Flooding tolerance: escape or quiescence

8/2/2019 Flooding tolerance: escape or quiescence

1/35

Pcly ^ocyclckBcteprjclr on Tfelr CiotmqyaofohqSrpcimr Sla~cpyarq& Rmc Lcrmcpfelby

mrrt7//xxx,`ao,ss,lf/ciotmqyaofohq/

Nfoobalh rofcpelic7 cyietc op usacyiclic


2/35

Nfoobalh oiispy xopfb*xabc elb rmc lsj`cpy alipceyc

xxx,beprjosrm,cbs/|nfooby/


3/35

Nfoobalh ay e jegop cl~apoljclref yrpcyy nop joyr tfelry

Tpo`fcjy7

Clcphq ipayay Iep`omqbperc yrep~eraol Pceira~c o}qhcl ytciacy Ro}aiarq Xercp yrersy

Iolycusclicy7

Yfox bannsyaol on heycy Pcbsicb fc~cfy on O0 elb IO0


4/35

Yclyalh on fox o}qhcl fc~cfy al tfelry

Rxo Psjc}ytciacy7 el ciofohaief jobcf al nfoobalh pcycepim

Pchsferaol on neyr tcraofc hpoxrm ro cyietc npoj ys`jcphclic

Iolrclry


5/35

Iotqpahmr 044> Ejcpaiel Yoiacrq on Tfelr @aofohayry

Lekelo& R,& cr ef, Tfelr Tmqyaof, 044>;5247255*290

El slpoorcb tmqfohclcrai rpcc onEpe`abotyay CPN tporcaly


6/35

O~cpc}tpcyyaol on PET0,50 pcysfry al alipceycb ys`jcphclic ysp~a~efelb clmelicb c}tpcyyaol on mqto}ae iopc hclcy

Eap Ys` Bc*ys`


7/35

Pcjo~ef on 59 ejalo eiaby er rmc L rcpjalef yabc on PET0,50 ennciryc}tpcyyaol on mqto}ae jepkcp hclcy elb ys`jcphclic ysp~a~ef

Eap Ys` Bc*ys`


8/35

Foiefayeraol on PET0,50 ay enncircb `q mqto}ae elb L rcpjalef jelatsferaol


9/35

PET0,50

Lopjo}ae Elo}ae

PET0,50

Jcrmaolalc ejalotctrabeyc

JI*

I*

Mqto}ae pcytolya~chclcy

O0

PET0,50I'*

PET0,50Eph*I'*

PET0,50

S`u

Ephalalc rpelyncpeycy

TPORCOFQYAY>

0>Y tporceyojc

O0#

Eph*I'*

O}qhcl yclyalh al tfelry ay jcbaercb `q el L*clbpsfc termxeq nop tporcal bcyre`afayeraol


10/35

O}qhcl yclyalh al tfelry jcbaercb `q el L*clb psfctermxeq nop tporcal bcyre`afayeraol


11/35




Iolrclry


12/35

Psjc}ytciacy epc c}toycb ro banncpclr nfoobalh alrclyaracy elb ymoxiolrpeyralh cfolheraol pcytolycy stol ys`jcphclic

)

nfooby/bsperaolnfooby-b!

Pe Pe PtPt

) nfooby

BsperaolNfooby -b!


13/35

Psjc}ytciacy c}tfoar rxo banncpclr ysp~a~ef yrperchacy stol nfoobalh


14/35

Ymopr nfooby Folh nfooby

Bcct

nfooby

Ymeffoxnfooby

Opbaleraol baehpej on ~chcreraolyal nfoob*tpolc pa~cp epcey

Pepcfq nfoobcb

^ocyclck cr ef,& 0442, Ciofohq =37 5>*08

Rpelyaclr*bcct

nfooby

Rpelyaclr*bcct

nfooby

Folh*ymeffox

nfooby

Folh*ymeffox

nfooby


15/35

Crmqfclc eiisjsfercy petabfq al ys`jcphcb Psjc}


16/35

Alyabc bayrelic npoj ebe}aef tcraofc yabc al j

4 044 244 >44 =44 5444

O}qhclioliclrperaolalkTe

4

3

54

53

04

03

Fahmr

Bepk

Ys`jcphcb tcraofcy on P, tefsyrpayelb P, eicroyebo lor me~c fox o}qhclioliclrperaoly

alyabc bayrelic npoj ebe}aef tcraofc yabc

4 044 244 >44 =44 5444 5044

O}qhclioliclrperao

l-kTe!

