vegetation

    changes

    occurred

    elsewhere, making

    it a

    worldwide

    phenomenon.

    We therefore

    use

    the

    term

    Younger Dryas,

    not

    just

    for the advance

    of

    Dryas-dominated

    tundra in

    the

    far

    north,

    but

    also

    for all the other

    changes

    in

    vegetation

    that were induced

    by

    the same climatic

    episode.

    It is

    the one

    period

    of

    cooling

    of

    sufficient

    intensity

    and duration to be seen

    clearly

    in

    the

    stage

    1

    Late

    Glacial

    section

    in

    deep-sea

    cores

    from

    the Pacific

    (Shackleton

    and

    Opdyke 1976).

    It also

    appears

    as a

    well-

    defined

    episode

    in the Greenland

    ice

    cores

    (Dansgaard

    et al.

    1982:Figure

    1).

    Studies

    of

    coral

    reefs

    off Barbados have

    provided

    direct

    evidence

    of the effect on sea levels of the

    Younger Dryas

    cooling.

    The rise in

    sea level slowed

    sharply

    between

    11,000

    and

    10,500 B.P.,

    and

    then increased

    slightly

    from

    10,500

    to

    10,000

    B.P.

    (Fairbanks

    1989:639).

    The

    Younger Dryas

    coincided

    with

    the

    episode

    of

    major

    mammal

    extinctions

    in

    Eurasia

    and North

    America, lending weight

    to the

    hypothesis

    that

    such

    rapid

    climatic

    fluctuations

    contributed

    to

    those

    events.

    Haynes

    (1991:447)

    has

    argued

    that

    a

    brief, intense period of drought, corresponding in time to the Younger Dryas, was a factor in the

    demise

    of

    many

    species

    of the

    Rancho

    La Brea fauna

    in North America.

    Given that the

    Younger Dryas

    episode

    of

    cooling

    was a

    worldwide

    phenomenon,

    its

    effects should

    have been felt

    in southwest

    Asia,

    but

    pollen

    cores

    and

    sedimentological analyses

    have

    provided

    little indication

    that

    it had a

    significant

    influence

    on

    the environment

    there. The

    main

    reason

    for

    this is that

    the

    environmental

    sequences

    for

    the different

    regions

    of southwest

    Asia are still

    very

    coarsely

    delineated.

    Relatively

    few

    pollen

    cores

    have been

    analyzed,

    and

    they

    have not

    provided

    such

    detailed

    replicated

    sequences

    of

    vegetation

    change

    as those

    from northern

    Europe,

    North

    America,

    and

    elsewhere.

    Furthermore,

    they

    are

    dated

    with

    very

    few

    radiocarbon

    determinations.

    Pollen cores

    extracted

    from locations

    in

    the

    Levant,

    the

    Ghab

    section

    of the Orontes

    Valley

    and

    Lake

    Huleh,

    much of which

    today

    is a

    marsh,

    for

    example (Figure

    1),

    showed that

    the forest

    cover

    expanded during the Late Glacial (Niklewski and van Zeist 1970; van Zeist and Bottema 1982:

    Figure

    14.6).

    The Ghab

    core

    suggested

    that

    the

    forest

    expanded

    quite

    steadily.

    The Huleh

    sequence,

    analyzed

    by

    Tsukada

    and reviewed

    by

    van

    Zeist

    and

    Bottema,

    did

    indicate

    that

    the

    vegetation

    cover

    fluctuated

    during

    the

    period

    in

    which the

    forest

    was

    expanding,

    but this

    section

    of

    the

    core lacked

    radiocarbon dates.

    A

    general

    increase

    in

    forest

    cover, especially

    cedar,

    until

    sometime

    between

    12,000

    and

    11,000

    B.P.

    is

    again

    apparent

    at

    Karamik

    Batakhgi

    in

    western

    Anatolia,

    and the

    same

    could

    be inferred

    for

    oak

    forest

    at

    Sogot

    Golii,

    although

    with

    only

    two

    dates

    available

    in each

    case,

    exact correlation

    is difficult

    (van

    Zeist and

    Bottema

    1982:Figures

    14.4,

    14.14;

    van

    Zeist

    et

    al.

    1975).

    A

    steady

    increase

    in

    forest

    cover

    could

    also

    be

    seen

    in the

    cores

    from

    Lake

    Zeribar

    in

    the

    Zagros

    Mountains,

    but

    there it

    was

    delayed

    until

    the

    mid

    Holocene

    (van

    Zeist and Bottema

    1977:Figures

    Ib and

    II, 1982:Figure 14.2),

    reflecting

    the arrival

    of trees

    migrating

    from forest

    refuges

    remote from

    Zeribar.

    The information available from geomorphological studies was even more sketchy. Fluctuations

    in

    the levels of lakes across

    southwest

    Asia

    provided

    some

    information

    for a

    very

    general

    recon-

    struction

    of climatic

    sequences (Roberts

    1982),

    but

    they

    were not detailed

    enough

    to

    detect the

    483

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    AMERICAN

    ANTIQUITY

    Figure

    1.

    Locations of

    Lake

    Huleh,

    the

    Ghab,

    and the

    prehistoric

    village

    of Abu

    Hureyra

    in

    the Levant.

    impact

    of the

    Younger Dryas

    episode. Sedimentological

    studies

    in

    the southern Levant and Sinai

    yielded very

    little relevant

    information for the Late Glacial

    (Goldberg

    1981).

    This

    evidence,

    limited

    though

    it

    was,

    seemed to

    suggest

    that,

    as the

    temperature

    rose

    during

    the

    Late

    Glacial,

    rainfall also

    increased,

    leading

    to an

    expansion

    of forest cover

    (van

    Zeist and Bottema

    1982).

