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8/6/2019 LOT 1 - Introduction
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Evolution of Language
Introduction
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Language and Speech
Are an important part of what makes
humans human
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Religion
Many cultures have
language origin myths
Either to explain why
we have language
Or to explain why we
have many languages
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Early speculation
Jespersens (1922) summarized:
Bow-wow theory
Pooh-pooh theory Ding-dong theory
Yo-he-ho theory
But his own alternative:
La-la theory
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In addition:
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The state-of-the-art in 1866
Evolution: Darwin 1858/1859
Neanderthal-skull: Schaaffhausen 1858
Linguistics: Stammbaum-theory, Schleicher ca.1853
Freud was 10 years old
Broca had discovered Brocas area in 1861
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Can we do better today?
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Topics
Evolution
Anatomy and Fossils
Genetics of Language Computer Models
Laboratory Experiments
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Carroll 2003
Gray &
Atkinson
2003
Darwin 1837
Evolution of Language and of
Languages
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Evolution
Both emergence of our capacity for
language and diversity of languages can
be seen as the outcome ofevolution
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Co-evolution
Co-evolution occurs when two systems evolve
and interact with each other
The systems influence each others fitness
Language and the capacity for language have co-evolved
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What exactly is evolution?
Three factors
Replication of information
Variation on this information
Selection on this variation
In the short term, these effects increase fitness Traits that increase replication are selected for
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An example
In the given
ecological context,
animals that can
eat higher food
have the advantage
transfer of information,
variation
selection
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Replication of Information (1)
Children must resemble their parents
Otherwise no direction to search
In language: need to be able to communicate with
parents
In biology: genes (DNA)
And perhaps to some extent: proteins and RNA
Difference between genotype (genes) and phenotype
(organism)
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Replication of information (2)
In biology genetic information is independent oflife experience Transfer from genotype to genotype
Phenotypic experience is not transferred
In language, only utterances are transferred, notlinguistic representations in the brain
This allows collective learning from experience Typical forcultural evolution
Lamarck (1809)
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Variation (1)
Variation is essential, otherwise one would not
explore the space of possibilities
In biology:
Recombination of genes
(diploid genes)
Crossover
Mutation
Essentially random
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Variation (2)
However, in language
(and culture)
variations can be
introducedconsciously, based on
experience
Making possible the
very rapid spread ofcultural innovations
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Selection (1)
Individuals that are in some respect betterare selected to create the next generation
But who determines what is better?
In biology fitness is by definition equal tonumber of offspring
But this is a post hoc measure, not an a priori
measure In biology, fitness depends on ecology
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Selection (2)
Selection is based on ability to createoffspring
Survival is a precondition for procreation
But other factors play a role
Sexual selection, among others
What does selection consist of in language? Acceptability to other language users?
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Variation/Selection
Functional changes Things that are easier to say are preferred by speakers
Things that are easier to understand are preferred by listeners
Contradictory forces
Ethnic identity Language is a very powerful factor in defining identity andsolidarity
Slang and jargon serve to distinguish oneself
Taboo Vocabularies can change rapidly when words can become taboo
Innovation Introduction ofnew words for new items
Or poetic innovation
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Complications
Multilingualism
Because a speaker uses multiple languages,these languages start to interfere
Famous example: Kupwar (India) Influences grammarand morphology
Loan words
W
ords for new objects, or words that havehigher prestige enterthe language
Influences lexicon
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More complications
Standards
Speakers adapt their speech to conform to acertain standard
Usually the standard has a certain prestige
Modern complications
Standardized education Mass media
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Evolution and tinkering
(Francois Jacob, Science 1977)
Evolution works with what is already there
This sometimes results in solutions that are less thanoptimal
Other terms are: Exaptation (cf. adaptation, the use of existing
structures for new purposes) Spandrels (Gould): Structures that have developed as
a side-effect of other adaptive functions, but that arenot necessarily adaptive themselves.
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Misconceptions about Evolution
Evolution always results in optimal
solutions
Evolution is goal-oriented
Evolution is an extremely slow process
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Evolution and teleology
Evolution is not teleological(goal directed) This is sometimes thought (e.g.
humans are the crown onevolution)
However there is somethingcalled the ratchet effect When a certain level of
complexity is reached, there isno turning back.
Example: once hard bodiedanimals evolve, soft bodiedones are at a strongdisadvantage (Ediacaran tolate pre-Cambrian transition)
Ediacaran fauna
Burgess shale fauna
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Evolution is slow?
Biological evolution tends to beslow
Because mutations are randomand most are bad
But in large populations,
adaptation can be quick Think antibiotic resistant bacteria
Some researchers claim evolutiongoes in fits and bursts
Punctuated equilibrium (Eldredgeand Gould 1972)
Cultural evolution can beextremely rapid
1977
2011
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Evolution Summary
Evolution is:
Transfer of information
Variation
Selection
Can be biological, cultural
There are important differences
Co-evolution also occurs The idea is simple, but the effects can be
very subtle.
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The evolutionary perspective (1)
Nothing in biology makes sense exceptin the light of evolution
Theodosius Dobzhansky (1973)
That does not mean that onlyan evolutionary perspective
is interesting Just that you need an evolutionary
perspective to understand ultimate causes
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The evolutionary perspective (2)
An evolutionary perspective is implicit in most
thinking about linguistic cognition
Consider:
A brain is necessary to
use language
But even flatworms have
brains
So what makes our brains
special, compared to otherbrains?
