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Community Ecology, BIOL 7083 – Fall 2018 Louisiana State University – Department of Biological Sciences

Reference List I have presented the references under topic headings; the order corresponds to the order in which I have presented the topics in previous versions of the course (the order is a bit different from the order in which topics are presented in the textbook we are using this semester, i.e., G. Mittelbach’s [2012] Community Ecology). For a reference that I used to prepare lectures under multiple topic headings, its citation information appears only once, so if you are looking for a particular reference you might have to search the document for the author if you do not find that reference under a particular topic. This is a work-in-progress, so if you find mistakes and want to send me the corrections, or if you can add the missing details to incomplete entries, please send me e-mail: kharms@lsu.edu 1. What is Community Ecology?

Additional references: Abrahamson, Warren G., ed. 1989. Plant-Animal Interactions. McGraw-Hill Publishing, New

York, NY. Agrawal, Anurag A., David D. Ackerly, Fred Adler, A. Elizabeth Arnold, Carla Cáceres,

Daniel F. Doak, Eric Post, Peter J. Hudson, John Maron, Kailen A. Mooney, Mary Power, Doug Schemske, Jay Stachowicz, Sharon Strauss, Monica G. Turner & Earl Werner. 2007. Filling key gaps in population and community ecology. Frontiers in Ecology and the Environment 5:145-152.

Barash, David P. 1973. The ecologist as Zen master. The American Midland Naturalist 89:214-217.

Bawa, Kamaljit S., W. John Kress, Nalini M. Nadkarni & Sharachchandra Lele. 2004. Beyond paradise – meeting the challenges in tropical biology in the 21st century. Biotropica 36:437-446.

Bray, J. Roger & J. T. Curtis. 1957. An ordination of the upland forest communities of southern Wisconsin. Ecological Monographs 27:325-349.

Brown, J. H. & B. A. Maurer. 1989. Macroecology: the division of food and space among species on continents. Science 243:1145-1150.

Carothers, John H. 1986. Homage to Huxley: on the conceptual origin of minimum size ratios among competing species. American Naturalist 128:440-442.

Carpenter, Stephen R., E. Virginia Armbrust, Peter W. Arzberger, F. Stuart Chapin III, James J. Elser, Edward J. Hackett, Anthony R. Ives, Peter M. Kareiva, Mathew A. Leibold, Per Lundberg, Marc Mangel, Nirav Merchant, William W. Murdoch, Margaret A. Palmer, Debra P. C. Peters, Steward T. A. Pickett, Kathleen K. Smith, Diana H. Wall & S.

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Zimmerman. 2009. Accelerate synthesis in ecology and environmental sciences. BioScience 59.

Clements, Frederic E. 1936. Nature and structure of the climax. Journal of Ecology 24:252-284.

Cody, M. L. & J. M. Diamond, eds. 1975. Ecology and Evolution of Communities. Belknap Press of Harvard University Press, Cambridge, MA. [Dedicated to Robert MacArthur]

Cody & Smallwood. Long-term Studies of Vertebrate Communities. Connell, Joseph H. 1961. Effects of competition, predation by Thais lapillus, and other

factors on natural populations of the barnacle Balanus balanoides. Ecological Monographs 31:61-104.

Connell, Joseph H. 1961. The influence of competition and other factors on the distribution of the barnacle Chthamalus stellatus. Ecology 42:710-723.

Cowles, Henry Chandler. 1899. The ecological relations of the vegetation on the sand dunes of Lake Michigan. The Botanical Gazette 27:95-117, 167-202, 281-308, 361-391.

Damschen et al. 2005. Visibility matters: increasing knowledge of women’s contributions to ecology. Front. Ecol. Environ. 3:212-219.

Darwin, Charles. 1859. The Origin of Species by Means of Natural Selection. Diamond, J. & T. J. Case, eds. 1986. Community Ecology. Harper and Row, NY, NY.

[Dedicated to G. Evelyn Hutchinson] Dobzhansky, Theodosius. 1973. Nothing in biology makes sense except in the light of

evolution. The American Biology Teacher 35:125-129. Fauth, J. E., J. Bernardo, M. Camara, W. J. Resetarits, Jr., J. Van Buskirk & S. A. McCollum.

1996. Simplifying the jargon of community ecology: A conceptual approach. The American Naturalist 147:282-286.

Forbes, Stephen A. 1887. The lake as a microcosm. [Reprinted in Bulletin of the Illinois State Natural History Survey 15 (1925):537-550]

Gleason, H. A. 1926. The individualistic concept of the plant association. Bulletin of the Torrey Botanical Club 53:7-26.

Gotelli, Nicholas J. 2001. A Primer of Ecology, 3rd ed. Sinauer Assocs., Inc., Sunderland, MA.

Gotelli, Nicholas J. & Gary R. Graves. 1996. Null Models in Ecology. Smithsonian Institution Press, Washington, DC.

Gotelli, N. J. & B. McGill. 2006. Null versus neutral models: What’s the difference? Ecography 29:793-800.

Grant, Peter R. & B. Rosemary Grant. 2008. How and Why Species Multiply: The Radiation of Darwin’s Finches. Princeton University Press, Princeton, NJ.

HilleRisLambers, J., P. B. Adler, W. S. Harpole, J. M. Levine & M. M. Mayfield. 2012. Rethinking community assembly through the lens of coexistence theory. Annual Review of Ecology, Evolution & Systematics 43:227-248.

Holdridge, L. R. 1947. Determination of world plant formations from simple climatic data. Science 105:367-368.

Howe, H. F. & L. C. Westley. 1988. Ecological Relationships of Plants and Animals. Oxford University Press, Oxford, UK.

Huston, Michael A. & Steve Wolverton. 2011. Regulation of animal size by eNPP, Bergmann’s rule, and related phenomena. Ecological Monographs 81:349-405.

Hutchinson, G. Evelyn. 1951. Copepodology for the ornithologist. Ecology 32:571-577.

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Hutchinson, G. Evelyn. 1965. The Ecological Theater and the Evolutionary Play. Yale University Press, New Haven, CT.

Hutchinson, G. Evelyn. 1967. A treatise on limnology. Vol. II. John Wiley & Sons, New York, NY.

Huxley, J. 1942. Evolution: the modern synthesis. Harper, New York, NY. Inchausti, Pablo. 1994. Reductionist approaches in community ecology. Am. Nat. 143:201-

221. Johnson, Marc T. J. & John R. Stinchcombe. 2007. An emerging synthesis between

community ecology and evolutionary biology. Trends in Ecology & Evolution 22:250-257. Kaspari, Michael. 2008. Knowing your warblers: thoughts on the 50th anniversary of

MacArthur (1958). Bulletin of the Ecological Society of America. October:448-458. Keddy, Paul & Evan Weiher. 1999. Introduction: The scope and goals of research on

assembly rules. Pp. 1-20 in E. Weiher & P. Keddy, eds., Ecological Assembly Rules: Perspectives, Advances, Retreats. Cambridge University Press, Cambridge, UK. [Useful background re the “agony of community ecology” and assembly rules]

Kingsland, Sharon E. 2005. The Evolution of American Ecology 1890-2000. The Johns Hopkins University Press, Baltimore, MD.

Lawton, John H. 1999. Are there general laws in ecology? Oikos 84:177-192. Leibold, M. A., M. Holyoak, N. Mouquet, P. Amarasekare, J. M. Chase, M. F. Hoopes, R. D.

Holt, J. B. Shurin, R. Law, D. Tilman, M. Loreau & A. Gonzalez. 2004. The metacommunity concept: a framework for multi-scale community ecology. Ecology Letters 7:601-613.

Leibold, Mathew A. & Mark A. McPeek. 2006. Coexistence of the niche and neutral perspectives in community ecology. Ecology 87:1399-1410.

Lewontin, R. C. 1974. The Genetic Basis of Evolutionary Change. Columbia U. Press, N.Y. [P. A. Keddy (1992; Keddy & Weiher chapt. 1999, pg. 19) refers to Lewontin’s description of “the agony of community ecology”]

Lindeman, Raymond L. 1942. The trophic-dynamics aspect of ecology. Ecology 23:399-418. MacArthur, Robert H. 1958. Population ecology of some warblers of northeastern coniferous

forests. Ecology 39:599-619. MacArthur, Robert H. 1971. Patterns of terrestrial bird communities. Pp. 189-221 in D. S.

Farner & J. R. King, eds. Avian Biology. Academic Press, New York, NY. MacArthur, Robert H. 1972. Geographical Ecology: Patterns in the Distribution of Species.

Princeton University Press, Princeton, NJ. Marquet, P. A., A. P. Allen, J. H. Brown, J. Dunne, B. J. Enquist, J. Gillooly, P. A. Gowaty, J.

L. Green, D. Storch, J. Harte, S. P. Hubbel, J. O'Dwyer, Jordan Oki, A. Ostling, M. Ritchie & G. West. 2014. On theory in ecology. Bioscience 64:701-710.

Martínez del Rio, Carlos & Arthur D. Middleton. 2010. Laws for ecology? Ecology 91:1244-1245. [Book review of Dodds, Walter K. 2009. Laws, theories, and patterns in ecology.]

May, Robert M. 1986. The search for patterns in the balance of nature: advances and retreats. Ecology 67:1115-1126.

Matthews, W. J. 1998. Patterns in Freshwater Fish Ecology. Chapman and Hall, NY, NY. McGill, Brian J., Brian J. Enquist, Evan Weiher & Mark Westoby. 2006. Rebuilding

community ecology from functional traits. TREE 21:178-185. McIntosh, R. P. 1985. The Background of Ecology. Cambridge University Press.

Cambridge, UK.

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McLachlan, Athol J. & Richard J. Ladle. 2010. Barriers to adaptive reasoning in community ecology. Biol. Rev.

McPeek, Mark A. 2017. Evolutionary Community Ecology. Princeton University Press, Princeton, NJ.

McShea, Daniel W. & Robert N. Brandon. 2010. Biology’s First Law: The Tendency for Diversity and Complexity to Increase in Evolutionary Systems. University of Chicago Press, Chicago, IL.

Merriam. 1889. Mittelbach, Gary G. 2012. Community Ecology. Sinauer, Sunderland, MA. Mittelbach, Gary G. & Douglas W. Schemske. 2015. Ecological and evolutionary

perspectives on community assembly. TREE… Morin, Peter J. 1999. Community Ecology. Blackwell Science, Inc., Oxford, U.K. Morin, Peter J. 2011. Community Ecology, 2nd ed. Wiley-Blackwell, Oxford, U.K. Myers, Jonathan A. & Joseph A. LaManna. 2016. The promise and pitfalls of β-diversity in

ecology and conservation. Journal of Vegetation Science 27:1081-1083. Paine, Robert T. 2010. Macroecology: Does it ignore or can it encourage further ecological

syntheses based on spatially local experimental manipulations? The American Naturalist 176:385-393.

Pianka, E. R. 1992. The state of the art in community ecology. Proceedings of the First World Congress of Herpetology. In Herpetology: Current Research on the Biology of Amphibians & Reptiles, ed. K. Adler, pp. 141-162. Society for the Study of Amphibians & Reptiles, Oxford, OH. [“community ecology has for too long been perceived as repugnant and intractably complex… the discipline has been neglected and now lags far behind the rest of ecology”]

Real, Leslie A. & James H. Brown, eds. 1991. Foundations of Ecology: Classic Papers with Commentaries. The University of Chicago Press, Chicago, IL.

Reiners, William A., Jeffrey A. Lockwood, Derek S. Reiners & Steven D. Prager. 2017. 100 years of ecology: what are our concepts and are they useful? Ecological Monographs 87:260-277.

Ricklefs, Robert E. 2004. A comprehensive framework for global patterns in biodiversity. Ecology Letters 7:1-15.

Ricklefs, Robert E. 2008. Disintegration of the ecological community. The American Naturalist 172:741-750.

Root, R. B. 1967. The niche exploitation pattern of the blue-gray gnatcatcher. Ecological Monographs 37:317-350.

Roughgarden, Joan. 2009. Is there a general theory of community ecology? Biol. Philos. 24:521-529.

Scheiner, Samuel M. 2010. Toward a conceptual framework for biology. The Quarterly Review of Biology 85:293-318.

Scheiner, Samuel M. & Michael R. Willig. 2008. A general theory of ecology. Theoretical Ecology 1:21-28.

Scheiner, Samuel M. & Michael R. Willig. 2011. A general theory of ecology. Pp. 3-18 in S. M. Scheiner & M. R. Willig, eds. The Theory of Ecology. The University of Chicago Press, Chicago, IL.

Schoener, Thomas W. 2011. The newest synthesis: understanding the interplay of evolutionary and ecological dynamics. Science 331:426-429.

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Shipley, Bill & Paul A. Keddy. 1987. The individualistic and community-unit concepts as falsifiable hypotheses. Vegetatio 69:47-55.

Simberloff, D. 1983. Sizes of coexisting species. Pp. 404-430 in D. J. Futuyma & M. Slakin, eds. Coevolution. Sinauer, Sunderland, MA.

Simberloff, Daniel. 2004. Community ecology: is it time to move on? The American Naturalist 163:787-799.

Smith, F. A., S. K. Lyons, S. K. M. Ernest & J. H. Brown. 2008. Macroecology: more than the division of food and space among species on continents. Progress in Physical Geography 32:115-138.

Strong, D. R., Jr. 1980. Null hypotheses in ecology. Synthese 43:271-285. Strong, D. R., Jr., D. Simberloff, L. G. Abele & A. B. Thistle, eds. 1984. Ecological

Communities: Conceptual Issues and the Evidence. Princeton University Press, Princeton, NJ. [This volume was in part a reaction to Cody & Diamond (1975), see pg. 4; Dedicated to Richard P. Seifert]

Tansley, A. G. 1935. The use and abuse of vegetational concepts and terms. Ecology 16:284-308. [Coined the term “ecosystem”; for commentary on the paper, see D. Urban (2014) Bulletin of the Ecological Society of America 95:124-125]

Tewksbury, Joshua J., John G. T. Anderson, Jonathan D. Bakker, Timothy J. Billo, Peter W. Dunwiddie, Marth J. Groom, Stephanie E. Hampton, Steven G. Herman, Douglas J. Levey, Noelle J. Machnicki, Carlos Martínez del Rio, Mary E. Power, Kirsten Rowell, Anne K. Salomon, Liam Stacey, Stephen C. Trombulak & Terry A. Wheeler. 2014. Natural history’s place in science and society. BioScience 64:300-310.

Thompson, John N., O. J. Reichman, Peter J. Morin, Gary A. Polis, Mary E. Power, Robert W. Sterner, Carol A. Couch, Laura Gough, Robert Holt, David U. Hooper, Felicia Keesing, Charles R. Lovell, Bruce T. Milne, Manuel C. Molles, David W. Roberts & Sharon Y. Strauss. 2011. Frontiers of Ecology. BioScience:51:15-24.

Thoreau, Henry David. ~1860. Faith in a Seed. Turchin, P. 2001. Does population ecology have general laws? Oikos 94:17-26. Turner, Monica G. 1989. Landscape ecology: The effect of pattern on process. Annual

Review of Ecology & Systematics 20:171-197. Turner, Monica G. 2005. Landscape ecology in North America: Past, present, and future.

Ecology 86:1967-1974. Turner, Monica G. 2005. Landscape ecology: What is the state of the science? Annual

Review of Ecology, Evolution & Systematics 36:319-344. Urban, D. L. et al. 2002. Extending community ecology to landscapes. Ecoscience 9:200-

212. Vellend, Mark. 2010. Conceptual synthesis in community ecology. The Quarterly Review of

Biology 85:183-206. Vellend, Mark & Monica A. Geber. 2005. Connections between species diversity and genetic

diversity. Ecology Letters 8:767-781. Vellend, Mark & John L. Orrock. 2010. Ecological and genetic models of diversity: Lessons

across disciplines. Pp. 439-461 in J. B. Losos & R. E. Ricklefs, eds. The Theory of Island Biogeography at 40: Impacts and Prospects. Princeton University Press, Princeton, NJ.

von Bertalanffy, Ludwig. 1968. General System Theory: Foundations, Development, Applications.

Vorhoef & Morin. 2010. Community Ecology: Processes, Models, and Applications.