    Thus

    it

    appeared

    that

    the transition to

    farming

    took

    place

    in

    quite

    favorable environmental

    conditions

    (Moore

    1985:12).

    THE

    NEW

    EVIDENCE

    Our

    impression

    of a

    steady improvement

    in

    environment over much of southwest Asia from

    the

    Late Glacial into the early Holocene needs to be revised in the light of two new lines of evidence

    that have been

    obtained

    in

    the Levant

    (Figure 1).

    Firstly,

    Baruch

    and

    Bottema

    (1991)

    have

    recently

    extracted

    and

    analyzed

    a new

    pollen

    core

    from

    the Huleh Basin

    that

    provides

    a more

    detailed,

    well-

    484

    [Vol.

    57,

    No.

    3,

    1992]

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    REPORTS

    depth

    -lm

    ^--

    9 270

    +

    120

    B.

    P

    GrN-17067

    I

    10,440

    ?

    120

    B.P

    GrN-1 068

    -11,540+

    100 B.R

    GrN-14986

    |

    17,140

    ?220

    B.P

    -1 .67m

    G*

    GrN-14463

    arboreal nonorboreal

    pollen

    0

    25 50 75 100%

    Figure

    2. The ratio

    of

    arboreal

    to nonarboreal

    pollen

    in

    the

    diagram

    from Lake

    Huleh

    (after

    Baruch

    and

    Bottema

    1991).

    dated

    sequence

    of

    vegetation change

    from

    the end

    of the

    Pleniglacial

    ca.

    17,000

    B.P. into

    the

    early

    Holocene

    (Figure 2).

    The

    information most

    pertinent

    for our

    discussion concerns the

    changing

    ratio

    between arboreal and nonarboreal

    pollen.

    At the end of

    the

    Pleniglacial

    the ratio of tree

    pollen

    to

    grasses

    and

    steppe plants

    was

    low,

    about 20

    percent.

    Then at

    an estimated date of ca.

    15,000

    B.P.

    the ratio of tree

    pollen

    increased

    steadily

    until it

    reached

    a

    maximum of 75

    percent

    at

    11,540

    ?

    100

    B.P.

    (GrN-14986).

    Baruch

    and Bottema

    suggest

    that the

    increase

    in

    tree cover was caused

    by

    a

    marked rise

    in

    precipitation

    because

    it

    happened during

    the

    period

    of

    Late

    Glacial

    warming.

    Others such

    as

    El-Moslimany (1986)

    reasonably argue

    that such a

    change

    can be attributed more

    specifically

    to

    increased

    availability

    of

    moisture

    during

    the

    growing

    season of

    spring

    and

    summer,

    regardless of precipitation during the autumn and winter. Thereafter the forest shrank and/or thinned

    rapidly

    until ca.

    10,650

    B.P. when

    the ratio

    was

    slightly

    less than

    25

    percent

    arboreal

    pollen.

    It

    recovered to

    nearly

    50

    percent

    at

    10,440

    +

    120

    B.P.

    (GrN-17068),

    and then declined

    again

    over

    485

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    AMERICAN

    ANTIQUITY

    the next few centuries. The same trends

    may

    be

    detected

    in

    the Tsukada

    core,

    but are more

    clearly

    visible

    in

    the Baruch and Bottema

    diagram

    where

    they

    are more

    closely

    dated.

    It

    appears

    from Baruch

    and Bottema's

    analysis

    that the

    improved

    conditions for forest

    growth

    that

    took

    place

    in

    the

    region during

    the

    Late Glacial were

    spread

    over several millennia. The

    improvement thus tends to correlate with the lengthy period of climatic warming to be seen in the

    Barbados coral

    reefs

    (Fairbanks 1989).

    The most

    important

    observation for

    us,

    however,

    is the

    sharp

    reversal

    in

    the arboreal-nonarboreal

    pollen

    ratio from ca.

    11,500

    to

    10,650

    B.P.

    that Baruch

    and

    Bottema

    believe marks the

    Younger Dryas. Evidently

    the cooler conditions that obtained

    then

    were

    associated with

    a

    sharp

    decline in

    precipitation during

    the

    growing

    season and thus

    to a

    substantial reduction

    in

    forest cover.

    The

    second

    important

    source of new evidence for environmental

    change

    in the Levant

    during

    the

    Younger Dryas

    is the

    food-plant

    remains recovered from the

    early village

    of Abu

    Hureyra

    on

    the

    Euphrates.

    The first settlement

    there,

    Abu

    Hureyra

    1,

    of

    Epipaleolithic

    2

    cultural

    affinities,

    was

    inhabited from ca.

    11,500

    to

    10,000

    B.P.

    (Moore 1991) by

    a settled

    population

    of hunters and

    gatherers

    (Hillman,

    Colledge,

    and Harris

    1989;

    Legge

    and

    Rowley-Conwy 1987).

    Charred seeds

    and

    fruits recovered through systematic flotation have provided a record both of vegetation change and

    human

    plant exploitation

    throughout

    the

    occupation

    of this

    village (Hillman,

    Colledge,

    and Harris

    1989:Figure 14.1).

    On later sites with

    clear

    patterns

    of context-related

    variation,

    diachronic

    change

    in

    plant

    use can

    generally

    be demonstrated

    only

    when there are

    large

    numbers

    of

    productive samples

    derived

    from

    equivalent

    context

    types

    from each

    phase

    of

    occupation (Charles

    and

    Hillman

    1992;

    Hillman

    1981).