0

2

>

=

54

50

52

5>

5=

04

00

Fahmr

Bepk

P, tefsyrpay

P, eicroye


17/35

Ymoor cjcphclic ay lor nsliraolef al P,eicroyebsc ro fox topoyarq


18/35




Iolrclry


19/35

Crmqfclc boxl*pchsfercy E@E alP, tefsyrpay`sr lor al P, eicroye

@clyimot cr ef, -0443! Tfelr Gosplef 227 83>*8>=

Psjc} tefsyrpay

Psjc} eicroye


20/35

Es}al yrajsfercy tcraofc cfolheraol stol ys`jcphclic

Rajc enrcp ys`jcphclic -m!

[AEEZth/jhNX

Ebe}aef

E`e}aef


21/35

4

54

04

94

24

34

Tcraofccfolheraol

-jj/2=m!

Iolrpof Teifo`srpewofi y Y Y + HE

HE5ioliclrperaol-lh/hBX!

Rajc enrcp ys`jcphclic -m!

Ys`jcphcb

Iolrpof

4

3

54

53

04

03

4 3 54 53 04 03 94 93 24 23 34

HE ay pcusapcb nop ys`jcphclic*albsicbtcraofc cfolheraol

HE5-lh/hBX!e

nrcp02m

Eap

Eap I0M2

I0M2

P, tefsyrpay

P, tefsyrpay


22/35

C}telyal rpelyipatraol elb tporcal e`slbelic alipceycy stol ys`jcphclic

P, tefsyrpay

P, eicroye


23/35


24/35

P, tefsyrpay

P, eicroye

eap . ys`jcphcb

tcraofcy

lopjefawcb iBLE fa`pepq

232 ycusclialh -234 `t!. bc lo~o eyycj`fq

Pcncpclic rpelyipatrojc

-`feyr ehealyr Epe`abotyaynop elloreraol!

tcraofcy7 2m eap& 2m ys`jcphcb

tcraofcy7 54m eap& 54m ys`jcphcb

lol*lopjefawcb iBLE fa`pepq

Affsjale/yofc}e ycusclialh -83`t!

eap ys`jcphcb

jet yofc}e pceby

eap

ys`jcphcb

Rpelyipatrojc elefqyay syalh PLE ycu rcimlofohq


25/35

0&11055&49>9&55>

P, tefsyrpayP, eicroye

P, tefsyrpay

P, tefsyrpay P, tefsyrpay

P, tefsyrpay

P, eicroye P, eicroye

P, eicroyeP, eicroye

2=

2=

18

18

54

54

91

91

55>2099

>=3

55

89

83 5>

53=4

T*~efsc : 4,45& vNIv1 epc banncpclraeffq c}tpcyycb!

Foh0-ITJ!/k` -ioslry tcp jaffaol tcp k`! Foh0-Nofb Imelhc! Nswwq K ifsyrcpalh on rpelyipatr e`slbelic


26/35

P, eicroye P, tefsyrpay

Eap Ys` Eap Ys` P, eicroye P,tefsyrpay

A

AA

AAA

A^

^

Â

ÂA

ÂAA

A]

]

Nswwq K ifsyrcpalh on rpelyipatr e`slbelicon 2>1 BCHy

A]RM 99ERC]FE9

4,324,1=

0,210,=3

AAKABEPATcpo}abeyc 90

*4,334,=4

9,8=2,14

AAA Ejjolasj Rp,0 *4,9> *0,5>

A^ ET0 rpely,neirop LE 0,25

^ Fato}qhcleyc5 LE 5,10

Â

PET0,50]RM 8CAL2M@5

4,>15,205,>5

*4,99

5,839,855,120,2=

ÂA LICB2 *4,=> *0,59

ÂAA Eftme*ejqfeyc5 9,89 LE

A] HE0O] *0,01 *0,8=

] ERC]T@5 LE *0,49

Foh nofb imelhc

P,eic P,tef


27/35

lejc E,

rmefaele

J,

rpsliersfe

T,

rpaimoiepte

P,iojjslay

@,

àyreimqol

O,yera~e

è yai mc fa}*foot*mcfa} - MFM! ''' '' ' '' ' ' '' '''

bc mq bp era ol pc yt ol ya~ c tp orc al - PB 00! ' '

tcpo}abeyc '' ''' ''' '' '''

lo marJETKK ''' '' '' ' '' ''

MYT/Imetcpolc ''' ''' ''' '