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Watt, A. S. 1947. Pattern and process in the plant community. Journal of Ecology 35:1-22. [For commentary on the paper, see D. Urban (2014) Bulletin of the Ecological Society of America 95:124-125]

Wethey, David. 1984. Biological Bulletin. [Repeat of Connell’s studies in New England.] Whittaker, Robert H. Ecol. Monogr. 26:1-69. Whittaker, Robert H. 1975. Communities and Ecosystems, 2nd ed. MacMillan, New York,

NY. Whittaker, Robert H. & W. A. Niering. 1965. Vegetation of the Santa Catalina Mountains,

Arizona: a gradient analysis of the south slope. Ecology 46:429-452. Whittaker, Robert H. & W. A. Niering. 1975. Vegetation of the Santa Catalina Mountains,

Arizona. V. Biomass, production, and diversity along the elevation gradient. Ecology 56:771-790.

Woodward, F. I., M. R. Lomas & C. K. Kelly. 2004. Global climate and the distribution of plant biomes. Phil. Trans. R. Soc. Lond. B 359:1465-1476.

2. Community-level patterns: Species richness; alpha, beta & gamma diversity; some geographic patterns (e.g., species-area curves, ecotones), the processes that cause them (e.g., Island Biogeography Theory), and considerations of scale; etc.

Additional references: Adams, Benjamin J., Stefan A. Schnitzer & Stephen P. Yanoviak. 2016. Trees as islands:

canopy ant species richness increases with the size of liana-free trees in a Neotropical forest. Ecography 39:1-9.

Allouche, Omri, Michael Kalyuzhny, Gregorio Moreno-Rueda, Manuel Pizarro & Ronen Kadmon. 2012. Area-heterogeneity tradeoff and the diversity of ecological communities. Proceedings of the National Academy of Sciences 109:17495-17500.

Brown, James H. 1995. Macroecology. The University of Chicago Press, Chicago, IL. Brown, James H. 2014. Why are there so many species in the tropics? Journal of

Biogeography 41:8-22. [Emphasizes kinetics – “the Red Queen runs fasters when she is hot;” see also: Field & Svenning (2014) Tropical diversity and the energetic ecology of the Red Queen. Journal of Biogeography 41:6-7.]

Caley, M. Julian & Dolph Schluter. 1997. Relationship between local and regional diversity. Ecology 78:70-80.

ChaseJM_TMKnight_2013_EcologyLetters_StandarizedSamplingIsNotEnough.pdf Chave, Jérôme. 2013. The problem of pattern and scale in ecology: what have we learned in

20 years? Ecology Letters. Chiarucci, Alessandro, Giovanni Bacaro & Samuel M. Scheiner. 2011. Old and new

challenges in using species diversity for assessing biodiversity. Philosophical Transactions of the Royal Society B 366:2426-2437.

Chown, Steven L., Brent J. Sinclair, Hans P. Leinaas & Kevin J. Gaston. 2004. Hemispheric asymmetries in biodiversity – a serious matter for ecology. PLoS Biology 2:1701-1707.

Colwell, R. EstimateS. [Freeware software for estimating a variety of variables useful for describing community-level patterns; available at: http://viceroy.eeb.uconn.edu/estimates]

Colwell, Robert K., Anne Chao, Nicholas J. Gotelli, Shang-Yi Lin, Chang Xuan Mao, Robin L.

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Chazdon & John T. Longino. 2012. Models and estimators linking invidividual-based and sample-based rarefaction, extrapolation and comparison of assemblages. Journal of Plant Ecology 5:3-21.

Condit, Richard, Robin B. Foster, Stephen P. Hubbell, R. Sukumar, Egbert G. Leigh, N. Manokaran, Suzanne Loo de Lao, James V. LaFrankie & Peter S. Ashton. 1998. Assessing forest diversity on small plots: Calibration using species-individual curves from 50-ha plots. Pp. 247-268 in F. Dallmeier & J. A. Comisky, eds., Forest Biodiversity: Research, Monitoring and Modeling. UNESCO, Paris, France.

Condit, Richard, Nigel Pitman, Egbert G. Leigh Jr., Jérôme Chave, John Terborgh, Robin B. Foster, Percy Nunez V., Salomon Aguilar, Renato Valencia, Gorky Villa, Helene C. Muller-Landau, Elizabeth Losos & Stephen P. Hubbell. 2002. Beta-diversity in tropical forest trees. Science 295:666-669.

Crist, Thomas O., Joseph A. Veech, Jon C. Gering & Keith S. Summerville. 2003. Partitioning species diversity across landscapes and regions: a hierarchical analysis of a, b, and g diversity. Am. Nat. 162:734-743.

Currie, D. J., G. G. Mittelbach, H. V. Cornell, R. Field, J.-F. Guégan, B. A. Hawkins, D. M. Kaufman, J. T. Kerr, T. Oberdorff, E. O’Brien, J. R. G. Turner. 2004 Predictions and tests of climate-based hypotheses of broad-scale variation in taxonomic richness. Ecology Letters 7(12): 1121-1134. [Sebastian pointed out their 3 hypotheses; along with Evans et al. (2005) – 9 hypotheses.]

Feest, Alan, Timothy D. Aldred & Katrin Jedamzik. 2010. Biodiversity quality: A paradigm for biodiversity. Ecological Indicators 10:1077-1082.

Gaston, K. J. & T. M. Blackburn. 1999. Macroecology: A critique. Oikos 84:353-368. Gaston, K. J., T. M. Blackburn & J. I. Spicer. 1998. Rapaport’s Rule: Time for an Epitaph?

Trends in Ecology and Evolution 13:70-74. Gilpin, Michael E. & Jared M. Diamond. 1976. Calculation of immigration and extinction

curves from the species-area-distance relation. Proceedings of the National Academy of Sciences USA 73:4130-4134.

Gotelli, N. EcoSim. [Freeware software for a variety of null-model analyses within the realm of community ecology; available at: http://homepages.together.net/~gentsmin/ecosim.htm]

Gotelli, Nicholas J. & Robert K. Colwell. 2001. Quantifying biodiversity: procedures and pitfalls in the measurement and comparison of species richness. Ecology Letters 4:379-391.

Gotelli, N. J. et al. 2009. Patterns and causes of species richness: a general model for macroecology. Ecology Letters 12:873-886.

Gross, Katherine L., Gary G. Mittelback & Heather L. Reynolds. 2005. Grassland invasibility and diversity: Responses to nutrients, seed input, and disturbance. Ecology 86:476-486.

Harrison, Susan & Howard Cornell. 2008. Toward a better understanding of the regional causes of local community richness. Ecology Letters 11:969-979.

Harrison, Susan, Hugh D. Safford, James B. Grace, Joshua H. Viers & Kendi F. Davies. 2006. Ecological Monographs 76:41-56.

Hatton, Ian A., Kevin S. McCann, John M. Fryxell, T. Jonathan Davies, Matteo Smerlak, Anthony R. E. Sinclair & Michel Loreau. 2015. The predator-prey power law: Biomass scaling across terrestrail and aquatic biomes. Science 349:1070.

Hawkins, Bradford A., Jose Alexandre Felizola Diniz-Filho, Carlos A. Jaramillo & Stephen A. Soeller. 2006. Post-Eocene climate change, niche conservatism, and the latitudinal

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diversity gradient of New World birds. J. Biogeography Hawkins, B. A., J. A. F. Diniz Filho, C. A. Jaramillo and S. A. Soeller. 2007. Climate, niche

conservatism, and the global bird diversity gradient. The American Naturalist 170:S16-S27. [“The Amazon’s large and ancient landmass coupled with a warm climate probably buffered birds from ice age extinctions and aided in species diversification…” – from J. Wolfe’s dissertation.]

He & Hubbell. 2011. Nature. He, Fangliang & Stephen Hubbell. 2013. Estimating extinction from species-area

relationships: why the numbers do not add up. Ecology 94:1905-1912. He, Fangliang & Pierre Legendre. 2002. Species diversity patterns derived from species-area

models. Ecology 83:1185-1198. Holyoak, Marcel, Mathew A. Leibold & Robert D. Holt, eds. 2005. Metacommunities:

Spatial Dynamics and Ecological Communities. University of Chicago Press, Chicago, IL. Hillebrand, Helmut. 2005. Regressions of local on regional diversity do not reflect the

importance of local interactions or saturation of local diversity. Oikos 110:195-198. Hubbell, Stephen P. 2010. Neutral theory and the theory of island biogeography. Pp. 264-292

in Jonathan B. Losos & Robert E. Ricklefs, eds., The Theory of Island Biogeography Revisited. Princeton University Press, Princeton, NJ.

Hurlbert, S. H. 1971. The nonconcept of species diversity: A critique and alternative parameters. Ecology 52:577-586.

Hurlbert - unicorns Karlson, Ronald H., Howard V. Cornell & Terence P. Hughes. 2004. Coral communities are

regionally enriched along an oceanic biodiversity gradient. Nature 429:867-870. [Compare and contrast Wright et al. (2003) with Karlson et al. (2004). Wright et al. (2003) conclude that there is limited large-scale effect, whereas Karlson et al. (2004) show strong large-scale effect on local diversity.]

Kraft et al. 2011. Beta diversity… Science. Kreft, Holger & Walter Jetz. 2007. Global patterns and determinants of vascular plant

diversity. PNAS 104:5925-5930. Lande, R. 1996. Statistics and partitioning of species diversity, and similarity among multiple

communities. Oikos 76:5-13. Laurance, William F. 2008. Theory meets reality: How habitat fragmentation research has

transcended island biogeography theory. Biological Conservation 141:1731-1744. Levin, Simon A. 1992. The problem of pattern and scale in ecology. Ecology 73:1943-1967.

[For commentary on the paper, see D. Urban (2014) Bulletin of the Ecological Society of America 95:124-125]

Loreau, Michael. 2000. Are communities saturated? On the relationship between α, β and γ diversity. Ecology Letters 3:73-76.

Losos, Jonathan B. & Robert E. Ricklefs, eds. 2010. The Theory of Island Biogeography Revisited. Princeton University Press, Princeton, NJ.

MacArthur, Robert H. & Edward O. Wilson. 1967. The Theory of Island Biogeography. Princeton University Press, Princeton, NJ.

Magurran, Anne E. 1988. Ecological Diversity and its Measurement. Princeton University Press, Princeton, USA.

Magurran, Anne E. 2004. Measuring Biological Diversity. Blackwell Publishing, Oxford, U.K.

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Myers, Jonathan A., Jonathan M. Chase, Iván Jiménez, Peter M. Jørgensen, Alejandro Araujo-Murakami, Narel Paniagua-Zambrana & Renate Seidel. 2013. Beta-diversity in temperate and tropical forests reflects dissimilar mechanisms of community assembly. Ecology Letters 16:151-157.

Nekola, J. C. & P. S. White. 1999. The distance decay of similarity in biogeography and ecology. J. Biogeogr. 26:867-878.

Niering, W., R. Whittaker & C. Lowe. 1963. The Saguaro: a population in relation to its environment. Science 142:15-23.

Peet, Robert K. 1974. The measurement of species diversity. Annual Review of Ecology and Systematics 5:285-307.

Plotkin, Joshua B., Matthew D. Potts, Nandi Leslie, N. Manokaran, James LaFrankie & Peter S. Ashton. 2000. Species-area curves, spatial aggregation, and habitat specialization in tropical forests. Journal of Theoretical Biology 207:81-99.

Purvis, Andy & Andy Hector. 2000. Getting the measure of biodiversity. Nature 405:212-219.

Rahel, F.J. 1990. The hierarchical nature of community persistence: a problem of scale. The American Naturalist 136:328-344.

Ricklefs, Robert E. 1987. Community diversity: Relative roles of local and regional processes. Science 235:167-171.

Ricklefs, Robert E. & Susanne S. Renner. 2012. Global correlations in tropical tree species richness and abundance reject neutrality. Science 335:464-467.

Rosenzweig, Michael. 1995. Species diversity in space and time. Cambridge University Press, Cambridge, UK.

Rossi, Richard E., David J. Mulla, André G. Journel & Eldon H. Franz. 1992. Geostatistical tools for modeling and interpreting ecological spatial dependence. Ecological Monographs 62:277-314.

Scheiner, S. M. 1992. Measuring pattern diversity. Ecology 73:1860-1867. Scheiner, Samuel M., Alessandro Chiarucci, Gordon A. Fox, Matthew R. Helmus, Daniel J.

McGlinn & Michael R. Willig. 2011. The underpinnings of the relationship of species richness with space and time. Ecological Monographs 81:195-213.

Scheiner, S. M., S. B. Cox, M. R. Willig, G. G. Mittelbach, C. Osenberg, M. Kaspari. 2000. Species richness, species-area curves, and Simpson’s paradox. Evol. Ecol. Res. 2:791-802.

Simberloff, Daniel S. 1973. Species turnover and equilibrium island biogeography. Science 194:572-578.

Simberloff, Daniel S. 1974. Equilibrium theory of island biogeography and ecology. Annual Review of Ecology and Systematics 5:161-182.

Stevens, Richard D. & Michael R. Willig. 2002. Geographical ecology at the community level: Perspectives on the diversity of New World bats. Ecology 83:545-560.

Turchin, P. 1996. Fractal analyses of animal movement: a critique. Ecology 77:2086-2090. Veech, Joseph A., Keith S. Summerville, Thomas O. Crist & Jon C. Gering. 2002. The

additive partitioning of species diversity: recent revival of an old idea. Oikos 99:3-9. WalkerS_etal_2003_Journal_of_Vegetation_Science_PropertiesOfEcotones.pdf Wardle, David A., Gregor W. Yeates, Gary M. Barker, Peter J. Bellingham, Karen I. Bonner &

Wendy M. Williamson. 2003. Island biology and ecosystem functioning in epiphytic soil communities. Science 301:1717-1720.

Whittaker, R. H. 1972. Evolution and measurement of species diversity. Taxon 21:213-251.

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Wiens, J. A. 1989. Spatial scaling in ecology. Functional Ecology 3:385-397. Wilson, J. Bastow & Barbara J. Anderson. 2001. Species-pool relations: like a wooden light

bulb? Folia Geobotanica 36:35-44. [“The species pool concept is stimulating, but it is probably operationally impossible to test. Like a wooden light bulb, it is beautiful and interesting, but of little use” (Abstract)]

Wright, Justin P., Alexander S. Flecker & Clive G. Jones. 2003. Local vs. landscape controls on plant species richness in beaver meadows. Ecology 84:3162-3173. [Compare and contrast Wright et al. (2003) with Karlson et al. (2004). Wright et al. (2003) conclude that there is limited large-scale effect, whereas Karlson et al. (2004) show strong large-scale effect on local diversity.]

3. Community-level patterns: Relative abundance distributions

Additional references: See Wikipedia “Relative Species Abundance” Bell, Graham. 2001. Neutral macroecology. Science 293:2413-2418. Chave, Jérôme, Helene C. Muller-Landau & Simon A. Levin. 2002. Comparing classical

community models: Theoretical consequences for patterns of diversity. Am. Nat. 159:1-23.

Chisholm, Ryan A. & Stephen W. Pacala. 2010. Niche and neutral models predict asymptotically equivalent species abundance distributions in high-diversity ecological communities.

Conlisk et al. 2009. Hubbell’s local abundance distribution: insights from a simple colonization rule. Oikos.

Connolly, Sean R., Terry P. Hughes, David R. Bellwood & Ronald H. Karlson. 2005. Community structure of corals and reef fishes at multiple scales. Science 309:1363-1365.

de Aguiar, M. A. M., M. Baranger, E. M. Baptestini, L. Kaufman & Y. Bar-Yam. 2009. Global patterns of speciation and diversity. Nature 460:384-387.

Gotelli, Nicholas J. 2001. A Primer of Ecology, 3rd ed. Sinauer Assocs., Inc., Sunderland, MA. [Excellent overview of simple population and multi-species models.]

Gotelli, Nicholas J. & Gary R. Graves. 1996. Null Models in Ecology. Smithsonian Institution Press, Washington, DC.

Hubbell. 1979. Hubbell. Coral reef symposium. Hubbell, Stephen P. 2001. The Unified Neutral Theory of Biodiversity and Biogeography.