    It is

    possible

    to use the floated

    samples

    from

    Epipaleolithic

    Abu

    Hureyra

    (from

    39

    of the 80 levels

    excavated)

    to

    explore

    diachronic

    change by

    virtue

    of the fact that

    (a)

    the source

    deposits

    were

    relatively

    uniform,

    and were dominated

    in

    most cases

    by

    mixed

    accumulations

    of

    ashes

    from

    many years

    of

    fires that

    incorporated

    numerous

    cycles

    of seasonal

    activities,

    and

    (b)

    most of the float

    samples

    were extracted

    from

    very large

    volumes

    of these

    deposits

    (ranging

    from

    370 to

    4,000

    liters

    in

    all but three

    cases),

    and

    each

    contained

    literally

    thousands of identifiable

    items

    of

    food

    plants.2

    The Abu Hureyra 1 sequence of occupation has been divided into three periods, of which Period

    1A

    (ca.

    11,500-11,000 B.P.)

    is

    the oldest.

    During

    1A the

    inhabitants

    gathered

    plant

    foods

    from

    three

    vegetation

    zones,

    the

    moist flood

    plain

    of

    the

    Euphrates,

    the

    adjacent

    steppe,

    and

    a

    broad

    forest-steppe

    ecotone

    that was

    within

    foraging

    distance

    of the site.

    The

    latter extended

    eastward

    from the

    edge

    of

    the

    oak-Rosaceae

    forest that

    lay

    an unknown distance

    to the west.

    The

    predominant

    vegetation

    of the Abu

    Hureyra region

    was

    steppe,

    just

    as

    it is

    today.

    However,

    three classes

    of food

    plants represented

    in

    the remains

    indicate that conditions

    were

    much moister

    during

    the

    spring

    and

    summer

    growing

    seasons than

    they

    are

    now

    (Figure 3). Firstly,

    remains

    of fruit

    stones

    and seeds

    of

    the

    hackberry

    tree Celtis

    tournefortii,

    plum, pear,

    and

    medlar,

    all characteristic

    of

    the

    Mediterranean

    oak-Rosaceae

    forest

    zone,

    together

    with seed remains

    of

    a white-flowered

    asphodel Asphodelus

    microcarpus,

    characteristic

    of the Mediterranean

    zone

    generally,

    indicate

    that the oak-Rosaceae

    forest fringe must have been a great deal closer than the ca. 120 km to the west to which it could

    theoretically

    extend

    under

    natural conditions

    today.

    Secondly,

    the

    presence

    of Pistacia

    fruitlet remains

    and the

    apparent

    absence of Pistacia

    wood

    charcoal

    suggests

    that,

    although

    this tree did not

    grow

    close

    enough

    for its

    twigs

    or wood

    to be

    gathered

    as

    fuel,

    it

    must have

    grown

    much closer than

    it

    does

    today

    (Hillman,

    Colledge,

    and

    Harris

    1989).3

    The nearest

    patches

    of

    Pistacia

    steppe-woodland

    are

    now

    high

    on the

    Jebel

    Abu

    Rujmein

    90

    km to the

    south,

    and

    on the

    Jebel

    Abdul

    Aziz

    180

    km to the east-northeast.

    In Period

    1

    A Pistacia

    probably

    penetrated

    the

    steppe

    in

    the form of

    lines of trees

    growing

    along

    low wadi

    terraces,

    perhaps

    to within a

    few kilometers

    of Abu

    Hureyra,

    just

    as

    it

    today penetrates

    the

    Azraq

    Desert

    Basin

    in

    eastern

    Jordan

    along

    the

    Wadi

    Butum.

    The

    third

    source of evidence

    that

    Period

    1A

    was characterized

    by

    relatively

    moist

    springs

    and/

    or summers

    comes from the

    remains

    of

    wild

    einkorn

    wheat and

    two wild

    ryes.

    Today

    these

    wild

    cereals are characteristic of the ecotone between oak-Rosaceae forest and steppe, and although two

    of them are able to extend

    well

    beyond

    the

    forest

    fringe

    on

    deep,

    fine-grained

    soils

    (Blumler

    1984,

    1992),

    recent

    surveys suggest

    that

    they

    cannot

    penetrate

    steppe

    as

    far as the

    Pistacia

    steppe

    woodland

    486

    [Vol.

    57,

    No.

    3,

    1992]

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    7/14

    FRUIT

    REES

    &

    HERBACEOUS

    PERENNIALS

    PISTACIA

    OF

    FOREST

    FOREST-STEPPE

    WILD

    EREALSODAY YPICAL

    OFOPENOAK OREST ND

    OAKFOREST-STEPPECOTONE

    PERENNIAL

    USS

    OFSTEPPE

    AND

    F

    ABUHUREYRA

    TRENCH

    m

    OCCUPATION E

    E

    . .

    SEQUENCE

    LEVELS

    X

    *

    10,000

    B.. ?

    400

    402

    AM~

    411

    10

    412

    420

    418

    419

    10,400

    B.P

    427

    425

    426

    430

    lB

    449

    454

    455

    457

    1

    1,000

    B.P

    467

    468

    473

    1A

    ^

    A

    474

    469

    470

    471

    11,500

    B.P

    I

    0

    50

    100

    I I I

    200

    I numbers of charred seeds or fruits per 200 I1f

    deposit

    Figure

    3.

    Trends

    in

    the

    exploitation

    of wild

    cereals, pulses,

    and

    other

    open-forest

    plants

    by

    the

    inhabitants

    of

    Abu

    occupation.