Dugesia sp.
Photo: Roberto Petracini
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Particulars of language evolution
Language needs strong cooperation
An interesting problem how this evolved
Co-evolution of culturally transmitted
language and biological adaptations crucial
A mutation for language is useless if no others
have it
Cultural system must already be present for
biological adaptation to work
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Evolution of language (2)
How can we study evolution of language?
No fossilized speech
and olderthan historical sources But data on: Modern human behavior
Fossils (Tuesdays class topic)
Animal behavior
Genetics (Wednesdays class topic) But there are questions we cannot answer
R. C. Lewontin (1998) The Evolution ofCognition: Questions wewill never answer
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Modern Language
Are there primitive languages?
No, not if native
No data on language evolution
But data on possibilities of language
But: pidgin-languages
Jargons, second language etc.
And creolisation
Oremergence of new language Nicaraguan sign language
Idea: proto-language
Bickerton
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When did language
emerge? Two extremes:
Late emergence (~30 000 years ago)
Early emergence (A. africanus)
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Symbolic explosion?
Symbolic culturalartifacts are found inNeandertals andH
. sapiens They are much olderthan previouslythought
T
hey appear anddisappearin the fossilrecord Second Bull, Lascaux cave
Photo: N. Aujoulat MCC-CNP
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Against late emergence
How can complex language evolve soquickly?
How does one explain biological
adaptations to language?
Homo sapiens started to spread muchearlierthan 5070 000 years ago
Would language have emerged in differentplaces?
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Deep history of language
Historical linguistics can reconstruct olderforms of a language (e.g. indo-european)
Traditional linguistics up to ~8000 years ago
But: Ruhlen claims proto-world
Very unlikely
Human expansion started about 150120 KAago
After this time all similarities are gone
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A summary of our knowledge
Language >100 000 years old
No primitive languages exist, and we know little abouthow they spread very long ago But we can observe emergence of new languages
That all have certain special properties
We can also observe incomplete languages
Language probably emerged as primitive proto-language Why is an interesting, but hard-to-answer question
How did it spread, how did it emerge? Cannot be reconstructed from fossils
But possible to model
And possible to compare to animal communication
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Comparative data (1)
Vocal behavior in other species can aid
understanding of speech
Precursors Primate vocal behavior the starting point
Parallel evolution
Song birds
Whales
Bats (?)
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Comparative data (2)
Interesting when trying to establish how
language can have evolved
Less interesting if only to establish
humans uniqueness
Sooner or later you will find similarbehavior in
other animals, or in other parts of cognition
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Hocketts
design-
features
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Ape Cognitive abilities
They can co-author
scientific papers
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They are, howeververy smart animals
They can learn impressive lexicons
Mostly directed towards requests Their abilities to combine words are disputed
No rules, no recursion
Ape Cognitive abilities
Sue Savage Rumbaugh & KanziPhoto: http://kanzi-superstar.blogspot.com/
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Chimpanzees use large sets of vocal signals
But mostly innate
Not discrete
Chimpanzees and gorillas use gestures
But dyadic
Some learned through
ritualization (not imitation)
Ape Cognitive abilities
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Gibbon song
Geissman, 2002
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Ape Cognitive abilities
Vocal culture in chimpanzees
(Crockford et al. 2004)
Some vocal learning in orangutans
Kuypers-Jrgens hypo-
thesis: control over mouth
but not over larynx
(Fitch 2010, 347-355)
Washington Post 4/3/2009
Bonnie
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Ability to cooperate is limited
But some cooperative hunting in chimps
And elaborate knowledge ofsocial hierarchy
Chimps appear to be much less altruistic
Some degree oftheory-of-mind
But much more limited than in humans
Ape Cognitive abilities
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Ape Cognitive abilities
Recursion Tamarin monkeys (Fitch & Hauser 2004): no
Starlings (Gentner et al. 2006): yes
Zebra finches (van Heijningen et al. 2009): no
Pirah: no
Brazilian Portuguese (university): yes
Brazilian Portuguese (illiterate): no
What is recursion anyway?
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Songbirds (1)
Doupe & Kuhl 1999
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Songbirds (2)
Many species Comparative studies possible
Crossing studies possible (wild and domesticatedbengalese finches)
Manipulation possible (fewer ethical concerns) Great diversity
Song complexity
Degree of learning
But different Neurologically Anatomically
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Humpback whale song
Payne & McVay (1971)
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Whales and dolphins
Humpback whales have complex, learned
songs
But these appear to be fixed per season
Some dolphin species show very complex
vocal behavior
Signature whistles
Hard to study, and relatively little is known
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Less well-known vocalizers
Pinnipeds
(Seals etc.)
Are capable ofimitation
For learning mating songs
Bats
Echolocation But also social vocalizations
Learned in some species
Phyllostomus hastatus
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Finally
Both biological and cultural evolution were
important for language evolution
Modern language tells us about ourabilities, notabout ourhistory (unfortunately)
Other species have complex vocal behavior,apes have many necessary precursors for
language, but no vocal imitation