Princeton University Press, Princeton, NJ. [For class – Chapt. 2: On current theories of relative species abundance, pp. 30-47; Chapt. 3: Dynamical models of the relative abundance of species, pp. 48-75; Chapt. 4: Local Community Dynamics Under Ecological Drift, see first two pages, i.e., pp. 76-77]

May, Robert M. 1975. Patterns of species abundance and diversity. Pp. 81-120 in M. L. Cody & J. M. Diamond, eds., Ecology and Evolution of Communities. Belknap Press of Harvard University Press, Cambridge, MA.

May, Robert M. 1986. The search for patterns in the balance of nature: advances and retreats.

11

Ecology 67:1115-1126. [See especially the final sections] May, Robert M. 1988. How many species are there on Earth? Science 241:1441-1449. McGill, Brian J. 2003. A test of the unified neutral theory of biodiversity. Nature 422:881-

885. McGill, Brian J., Rampal S. Etienne, John S. Gray, David Alonso, Marti J. Anderson, Habtamu

Kassa Benecha, Maria Dornelas, Brian J. Enquist, Jessica L. Green, Fangliang He, Allen H. Hurlbert, Anne E. Magurran, Pablo A. Marquet, Brian A. Maurer, Annette Ostling, Candan U. Soykan, Karl I. Ugland & Ethan P. White. 2007. Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework. Ecology Letters 10:995-1015.

Nekola_Brown_2007_EcolLetters.pdf Preston, F. W. 1948. The commonness, and rarity, of species. Ecology 29:254-283. Preston, F. W. 1962. The canonical distribution of commonness and rarity. Ecology 43:185-

215, 410-432. Rosindell, James, Stephen P. Hubbell & Rampal S. Etienne. 2011. The Unified Neutral

Theory of Biodiversity and Biogeography at Age Ten. Trends in Ecology & Evolution 26:340-348.

Rosindell, James, Stephen P. Hubbell, Fangliang He, Luke J. Harmon & Rampal S. Etienne. 2012. The case for ecological neutral theory. Trends in Ecology & Evolution 27:203-208.

Volkov, Igor, Jayanth R. Banavar, Fangliang He, Stephen P. Hubbell & Amos Maritan. 2005. Density dependence explains tree species abundance and diversity in tropical forests. Nature 438:658-661.

Volkov, Igor, Jayanth R. Banavar, Stephen P. Hubbell & Amos Maritan. 2003. Neutral theory and relative species abundance in ecology. Nature 424:1035-1037.

Volkov, Igor, Jayanth R. Banavar, Stephen P. Hubbell & Amos Maritan. 2007. Patterns of relative species abundance in rainforests and coral reefs. Nature 450:45-49.

Volkov, Igor, Jayanth R. Banavar, Stephen P. Hubbell & Amos Maritan. 2009. Inferring species interactions in tropical forests. PNAS.

Whittaker, R. H. 1965. Dominance and diversity in land plant communities. Science 147:250-260.

Zillio, Tommaso & Richard Condit. 2007. The impact of neutrality, niche differentiation and species input on diversity and abundance distributions. Oikos 116:931-940.

4. Pair-wise species interactions: Competition. I. Theory Additional references: Abrams, Peter A. 1984. Recruitment, lotteries, and coexistence in coral reef fish. The

American Naturalist 123:44-55. Aschehoug, Erik T., Rob Brooker, Daniel Z. Atwater, John L. Maron & Ragan M. Callaway.

2016. The mechanisms and consequences of interspecific competition among plants. Annual Review of Ecology, Evolution & Systematics 47:263-281.

Bolker Brown, W. L., Jr. & E. O. Wilson. 1956. Character displacement. Systematic Zoology 5:49-

64.

12

Case, Ted J. & Michael E. Gilpin. 1974. Interference competition and niche theory. Proceedings of the National Academy of Science, USA 71:3073-3077.

Chase, Jonathan M. & Mathew A. Leibold. 2003. Ecological Niches: Linking Classial and Contemporary Approaches. Chicago University Press, Chicago, IL.

Connell, Joseph H. 1980. Diversity and the coevolution of competitors, or the ghosts of competition past. Oikos 35:131-138. [Get also J. Roughgarden’s criticism of the paper and Connell’s reply!]

Goldberg, D. E. 1996. Competitive ability: definitions, contingency and correlated traits. Philos. Trans. R. Soc. B 351:1377-1385.

Grace, James B. & David Tilman, eds. 1990. Perspectives on Plant Competition. Academic Press, Inc., San Diego, CA.

Grether, Gregory F., Neil Losin, Christopher N. Anderson & Kenichi Okamoto. 2009. The role of interspecific interference competition in character displacement and the evolution of competitor recognition. Biological Reviews.

Hardin, Garrett. 1960. The competitive exclusion principle. Science 131:1292-1297. [Coined the term “competitive exclusion principle”]

Horn, H. & R. MacArther. 1972. Competition among fugitive species in a harlequin environment. Ecology 53:749-752.

Horn & May. 1977. Hutchinson, G. Evelyn. 1957. Concluding remarks. Cold Spring Harbor Symposium on

Quantitative Biology 22:415-427. Keddy. Competition, 1st ed. Keddy, Paul A. 2001. Competition, 2nd ed. Kluwer Academic Pubs., Dordrecht, The

Netherlands. Lewin, Roger. 1983. Santa Rosalia was a goat. Science 221:636-639. MacArthur & Levins. 1964. McPeek, Mark A. 2012. Intraspecific density dependence and a guild of consumers coexisting

on one resource. Ecology 93:2728-2735. Miller, T. E., C. P. terHorst & J. H. Burns. 2009. The ghost of competition present. The

American Naturalist 173:347-353. Pacala, Stephen W. 1986. Neighborhood models of plant population dynamics. II. Multi-

species models of annuals. Theoretical Population Biology 29:262-292. Pacala, Stephen W. & David Tilman. 1994. Limiting similarity in mechanistic and spatial

models of plant competition in heterogeneous environments. The American Naturalist 143:222-257.

Pfennig, David W. & Karin S. Pfennig. 2012. Evolution’s Wedge: Competition and the Origins of Diversity. University of California Press, Berkeley, CA.

Pfennig, Karin S. & David W. Pfennig. 2009. Character displacement: ecological and reproductive responses to a common evolutionary problem. The Quarterly Review of Biology 84:253-276.

Rees, Mark & Joy Bergelson. 1997. Asymmetric light competition and founder control in plant communities. Journal of Theoretical Biology 184:353-358.

Riebesell, John F. 1974. Paradox of enrichment in competitive systems. Ecology 55:183-187. Roughgarden, Jonathan. 1983. Competition and theory in community ecology. American

Naturalist 122:583-601. [Criticism of: Connell (1980); Strong et al. (1979); Connor & Simberloff (1979)]

13

Schaffer, William M. & E. G. Leigh. 1976. The prospective role of mathematic theory in plant ecology. Systematic Botany 1:209-232. [Lotka-Volterra is inappropriate for sessile plants!].

Scheffer_vanNes_2006_PNAS.pdf Taper, Mark. 1993. A new look at competition theory and the structure of ecological

communities. Trends in Ecology and Evolution 8:308-309. Tilman, David. 1982. Resource Competition and Community Structure. Princeton University

Press, Princeton, NJ. Tilman, David. 1987. The importance of the mechanisms of interspecific competition. The

American Naturalist 129:769-774. Tilman, David. 1988. Dynamics and Structure of Plant Communities. Princeton University

Press, Princeton, NJ. Van Allen, Benjamin G. & Volker H. W. Rudolf. 2016. Carryover effects drive competitive

dominance in spatially structured environments. Proceedings of the National Academy of Sciences USA [carry-over effects can facilitate spatial coexistence]

Weiner, Jacob. 1990. Asymmetric competition in plant populations. Trends in Ecology and Evolution 5:360-364.

5. Pair-wise species interactions: Competition. II. Empirical underpinnings Additional references: Adler, Peter B., Stephen P. Ellner & Jonathan M. Levine. 2010. Coexistence of perennial

plants: an embarrassment of niches. Ecology Letters 13:1019-1029. Antonovics, Janis & Donald A. Levin. 1980. The ecological and genetic consequences of

density-dependent regulation in plants. Annual Review of Ecology and Systematics 11:411-452.

Ashton, Isabel, Amy E. Miller, William D. Bowman & Katherine N. Suding. 2010. Niche complementarity due to plasticity in resource use: plant partitioning of chemical N forms. Ecology 91:3252-3260.

Bolnick, Daniel I., Travis Ingram, William E. Stutz, Lisa K. Snowberg, On Lee & Jeff S. Paull. 2010. Ecological release from interspecific competition leads to decoupled changes in population and individual niche width. Proceedings of the Royal Society B 277:1789-1797.

Brown, James H. & Diane W. Davidson. 1977. Competition between seed-eating rodents and ants in desert ecosystems. Science 196:880-882. [Get also the newer paper – from Sebastian Tello’s presentation – in which rodent-ant interaction is actually indirect facilitation!]

Byers, James E. 2000. Competition between two estuarine snails: Implications for invasions of exotic species. Ecology 81:1225-1239.

Carrascal. L. M., J. L. Terrería & A. Valido. 1992. Habitat distribution of canary chaffinches among islands: Competitive exclusion or species-specific habitat preferences? Journal of Biogeography 19:383-390.

Connell, Joseph H. 1983. On the prevalence and relative importance of interspecific competition: Evidence from field experiments. The American Naturalist 122:661-696.

14

Coomes, David A. & Peter J. Grubb. 2000. Impacts of root competition in forests and woodlands: a theoretical framework and review of experiments. Ecological Monographs 70:171-207.

Damgaard, Christian & Adeline Fayolle. 2010. Measuring the importance of competition: a new formulation of the problem. J. Ecology 98:1-6.

Dhondt,André A. 2012. Interspecific Competition in Birds. Oxford U. Press, Oxford, UK. Duckworth, Renée A., Virginia Belloni & Samantha R. Anderson. 2015. Cycles of species

replacement emerge from locally induced maternal effects on offspring behavior in a passerine bird. Science 347:875-877. [Pg. 875 – “Interactions among individuals often drive population- and community-level patterns… and in turn, community dynamics and species interactions can affect individual variation in traits…”]

Dudley, Tom L., Carla M. D’Antonio & Scott D. Cooper. 1990. Mechanisms and consequences of interspecific competition between two stream insects. Journal of Animal Ecology 59:849-866.

Eberhard, William G. 1989. Niche expansion in the spider Wendilgarda galapagensis (Araneae, Theridiosomatidae) on Cocos Island. Rev. Biol. Trop. 37:163-168.

Ferson, Scott, Pat Downey, Paul Klerks, Marc Weissburg, Irwin Kroot, Steven Stewart, Geoff Jacquez, James Ssemakula, Richard Malenky & Karl Anderson. 1986. Competing reviews, or why do Connell and Schoener disagree? The American Naturalist 127:571-576.

Gause, G. F. 1932. Experimental studies on the struggle for existence. Journal of Experimental Biology 9:389-402.

Gause. 1934. [See Begon, Harper, Townsend] Goldberg, D. E. & A. M. Barton. 1992. Patterns and consequences of interspecific

competition in natural communities: A review of field experiments with plants. The American Naturalist 139:771-801.

Grace, James B. & David Tilman, eds. 1990. Perspectives on Plant Competition. Academic Press, Inc., San Diego, CA.

Grant, P. R. 1968. Bill size, body size, and the ecological adaptations of bird species to competitive situations on islands. Systematic Zoology 17:319-333.

Grant, P. R. 1972. Interspecific competition among rodents. Annual Reviews of Ecology & Systematics 3:79-106.

Grant, P. R. 1972. Convergent & divergent character displacement. Biological Journal of the Linnean Society 4:39-68.

Grant, P. R. 1975. The classical case of character displacement. Evolutionary Biology 8:237-337.

Grant, Peter R. 1986. Ecology & Evolution of Darwin’s Finches. Princeton University Press, Princeton, NJ.

Grant, P. R. & I. Abbott. 1980. Interspecific competition, null hypotheses and island biogeography. Evolution 34:332-341.

Hoogland, John L. & Charles R. Brown. 2016. Prairie dogs increase fitness by killing interspecific competitors. Proceedings of the Royal Society B…

Lawton, John. 1991. Warbling in different ways. Oikos 60:273-274. Lewis, Simon L. & Edmund V. J. Tanner. 2000. Effects of above- and belowground

competition on growth and survival of rain forest tree seedlings. Ecology 81:2525-2538. Maor, Roi, Tamar Dayan, Henry Ferguson-Gow & Kate E. Jones. 2017. Temporal niche

15

expansion in mammals from a nocturnal ancestor after dinosaur extinction. Nature Ecology & Evolution…

Pacala, Stephen & Jonathan Roughgarden. 1982. Resource partitioning and interspecific competition in two two-species insular Anolis lizard communities. Science 217:444-446.

Pacala, Stephen W. & Jonathan Roughgarden. 1985. Population experiments with the Anolis lizards of St. Maarten and St. Eustatius. Ecology 66:129-141.

Petren, Kenneth, Douglas T. Bolger & Ted J. Case. 1993. Mechanisms in the competitive success of an invading sexual gecko species over an asexual native. Science 259:354-358.

Platt, W. J. & I. M. Weiss. 1977. Resource partitioning and competition within a guild of fugitive prairie plants. The American Naturalist 111:479-513.

Richman, A. D. & T. Price. 1992. Evolution of ecological differences in the Old World leaf warblers. Nature 355:817-821.

Schluter, Dolph. 1994. Experimental evidence that competition promotes divergence in adaptive radiation. Science 266:798-801.

Schluter, Dolph & John Donald McPhail. 1992. Ecological character displacement and speciation in sticklebacks. The American Naturalist 140:85-108.

Schoener, T. W. 1983. Field experiments on interspecific competition. The American Naturalist 122:240-285.

Schoener, T. W. 1985. Some comments on Connell’s and my reviews of field experiments on interspecific competition. The American Naturalist 125:730-740.

Silander & Antonovics. 1982. Analysis of interspecific interactions in coastal plant community – a perturbation approach. Nature 298:557-560.

Silander, John A., Jr. & Stephen W. Pacala. 1985. Neighborhood predictors of plant performance. Oecologia 66:256-263.

Spiller, David A. 1984. Competition between two spider species: experimental field study. Ecology 65:909-919.

Tansley. 1917. [See Begon, Harper & Townsend] Townsend, Howard M., Kathryn P. Huyvaert, Peter J. Hodum & David J. Anderson. 2002.

Nesting distributions of Galápagos boobies (Aves: Sulidae): an apparent case of amensalism. Oecologia 132:419-427.

Welden, C. W. & W. L. Slauson. 1986. The intensity of competition versus its importance: An overlooked distinction and some implications. Quarterly Review of Biology 61:23-44.

6. Pair-wise species interactions: Predator-prey, parasitoid-host & parasite-host interactions

Additional references: Abrams, P. A. & C. J. Walters. 1996. Invulnerable prey and the paradox of enrichment.

Ecology 77:1125-1133. Agrawal, Anurag A. 2001. Phenotypic plasticity in the interactions and evolution of species.

Science 294:321-326. Altizer, Sonia, Rebecca Bartel & Barbara A. Han. 2011. Animal migration and infectious

disease risk. Science 331:296-302. Anderson, Roy M. & Robert M. May. 1979. Population biology of infectious diseases: Part I.

16

Nature 280:361-367. Anderson, Roy M. & Robert M. May. 1979. Population biology of infectious diseases: Part

II. Nature 280:455-461. Benincà, Klaus D. Jöhnk, Reinhard Heerkloss & Jef Huisman. 2009. Coupled predator-prey

oscillations in a chaotic food web. Ecology Letters 12:1-12. Berger, The Better to Eat You With. Berger et al. 2001. Science 291:1036-1039. Berryman, A. A. 1992. The origins and evolution of predator-prey theory. Ecology 73:1530-

1535. Brooks, J. L. & S. I. Dodson. 1965. Predation, body size, and composition of plankton.

Science 150:28-35. BuckJC_KDLafferty_AmNatRevision1_ScaleDependentSafetyInNumbers.pdf Byers, John A. 1998. American Pronghorn: Social Adaptations and the Ghosts of Predators

Past. University of Chicago Press, Chicago, IL. Canestrari, Daniela, Diana Bolopo, Ted C. J. Turlings, Gregory Röder, José M. Marcos &

Vittorio Baglione. 2014. From parasitism to mutualism: unexpected interactions between a cuckoo and its host. Science 343:1350-1352.