    The

    diagram

    is based

    on a small

    portion

    of the evidence recovered

    (compare

    Hillman,

    Colledge,

    and Harris

    19

    L

    I

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    8/14

    AMERICAN

    ANTIQUITY

    (Hillman, Colledge,

    and

    Harris

    1989). Today,

    even without

    grazing

    and

    cultivation,

    it is

    unlikely

    that

    they

    could

    grow any

    closer

    than

    100

    km

    to the

    west and north.

    However,

    to

    have been

    gathered

    by

    the

    people

    of

    Abu

    Hureyra

    1,

    they

    must have been

    growing very

    much

    nearer than that.

    This

    pattern

    of

    gathering

    persisted

    throughout

    Period 1A. Then

    an

    abrupt change

    took

    place.

    The

    inhabitants

    appear

    to have ceased

    all

    gathering

    of tree

    fruits

    of

    the

    forest

    or forest

    fringe.

    One

    explanation

    is that

    the

    fruits

    were

    now

    out

    of

    range

    of

    foragers

    from Abu

    Hureyra simply

    because

    increasing aridity

    was

    preventing

    fruit formation

    on trees

    in

    the

    nearest areas

    of

    the

    forest

    fringe,

    and

    was

    thereby

    inducing

    the start

    of a

    forest

    retreat.

    This

    explanation

    is

    arguably

    supported by

    the

    ensuing

    set of

    changes

    at the

    beginning

    of

    Period 1B

    (ca.

    11,000-10,400 B.P.),

    which show a

    brief

    episode

    of

    sharply

    increased

    exploitation

    of the wild

    cereals,

    grains

    of

    feather-grass

    (Stipa spp.),

    and

    seeds

    of

    asphodel (Figure

    3; Hillman,

    Colledge,

    and Harris

    1989:Figure 14.1).

    This fits

    the

    temporary

    increase

    in

    yields

    from

    forest-fringe

    grasses

    and other herbs

    such as

    asphodel

    that

    might

    be

    expected

    when the trees started

    dying

    back

    and

    cast

    less shade on

    the herb

    layer.4

    However,

    our evidence

    for

    this brief

    episode early

    in

    1

    B

    comes

    from two

    rich

    samples

    and, despite

    the essential

    similarity

    of

    the

    formation

    processes

    reflected

    in

    most

    of the Abu

    Hureyra

    1

    levels and the

    huge

    amounts

    of

    deposit sampled, a difference in two such samples could theoretically represent no more than aberrant

    taphonomy.

    Very

    soon thereafter

    we

    see a

    more

    abrupt

    and

    unequivocal

    change,

    a

    complete

    cessation

    of the

    use

    of

    asphodel

    seed,

    perhaps

    for

    food

    or

    medicine,

    and

    a dramatic decline

    in

    the use

    of the three

    wild

    cereals and at

    least some

    of

    the

    feather

    grass species.

    These

    changes,

    combined

    with

    declining

    exploitation

    of Pistacia

    fruitlets,

    suggest

    that

    advancing aridity

    was now also

    causing

    a retreat

    of

    the herbaceous

    plants

    of the forest

    fringe, following

    the earlier

    dieback

    of

    the

    trees.

    This

    view

    is

    supported by

    the

    increased

    use

    of

    small-seeded

    legumes

    such

    as

    the

    clovers

    and

    medicks

    (Trifolium,

    Trigonella,

    and

    Medicago spp.),

    which

    require

    careful

    detoxification,

    and

    that

    we

    regard

    as "fallback

    foods,"

    which would

    generally

    have served as

    staples

    only

    when other

    major

    plant

    foods

    were

    becoming

    scarcer.

    In

    addition,

    a

    number

    of these small-seeded

    legumes

    can

    tolerate

    very

    arid

    conditions,

    and some of them

    would

    have continued

    to

    be available

    in undiminished

    abundance.

    We

    estimate

    from the 12 radiocarbon dates for Period 1B that this

    abrupt

    change

    began

    about

    10,600

    B.P.

    (Moore

    1992).

    These

    trends

    became even

    more

    marked

    in Period 1C

    (ca.

    10,400-

    10,000 B.P.),

    when

    we also

    see a decline

    in the use

    of

    valley-bottom

    foods,

    perhaps reflecting

    reduced

    overbank

    flooding

    as a result

    of

    lower levels

    of

    precipitation

    over

    the Anatolian

    catchment of

    the

    headwaters

    of

    the

    Euphrates.

    Abu

    Hureyra

    is

    in

    a semiarid

    region

    where

    slight changes

    in

    climate

    can lead to

    major

    adjustments

    in

    the

    composition

    and extent

    of

    vegetation

    zones

    and

    their

    component

    communities

    (cf.

    Davis

    1986;

    Webb

    1986).

    We

    argue

    that

    the most

    economical

    explanation

    of

    such

    a

    series of

    shifts

    in

    the

    pattern

    of

    plant collecting

    is an alteration

    in

    the

    composition

    of

    plant

    communities

    in

    the

    Abu

    Hureyra

    catchment

    brought

    about

    by

    climatic

    change. Certainly,

    Abu

    Hureyra

    1

    was inhabited

    long

    enough

    for

    the effects of

    the Late

    Glacial

    climatic

    fluctuations

    to be

    reflected

    in

    the

    vegetation

    record, and Periods 1B and 1C coincided with the Younger Dryas when cooler and/or more arid

    conditions

    prevailed

    in

    many regions.