YEJ*bctclbclr jcrmqfrpelyncpeyc ' ''

rpelytoyol tporcal ''

`MFM/kabepa ''' '' '' ' ''

tcpo}abeyc ''' ''' ''' ''' ''

HRT albalh -PMB! '' '

tporcal kaleyc ''' ''' ''' ''' ''' '''

ES]/AEE ''' ''' ''' ''' '' ''

tporcal kaleyc -XEH5! ''' '' ''' '' '' ''

fatab rpelyncp tporcal/tporceyc alma arop ''' ''' '' ''' '' ''

wali nalhcp -IOT5! ''' ''' ''' ''' ''' '''

hcpjal ''' ' ' '' '' ''lo mar

LE B+ E BT*paòyq frpelyn cpe yc -YPO3! '' '' '' ' '

lol yqjàorai mecjohfoàl ''' ''' ''' ''' '''

imaraleyc ' ' '

0OH*Nc-AA! o}qhcleyc '' '''

ys`rafeyc '

YlPK5 cre ys`slar ''' ''' ''' ''' '' ''

clbotctrabeyc almaàrop '

slkloxl '' '' '' '' ''

bcmqbpohcleyc/pcbsireyc '' '' ' '

slkloxl tporcal ' '' ' ' '

JQ@ -31! ''' ''' ''' ''' ''' '''

fato}qhcleyc ''' ''' ''' '' ''

hfsrermaol Y*rpelyncpeyc '' ' '' '' '''

bcmqbperaol pcytol~a~c pcfercb '

hfsrermaol Y*rpelyncpeyc '' '' '' '

m ef oeia b b cm ef ohcl eyc m qb pofeyc ''' ''' ''' ''' ''' '''

hfqioyabc mqbpofeyc/imaraleyc ''' ' ''' ''' '''

XB24 cre tpotcffcp ''' ''' ''' ''' '''

ysfnorpelyncpeyc ' '' ''' ''' '' ''

iqroimpojc T234 '' '' '' '' ' ''

eyteprqf tporceyc ''' '' ''' ''' ''

slkloxl tporcal ' ''

lo mar

SBT*hfqioyqfrpelyncpeyc ' ' ' '

tciral jcrmqfcyrcpeyc almaàrop '' ' ''' '' '

XPKQ -32! '' '' ' 'ellc}al '' ''' ''' '''

èyai mcfa}*foot*mcfa} -`MFM! '' ' '' ''' '''

P, eicroyeP, tefsyrpay

Hclc nejafacy xarm e yahlanaielr ytciacy ] rpcerjclr alrcpeiraol -T64,45!


28/35

lejc E,

rmefaele

J,

rpsliersfe

T,

rpaimoiepte

P,iojjslay

@,

àyreimqol

O,yera~e

èyai mcfa }* foot*mcfa } -`MFM! ''' ''' ''' ''' '''

bcmqbperaol pcytolya~ c tporca l -PB00! ' '

tcpo}abeyc '' ''' ''' '' '''

lo mar

JETKK ''' '' '' ' '' ''

MYT/Imetcpolc ''' ''' ''' '

YEJ*bctclbclr jcrmqfrpelyncpeyc ' ''

rpelytoyol tporcal ''

`MFM/kabepa ''' '' '' ' ''

tcpo}abeyc ''' ''' ''' ''' ''

HRT albalh -PMB! '' '

tporcal kaleyc ''' ''' ''' ''' ''' '''

ES]/AEE ''' ''' ''' ''' '' ''

tporcal kaleyc -XEH5! ''' '' ''' '' '' ''

fatab rpelyncp tporcal/tporceyc alma arop ''' ''' '' ''' '' ''

wali nalhcp -IOT5! ''' ''' ''' ''' ''' '''

hcpjal ''' ' ' '' '' ''

lo marLEB+ EBT*paòyq frpel yncpeyc -YPO3! '' '' '' ' '

lol yqjàorai mecjohfoàl ''' ''' ''' ''' '''

imaraleyc ' ' '

0OH*Nc-AA! o}qhcleyc '' '''

ys`rafeyc '

YlPK5 cre ys`slar ''' ''' ''' ''' '' ''

clbotctrabeyc almaàrop '

slkloxl '' '' '' '' ''

bcmqbpohcleyc/pcbsireyc '' '' ' '

slkloxl tporcal ' '' ' ' '