Davies, N. B. 1992. Dunnock Behavior and Social Evolution. Oxford University Press, Oxford, U.K.

DeLong, John P., Zeina Al-Ameeli, Garry Duncan, James L. Van Etten & David D. Dunigan. 2016. Predators catalyze an increase in chloroviruses by foraging on the symbiotic hosts of zoochlorellae. Proceedings of the National Academy of Sciences 113:13780-13784.

Dobson, Andy & Mick Crawley. 1994. Pathogens and the structure of plant communities. Trends in Ecology and Evolution 9:393-398.

Dobson, A. P. & P. J. Hudson. 1992. Regulation and stability of a free-living host-parasite system: Trichostrongylus tenuis in red grouse. II. Population models. Journal of Animal Ecology 61:487-498.

Eberhard, W. G. 1975. The ecology and behavior of a subsocial pentatomid bug and two scelionid wasps: strategy and counterstrategy in a host and its parasites. Smith. Contrib. Zool. 205:1-39.

Eberhard, William G. 2000. Spider manipulation by a wasp larva. Nature 406:255-256. Elton, C. & M. Nicholson. 1942. The ten-year cycle in numbers of the lynx in Canada.

Journal of Animal Ecology 11:215-244. Ewald, Paul W. 1994. Evolution of Infectious Disease. Oxford University Press, Oxford,

U.K. Feder, Martin E. & George V. Lauder. 1986. Predator-Prey Relationships: Perspectives and

Approaches from the Study of Lower Vertebrates. Chicago University Press, Chicago, IL. Gilbert, Gregory S. 2002. Evolutionary ecology of plant diseases in natural ecosystems. Ann.

Rev. Phytopathol. 40:13-43. Gilbert, Gregory S. & Campbell O. Webb. 2007. Phylogenetic signal in plant pathogen-host

range. PNAS 104:4979-4983. Gilg, Olivier, Ilkka Hanski & Benoît Sittler. 2003. Cyclic dynamics in a simple vertebrate

predator-prey community. Science 302:866-868. Glasser, J. W. 1979. The role of predation in shaping and maintaining the structure of

communities. The American Naturalist 113:631-641. Godfray, H. C. J. 1994. Parasitoids: Behavioral and Evolutionary Ecology. Princeton

17

University Press, Princeton, NJ. Godfray, H. C. J. & S. W. Pacala. 1992. Aggregation and the population dynamics of

parasitoids and predators. The American Naturalist 140:30-40. Gurney, W. S. C., S. P. Blythe & R. M. Nisbet. 1980. Nicholson’s blowflies revisited. Nature

287:17-21. Halfwerk, W., P. L. Jones, R. C. Taylor, M. J. Ryan & R. A. Page. 2014. Risky ripples allow

bats and frogs to eavesdrop on a multisensory sexual display. Science 343:413-416. [Could be good for discussion paper!]

Harrison, Gary W. 1995. Comparing predator-prey models to Luckinbill’s experiment with Didinium and Paramecium. Ecology 76:357-374.

Herre, Edward Allen. 1993. Population structure and the evolution of virulence in nematode parasites of fig wasps. Science 259:1442-1445.

Holling, C. S. 1959. The components of predation as revealed by a study of small mammal predation of the European pine sawfly. The Canadian Entomologist 91:293-320.

Hudson, Peter J., Andy P. Dobson & Dave Newborn. 1998. Prevention of population cycles by parasite removal. Science 282:2256-2258.

Huffaker, C. B. 1958. Experimental studies on predation: dispersion factors and predator-prey oscillations. Hilgardia 27:343-383 (or 795-834???).

Ims_etal_2011_PNAS_Lemmings.pdf Janzen, Daniel H. 1976. Why bamboos wait so long to flower. Annual Review of Ecology

and Systematics. 7:347-391. Jost, C. 2000. Predator-prey theory: Hidden twins in ecology and microbiology. Oikos

90:202-208. Keith, L. B. 1983. Role of food in hare population cycles. Oikos 40:385-395. Krebs, Charles J., Rudy Boonstra, Stan Boutin & A. R. E. Sinclair. 2001. What drives the 10-

year cycle of snowshoe hares? BioScience 51:25-35. Krebs, J. R. & N. B. Davies. 1984. Behavioural Ecology: An Evolutionary Approach. 2nd

Edition. Blackwell Scientific Publications, Oxford, UK. Krebs, J. R. & N. B. Davies. 1987. An Introduction to Behavioural Ecology. 2nd Edition.

Blackwell Scientific Publications, Oxford, UK. LaffertyKD_etal_2008_EcologyLetters_ParasitesInFoodWebs.pdf Light, Jessica E. & Mark S. Hafner. 2008. Codivergence in heteromyid rodents (Rodentia:

Heteromyidae) and their sucking lice of the genus Fahrenholzia (Phthiraptera: Anoplura). Systematic Biology 57:449-465.

Lively & Dybdahl [get refs. from Cain et al. Ecology 3rd ed.] Loyn, Richard H., Ross G. Runnalls & Gail Y. Forward. 1983. Territorial Bell Miners and

other birds affecting populations of insect prey. Science 221:1411-1413. Luckinbill, L. S. 1973. Coexistence in laboratory populations of Paramecium aurelia and its

predator Didinium nasutum. Ecology 54:1320-1327. Luckinbill, L. S. 1974. The effects of space and enrichment on a predator-prey system.

Ecology 55:1142-1147. Maron, John L. & Susan Harrison. 1997. Spatial pattern formation in an insect host-parasitoid

system. Science 278:1619-1621. May, Robert M. 1972. Limit cycles in predator-prey communities. Science 177:900-902. May, Robert M. 1983. Parasitic infections as regulators of animal populations. American

Scientist 71:36-45.

18

McCallum, Hamish, Nigel Barlow & Jim Hone. 2001. How should pathogen transmission be modelled? Trends in Ecology & Evolution 16:295-300. [From Bret Elderd – Feb. 2014]

McCauley, Shannon J., Locke Rowe & Marie-Josee Fortin. 2011. The deadly effects of “nonlethal” predators. Ecology 92:2043-2048.

McPeek, Mark A. 2014. Keystone and intraguild predation, intraspecific density dependence, and a guild of coexisting consumers. The American Naturalist 183:E1-E16.

McPeek, Mark A. 2014. Limiting factors, competitive exclusion, and a more expansive view of species coexistence. The American Naturalist 183:iii-iv. [Re Levin (1970); see also Chase & Leibold (2001)]

Mitchell, Charles E. & Alison G. Power. 2003. Release of invasive plants from fungal and viral pathogens. Nature 421:625-627.

Mitchell, S. N. et al. 2015. Evolution of sexual traits influencing vectorial capacity in anopheline mosquitoes. Science 347:985-988.

Murdoch, William W. 1969. Switching in general predators: Experiments on predator specificity and stability of prey populations. Ecological Monographs 39:335-354.

Murdoch, W. W., S. Avery & M. E. B. Smith. 1975. Switching in a predatory fish. Ecology 56:1094-1105.

Murdoch, W. W., R. M. Nisbet, E. McCauley, A. M. DeRoos & W. S. C. Gurney. 1998. Plankton abundance and dynamics across nutrient levels: Tests of hypotheses. Ecology 79:1339-1356.

Page, Rachel A. & Michael J. Ryan. 2005. Flexibility in assessment of prey cues: frog-eating bats and frog calls. Proc. R. Soc. B 272:841-847.

Parker, Ingrid M. & Gregory S. Gilbert. 2004. The evolutionary ecology of novel plant-pathogen interactions. Annual Review of Ecology, Evolution & Systematics 35:675-700.

Reeve, J. D., J. T. Cronin and D. R. Strong. 1994. Parasitism and generation cycles in a salt-marsh planthopper. Journal of Animal Ecology 63: 912-920.

Rosenzweig, Michael L. 1969. Why the prey curve has a hump. The American Naturalist 103:81-87.

Rosenzweig, Michael L. 1971. Paradox of enrichment: Destabilization of exploitation ecosystems in ecological time. Science 171:385-387.

Rosenzweig, Michael L. & Robert H. MacArthur. 1963. Graphical representation and stability conditions of predator-prey interactions. The American Naturalist 97:209-223.

Sait, Steven M., Wei-Chung Liu, David J. Thompson, H. Charles J. Godfray & Michael Begon. 2000. Invasion sequence affects predator-prey dynamics in a multi-species interaction. Nature 405:448-450.

Sanford, Eric, Melissa S. Roth, Glenn C. Johns, John P. Wares & George N. Somero. 2003. Local selection and latitudinal variation in a marine predator-prey interaction. Science 300:1135-1137.

Sih, Andrew, Philip Crowley, Mark McPeek, James Petranka & Kevin Strohmeier. 1985. Predation, competition, and prey communities: A review of field experiments. Annual Review of Ecology and Systematics 16:269:311.

Sinclair, A. R. E., Simon Mduma & Justin S. Brashares. 2003. Patterns of predation in a diverse predator-prey system. Nature 425:288-290.

Vaughn, Caryn C. & Christopher M. Taylor. 2000. Macroecology of a host-parasite relationship. Ecography 23:11-20.

Vermeij, Geerat J. 1987. Evolution and Escalation: An Ecological History of Life. Princeton

19

University Press. Westover, K. M. & J. D. Bever. 2001. Mechanisms of plant species coexistence:

complementary roles of rhizosphere bacteria and root fungal pathogens. Ecology 82:3285-3294.

Williams, G. C. & R. M. Nesse. 1991. The dawn of Darwinian medicine. Quarterly Review of Biology 66:1–22.

Williams, Kathy, Kimberly G. Smith & Frederick M. Stephen. 1993. Emergence of 13-yr periodical cicadas (Cicadidae: Magicicada): Phenology, mortality, and predator satiation. Ecology 74:1143-1152.

Wilson, Erin E., Lynne M. Mullen & David A. Holway. 2009. Life history plasticity magnifies the ecological effects of a social wasp invasion. PNAS.

Wood, Chelsea L. & Pieter T. J. Johnson. 2015. A world without parasites: exploring the hidden ecology of infection. Frontiers in Ecology and the Environment 13:425-434.

7. Pair-wise species interactions: Herbivory Additional references: Abrahamson, Warren G., ed. 1989. Plant-Animal Interactions. McGraw-Hill Publishing, New

York, NY. Adler, Lynn S. 2000. Alkaloid uptake increases fitness in a hemiparasitic plant via reduced

herbivory and increased pollination. American Naturalist 156:92-99. Adler, Lynn S. 2003. Ecology 2083-2091. Agrawal, Anurag A. 1998. Leaf damage and associated cues induce aggressive ant

recruitment in a Neotropical ant-plant. Ecology 79:2100-2112. Agrawal, Anurag A., Amy P. Hastings, Marc T. J. Johnson, John L. Maron & Juha-Pekka

Salminen. 2012. Insect herbivores drive real-time ecological and evolutionary change in plant populations. Science 338:113-116.

Agrawal, Anurag A., Mark Fishbein, Rayko Halitschke, Amy P. Hastings, Daniel L. Rabosky & Sergio Rasmann. 2009. Evidence for adaptive radiation from a phylogenetic study of plant defenses. PNAS.

Belsky, A. J., W. P. Carson, C. L. Jensen & G. A. Gox. 1993. Overcompensation by plants: Herbivore optimization or red herring? Evolutionary Ecology 7:109-121.

Bennett, A. & J. D. Bever. 2007. Mycorrhizal mediation of plant response to herbivory. Ecology 88:210–218.

Benrey, Betty & Robert F. Denno. 1997. The slow-growth-high-mortality hypothesis: A test using the cabbage butterfly. Ecology 78:987-999.

Bond W. J. & J. A. Silander. 2007. Springs and wire plants: anachronistic defences against Madagascar's extinct elephant birds. Proc. Roy. Soc. B 274:1985-1992. [Notice the ghosts of herbivory past!]

Bruinsma, M., and M. Dicke. 2008. Herbivore-induced indirect defense: from induction mechanisms to community ecology. Pages 31-60 in A. Schaller, editor. Induced plant resistance to herbivory. Springer Science+Business Media B.V. [Add fig. from Amanda Accamando’s presentation!]

Bryant, J.

20

Cárdenas, Rafael E., Renato Valencia, Nathan J. B. Kraft, Adriana Argoti & Olivier Dangles. 2014. Plant traits predict inter- and intraspecific variation in susceptibility to herbivory in a hyperdiverse Neotropical rain forest tree community. Journal of Ecology 102:939-952.

Coley, P. D. 1983. Herbivory and defensive characteristics of tree species in a lowland tropical forest. Ecological Monographs 53:209-233.

Coley, P. D. & J. A. Barone. 1996. Herbivory and plant defenses in tropical forests. Annual Review of Ecology and Systematics. 27:305-335.

Coley, P. D., J. P. Bryant & F. S. Chapin, III. 1985. Resource availability and plant anti-herbivore defense. Science 230:895-899.

Coley, Phyllis D., John Lokvam, Kathleen Rudolph, Keryn Bromberg, Tara E. Sackett, Leslie Wright, Tania Brenes-Arguedas, Dan Dvorett, Seth Ring, Alex Clark, Caroline Baptiste, R. Toby Pennington & Thomas A. Kursar. 2005. Divergent defensive strategies of young leaves in two species of Inga. Ecology 86:2633-2643.

Dietz, Hansjörg, Lea R. Wirth & Holger Buschmann. 2004. Variation in herbivore damage to invasive and native woody plant species in open forest vegetation on Mahé, Seychelles. Biological Invasions 6:511-521.

Dyer, L. A., M. S. Singer, J. T. Lill, J. O. Stireman, G. L. Gentry, R. J. Marquis, R. E. Ricklefs, H. F. Greeney, D. L. Wagner, H. C. Morais, I. R. Diniz, T. A. Kursar & P. D. Coley. 2007. Host specificity of Lepidoptera in tropical and temperate forests. Nature 448:696-699.

Ehrlich, P. & P. Raven. 1964. Butterflies and plants: A study in coevolution. Evolution 18:586-608.

EndaraMJ_etal_2017_PNAS_CoevolArmsRace_vs_HostDefenseChase.pdf Fadzly, Nik, Cameron Jack, H. Martin Schaefer & K. C. Burns. 2009. Ontogenetic colour

changes in an insular tree species: signalling to extinct browsing birds? New Phytologist. [Notice the ghosts of herbivory past!]

Farrell & Mitter. 1998. The timing of insect/plant diversification: might Tetraopes (Coleoptera: Cerambycidae) and Asclepias (Asclepiadaceae) have co-evolved? Biol. J. Linn. Soc. 63:553-577.

Feeny, P. P. 1976. Plant apparency and chemical defense. Pp. 1-40 in J. Wallace & R. L. Mansell, eds., Biochemical Interactions Between Plants and Insects. Recent Advances in Phytochemistry 10. Plenum Press, USA.

Fine, Paul V. A., Italo Mesones & Phyllis D. Coley. 2004. Herbivores promote habitat specialization by trees in Amazonian forests. Science 305:663-665.

Fine et al. Ecology… Futuyma, D. J. 1976. Food plant specialization and environmental predictability in

Lepidoptera. The American Naturalist 110:285-292. Futuyma, Douglas J. & Anurag A. Agrawal. 2009. Macroevolution and the biological

diversity of plants and herbivores. Proceedings of the National Academy of Sciences 106:18054-18061.

Goheen, Jake – from seminar at LSU 11/9/09, thorniest acacias on richest soils at Mpala (counter Coley predictions)

Greenwood & Atkinson (1980) [Moa-browsing hypothesis] Grubb, Peter J. 1992. A positive distrust of simplicity – lessons from plant defences and from

competition among plants and animals. Journal of Ecology 80:585-610. Hay, Mark E. 1997. The ecology and evolution of seaweed-herbivore interactions on coral

reefs. Proceedings of the 8th International Coral Reef Symposium Vol. I:23-32.