    Reduction

    in

    moisture

    availability during

    the

    spring

    and/or

    summer

    growing

    seasons

    in lowland

    and

    some

    upland

    areas of southwest

    Asia

    is

    strongly

    indicated

    by

    the

    forest

    retreat

    seen

    in the

    pollen

    cores from

    Lakes

    Huleh and

    Zeribar,

    and Karamik

    Batakligi

    (van

    Zeist

    and Bottema

    1977, 1982;

    van

    Zeist et

    al.

    1975),

    and this can

    arguably

    be

    extended

    to

    the

    Ghab as

    well.

    It

    is

    precisely

    this

    reduction

    in the

    availability

    of

    growing-season

    moisture

    that

    we

    see reflected

    in

    the

    record

    of

    vegetation change

    at Abu

    Hureyra.

    As

    aridity

    increased

    in

    the

    Younger

    Dryas,

    the

    forest

    and

    forest-steppe

    ecotone

    retreated

    westward,

    to be

    replaced by

    more

    drought-resistant

    types

    of

    steppe.

    In

    consequence,

    the

    availability

    of

    many

    former

    foods

    was

    progressively

    reduced,

    and

    in

    compensation

    the

    inhabitants

    of

    Abu

    Hureyra

    seem

    to

    have increased

    their

    consumption

    of

    other

    foods

    such

    as the

    small-seeded

    legumes,

    thus

    allowing

    them

    to continue

    occupation

    for

    several

    centuries more.5 However, the population was already sedentary by Period 1A, and the constraints

    on

    population

    growth

    occasioned

    by

    a

    mobile existence

    would

    probably

    have

    already

    been

    relaxed

    (Hillman 1987).

    The

    resulting

    combination

    of

    increasing

    population

    and

    declining

    availability

    of

    488

    [Vol.

    57,

    No.

    3,

    1992]

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    9/14

    REPORTS

    previously preferred plant staples

    in

    the

    vicinity

    of Abu

    Hureyra

    may

    have

    imposed

    increasing

    stresses on the

    carrying

    capacity

    of

    the local

    environment,

    and so

    contributed to the

    temporary

    abandonment of

    the

    settlement. When

    Abu

    Hureyra

    was

    reoccupied

    a few

    centuries

    later,

    its

    new

    inhabitants

    were

    already

    farmers.

    The records of Late Glacial

    vegetation

    change

    from the

    new

    Huleh core and Abu

    Hureyra

    show

    analogous

    trends: an

    initial

    flourishing

    of

    forest and

    forest-fringe

    vegetation

    reflecting

    a

    relative

    abundance of

    growing-season

    moisture,

    followed

    by

    a

    sharp

    reversal as drier

    conditions set

    in

    during

    the

    Younger

    Dryas.

    Direct

    comparison

    of

    radiocarbon

    dates would

    suggest

    that the effects were

    experienced

    slightly

    earlier in

    the southern

    Levant than farther

    north. That

    may

    have been

    so,

    but

    we

    also need

    to allow

    for the

    possibility

    of

    interlaboratory

    error

    when

    comparing

    sequences

    of dates

    obtained

    by

    different

    laboratories.

    Recent

    studies

    have

    shown

    that the dates

    obtained

    by

    different

    laboratories for the

    same

    samples

    may

    vary

    systematically by

    several

    hundred

    years

    (Scott

    et al.

    1990).

    In

    any

    case,

    the

    abrupt

    change

    in

    vegetation

    at

    both

    localities

    correlates with the

    worldwide

    Younger

    Dryas episode

    of

    cooling.

    Having

    established that

    the

    Younger

    Dryas

    had a

    significant

    effect on the

    environment of

    the

    Levant, we should examine some of the other pollen cores obtained years ago to see if its impact

    can

    be

    discerned

    elsewhere in

    southwest Asia. The

    section of the

    original

    Ghab core

    covering

    the

    Late

    Glacial and

    earlier

    Holocene

    (pollen

    zone

    Z)

    exhibited

    much the

    same trends as

    the new Huleh

    core: a

    major

    expansion

    of

    forest

    followed

    by

    a

    decrease,

    then

    a modest

    growth

    of tree cover

    once

    more

    (Niklewski

    and van

    Zeist

    1970:Figure 3).

    The

    core

    had three

    radiocarbon

    dates,

    but

    only

    one

    of them

    related

    to the

    last

    40,000

    years.

    This date

    of

    10,080

    ?

    55 B.P.

    (GrN-5810)

    seemed

    to

    correspond

    to the

    climax of

    the main

    period

    of

    forest

    expansion,

    suggesting

    that the

    Ghab

    vegetation

    sequence

    was

    out of

    phase

    with the

    rest of the

    Levant.

    Given that

    the

    vegetation

    sequences

    from

    Lake

    Huleh and

    Abu

    Hureyra

    seem to

    correlate

    quite well,

    and that

    Abu

    Hureyra

    is

    just

    180

    km

    downwind

    of the

    Ghab,

    the

    most

    likely explanation

    for this

    discrepancy

    is

    that the

    radiocarbon

    date,

    obtained from

    a

    sample

    of

    shells,

    is

    discordant with

    its

    stratigraphic

    position.

    Baruch

    and

    Bottema

    (1991)

    have

    also

    allowed the

    possibility

    that

    the date

    may

    be

    in

    error.

    If

    that is

    the

    case,

    then the evidence from the Ghab core would indicate that the cooler conditions of the

    Younger

    Dryas

    were

    also felt in

    northwest

    Syria

    where

    they

    caused a

    synchronous

    decline

    in

    tree

    cover.

