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    Copyright 2003 Pearson Education, Inc. publishing as Benjamin Cummings

    All the organisms in a particular area make upa community

    A number of factors characterize everycommunity

    Biodiversity

    The prevalent form ofvegetation

    Response to disturbances

    Trophic structure(feeding relationships)

    36.1 A community is all the organisms inhabiting aparticular area

    Figure 36.1

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    Biodiversity is the variety of different kinds oforganisms that make up a community

    Biodiversity has two components

    Species richness, or the total number of

    different species in the community

    The relative abundance of different species

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    Interspecific competition occursbetween twopopulations if they both require the same

    limited resource

    A population's niche is its role in thecommunity

    The sum total of its use of the biotic and abioticresources of its habitat

    36.2 Competition may occur when a sharedresource is limited

    STRUCTURAL FEATURES OF COMMUNITIES

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    Predation is an interaction where one specieseats another

    The consumer is called the predator and the

    food species is known as the prey

    Parasitism can be considered a form ofpredation

    36.3 Predation leads to diverse adaptations in bothpredator and prey

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    As predators adapt to prey, sometimes naturalselection also shapes the prey's defenses

    This process ofreciprocaladaptation is

    known ascoevolution

    Example:

    Heliconius andthe passionflowervine

    Figure 36.3A

    Eggs

    Sugardeposits

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    Prey gain protection against predators througha variety of defense mechanisms

    Mechanical defenses, such as the quills of aporcupine

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    Chemical defenses are widespread and veryeffective

    Animals with effective chemical defenses areoften brightly colored to warn predators

    Example: the poison-arrow frog

    Figure 36.3B

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    Camouflage is a very common defense in theanimal kingdom

    Example: the gray tree frog

    Figure 36.3C

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    A keystone species exerts strong control oncommunity structure because of its ecologicalrole

    A keystone predator may maintain communitydiversity by reducingthe numbers of thestrongest competitors

    in a communityThis sea star is a

    keystone predator

    36.4 Predation can maintain diversity in acommunity

    Figure 36.4A

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    Predation by killer whaleson sea otters, allowing sea

    urchins to overgraze on kelp

    Sea otters represent thekeystone species

    Figure 36.4B

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    A symbiotic relationship is an interactionbetween two or more species that live togetherin direct contact

    There are three main types of symbioticrelationships within communities

    Parasitism

    Commensalism

    Mutualism

    36.5 Symbiotic relationships help structurecommunities

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    Parasitism is a kind of predator-preyrelationship

    The parasite benefits and the host is harmed inthis symbiotic relationship

    A parasite obtains food at the expense of its host

    Parasites are typically smaller than their hosts

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    In the 1940s, Australia was overrun byhundreds of millions of European rabbits

    The rabbits destroyed huge expanses of Australia

    They threatened the sheep and cattle industries

    In 1950, a parasitethat infects rabbits(myxoma virus)

    was deliberatelyintroduced tocontrol the rabbitpopulation

    Figure 36.5A

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    Commensalism is a symbiotic relationshipwhere one partner benefits and the other is

    unaffected

    Examples of commensalism

    Algae that grow on the shells of sea turtles

    Barnacles that attach to whales

    Birds that feed on insects flushed out of thegrass by grazing cattle

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    A community interacts with abiotic factors,forming an ecosystem

    Energy flows from the sun, through plants,animals, and decomposers, and is lost as heat

    Chemicals are recycled between air, water,soil, and organisms

    36.8 Energy flow and chemical cycling are the twofundamental processes in ecosystems

    ECOSYSTEM STRUCTURE AND DYNAMICS

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    A terrarium ecosystem

    Figure 36.8

    Chemical cycling(C, N, etc.)

    Lightenergy

    Chemicalenergy

    Heatenergy

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    A food chain is the stepwise flow of energy andnutrients

    from plants (producers)

    to herbivores (primary consumers)

    to carnivores (secondary and higher-levelconsumers)

    36.9 Trophic structure is a key factor in ecosystemdynamics

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    Figure 36.9A

    TROPHIC LEVEL

    Quaternaryconsumers

    Tertiaryconsumers

    CarnivoreCarnivore

    Carnivore Carnivore

    Carnivore Carnivore

    Herbivore Zooplankton

    Plant Phytoplankton

    Secondary

    consumers

    Primaryconsumers

    Producers

    A TERRESTRIAL FOOD CHAIN AN AQUATIC FOOD CHAIN

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    Decomposition is the breakdown of organiccompounds into inorganic compounds

    Decomposition is essential for the continuationof life on Earth

    Detritivoresdecompose wastematter and recyclenutrients

    Examples: animalscavengers, fungi,and prokaryotes

    Figure 36.9B

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    A food web is a network of interconnectingfood chains

    It is a more realistic view of the trophicstructure of an ecosystem than a food chain

    36.10 Food chains interconnect, forming food webs

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    Figure 36.10

    Tertiaryandsecondaryconsumers

    Secondaryandprimaryconsumers

    Primaryconsumers

    Producers

    (Plants, algae,phytoplankton)

    Detritivores

    (Prokaryotes, fungi,certain animals)

    Wastes anddead organisms

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    Biomass is the amount of living organicmaterial in an ecosystem

    Primary production is the rate at which

    producers convert sunlight to chemical energy

    The primary production of the entire biosphereis about 170 billion tons of biomass per year

    36.11 Energy supply limits the length of food chains

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    A pyramid of production reveals the flow ofenergy from producers to primary consumers

    and to higher trophic levels

    Figure 36.11

    Tertiaryconsumers

    Secondaryconsumers

    Primaryconsumers

    Producers

    10 kcal

    100 kcal

    1,000kcal

    10,000 kcal

    1,000,000 kcal of sunlight

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    Only about 10% of the energy in food is storedat each trophic level and available to the next

    level

    This stepwise energy loss limits most foodchains to 3 - 5 levels

    There is simply not enough energy at the verytop of an ecological pyramid to support anothertrophic level