21

Herms, D. A. & W. J. Mattson. 1992. The dilemma of plants: To grow or defend. The Quarterly Review of Biology 67:283-335.

Howe, H. F. & J. S. Brown. 2001. The ghost of granivory past. Ecology Letters 4:371-378. Hunter, Mark D. 2016. The Phytochemical Landscape: Linking Trophic Interactions and

Nutrient Dynamics. Princeton University Press, Princeton, NJ. Janzen, D. H. 1980. When is it coevolution? Evolution 34:611-612. Karban, Richard. 1989. Community organization of Erigeron glaucus folivores: effects of

competition, predation and host plant. Ecology 70:1028-1039. Karban, R. & I. T. Baldwin. 1997. Induced Responses to Herbivory. University of Chicago

Press, Chicago, IL. Kitajima, Kaoru. 1994. Relative importance of photosynthetic traits and allocation patterns as

correlates of seedling shade tolerance of 13 tropical trees. Oecologia 98:419-428. LoPresti, Eric F. & Richard Karban. 2016. Chewing sandpaper: grit, plant apparency, and

plant defense in sand-entrapping plants. Ecology 97:826-833. Marquis, Robert J. 1984. Leaf herbivores decrease fitness of a tropical plant. Science

226:537-539. Marquis, Robert J. & Helen Miller Alexander. 1992. Evolution of resistance and virulence in

plant-herbivore and plant-pathogen interactions. Trends in Ecology and Evolution 7:126-129.

McNaughton, S. J. 1983. Serengeti grassland ecology: The role of composite environmental factors and contingency in community organization. Ecological Monographs 53:291-320.

McNaughton, S. J. 1985. Ecology of a grazing ecosystem: The Serengeti. Ecological Monographs 55:259-294.

Moles, Angela T. et al. 2010. Functional Ecology. [Lat. gradients in defense & herbivory] Novotny, Vojtech, Scott. E. Miller, Jiri Hulcr, Richard A. I. Drew, Yves Basset, Milan Janda,

Gregory P. Setliff, Karolyn Darrow, Alan J. A. Stewart, John Auga, Brus Isua, Kenneth Molem, Markus Manumbor, Elvis Tamtiai, Martin Mogia & George D. Weiblen. 2007. Low beta diversity of herbivorous insects in tropical forests. Nature 448:692-695.

O’Dowd, D. J. & M. F. Wilsson. 1989. Leaf domatia and mites on Australian plants: Ecological and evolutionary implications. Biological Journal of the Linnean Society 37:191-236.

Parker, John D., Deron E. Burkepile & Mark E. Hay. 2006. Opposing effects of native and exotic herbivores on plant invasions. Science 311:1459-1461.

Parker, John D. & Mark E. Hay. 2005. Biotic resistance to plant invasions? Native herbivores prefer non-native plants. Ecology Letters 8:959-967.

Pennings et al. 2009. Ecol. – lat. gradient in herbivory (see Ganesh’s proposal) ReisenmanCE_etal_2010_ProcRoySoc_AntagonisticEffectsOfFloralScent.pdf Rhoades, D. F. & R. G. Cates. 1976. Toward a general theory of plant antiherbivore

chemistry. Pp. 168-213 in J. Wallace & R. L. Mansell, eds., Biochemical Interactions Between Plants and Insects. Recent Advances in Phytochemistry 10. Plenum Press, USA.

Sedio, Brian Stamp, Nancy. 2003. Out of the quagmire of plant defense hypotheses. The Quarterly

Review of Biology 78:23-55. Steinberg, Peter D., James A. Estes & Frank C. Winter. 1995. Evolutionary consequences of

food chain length in kelp forest communities. Proceedings of the National Academy of Sciences, USA 92:8145-8148.

22

Strong, Donald R. 1988. Insect host range. Ecology 69:885. Tiffin, Peter. 2000. Are tolerance, avoidance, and antibiosis evolutionarily and ecologically

equivalent responses of plants to herbivores? American Naturalist 155:128-138. Turlings T. C. J., Tumlinson, J. H., and W. J. Lewis. 1990. Exploitation of herbivore-induced

plant odors by host-seeking parasitic wasps. Science 250:1251-1253. [Add fig. from Amanda Accamando’s presentation!]

UnderwoodN_etal_2014_QRevBio_AssociationalEffects.pdf Van Bael, Sunshine A., Jeffrey D. Brawn & Scott K. Robinson. 2003. Birds defend trees from

herbivores in a Neotropical forest canopy. Proceedings of the National Academy of Sciences, USA 100:8304-8307.

Walter, D. E. & D. J. O’Dowd. 1992. Leaves with domatia have more mites. Ecology 73:1514-1518.

Wetzel, William C., Heather M. Kharouba, Moria Robinson, Marcel Holyoak & Richard Karban. 2016. Variability in plant nutrients reduces insect herbivore performance. Nature 539:425-427. [“plants contribute to the suppression of herbivore populations through variable [intraspecific] nutrient levels, not just by having low average quality”; “nutrient heterogeneity within agricultural crops could contribute to the sustainable control of insect pest in agroecosystems”; “Jensen’s inequality” is the key to relating intraspecific trait variance to herbivore performance]

Whitehead, Susan. Windle, Phyllis N. & Eldon H. Franz. 1979. The effects of insect parasitism on plant

competition: greenbugs and barley. Ecology 60:521-529. Wold, E. N. & R. J. Marquis. 1997. Induced responses to early-season herbivory in white oak:

Effects on late-season herbivores and benefits to the plant. Ecology 78:1356-1369. Young, T. P. 1987. Increased thorn length in Acacia depranolobium - an induced response to

browsing. Oecologia 71:436-438.

8. Pair-wise species interactions: Mutualisms (pollination, seed dispersal, trophic & defense symbioses, etc.) and other types of Facilitation

Additional references: Adler, Lynn S. 2000. Alkaloid uptake increases fitness in a hemiparasitic plant via reduced

herbivory and increased pollination. American Naturalist 156:92-99. [Same as for herbivory this year!]

Arnold, A. E., Z. Maynard, G. Gilbert, P. D. Coley & T.A. Kursar. 2000. Are tropical fungal endophytes hyperdiverse? Ecology Letters 3:267-274.

Axelrod, R. & W. D. Hamilton. 1981. The evolution of cooperation. Science 211:1390–1396. Barlow, Connie. 2000. The Ghosts of Evolution. Basic Books, New York, NY. Bergstrom, Carl T. & Michael Lachmann. 2003. The Red King effect: When the slowest

runner wins the coevolutionary race. Proceedings of the National Academy of Sciences, USA 100:593-598.

Bertness, Mark D. & Ragan Callaway. 1994. Positive interactions in communities. Trends in Ecology and Evolution 9:191-193. [Note – their graphical model is similar to Menge & Sutherland (1987; Am. Nat.)]

23

Bever, J. D. 2002. Negative feedback within a mutualism: Host-specific growth of mycorrhizal fungi reduces plant benefit. Proceedings of the Royal Society of London. 269: 2595-2601.

Bever, J. D. 2003. Soil community dynamics and the coexistence of competitors: Conceptual frameworks and empirical tests. New Phytologist 157:465-473.

Bever, J. D., S. Richardson, B. M. Lawrence, J. Holmes & M. Watson. 2009. Preferential allocation to beneficial symbiont with spatial structure maintains mycorrhizal mutualism. Ecology Letters 12:13–21.

Boucher, D. H., ed. 1985. The Biology of Mutualism. Oxford University Press, Oxford, UK. Boucher, D. H., S. James & K. H. Keeler. 1982. The ecology of mutualism. Annual Review

of Ecology and Systematics 13:315-347. Bronstein, J. L. 1994. Our current understanding of mutualism. The Quarterly Review of

Biology 69:31-51. Bronstein J. L. 2001. The exploitation of mutualisms. Ecology Letters 4:277-287. Bronstein J. L. 2001. The costs of mutualism. American Zoologist 41:825-839. Bronstein J. L., R. Alarcon & M. Geber. 2006. The evolution of plant-insect mutualisms.

New Phytologist 172:412-428. Burns, K. C. 2003. Broad-scale reciprocity in an avian seed dispersal mutualism. Global

Ecology & Biogeography 12:421-426. Callaway, Ragan M. 2007. Positive Interactions and Interdependence in Plant Communities.

Springer, Dordrecht, The Netherlands. Callaway, Ragan M., R. W. Brooker, Phillippe Choler, Zaal Kikvidze, Christopher J. Lortie,

Richard Michalet, Leonardo Paolini, Francisco I. Pugnaire, Beth Newingham, Erik T. Aschehoug, Cristina Armas, David Kikodze & Bradley J. Cook. 2002. Positive interactions among alpine plants increase with stress. Nature 417:844-848.

Clay, K. & J. Holah. 1999. Fungal endophyte symbiosis and plant diversity in successional fields. Science 285:1742-1744.

Connor, R. C. 1995. The benefits of mutualism: A conceptual framework. Biological Reviews of the Cambridge Philosophical Society 70:427-457.

Currie, Cameron R., Bess Wong, Alison E. Stuart, Ted R. Schultz, Stephen A. Rehner, Ulrich G. Mueller, Gi-Ho Sung, Joseph W. Spatafora & Neil A. Straus. 2003. Ancient tripartite coevolution in the attine ant-microbe symbiosis. Science 299:386-388.

Denison, R. Ford, E. Toby Kiers & Stuart A. West. 2003. Darwinian agriculture: when can humans find solutions beyond the reach of natural selection? Quart. Review Biol. 78:145-168.

Freeman, S. & Rodriguez, R. J. 1993. Genetic conversion of a fungal plant pathogen to an nonpathogenic, endophytic mutualist. Science 260:75-78.

Futuyma, Douglas J. & Montgomery Slatkin, eds. 1983. Coevolution. Sinauer Assoc. Inc., Sunderland, MA.

GehringCA_etal_2017_TreeGeneticsMicrobiomeDroughtTolerance.pdf Grime, P. J., J. M. L. Mackey, S. H. Hillier & D. J. Read. 1987. Floristic diversity in a model

system using experimental microcosms. Nature 328:420-422. Harder, Lawrence D. & Steven D. Johnson. 2009. Darwin’s beautiful contrivances:

evolutionary and functional evidence for floral adaptation. NewPhytologist 183:530-545. Herre, Edward Allen. 1996. An overview of studies on a community of Panamanian figs.

Journal of Biogeography 23:593-607.

24

Herre, E. A., N. Knowlton, U. G. Mueller & S. A. Rehner. 1999. The evolution of mutualisms: Exploring the paths between conflict and cooperation. Trends in Ecology and Evolution 14:49-53.

Hoeksema, Jason D. & Emilio M. Bruna. 2000. Pursuing the big questions about interspecific mutualism: A review of theoretical approaches. Oecologia 125:321-330.

Howe, Henry F. 1985. Gomphothere fruits: a critique. American Naturalist 125:853-865. Howe, Henry F. & Maria N. Miriti. 2004. When seed dispersal matters. BioScience 54:651-

660. Jaenike, J. & T. D. Brekke. 2011. Defensive endosymbionts: A cryptic trophic level in

community ecology. Ecology Letters 14:150-155. Jandér, K. Charlotte & Edward Allen Herre. 2010. Host sanctions and pollinator cheating in

the fig tree-fig wasp mutualism. Proc. Roy. Soc. B Janzen, Daniel H. 1966. Interaction of the bull’s horn acacia (Acacia cornigera L.) with an

ant inhabitant (Pseudomyrmex ferrugineq F. Smith) in eastern Mexico. University of Kansas Science Bulletin 47:315-558.

Janzen, Daniel H. 1985. Spondias mombin is culturally deprived in megafauna-free forest. Journal of Tropical Ecology 1:131-155. [Notice the ghosts of seed-dispersal mutualism past!]

Janzen. 1985. xxx. In D. Boucher, ed. The Biology of Mutualism. Oxford University Press, Oxford, UK.

Janzen, Daniel H. & Paul S. Martin. 1982. Neotropical anachronisms: the fruits the gomphotheres ate. Science 215:19-27.

Johnson, N. C., P. J. Copeland, R. K. Crookston & F. L. Pfleger. 1992. Mycorrhizae: Possible explanation for yield decline with continuous corn and soybean. Agronomy Journal 84:387-390.

Johnson, N. C., J. H. Graham & F. A. Smith 1997. Functioning of mycorrhizas along the mutualism-parasitism continuum. New Phytologist 135:1-12.

Johnson, N. C., G. D. Tilman & D. A. Wedin. 1992. Plant and soil controls on mycorrhizal fungal communities. Ecology 73:2034-2042.

Jordano, Pedro. 1995. Frugivore-mediated selection on fruit and seed size: birds and St. Lucie's cherry, Prunus mahaleb. Ecology 76:2627-2639.

Kersch, Monica Frank & Carlos Roberto Fonseca. 2005. Abiotic factors and the conditional outcome of an ant-plant mutualism. Ecology 86:2117-2126.

Kiers, E. Toby, Robert A. Rousseau, Stuart A. West & R. Ford Denison. 2003. Host sanctions and the legume-rhizobium mutualism. Nature 425:78-81.

Kiers, E. Toby & Stuart A. West. 2015. Evolving new organisms via symbiosis. Science 348:392-394.

Knight, T. M., J. A. Steets, J. C. Vamosi, S. J. Mazer, M. Burd, D. R. Campbell, M. R. Dudash, M. O. Johnston, R. J. Mitchell & T.-L. Ashman. 2005. Pollen limitation of plant reproduction: pattern and process. Annual Reviews of Ecology, Evolution and Systematics 36:467–497.

Lanner, Ronald M. 1996. Made for Each Other: A Symbiosis of Birds and Pines. Oxford University Press, Oxford, UK.

Leigh – Genes, bees & ecosystems Leigh, Egbert, Jr. 1994. Do insect pests promote mutualism among tropical trees? Journal of

Ecology 82:677-680.

25

Leigh, Egbert Giles, Jr. 1999. Tropical Forest Ecology. Oxford University Press, Oxford, UK.

Leigh, Egbert Giles, Jr. & Thelma Elizabeth Rowell. 1995. The evolution of mutualism and other forms of harmony at various levels of biological organization. Ecologie 26:131-158.

Mainero, Jorge S. & Carlos Martinez del Rio. 1985. Cheating and taking advantage in mutualistic associations. In D. Boucher, ed., The Biology of Mutualism. Oxford University Press, Oxford, UK.

Memmott, Jane and Nickolas M. Waser. 2002. Integration of alien plants into a native flower-pollinator visitation web. Proceedings of the Royal Society of Londond, Series B 269:2395-2399.

Ollerton, J. 2006. "Biological Barter": Patterns of Specialization Compared across Different Mutualisms. pp. 411-435 in: Waser, N.M. & Ollerton, J. (Eds) Plant-Pollinator Interactions: From Specialization to Generalization. University of Chicago Press.

Paszkowski, U. 2006. Mutualism and parasitism: the yin and yang of plant symbioses. Current Opinion on Plant Biology 9:364-370.

Roy, B.A. and A. Widmer. 1999. Floral mimicry: a fascinating yet poorly understood phenomenon. Trends in Plant Science 4:325-330. [See also Bitty Roy’s work on Dracula orchid floral mimicry of fungi.]

Seifert, E. K., J. D. Bever & J. Maron. 2009. Evidence for evolution of reduced mycorrhizal dependence during plant invasion. Ecology 90:1055–1062.

Silliman, Brian R. & Steven Y. Newell. 2003. Fungal farming in a snail. PNAS 100:15643-15648.

Stachowicz, John J. 2001. Mutualism, facilitation, and the structure of ecological communities. BioScience 51:235-246.

Stachowicz, John J. & Robert B. Whitlatch. 2005. Multiple mutualists provide complementary benefits to their seaweed host. Ecology 86:2418-2427.