    Away

    to

    the

    east at

    Lake

    Zeribar in

    the

    Zagros Mountains,

    the Late

    Glacial

    was

    marked

    by

    an

    increase in the

    pollen

    of

    herbaceous

    plants

    at the

    expense

    of

    chenopods

    and

    Artemisia

    (van

    Zeist

    and

    Bottema

    1982:Figure 14.2),

    implying

    an

    increase

    in

    moisture

    during

    the

    growing

    season.

    That

    trend

    was

    sharply

    reversed

    for

    several

    centuries after ca.

    11,500

    B.P.,

    an

    event

    that

    again

    correlates

    chronologically

    with

    the

    Younger

    Dryas.

    It

    implies

    that

    there

    was a

    decrease

    in

    moisture

    during

    this

    period,

    just

    as we

    have seen in

    the

    Levant.

    A

    similar

    reduction

    in

    moisture

    availability

    during

    the

    growing

    season

    can

    likewise

    be

    inferred from

    the

    dramatic

    decline in

    cedar

    pollen

    after ca.

    11,500

    B.P. at

    Karamik

    Batakligi in

    western

    Anatolia,

    and

    from its

    continued

    depression

    until

    the

    start

    of the

    Holocene

    (van

    Zeist et al.

    1975).

    Another contiguous region with a good pollen record from the Pleniglacial through the Holocene

    is

    northern

    Greece

    (van

    Zeist

    and

    Bottema

    1982).

    The

    general

    trend

    in

    the

    published

    pollen

    curves

    is

    similar:

    dry, steppic

    conditions

    in

    the

    Pleniglacial

    that

    persisted

    into

    the Late

    Glacial.

    Then

    oak

    and

    pine

    forest

    spread

    throughout

    the

    region

    as

    temperature

    and

    growing-season

    moisture

    increased.

    The

    initial

    phase

    of

    tree

    growth

    at

    Tenaghi Philippon

    on the

    Plain

    of

    Drama took

    place

    during

    the

    Late

    Glacial,

    and

    was then

    sharply

    reversed at

    an

    estimated

    date of

    ca.

    10,500

    B.P.,

    before

    resuming

    once

    more in

    the

    early

    Holocene

    (Wijmstra

    1969:523).

    This

    return to

    dry, steppic

    conditions

    appears

    to

    correspond

    approximately

    to the

    Younger

    Dryas,

    and

    correlates

    with the

    vegetation

    record

    from

    southwest

    Asia.

    These

    pollen

    sequences

    all

    suggest

    that

    the

    colder

    conditions

    of the

    Younger

    Dryas

    had a

    significant

    impact

    on

    vegetation

    throughout

    southwest Asia

    and

    also in

    the

    extreme

    southeast

    of

    Europe.

    Everywhere

    the

    Younger

    Dryas

    in

    southwest

    Asia was

    accompanied

    by

    a

    decrease in

    moisture

    that

    caused a temporary reversion to partially steppic conditions more typical of the Pleniglacial.

    It

    should

    also be

    noted

    that

    the five

    pollen

    cores

    cited

    above

    exhibit

    synchrony

    not

    only

    with

    the

    Younger

    Dryas episode,

    if

    the

    Ghab date

    is

    adjusted

    as

    proposed,

    but

    also with

    the two

    preceding

    489

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    AMERICAN

    ANTIQUITY

    changes

    in

    vegetation

    cover.

    All

    five

    indicate a

    period

    of intense

    aridity

    from

    approximately

    18,000

    to

    15,000

    B.P. This was

    followed,

    ca.

    15,000

    to

    11,500 B.P.,

    by

    a

    sharp

    increase

    in

    growing-season

    moisture

    reflected

    in dramatic

    forest

    expansion

    at Lake

    Huleh,

    the

    Ghab,

    Karamik

    Batakligi,

    and

    Tenaghi Philippon,

    and

    an increase

    in

    grasses

    and

    other

    herbs,

    at

    the

    expense

    of the

    more

    arid-

    tolerant Artemisia and

    chenopods,

    in the then

    mountain-steppe

    flora around Lake Zeribar.

    ARCHAEOLOGICAL IMPLICATIONS

    The new evidence has a number of

    significant

    implications

    for

    our

    understanding

    of human

    adaptations during

    the late

    Epipaleolithic

    in the Levant and the

    subsequent adoption

    of

    agriculture.

    The

    improvement

    in

    the environment

    that

    began

    in the

    Late Glacial

    ca.

    15,000

    B.P.

    provided

    increasingly

    favorable conditions for

    hunter-gatherers throughout

    the

    region

    from late

    in

    Epipaleo-

    lithic

    1

    into

    early

    Epipaleolithic

    2. This

    helps

    to

    explain

    the florescence of such

    groups,

    for

    example

    those

    labeled Geometric

    Kebaran and

    early

    Natufian

    in

    the southern Levant

    (Bar-Yosef

    and Belfer-

    Cohen

    1989),

    and the

    Epipaleolithic

    inhabitants

    of

    Abu

    Hureyra

    1A

    on

    the

    Euphrates.

    The climatic

    reversal that followed profoundly altered the environment in which such groups lived. The return

    of drier conditions

    sharply

    reduced

    the extent of the forest and

    caused the rich

    zone

    of

    open

    forest-

    steppe along

    its

    edge

    to

    retreat

    westward,

    and

    probably

    to diminish

    in

    width.

    Thus there was

    a

    sharp

    reduction

    in

    the extent

    of those zones most

    favorable

    for

    the,

    by

    now

    quite

    numerous,

    groups

    of

    Epipaleolithic

    2

    hunter-gatherers.

    Those

    conditions

    appear

    to have

    lasted about a

    millennium,

    coincident

    with the duration

    of the

    Younger Dryas

    elsewhere.