JQ@ -31! ''' ''' ''' ''' ''' '''

fato}qhcleyc ''' ''' ''' '' ''hfsrermaol Y*rpelyncpeyc '' ' '' '' '''

bcmqbperaol pcytol~a~c pcfercb '

hfsrermaol Y*rpelyncpeyc '' '' '' '

mefoeiab bcmefohcleyc mqbpofeyc ''' ''' ''' ''' ''' '''

hfqioyabc mqbpofeyc/imaraleyc ''' ' ''' ''' '''

XB24 cre tpotcffcp ''' ''' ''' ''' '''

ysfnorpelyncpeyc ' '' ''' ''' '' ''

iqroimpojc T234 '' '' '' '' ' ''

eyteprqf tporceyc ''' '' ''' ''' ''

slkloxl tporcal ' ''

lo mar

SBT*hfqioyqfrpelyncpeyc ' ' ' 'tciral jcrmqfcyrcpeyc almaàrop '' ' ''' '' '

XPKQ -32! '' '' ' '

'' ''' ''' '''

P, eicroye P, tefsyrpay

Tcp>2Tcp90 M`5 IOT5 XEH5 KBP YMQ0


29/35

Fahmr yahlefy -P& @!

MNP5MQ3FEN5

Fahmr tmclorqtc-lol*cfolhercb!

KABEPABchpeberaol

IOT5

KABEPA elb IOT57pchsferopy on slbcpxercp ymoor cfolheraol?


30/35

Jegop nalbalhy7

0, P, tefsyrpayay imepeircpawcb `q crmqfclc*jcbaercb tcraofc cfolheraol stolys`jcphclic& P, eicroye`q crmqfclc*bpa~cl alma`araol on cfolheraol,

9, Pchsferaol on clbohclosy E@E tfeqy el ajtoprelr pofc al rmc iolrpeyralh cfolheraol

pcytolycy on rmc rxo yrsbacb Psjc}ytciacy,

5, Psjc} tefsyrpayelb P, eicroyeme~c banncpclr nacfb foieraoly imepeircpawcb`q iolrpeyralh nfoobalh pchajcy,

2, PLE ycu rpelyipatrojc elefqyay pc~cefcb ettpo}, 52&444 hclc nejafacy al `orm ytciacy,

3, Al P, eicroye524 nejafacy epc banncpclraeffq c}tpcyycb stol ys`jcphclical fahmr& 98> al P, tefsyrpay,

>, PLE ycu pcysfry iolnapj ajtoprelr pofcy on E@E& es}al elb icff xeffy al ys`jcphclic*albsicb tcraofc cfolheraol,

8, Ajtoprelr lo~cf hclcy epc bayio~cpcb rmer bcjolyrperc ~cpq yrpolh c}tpcyyaolbanncpclicy `crxccl rmc rxo ytciacy,


31/35

Eikloxfcbhcjclry

Npelicyio FaiesyaTacpbojclaio TcpereGooyr ~el Bolhcl

Gsfae @eafcq*YcppcyCpai ^ayycpPeymja YeyabmepelPolefb TacpakBaik ^pcshbclmafOfc Tcbcpycl

Mely ~el ^cclBa~qe ^eymayrPo`cpr ^pcc`sphGopay @clyimotJepragl ~el WelrclElkac Ejjcpfeel

Po` Xcfyimcl

Iolncpclic AYTE 8*55 Oiro`cp 0459 APPA& Foy @eoy


32/35

C}tpcyyaol on Psjc}oprmofohscy on PET0,50ioppcfercy xarm nfoobalh rofcpelic

Faiesya cr ef,& 0455& Lerspc


33/35

Rmc Jcrmaolalc*Iqyrcalc ejalo eiab ycusclic er rmc L*rcpjalef tepr onPET0,50 usefanacy ar ey e ielbaberc ys`yrperc on rmc L*clb psfc termxeq

Jcrmaolalc ejalotctrabeyc

Ephalalc rpelyncpeyc

Tporcofqyay >


34/35

Tc

raofcfclhrm

-#oneapiolrpof!

Crmqfclc yrajsfercy tcraofc cfolheraol al P, tefsyrpayelb alma`ary ar al P, eicroyeTcraofc cfolheraol ay iesycb `q icff cfolheraol elb ay cuseffq bayrpa`srcb o~cp rmc tcraofc

P,eicroye

P,tefsyrpay


35/35

ayorahy xarm 6 34 pceby xcpc bayiepbcb elb ayorahyxarm 6 94 pceby tcp k` xcpc bayiepbcb

yahlanaielr pchsferaol T 6 4,445

Documents

Flooding tolerance: escape or quiescence