Thompson, John N. 1982. Interaction and Coevolution. John Wiley and Sons, NY, NY. Thompson, John N. 1994. The Coevolutionary Process. University of Chicago Press,

Chicago, IL. Thompson, John N. Geographic Mosaic… apparently a book… Wagner, Diane. 2000. Pollen viability reduction as a potential cost of ant association for

Acacia constricta (Fabaceae). American Journal of Botany 87:711-715. Wang, B. & Y. L. Qiu. 2006. Phylogenetic distribution and evolution of mycorrhizas in land

plants. Mycorrhiza 16:299–363. Wheelwright, Nathaniel T. & Gordon H. Orians. 1982. Seed dispersal by animals: Contrasts

with pollen dispersal, problems of terminology, and constraints on coevolution. The American Naturalist 119:402-413.

Willmer, P. B. & G. N. Stone. 1997. How aggressive ant-guards assist seed-set in Acacia flowers. Nature 388:165-167.

Wright, David Hamilton. 1989. A simple, stable model of mutualism incorporating handling time. The American Naturalist 134:664-667.

9. Multiple-species interactions: community modules; food webs; foundation species, interaction networks, keystone species & ecosystem engineers; higher-order interactions; indirect effects (e.g., apparent competition); top-down & bottom-up effects; tri-trophic

26

interactions; trophic cascades; assembly rules (including priority effects, and contrasts with ecological filters or sieves); extended phenotypes; etc.

Additional references: Allesina, Stefano, David Alonso & Mercedes Pascual. 2008. A general model for food web

structure. Science 320:658-661. Angelini, Christine & Brian R. Silliman. 2014. Secondary foundation species as drivers of

trophic and functional diversity: evidence from a tree-epiphyte system. Ecology 95:185-196.

Aschehoug, Erik T. & Ragan M. Callaway. 2015. Diversity increases indirect interactions, attenuates the intensity of competition, and promotes coexistence. The American Naturalist 186:452-459.

Bakun, Andrew. 2006. Wasp-waist populations and marine ecosystem dynamics: Navigating the “predator pit” topographies. Progress in Oceanography 69:271-288. [“typical community configuration, featuring many species at the bottom, many at the top, but contstricted to a very few dominant species at a mid-level…”; “the major control in these ecosystems may be neither ‘btoom up’ nor ‘top down’ but rather ‘both up and down from the middle’” pg. 272.]

Bascompte, Jordi. 2009. Mutualistic networks. Front. Ecol. Environ. 7:429-436. Beckerman, Andrew P., Maria Uriarte & Oswald J. Schmitz. 1997. Experimental evidence for

a behavior-mediated trophic cascade in a terrestrial food chain. Proceedings of the National Academy of Sciences, USA. 94:10735-10738.

Bennett, A., J. Alers-Garcia & J. D. Bever. 2006. Mechanisms of tritrophic interactions between plants, herbivores and AM fungi. American Naturalist 167:141–152.

Billick, Ian & Ted. J. Case. 1994. Higher order interactions in ecological communities: What are they and how can they be detected? Ecology 75:1529-1543.

Bond, W. J. 1994. Keystone Species. Pp. 237-253 in Ernst-Detlef Shulze & Harold A. Mooney (eds.). Biodiversity & Ecosystem Function. Springer, NY, NY.

Bond, W. J. 2005. Large parts of the world are brown or black: a different view on the ‘Green World’ hypothesis. J. Veg. Sci. 16:261-266.

Borer, E. T., B. S. Halpern & E. W. Seabloom. 2006. Asymmetry in community regulation: effects of predators and productivity. Ecology 87:2813-2820.

Briand, F. 1983. Environmental control of food web structure. Ecology 64:253-263. Bender, Edward A., Ted J. Case & Michael E. Gilpin. 1984. Perturbation experiments in

community ecology: theory and practice. Ecology 65:1-13. Callaway, Ragan M. & Lawrence R. Walker. 1997. Competition and facilitation: A synthetic

approach to interactions in plant communities. Ecology 78:1958-1965. Chase, J. M. 2007. Drought mediates the importance of stochastic community assembly.

Proc. Natl. Acad. Sci. USA 104:17430-17434. Chase, Jonathan M., Peter A. Abrams, James P. Grover, Sebastian Diehl, Peter Chesson,

Robert D. Holt, Shane A. Richards, Roger M. Nisbet & Ted J. Case. 2002. The interaction between predation and competition: A review and synthesis. Ecology Letters 5:302-315.

Cohen, Joel E. 1978. Food Webs and Niche Space. Princeton University Press, Princeton, NJ. Cohen, Joel. 1989. Ecology: Just proportions in food webs. Nature 341:104-105.

27

Cohen, Joel E., Tomas Jonsson and Stephen R. Carpenter. 2003. Ecological community description using the food web, species abundance, and body size. Proceedings of the National Academy of Science, USA 100:1781-1786.

Condon, Marty A., Sonja J. Scheffer, Matthew L. Lewis, Robert Wharton, Dean C. Adams & Andrew A. Forbes. 2014. Lethal interactions between parasites and prey increase niche diversity in a tropical community. Science 343:1240-1244. [See also: H. Charles J. Godfray. 2014. Society, where none intrudes. Science 343:1213-1214.]

Connell, Joseph H. 1990. Apparent versus “Real” Competition in Plants. Pp. 9-26 in James B. Grace & David Tilman, eds. Perspectives in Plant Competition, Academic Press, Inc., New York, NY.

Connor, E. F. & D. Simberloff. 1979. The assembly of species communities: Chance or competition? Ecology 60:1132-1140.

Croll, D. A., J. L. Maron, J. A. Estes, E. M. Danner & G. V. Byrd. 2005. Introduced predators transform subarctic islands from grasslands to tundra. Science 307:1959-1961.

Crooks, Kevin R. & Michael E. Soulé. 1999. Mesopredator release and avifaunal extinctions in a fragmented system. Nature 400:563-566.

Dayton P. K. 1972. Toward an understanding of community resilience and the potential effects of enrichments to the benthos at McMurdo Sound, Antarctica. In: Parker BC (Ed). Proceedings of the colloquium on conservation problems in Antarctica. Lawrence, KS: Allen Press.

Diamond, J. M. 1975. Assembly of species communities. Pp. 342-444 in M. L. Cody & J. M. Diamond, eds., Ecology and Evolution of Communities. Harvard University Press, Cambridge, MA.

Dodson, S. I. 1970. Complementary feeding niches sustained by size-selective predation. Limnology and Oceanography 15:131-137.

Drake, James A. 1990. Communities as assembled structures: Do rules govern pattern? TREE 5:159-164.

Drake, James A. 1991. Community-assembly mechanisms and the structure of an experimental species ensemble. The American Naturalist 137:1-26.

Dyer, Lee A. & Deborah Letourneau. 2003. Top-down and bottom-up diversity cascades in detrital vs. living food webs. Ecology Letters 6:60-68.

Dyer, L. A., T. R. Walla, H. F. Greeney, J. O. Stireman & R. F. Hazen. 2010. Diversity of interactions: a metric for studies of biodiversity. Biotropica 42:281-289.

Ellison, Aaron M., Michael S. Bank, Barton D. Clinton, Elizabeth A. Colburn, Katherine Elliott, Chelcy R. Ford, David R. Foster, Brian D. Kloeppel, Jennifer D. Knoepp, Gary M. Lovett, Jacqueline Mohan, David A. Orwig, Nicholas L. Rodenhouse, William V. Sobczak, Kristina A. Stinson, Jeffrey K. Stone, Christopher M. Swan, Jill Thompson, Betsy Von Holle & Jackson R. Webster. 2005. Loss of foundation species: consequences for the structure and dynamics of forested ecosystems. Frontiers in Ecology and the Environment 3:479-486.

Elton, Charles. 1927. Animal Ecology. Sidgwick & Jackson Pubs., London, UK. Estes, J. A. & J. F. Palmisano. 1974. Sea otters: Their role in structuring nearshore

communities. Science 185:1058-1060. Estes, J. A., M. T. Tinker, T. M. Williams & D. F. Doak. 1998. Killer whale predation on sea

otters linking oceanic and nearshore ecosystems. Science 282:473-476. Estes, J. A. et al. 2011. Trophic downgrading of planet Earth. Science.

28

Fretwell, Stephen D. 1977. The regulation of plant communities by food chains exploiting them. Perspectives in Biology and Medicine 20:169-185.

Fretwell, S. D. 1987. Food chain dynamics: The central theory of ecology? Oikos 50:291-301.

Giguere, L. 1979. An experimental test of Dodson’s hypothesis that Ambystoma (a salamander) and Chaoborus (a phantom midge) have complementary feeding niches. Canadian Journal of Zoology 57:1091-1097.

Goheen, Jake – from seminar at LSU 11/9/09, seeming indirect mutualism between mice and megaherbivores, since when megaherbivores excluded, mice suffer (???).

Gratton, Claudio & Robert F. Denno. 2003. Seasonal shift from bottom-up to top-down impact in phytophagous insect populations. Oecologia 134:487-495. [“It is now generally accepted that bottom-up and top-down forces act in concert to influence populations of most phytophagous insects” pg. 134]

Gross, Katherine. 2008. Positive interactions among competitors can produce species-rich communities. Ecology Letters 11:929-936.

Hairston, Nelson G., Frederick E. Smith & Lawrence B. Slobodkin. 1960. Community structure, population control, and competition. The American Naturalist 94:421-425.

Halpern, Benjamin S., Karl Cottenie & Bernardo R. Broitman. 2006. Strong top-down control in southern California kelp forest ecosystems. Science 312:1230-1232.

HartnettDC_GWTWilson_1999_Ecology_MycorrhizaeHelpDominantGrassesSuppressSppDivTallgrassPrairie.pdf

Hixon, Mark A. & Geoffrey P. Jones. 2005. Competition, predation, and density-dependent mortality in demersal marine fishes. Ecology 86:2847-2859.

Holt, R. D. 1977. Predation, apparent competition, and the structure of prey communities. Theoretical Population Biology 12:197-229.

Holt, R. D. 1997. Community modules. Pp. 333-349 in A. C. Gange & V. K. Brown, eds. Multitrophic Interactions in Terrestrial Ecosystems, Blackwell Science, London, UK.

Holt, R. D. & J. H. Lawton. 1994. The ecological consequences of shared natural enemies. Annual Review of Ecology and Systematics 25:495-520.

Huisman, J., Weissing, F.J. 2002. Oscillations and chaos generated by competition for interactively essential resources. Ecol. Res. 17, 175-181.

Huisman, J. & Weissing, F.J. 2001. Biological conditions for oscillations and chaos generated by multispecies competition. Ecol. 82, 2682-2695.

Huisman, J. & Weissing, F. J. 2001. Fundamental unpredictability in multispecies competition. Amer. Nat. 157, 488-494.

Hunter, Mark D. & Peter W. Price. 1992. Playing chutes and ladders: Heterogeneity and the relative roles of bottom-up and top-down forces in natural communities. Ecology 73:724-732.

Jeffries, M. J. & J. H. Lawton. 1984. Enemy free space and the structure of ecological communities. Biological Journal of the Linnaean Society 23:269-286.

Jenkins, B., R. L. Kitching & S. L. Pimm. 1992. Productivity, disturbance, and food web structure at a local spatial scale in experimental container habitats. Oikos 65:249-255.

Jones, Clive G., J. H. Lawton & M. Shachak. 1994. Organisms as ecosystem engineers. Oikos 69:373-386.

Jones, Clive G., Richard S. Ostfeld, Michele P. Richard, Eric M. Schauber & Jerry O. Wolff. 1998. Chain reactions linking acorns to gypsy moth outbreaks and lyme disease risk.

29

Science 279:1023-1026. Kareiva, Peter M. & Mark D. Bertness. 1997. Re-examining the role of positive interactions

in communities. Ecology 78:1945. [See the articles introduced by Kareiva & Bertness in this volume’s Special Feature section, many of which compare and contrast positive and negative interactions]

Karst, J., B. Gilbert & M. J. Lechowicz. 2005. Fern community assembly: the roles of chance and the environment at local and intermediate scales. Ecology 86:2473-2486.

Kilronomos, J. N. 2002. Feedback with soil biota contributes to plant rarity and invasivesness in communities. Nature 417:67-70.

Knight, Tiffany M., Michael W. McCoy, Jonathan M. Chase, Krista A. McCoy & Robert D. Holt. 2005. Trophic cascades across ecosystems. Nature 437:880-883.

Křivan, Vlastimil & Oswald J. Schmitz. 2004. Trait and density mediated indirect interactions in simple food webs. Oikos 107:239-250.

Lafferty, Kevin D. & Thomas H. Suchanek. 2016. Revisiting Paine’s 1966 sea star removal experiment, the most-cited empirical article in the American Naturalist. The American Naturalist 188:365-378.

Laska, Mark S. & J. Timothy Wootton. 1998. Theoretical concepts and empirical approaches to measuring interaction strength. Ecology 79:461-476.

Leibold. 1989. [Herbivory & defense idea presented by Jake Goheen at LSU on 11/9/09.] Leibold, M. A. & H. M. Wilbur. 1992. Interactions between food web structure and nutrients

on pond organisms. Nature 360:341-343. Letnic, Mike, Freya Koch, Chris Gordon, Mathew S. Crowther & Christopher R. Dickman.

2009. Keystone effects of an alien top-predator stem extinctions of native mammals. Proc. Roy. Soc. B 276:3249-3256.

Levi, Taal & Christopher C. Wilmers. 2012. Wolves-coyotes-foxes: a cascade among carnivores. Ecology 93:921-929.

Matson, P. A. & M. D. Hunter. 1992. The relative contributions of top-down and bottom-up forces in population and community ecology. Ecology 73:723 [Introduction to a special feature on top-down and bottom-up forces, with articles by Hunter & Price, Menge, Power, and Strong]

May, Robert M. 1973. Stability and Complexity in Model Ecosystems. Princeton University Press, Princeton, NJ.

McCann, Kevin, Alan Hastings & Gary R. Huxel. 1998. Weak trophic interactions and the balance of nature. Nature 395:794-798.

McLaren, B. E. & R. O. Peterson. 1994. Wolves, moose, and tree rings on Isle Royale. Science 266:1555-1558.

Menge, Bruce A. 1992. Community regulation: Under what conditions are bottom-up factors important on rocky shores? Ecology 73:755-765.

Menge, Bruce A. 1995. Indirect effects in marine rocky intertidal interaction webs: Patterns and importance. Ecological Monographs 65:21-74.

Menge, Bruce A., J. Lubchenco, M. E. S. Bracken, F. Chan, M. M. Foley, T. L. Freidenburg, S. D. Gaines, G. Hudson, C. Krenz, H. Leslie, D. N. L. Menge, R. Russell & M. S. Webster. 2003. Coastal oceanography sets the pace of rocky intertidal community dynamics. PNAS 100:12229-12234.

Michel, Nicole L., Thomas W. Sherry & Walter P. Carson. 2014. The omnivorous collared peccary negates an insectivore-generated trophic cascade in Costa Rican wet tropical forest

30

understory. Journal of Tropical Ecology 30:1-11. Miller, Thomas E., Emma R. Moran & Casey P. terHorst. 2014. Rethinking niche evolution:

experiments with natural communities of protozoa in pitcher plants. The American Naturalist 184:xx-xx.

Murdoch, W. W. 1966. “Community structure, population control, and competition” – a critique. The American Naturalist 100:219-226.

Murdoch, W. W., B. E. Kendall, R. M. Nisbet, C. J. Briggs, E. McCauley & R. Bolser. 2002. Single-species models for many-species food webs. Nature 417:541-543.

Oksanen, L., S. D. Fretwell, J. Arruda & P. Niemela. 1981. Exploitation ecosystems in gradients of primary productivity. The American Naturalist 118:240-261.

Orrock, J. L., R. D. Holt & M. L. Baskett. 2010. Refuge-mediated apparent competition in plant–consumer interactions. Ecology letters 13:11-20.

Paine, Robert T. 1966. Food web complexity and species diversity. The American Naturalist 100:65-75.

Palumbi, Stephen R. 2003. Ecological subsidies alter the structure of marine communities. PNAS 100:11927-11928. [Commentary on Menge et al. (2003)]

Parker, Ingrid M., Megan Saunders, Megan Bontrager, Andrew P. Weitz, Rebecca Hendricks, Roger Magarey, Karl Suiter & Gregory S. Gilbert. 2015. Phylogenetic structure and host abundance drive disease pressure in communities. Nature 520:542-544.