    The environment

    improved

    towards

    the end of the

    period,

    on the

    evidence of the new Huleh

    core and that

    from the

    Ghab,

    but

    the ratio

    of arboreal

    pollen

    never reached the

    level it had attained

    in

    the

    Late Glacial before

    the onset of

    the

    Younger Dryas.

    It has

    always

    seemed

    anomalous

    that the

    pollen

    cores

    suggested

    a

    steady

    improvement

    of con-

    ditions

    from

    the Late Glacial into

    the

    early

    Holocene,

    while

    Leroi-Gourhan's

    studies of

    the

    pollen

    from

    Epipaleolithic

    2

    sites

    indicated that

    their environs were

    often

    quite steppic (Darmon

    and Leroi-

    Gourhan 1991; Henry 1989:73; Leroi-Gourhan 1984). That anomaly is now resolved; evidently

    steppe species

    of

    plants

    increased

    around those sites

    during

    the

    eleventh

    millennium

    B.P. because

    the

    climate

    became

    drier.

    The

    decrease

    in moisture

    availability

    set

    in

    during

    Epipaleolithic

    2

    and

    would

    have had

    a

    con-

    siderable

    impact

    on

    prevailing

    human

    patterns

    of

    foraging.

    It

    may

    have

    taken several centuries

    for

    the

    full

    effects

    to be

    felt,

    especially

    on sites

    in

    the better-watered

    zones,

    but

    they

    would

    have been

    experienced by people throughout

    the

    Levant,

    and

    adjustments

    in

    subsistence

    would

    have been

    necessary.

    The

    pattern

    of

    hunting

    remained

    the same

    throughout

    the

    sequence

    of

    occupation

    at Abu

    Hureyra

    1,

    indicating

    that the

    deterioration

    in

    climate

    had no

    adverse effect

    on

    the

    density

    of

    the

    herds of

    Persian

    gazelle

    (Gazella

    subgutturosa),

    a

    steppe species

    that was

    the main

    source

    of meat

    (Legge

    and Rowley-Conwy 1987). That may have been

    the case

    elsewhere, although

    at

    other

    Epipaleolithic

    2 sites

    in

    the forest

    zone the increase

    in

    the

    proportions

    of the

    various

    gazelle

    subspecies

    killed

    compared

    with

    Epipaleolithic

    1

    (Moore

    1982:227) may partly

    reflect

    the onset

    of more

    steppic

    conditions.

    We

    have

    seen,

    however,

    that

    the

    inhabitants

    of

    Abu

    Hureyra apparently

    modified

    their

    gathering

    of

    plants

    in

    response

    to

    the

    alterations

    in

    the

    vegetation

    in

    the site catchment.

    Abu

    Hureyra

    is the

    only

    Levantine

    site excavated

    so far that was

    inhabited

    throughout

    the

    Younger Dryas,

    and

    is also

    the

    only

    one

    to have

    yielded

    a

    long

    sequence

    of

    plant

    remains.

    Thus we

    should

    not

    expect

    to

    see such

    direct

    evidence

    of

    changes

    in

    subsistence

    from other sites

    occupied

    for

    shorter

    lengths

    of

    time.

    There is

    much other

    evidence,

    however,

    of

    major changes

    in

    culture

    and the

    pattern

    of

    settlement

    as

    the Levantine

    environment

    deteriorated.

    Most of

    the more

    substantial

    sites

    in

    the

    Natufian heartland were

    inhabited

    during

    the earlier

    stages

    of

    that culture.

    It is on

    those

    sites,

    Mugharet

    el Wad

    (Garrod 1957),

    Ain

    Mallaha

    (Perrot

    1966), Wadi Hammeh 27 (Edwards et al. 1988), and several others, that the Natufian culture has

    been

    found

    in

    its

    most

    developed

    form.

    They

    were substantial sites with huts

    and

    other

    structures,

    rich

    assemblages

    of

    bone and

    ground-stone

    artifacts,

    and also

    exquisite

    naturalistic

    carvings

    in

    bone

    490

    [Vol.

    57,

    No.

    3,

    1992]

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    REPORTS

    and

    stone. As Garrod

    (1957:224)

    noted

    in

    her

    original

    survey

    of the

    culture,

    and Valla and

    Henry

    have since

    reaffirmed

    (Henry

    1989:181;

    Valla

    1988:582),

    most of

    those elements ceased to be

    made

    in

    the

    later

    Natufian,

    and

    occupation

    on sites in

    the

    heartland was

    disrupted.

    It

    now

    appears

    that

    this

    disturbance in

    the

    pattern

    of settlement

    in

    the later Natufian

    coincided with the

    environmental

    deterioration of

    the

    Younger

    Dryas.

    It would seem

    that

    conditions worsened

    sufficiently

    to

    upset

    the

    pattern

    of

    subsistence

    plant

    gathering

    at sites

    in

    the

    Natufian heartland

    and with it

    the

    relatively

    sedentary

    life of

    their inhabitants.

    If

    supplies

    of

    wild

    plants

    and

    animals were no

    longer

    so

    abundant

    close

    by

    those

    sites

    because of

    environmental

    deterioration,

    their

    inhabitants would have

    been

    obliged

    to

    modify

    their

    subsistence activities.

    It

    appears

    that

    heir

    initial

    response

    was

    to resume

    a

    more

    mobile

    pattern

    of

    hunting

    and

    gathering,

    which

    had

    probably

    characterized life

    in

    the

    Plen-

    iglacial.