    36 12 C i A i i i

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    The dynamics of energy flow apply to thehuman population as much as to otherorganisms

    When we eat grain or fruit, we are primaryconsumers

    When we eat beef or other meat from herbivores,we are secondary consumers

    When we eat fish like trout or salmon (which eatinsects and other small animals), we are tertiaryor quaternary consumers

    36.12 Connection: A production pyramid explainswhy meat is a luxury for humans

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    Because the production pyramid tapers sosharply, a field of corn or other plant crops can

    support many more vegetarians than meat-eaters

    Figure 36.12

    Secondaryconsumers

    Primaryconsumers

    Producers

    Humanvegetarians

    Corn

    Humanmeat-eaters

    Cattle

    Corn

    TROPHIC LEVEL

    36 13 Ch i l l d b t i

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    Ecosystems require daily infusions of energy

    The sun supplies the Earth with energy

    But there are no extraterrestrial sources ofwater or other chemical nutrients

    Nutrients must be recycled between organismsand abiotic reservoirs

    Abiotic reservoirs are parts of the ecosystemwhere a chemical accumulates

    36.13 Chemicals are recycled between organicmatter and abiotic reservoirs

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    There are four main abiotic reservoirs

    Water cycleCarbon cycle

    Nitrogen cycle

    Phosphorus cycle

    35 14 W t th h th bi h i

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    Heat from the sun drives the global water cycle

    Precipitation

    Evaporation

    Transpiration

    35.14 Water moves through the biosphere in aglobal cycle

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    Figure 36.14

    Solarheat

    Precipitationover the sea

    (283)

    Net movement

    of water vaporby wind (36)

    Flow of waterfrom land to sea(36)

    Water vaporover the sea

    Oceans

    Evaporationfrom the sea

    (319)

    Evaporationandtranspiration(59)

    Water vaporover the land

    Precipitationover the land(95)

    Surface waterand groundwater

    36 15 Th b l d d h t th i

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    Carbon is taken from the atmosphere byphotosynthesis

    It is used to make organic molecules

    It is returned to the atmosphere by cellularrespiration

    36.15 The carbon cycle depends on photosynthesisand respiration

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    Figure 36.15

    CO2 in atmosphere

    Cellular respiration

    Higher-levelconsumers

    Primaryconsumers

    Plants,algae,

    cyanobacteria

    Photosynthesis

    Wood andfossil fuels

    Detritivores(soil microbes

    and others) Detritus

    Decomposition

    Burning

    36 16 Th it l li h il b t i

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    Nitrogen is plentiful in the atmosphere as N2

    But plants cannot use N2

    Various bacteria in soil (and legume rootnodules) convert N2 to nitrogen compoundsthat plants can use

    Ammonium (NH4

    +) and nitrate (NO3

    )

    36.16 The nitrogen cycle relies heavily on bacteria

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    Some bacteria break down organic matter andrecycle nitrogen as ammonium or nitrate to

    plants

    Other bacteria return N2 to the atmosphere

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    Figure 36.16

    Nitrogen (N2) in atmosphere

    Amino acidsand proteins in

    plants and animalsAssimilationby plants

    Denitrifyingbacteria

    Nitrates(NO3

    )

    Nitrifyingbacteria

    Detritus

    Detritivores

    Decomposition

    Ammonium (NH4+)

    Nitrogenfixation

    Nitrogen-fixingbacteria in soil

    Nitrogen-fixingbacteria in root

    nodules of legumes

    Nitrogenfixation

    36 17 The phosphorus cycle depends on the

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    Phosphates (compounds containing PO43-

    ) andother minerals are added to the soil by thegradual weathering of rock

    Consumers obtain phosphorus in organic formfrom plants

    Phosphates are returned to the soil through

    excretion by animals and the actions ofdecomposers

    36.17 The phosphorus cycle depends on theweathering of rock

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    Figure 36.17

    Upliftingof rock

    Phosphatesin solution

    Weatheringof rock

    Phosphates

    in rock

    Phosphatesin organic

    compounds

    Detritus

    Detritivoresin soil

    Phosphatesin soil(inorganic)

    Rock Precipitated(solid) phosphates

    Plants

    Animals

    Decomposition

    Runoff

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    Dams were builtacross streams at

    the bottom of eachwatershed tomonitor water andnutrient losses

    Figure 36.18A

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    In 1966, one of the valleys was completelylogged

    It was thensprayed withherbicides for3 years to

    prevent plantregrowth

    All the original

    plant materialwas left inplace todecompose Figure 36.18B

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    36 19 Talking About Science: David Schindler

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    Eutrophication is a process in which nutrientrunoff from agricultural lands or livestockoperations causes photosynthetic organisms in

    ponds and lakes to multiply rapidly

    The result is algal bloom

    36.19 Talking About Science: David Schindlertalks about the effects of nutrients onfreshwater ecosystems

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    Algal bloom can cause a pond or lake to losemuch of its species diversity

    Human-caused eutrophication wiped outfisheries in Lake Erie in the 1950s and 1960s

    Figure 36.19B

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    Dr. David Schindler is an ecologist who workedat the Experimental Lakes Project in northern

    Ontario

    He performedseveral classic

    experiments oneutrophicationthat led to the banon phosphates in

    detergents

    Figure 36.19A

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    According to Dr. Schindler, there are threeserious threats to freshwater ecosystems

    Acid precipitation

    Climate warming

    Changes in land use