Peacor, Scott D. & Earl E. Werner. 2001. The contribution of trait-mediated indirect effects to the net effects of a predator. Proceedings of the National Academy of Science, USA 98:3904-3908.

Pearson, Dean E. 2010. Trait- and density-mediated indirect interactions initiated by an exotic invasive plant autogenic ecosystem engineer. The American Naturalist 176:394-403.

Persson, Lennart. 1991. Behavioral response to predators reverses the outcome of competition between prey species. Behavioral Ecology and Sociobiology 28:101-105.

Persson, A., L. A. Hansson, C. Bronmark, P. Lundberg, L. B. Pettersson, L. Greenberg, P. A. Nilsson, P. Nystrom, P. Romare & L. Tranvik. 2001. Effects of enrichment on simple aquatic food webs. The American Naturalist 157:654-669.

Peterson, C. H. 1984. Does a rigorous criterion for environmental identity preclude the existence of multiple stable points? Am. Nat. 124:127-133. [Four criteria for multiple stable states; see Petraitis et al. (2009)]

Petraitis, Peter S., Elizabeth T. Methratta, Erika C. Rhile, Nicholas A. Vidargas & Steve R. Dudgeon. 2009. Experimental confirmation of multiple community states in a marine ecosystem. Oecologia 161:139-148.

Pimm, Stuart L. 1982. Food Webs. Chapman and Hall, London, UK. Pimm, Stuart L. 1991. The Balance of Nature? Chicago University Press, Chicago, IL. Pimm, Stuart L. 2001. The World According to Pimm: A Scientist Audits the Earth.

McGraw-Hill, New York, NY. Pimm, S. L. & J. H. Lawton. 1977. Number of trophic levels in ecological communities.

Nature 268:329-331. Pimm, S. L. & J. H. Lawton. 1978. On feeding on more than one trophic level. Nature

275:542-544. Pimm, S., J. Lawton & J. Cohen. 1991. Food web patterns and their consequences. Nature

350:669-674. Polis, Gary A. 1991. Complex interactions in deserts: An empirical critique of food-web

31

theory. The American Naturalist 138:123:155. Polis, G. A. & R. D. Holt. 1992. Intraguild predation: The dynamics of complex trophic

interactions. Trends in Ecology and Evolution 7:151-154. Polis, G. & D. Strong. 1996. Food web complexity and community dynamics. The American

Naturalist 147:813-846. Polis, G. A. & K. O. Winemiller, eds. 1996. Food Webs: Integration of Patterns and

Dynamics. Chapman and Hall, New York, NY. Post, Eric, Rolf O. Peterson, Nils Chr. Stenseth & Brian E. McLaren. 1999. Ecosystem

consequences of wolf behavioural response to climate. Nature 401:905-907. Power, M. E. 1992. Top down and bottom up forces in food webs: Do plants have primacy?

Ecology 73:733-746. Reynolds, Heather L., A. Packer, J. D. Bever & K. Clay. 2003. Grassroots ecology: plant-

microbe-soil interactions as drivers of plant community structure and dynamics. Ecology 84:2281-2291.

Sanderson, James G., Jared M. Diamond & Stuart L. Pimm. 2009. Pairwise co-existence of Bismark and Solomon landbird species. Evolutionary Ecology Research 11:771-786.

Scheffer, Marten, Sándor Szabó, Alessandra Gragnani, Egbert H. van Nes, Sergio Rinaldi, Nils Kautsky, Jon Norberg, Rudi M. M. Roijackers & Rob J. M. Franken. 2003. Floating plant dominance as a stable state. Proceedings of the National Academy of Sciences 100:4040-4045.

Schemske et al. 2009. Ann. Rev. Ecol. Evol. Syst. [Lat. gradients in biotic interactions.] Schmitt, R. J. 1987. Indirect interactions between prey: Apparent competition, predator

aggregation, and habitat segregation. Ecology 68:1887-1897. Schmitt, Russell J., Sally J. Holbrook, Andrew J. Brooks & Jennifer C. P. Lape. 2009.

Intraguild predation in a structured habitat: distringuishing multiple-predator effects from competitor effects. Ecology 90:2434-2443.

Schmitz, O. J. 2008. Herbivory from individuals to ecosystems. Annual Review of Ecology, Evolution & Systematics 39:133-152.

Schoener, T. W. 1989. Ecology… [Good ref. for HSS] Shurin, Jonathan B., Daniel S. Gruner & Helmut Hillebrand. 2006. All wet or dried up? Real

differences between aquatic and terrestrial food webs. Proc. Roy. Soc. B 273:1-9. Siepielski, Adam M. & Craig W. Benkman. 2008. A seed predator drives the evolution of a

seed dispersal mutualism. Proceedings of the Royal Society B 275:1917-1925. Silliman & Zieman. 2001. Ecol. 82… Stokstad, Erik. 2009. On the origin of ecological structure. Science 326:33-35. Strauss, Sharon Y. 1991. Indiret effects in community ecology: their definition, study and

importance. Trends in Ecology & Evolution 6:206-210. Strong, Donald R. 1992. Are trophic cascades all wet? Differentiation and donor-control in

speciose ecosystems. Ecology 73:747-754. Terborgh, John. 1986. Keystone plant resources in the tropical forest. Pp. 330-344 in Michael

E. Soulé, ed., Conservation Biology. Sinauer, Sunderland, MA. Terborgh, John. 1988. The big things that run the world – a sequel to E. O. Wilson.

Conservation Biology 2:402-403. Terborgh, John. 2010. The trophic cascade on islands. Pp. 116-142 in Jonathan B. Losos &

Robert E. Ricklefs, eds., The Theory of Island Biogeography Revisited. Princeton University Press, Princeton, NJ.

32

Weiher E. & P. Keddy, eds. 1999. Ecological Assembly Rules: Perspectives, Advances, Retreats. Cambridge University Press, Cambridge, UK.

Wellborn, Gary A., David K. Skelly & Earl E. Werner. 1996. Mechanisms creating community structure across a freshwater habitat gradient. Ann. Rev. Ecol. Syst. 27:337-363.

Werner, Earl E. & Scott D. Peacor. 2003. A review of trait-mediated indirect interactions in ecological communities. Ecology 84:1083-1100.

Wiens, John A. 1989. The Ecology of Bird Communities. Volume 1. Foundations and Patterns. Cambridge University Press, Cambridge, UK.

Wiens, John A. 1989. The Ecology of Bird Communities. Volume 2. Processes and Variations. Cambridge University Press, Cambridge, UK.

Williams, Richard J. & Neo D. Martinez. 2000. Simple rules yield complex food webs. Nature 404:180-183.

Wilson, Edward O. 1987. The little things that run the world (the importance and conservation of invertebrates). Conservation Biology 1:344-346.

Wise, D. H. 1993. Spiders in Ecological Webs. Cambridge University Press, Cambridge, UK. Wootton, J. Timothy. 1993. Indirect effects and habitat use in an intertidal community:

Interaction chains and interaction modifications. The American Naturalist 141:71-89. Yodzis, Peter. 1984. Energy flow and the vertical structure of real ecosystems. Oecologia

65:86-88. Zimmer, Richard K. et al. 2016. A multifunctional chemical cue drives opposing

demographic processes and structures ecological communities. Ecology 97:2232-2239.

10. Origins and maintenance of species diversity and diversity gradients. I. Theory and selected empirical examples

Additional references: Abrams, Peter A. 1988. Resource productivity-consumer species diversity: simple models of

competition in spatially heterogeneous environments. Ecology 69:1418-1433. Adams, J. M. & F. I. Woodward. 1989. Patterns in tree species richness as a test of the glacial

extinction hypothesis. Nature 339:699-701. Adler, Peter B., Janneke HilleRisLambers & Jonathan M. Levine. 2007. A niche for

neutrality. Ecology Letters 10:95-104. Anderson, G. R. V., A. H. Ehrlich, P. R. Ehrlich, J. Roughgarden, B. C. Russell & F. H.

Talbot. 1981. The community structure of coral reef fishes. The American Naturalist 117:476-495.

Armbrecht, Inge, Ivette Perfecto & John Vandermeer. 2004. Enigmatic biodiversity correlations: ant diversity responds to diverse resources. Science 304:284-286.

Ashton, Peter S. 1969. Speciation among tropical forest trees: some deductions in the light of recent evidence. Biological Journal of the Linnean Society 1:155-196.

Ashton, Peter S. 1993. Species richness in plant communities. Pp. 4-22 in P. L. Fiedler & S. K. Jain, eds. Conservation Biology. Chapman and Hall, New York, NY.

33

Baer, Sara G., John M. Blair & Scott L. Collins. 2015. Environmental heterogeneity has a weak effect on diversity during community assembly in tallgrass prairie. Ecological Monographs 86:94-106.

Barabas_etal_2018_EcologicalMonographs_ChessonCoexistenceTheory.pdf Belmaker, Jonathan & Walter Jetz. 2015. Relative roles of ecological and energetic

constraints, diversification rates and region history on global species richness gradients. Ecology Letters 18:563-571.

Biswas, Shekhar R. & Helene H. Wagner. 2012. Landscape contrast: a solution to hidden assumptions in the metacommunity concept? Landscape Ecology 27:621-631.

Bolnick, Daniel I., Priyanga Amarasekare, Márcio S. Araújo, Reinhard Bürger, Jonathan M. Levine, Mark Novak, Volker H. W. Rudolf, Sebastian J. Schreiber, Mark C. Urban & David A. Vasseur. 2011. Why intraspecific trait variation matters in community ecology. Trends in Ecology & Evolution 26:183-192.

Brown, J. H. 1988. Species diversity. Pp. 57-89 in A. A. Myers & P. S. Giller, eds. Analytical Biogeography. Chapman and Hall, London, UK.

Bush, M. B & P. E. Oliveira. 2006. The rise and fall of the Refugial Hypothesis of Amazonian Speciation: a paleoecological perspective. Biota Neotrop. vol. 6.

Chase, Jonathan M. 2005. Towards a really unified theory for metacommunities. Functional Ecology 19:182-186.

Chase, Jonathan M. 2014. Spatial scale resolves the niche versus neutral theory debate. Journal of Vegetation Science 25:319-322. [See also McPeek re niche-neutral]

Chesson, Peter. 2000. Mechanisms of maintenance of species diversity. Annual Review of Ecology and Systematics 31:343-366.

Chesson, P. L. & R. R. Warner. 1981. Environmental variability promotes coexistence in lottery competitive systems. The American Naturalist 117:923-943.

Chisholm, Ryan A., Richard Condit, K. Adb. Rahman, Patrick J. Baker, Sarayudh Bunyavejchewin, Yu-Yun Chen, George Chuyong, H. S. Dattaraja, Stuart Davies, Corneille E. N. Ewango, C. V. S. Gunatilleke, I. A. U. Nimal Gunatilleke, Stephen Hubbell, David Kenfack, Somboon Kiratiprayoon, Yiching Lin, Jean-Remy Makana, Nantachai Pongpattananurak, Sandeep Pulla, Ruwan Punchi-Manage, Raman Sukumar, Sheng-Hsin Su, I-Fang Sun, H. S. Suresh, Sylvester Tan, Duncan Thomas & Sandra Yap. 2014. Temporal variability of forest communities: empirical estimates of population change in 4000 tree species. Ecology Letters x:xx-xx.

Chown, Steven L. & Kevin J. Gaston. 2000. Areas, cradles and museums: The latitudinal gradient in species richness. Trends in Ecology and Evolution 15:311-315.

Chu, Chengjin & Peter B. Adler. 2015. Large niche differences emerge at the recruitment stage to stabilize grassland coexistence. Ecological Monographs 85:373-392.

Clark, James S. et al. 2010. High-dimensional coexistence based on individual variation: a synthesis of evidence. Ecological Monographs 80:569-608.

Clark, James S. & Jason S. McLachlan. 2003. Stability of forest biodiversity. Nature 423:635-638. [Contrasts “stabilizing mechanisms” with “equalizing mechanisms”]

Clark, J. S., M. Dietze, S. Chakraborty, P. K. Agarwal, I. Ibanez, S. LaDeau & M. Wolosin. 2007. Resolving the biodiversity paradox. Ecology Letters 10:647-659.

Clark, J. S. 2009. TREE Cody, M. L. & J. M. Diamond, eds. 1975. Ecology and Evolution of Communities. Belknap

Press of Harvard University Press, Cambridge, MA.

34

Colwell, Robert K. & David C. Lees. 2000. The mid-domain effect: geometric constraints on the geography of species richness. Trends in Ecology & Evolution 15:70-76.

Condit, Chisholm, Hubbell. 2012. Plos One. Connell, J. H. 1971. On the role of natural enemies in preventing competitive exclusion in

some marine animals and in rain forest trees. Pp. 298-313 in P. J. den Boer & G. R. Gradwell, eds. Dynamics of Populations. Centre for Agricultural Publishing and Documentation, Wageningen, The Netherlands.

Connell, J. H. 1978. Diversity in tropical rain forests and coral reefs. Science 199:1302-1310. Cornwell, William K., Dylan W. Schwilk & David D. Ackerly. 2006. A trait-based test for

habitat filtering: convex hull volume. Ecology 87:1465-1471. Currie, David J. & Viviane Paquin. 1987. Large-scale biogeographical patterns of species

richness of trees. Nature 329:326-327. Diamond, J. & T. J. Case, eds. 1986. Community Ecology. Harper and Row, NY, NY. Ellwood, M. D. Farnon, Andrea Manica & William A. Foster. 2009. Stochastic and

deterministic processes jointly structure tropical arthropod communities. Ecology Letters 12:277-284.

Felsenstein, J. 1981. Skepticism towards Santa Rosalia, or why are there so few kinds of animals? Evolution 35:124-138.

Field, Field, R., B. A. Hawkins, H. V. Cornell, D. J. Currie, J. A. F. Diniz-Filho, J. F. Guegan, D. M. Kaufman, J. T. Kerr, G. G. Mittelbach, T. Oberdorff, O, ', E. M. Brien, and J. R. G. Turner. 2009. Spatial species-richness gradients across scales: a meta-analysis. Journal of Biogeography 36:132-147.

Fine, Paul V. A. 2001. An evaluation of the geographic area hypothesis using the latitudinal gradient in North American tree diversity. Evolutionary Ecology Research 3:413-428.

Fine, Paul V. A. 2015. Ecological and evolutionary drivers of geographic variation in species diversity. Annual Review of Ecology, Evolution & Systematics 46:369-392.

Fine, Paul V. A. & Richard H. Ree. 2006. Evidence for a time-integrated species-area effect on the latitudinal gradient in tree diversity. American Naturalist 168:796-804.

Fine, Paul V. A., R. H. Ree & R. J. Burnham. 2008. The disparity in tree species richness between tropical, temperate ad boreal biomes: the geographic area and age hypothesis. In: W. P. Carson and S. Schnitzer (eds.) Tropical Forest Community Ecology. Blackwell Scientific, London.

Fischer, A. G. 1960. Latitudinal variations in organic diversity. Evolution 14:64-81. Fox, Jeremy W. 2013. The intermediate disturbance hypothesis should be abandoned. Trends

in Ecology & Evolution 28:86-92. Gentry, Alwyn H. 1982. Patterns of neotropical plant species diversity. Evolutionary Biology

15:1-84. Gentry, Alwyn H. 1988. Changes in plant community diversity and floristic composition on

environmental and geographical gradients. Annals of the Missouri Botanical Garden 75:1-34.

Gentry, Alwyn H. 1992. Tropical forest biodiversity: Distributional patterns and their conservational significance. Oikos 63:19-28.

Gilbert, L.E. 1980. Food web organization and the conservation of neotropical diversity. In: M. E. Soule and B. A. Wilcox, eds. Conservation Biology. Sinauer.

Givnish, Thomas J. 1999. On the causes of gradients in tropical tree diversity. J. Ecol. 87:193-210.

35

Grace, J. B. 1995. In search of the Holy Grail: Explanations for the coexistence of plant species. Trends in Ecology and Evolution 10:263-264.