    Farther

    south,

    in the

    Negev

    and

    Sinai,

    there

    were

    quite

    rapid

    changes

    in

    culture

    and the

    pattern

    of

    settlement,

    but there

    was

    continuity

    of

    occupation

    (Goring-Morris

    1987:436-439). Thus,

    the

    deterioration

    in

    climate

    apparently

    had less

    impact

    there.

    The

    transition

    from

    Epipaleolithic

    to

    Neolithic

    ca.

    10,000

    B.P.

    coincided with the

    end of

    the

    Younger

    Dryas

    climatic

    episode.

    It

    was a time of

    major

    readjustment

    in

    culture

    and

    patterns

    of

    settlement throughout the Levant. Settlements in the steppe zone were abandoned. Thus, the Azraq

    Basin

    and the

    oasis

    of El

    Kum

    were

    deserted

    (Cauvin

    1981:387;

    Garrard et

    al.

    1988),

    not to be

    reoccupied

    for

    at least a

    millennium. Even

    in

    the

    better-watered

    zones

    occupation

    ceased

    at

    nearly

    all

    sites.

    At the few

    settlements like

    Abu

    Hureyra, Jericho,

    and Beidha

    that were

    inhabited

    both

    in

    the

    Epipaleolithic

    and

    the

    succeeding Neolithic,

    occupation

    was

    briefly disrupted

    at the

    end of

    the

    Epipaleolithic.

    The

    new

    Neolithic

    pattern

    of

    settlement

    was

    based

    initially

    on

    a

    relatively

    few

    large

    sites

    in

    locations with

    rich

    soils

    and

    ample

    surface

    water,

    that is in

    locations that

    were

    suitable for

    agriculture.

    Remains of

    domesticated

    cereals and

    pulses

    have been

    found

    at

    Jericho and Tell

    Aswad,

    dating

    from

    about

    10,000

    B.P.

    (Hopf 1983;

    van

    Zeist and

    Bakker-Heeres

    1979),

    and a little

    later

    at Abu

    Hureyra.

    The

    economic and

    cultural

    transformations

    that

    marked the

    transition

    from

    Epi-

    paleolithic

    to

    Neolithic

    therefore

    appear

    to have

    been

    rapid.

    COMMENTARY

    The

    Younger

    Dryas

    climatic

    episode

    evidently

    had a

    significant

    impact

    on

    the

    environment of

    southwest

    Asia. It

    interrupted

    the Late

    Glacial

    improvement

    in

    climate

    and

    vegetation,

    and

    caused

    a brief

    return

    to

    the

    conditions

    of the

    Pleniglacial.

    The

    consequences

    were

    severe for

    the

    Epipaleolithic

    2

    peoples

    of the

    Levant.

    Their

    modes of

    gathering

    were

    disrupted

    and the

    resulting

    stresses

    led to

    widespread

    dislocation in

    patterns

    of

    settlement. It is

    surely

    no

    coincidence that

    the

    transition

    from

    Epipaleolithic

    to

    Neolithic and

    from

    hunting

    and

    gathering

    to

    farming happened

    at

    about

    the

    same

    time,

    just

    as

    the

    Younger

    Dryas

    had run

    its

    course. We

    suggest

    that

    the

    disruption

    that took

    place

    in

    Epipaleolithic

    2

    patterns

    of

    adaptation

    acted as a

    powerful

    incentive

    for

    the

    peoples

    of

    the

    Levant

    to

    develop

    new

    modes of

    subsistence.

    The

    Younger

    Dryas episode

    was

    not the

    only

    factor

    that led

    to this result. Among other processes, the advent of sedentary life, demonstrated at Abu Hureyra

    at

    least,

    and

    population

    growth

    during

    the

    Epipaleolithic

    undoubtedly

    contributed to

    the

    outcome

    (Hillman

    1987;

    Moore

    1985:13).

    Nevertheless,

    it was

    probably

    a

    significant

    catalyst

    in

    each of

    the

    areas within

    southwest Asia

    where

    cultivation is

    likely

    to have

    begun.

    Acknowledgments.

    We

    wish to

    thank

    Sytze

    Bottema or

    helpful

    discussions

    concerning

    he new

    Huleh

    pollen

    core,

    and for

    kindly

    allowing

    us

    to

    reproduce

    here

    part

    of

    the

    pollen

    diagram prepared

    by

    Uri

    Baruch

    and

    Bottema.We

    also

    thankthe

    reviewers or

    American

    Antiquity

    and other

    colleagues

    who

    have

    offered

    onstructive

    comments on

    earlier

    draftsof

    this

    paper.

    The

    hypothesis

    explored

    herewas

    developed

    during

    the

    1990-1991

    academic

    year

    when

    Moore

    held a

    National

    Endowment

    for the

    Humanities

    Fellowship

    for

    University

    Teachers;

    the

    support

    of

    the

    National

    Endowment is

    gratefully

    acknowledged.

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    2

    The

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    The Pistacia

    species

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    P. atlantica and

    P.

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    vera.

    4

    Although many Stipa species thrive in arid steppe and desert steppe, some species such as S. lagascai seem

    to form their densest stands closer

    to the forest

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    of

    particular plant

    foods

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    Certainly,

    ethnobotanical

    studies of recent

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    that

    they

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    Lee

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    Lee

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    The

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    by

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    unlikely

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    many parallels among

    food-

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    strategies

    can be devised

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    implemented

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    greater

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    (Hillman,

    Madeyska,

    and Hather

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    Received December

    12,

    1991;

    accepted

    February

    21,

    1992

    494

    [Vol.

    57,

    No.

    3,

    1992]