Grace, James B., Susan Harrison & Ellen I. Damschen. 2011. Local richness along gradients in the Siskiyou herb flora: R. H. Whittaker revisited. Ecology 92:108-120. [Abstract – “We find that regional pool size, combined with a modest number of other interacting abiotic and biotic factors, explains most of the variation in local herb richness in the Siskiyou biodiversity hotspot.”]

GrilliJ_etal_2017_Nature_HigherOrderInteractionsStabilityDynamicsCompNetworkModels.pdf

Grubb, P. J. 1977. The maintenance of species-richness in plant communities: the importance of the regeneration niche. Biological Review 52:107-145.

Grubb, P. J. 1986. Global trends in species-richness in terrestrial vegetation: a view from the northern hemisphere. Pp. 99-118 in J. H. R. Gee & P. S. Giller, eds. Organization of Communities Past and Present. Blackwell Scientific Publications, London, UK.

Grutter, Alexandra S., Jan Maree Murphy & J. Howard Choat. 2003. Cleaner fish drives local fish diversity on coral reefs. Current Biology 13:64-67.

Hacker, S. D. & S. D. Gaines. 1997. Some implications of direct positive interactions for community species diversity. Ecology 78:1990-2003.

Harms, Kyle E., Paul R. Gagnon, Heather A. Passmore, Jonathan A. Myers & William J. Platt. 2017. Groundcover community assembly in high-diversity pine savannas: Seed arrival and fire-generated environmental filtering. Ecosphere 8(3):e01716.10.1002/ecs2.1716.

Harms, Kyle E., Richard Condit, Stephen P. Hubbell & Robin B. Foster. 2001. Habitat associations of trees and shrubs in a 50-ha Neotropical forest plot. Journal of Ecology 89:947-959.

Harms, Kyle E., S. Joseph Wright, Osvaldo Calderón, Andrés Hernández & Edward Allen Herre. 2000. Pervasive density-dependent recruitment enhances seedling diversity in a tropical forest. Nature 404:493-495.

Harpole, W. Stanley & David Tilman. 2007. Grassland species loss resulting from reduced niche dimension. Nature…

Houlahan, J. E., D. J. Currie, K. Cottenie, G. S. Cumming, S. K. M. Ernest, C. S. Findlay, S. D. Fuhlendorf, U. Gaedke, P. Legendre, J. J. Magnuson, B. H. McArdle, E. H. Muldavin, D. Noble, R. Russell, R. D. Stevens, T. J. Willis, I. P. Woiwod & S. M. Wondzell. 2007. Compensatory dynamics are rare in natural ecological communities. PNAS 104:3273-3277.

Hubbell, Stephen P. 2005. Neutral theory in community ecology and the hypothesis of functional equivalence. XXXX 19:166-172.

Hubbell, Stephen P. & Robin B. Foster. 1986. Biology, chance and history and the structure of tropical rain forest tree communities. Pp. 314-329 in J. Diamond & T. J. Case, eds., Community Ecology. Harper and Row, NY, NY. [Hubbell and Foster (1986) – Biology, chance and history are not so much three separate types of explanation for patterns, but rather history gave rise to contemporary patterns, and our challenge is to determine the relative contributions of biology vs. chance in that history.]

Hurtt, George C. & Stephen W. Pacala. 1995. The consequences of recruitment limitation: reconciling chance, history and competitive differences between plants. J. Theor. Biol. 176:1-12.

Huston, Michael. 1979. A general hypothesis of species diversity. The American Naturalist

36

113:81-101. Huston, Michael A. 1994. Biological diversity: the coexistence of species on changing

landscapes. Cambridge University Press, Cambridge, UK. Hutchinson, G. E. 1959. Homage to Santa Rosalia or Why are there so many kinds of

animals? The American Naturalist 93:145-159. Hutchinson, G. E. 1961. The paradox of the plankton. The American Naturalist 95:137-145. Janzen, D. H. 1970. Herbivores and the number of tree species in tropical forests. The

American Naturalist 104:501-528. Jetz, Walter & Paul V. A. Fine. 2012. Global gradients in vertebrate diversity predicted by

historical area-productivity dynamics and contemporary environment. PLoS Biology 10:1-11. [Time-Area-Productivity & Temperature]

Kaspari, Michael, Sean O’Donnell & James R. Kercher. 2000. Energy, density, and constraints to species richness: Ant assemblages along a productivity gradient. The American Naturalist 155:280-293.

Kaspari, Michael, May Yuan & Leeanne Alonso. 2003. Spatial grain and the causes of regional diversity gradients in ants. The American Naturalist 161:459-477.

Klopfer, Peter H. 1959. Environmental determinants of faunal diversity. The Am. Nat. 93:337-342.

Kraft, Nathan J. B., Peter B. Adler, Oscar Godoy, Emily C. James, Steve Fuller & Jonathan M. Levine. 2015. Community assembly, coexistence and the environmental filtering metaphor. Functional Ecology 29:592-599.

Kraft, Nathan J. B., Oscar Godoy & Jonathan M. Levine. 2015. Plant functional traits and the multidimensional nature of species coexistence. Proceedings of the National Academy of Sciences 112:797-802.

Latham, Roger Earl. 1992. Co-occurring tree species change rank in seedling performance with resources varied experimentally. Ecology 73:2129-2144. [See for discussion of differences between plant vs. animal ecology]

Latham, Roger Earl and Robert E. Ricklefs. 1993. Continental comparisons of temperate-zone tree species diversity. Pp. 294-314 in R. E. Ricklefs & D. Schluter, eds. Species Diversity in Ecological Communities. University of Chicago Press, Chicago, IL, USA.

Letten, Andrew D., Po-Ju Ke & Tadashi Fukami. 2017. Linking modern coexistence theory and contemporary niche theory. Ecological Monographs 87:161-177.

Levin, S. A. 1981. Mechanisms for the generation and maintenance of diversity. Pp. 173-194 in R. W. Hiorns & D. Cooke, eds. The mathematical theory of dynamics of biological populations. Academic Press, London, UK.

Levine, Jonathan M. & Janneke HilleRisLambers. 2009. The importance of niches for the maintenance of species diversity. Nature 461:254-257.

LevineJM_etal_2017_Nature_BeyondPairwiseMechanismsSpeciesCoexistence.pdf Logue, Jürg B., Nicolas Mouquet, Hannes Peter, Helmut Hillebrand & The Metacommunity

Working Group. 2011. Empirical approaches to metacommunities: a review and comparison with theory. Trends in Ecology & Evolution 26:482-491.

LokeLHL_PATodd_2016_Ecology_StructuralComplexityIncreasesIntertidalDiversity.pdf Lomolino, Mark V. 2001. Elevation gradients of species-density: historical and prospective

views. Global Ecology & Biogeography 10:3-13. MackKML_JDBever_2014_JournalEcology.pdf – local-scale feedbacks and coexistence! Marra, P. P. & J. V. Remsen. 1997. Insights into the maintenance of high species diversity in

37

the Neotropics. Ornithological Monographs 48:445-483. Marshall, Katie E. & Jennifer L. Baltzer. 2015. Decreased competitive interactions drive a

reverse species richness latitudinal gradient in subarctic forests. Ecology 96:461-470. Menge, Bruce A. & John P. Sutherland. 1976. Species diversity gradients: synthesis of the

roles of predation, competition, and temporal heterogeneity. The American Naturalist 110:351-369.

Menge, Bruce A. & John P. Sutherland. 1987. Community regulation: variation in disturbance, competition, and predation in relation to environmental stress and recruitment. The American Naturalist 130:730-757. [Note – their graphical model is similar to Bertness & Callaway (1994; TREE)]

Muller-Landau. 2010. PNAS. Palmer, Michael W. 1994. Variation in species richness: towards unification of hypotheses.

Folia Geobotanica & Phytotaxonomica 29:511-530. Petraitis, P. S., R. E. Latham & R. A. Niesenbaum. 1989. The maintenance of species

diversity by disturbance. The Quarterly Review of Biology 64:393-418. Pianka, E. R. 1966. Latitudinal gradients in species diversity: A review of concepts. Am.

Nat. 100:33-46. Pimm, Stuart L. & James H. Brown. 2004. Domains of diversity. Science 304:831-833.

[Commentary on mid-domain effects] Power, Mary E., David Tilman, James A. Estes, Bruce A. Menge, William J. Bond, L. Scott

Mills, Gretchen Daily, Juan Carlos Castilla, Jane Lubchenco & Robert T. Paine. 1996. Challenges in the quest for keystones. BioScience 46:609-620.

Prance, G. T. 1982. A review of the phytogeographic evidences for Pleistocene climate changes in the Neotropics. Ann. Missouri Bot. Gard. 69:594-624.

Presley, Steven J., Laura M. Cisneros, Bruce D. Patterson & Michael R. Willig. 2011. Vertebrate metacommunity structure along an extensive elevational gradient in the tropics: a comparison of bats, rodents and birds. Global Ecology and Biogeography…

Presley, Steven J., Christopher L. Higgins & Michael R. Willig. 2010. A comprehensive framework for the evaluation of metacommunity structure. Oikos 119:908-917.

Rahbek, C. 1997. The relationship among area, elevation and regional species richness in Neotropical birds. American Naturalist 149:875-902.

Rahel, F.J., J.D. Lyons and P.A. Cochran. 1984. Stochastic or deterministic regulation of assemblage structure. It may depend on how the assemblage is defined. The American Naturalist 124:583-589.

Rangel, Thiago F. et al. 2018. Modeling the ecology and evolution of biodiversity: Biogeographical cradles, museums, and graves. Science 361:244.

Remsen, Jr., J. V. 1990. Community ecology of Neotropical kingfishers. Univ. Calif. Publ. Zool. 124:1-116.

Ricklefs, R. E. & D. Schluter, eds. 1993. Species Diversity in Ecological Communities. University of Chicago Press, Chicago, IL, USA.

Rohde, K. 1997. The larger area of the tropics does not explain the latitudinal gradient in species diversity. Oikos 79:169-172.

Rosenzweig. 1995. Species diversity in space and time. Cambridge University Press, New York, NY, USA.

Rosenzweig, Michael L. 1992. Species diversity gradients: We know more and less than we thought. Journal of Mammalogy 73:715-730.

38

Rull_2011_TREE_NeotropicalBiodiversity.pdf Saavedra, Serguei, Rudolf P. Rohr, Jordi Bascompte, Oscar Godoy, Nathan J. B. Kraft &

Jonathan M. Levine. 2017. A structural approach for understanding multispecies coexistence. Ecological Monographs 87:470-486.

Sale, P. F. 1979. Recruitment, loss and coexistence in a guild of territorial coral reef fishes. Oecologia 42:159-177.

Sale, P. F. 1985. The structure of communities of fish on coral reefs and the merit of a hypothesis-testing, manipulative approach to ecology. Pp. 478-490 in D. R. Strong, Jr., D. Simberloff, L. G. Abele & A.B. Thistle, eds., Ecological Communities: Conceptual Issues and the Evidence. Princeton University Press, Princeton, NJ.

Sale, P. F. & R. Dybdhal. 1978. Determinants of community structure for coral reef fishes in isolated coral heads at lagoonal and reef slope sites. Oecologia 34:57-74.

Sanders, H.L. 1969. Benthic marine diversity and the stability-time hypothesis. Brookhaven Symposium in Biology. Vol. 22:71-80.

Scheffer, M., Rinaldi, S., Huisman, J., Weissing, F.J. 2003. Why plankton communities have no equilibrium: solutions to the paradox. Hydrobiol. 491, 9-18.

SchluterD_MWPennell_2017_Nature_SpeciationGradients.pdf Shmida, A. and S. Ellner. 1984. Coexistence of plant species with similar niches. Vegetatio

58:29-55. Shmida, A. and M. V. Willson. 1985. Biological determinants of species diversity. Journal of

Biogeography 12:1-20. Siepielski, Adam M. & Mark A. McPeek. 2010. On the evidence for species coexistence: a

critique of the coexistence program. Ecology 91:3153-3164. Silvertown, Jonathan. 2005. Demons in Eden: The Paradox of Plant Diversity. University of

Chicago Press, Chicago, IL. Simonis, Joseph L. & Julie C. Ellis. 2014. Bathing birds bias b-diversity: Frequent dispersal

by gulls homogenizes fauna in a rock-pool metacommunity. Ecology 95:1545-1555. StevensMHH_WPCarson_1999_Ecology.pdf Stevens, Richard D., J. Sebastián Tello & Maria Mercedes Gavilanez. 2013. Stronger tests of

mechanisms underlying geographic gradients in biodiversity: insights from the dimensionality of biodiversity. PLoS ONE 8:1-11.

Strong, D. R., Jr., D. Simberloff, L. G. Abele & A. B. Thistle, eds. 1984. Ecological Communities: Conceptual Issues and the Evidence. Princeton University Press, Princeton, NJ.

Terborgh, J. 1973. On the notion of favorableness in plant ecology. The American Naturalist 107:481-501.

Terborgh, J. W., S. K. Robinson, T. A. Parker III, C. A. Munn & N. Pierpont. 1990. Structure and organization of an Amazonian forest bird community. Ecological Monographs 60:213-238.

Thompson, K. 1987. The resource ratio hypothesis and the meaning of competition. Functional Ecology 1:297-303.

Thompson, K. M & J. P. Grime. 1988. Competition reconsidered – a reply to Tilman. Functional Ecology 2:114-116.

Tilman, David. 2004. Niche tradeoffs, neutrality, and community structure: a stochastic theory of resource competition, invasion, and community assembly. PNAS 101:10854-10861.

39

Tilman, David & Stephen Pacala. 1993. The maintenance of species richness in plant communities. Pp. 13-25 in Robert E. Ricklefs & Dolph Schulter, eds., Species Diversity in Ecological Communities: Historical and Geographical Perspectives. The University of Chicago Press, Chicago, IL.

Van Der Plas, Fons, Thijs Janzen, Alejandro Ordonez, Wimke Fokkema, Josephine Reinders, Rampal S. Etienne & Han Olff. 2015. A new modeling approach estimates the relative importance of different community assembly processes. Ecology 96:1502-1515.

[i.e., trait-based assessment to examine relative contributions of dispersal, environmental filtering, and limiting similarity]

Vellend, Mark, Diane S. Srivastava, Kathryn M. Anderson, Carissa D. Brown, Jill E. Jankowski, Elizabeth J. Kleynhans, Nathan J. B. Kraft, Alathea D. Letaw, A. Andrew M. Macdonald, Janet E. Maclean, Isla H. Myers-Smith, Andrea B. Norris & Xinxin Xue. 2014. Assessing the relative importance of neutral stochasticity in ecological communities. Oikos 123:1420-1430.

Wade, Lizzie. 2015. Cradle of life. Science 350:496-501. WangX_etal_2016_Ecology_StochasticDilutionEffectsWeakenDeterministicEffectsSpRichFor

ests.pdf Warner, R. R. & P. L. Chesson. 1985. Coexistence mediated by recruitment fluctuations: A

field guide to the storage effect. The American Naturalist 125:769-787. Weigelt, Patrick, Manuel Jonas Steinbauer, Juliano Sarmento Cabral & Holger Kreft. 2016.

Late Quaternary climate change shapes island biodiversity. Nature 532:99-102. WeiserMD_etal_2017_Ecography_DivGradients_AbundAdaptationHypoth.pdf Willig, M. R., D. M. Kaufman & R. D. Stevens. 2003. Latitudinal gradients of biodiversity:

Pattern, process, scale, and synthesis. Ann. Rev. Ecol. Evol. Syst. 34:273-309. Wilson, J. Bastow. 1990. Mechanisms of species coexistence: Twelve explanations for

Hutchinson’s ‘paradox of the plankton’: Evidence from New Zealand plant communities. New Zealand Journal of Ecology 13:17-42.

Wilson, J. Bastow. 2011. The twelve theories of co-existence in plant communities: the doubtful, the important and the unexplored. Journal of Vegetation Science 22:184-195.

Winegardner, Amanda K., Brittany K. Jones, Ingrid S. Y. Ng, Tadeu Siqueira & Karl Cottenie. 2012. The terminology of metacommunity ecology. Trends in Ecology & Evolution 27:253-254.

Witman, Jon D., Ron J. Etter & Franz Smith. 2004. The relationship between regional and local species diversity in marine benthic communities: a global perspective. Proceedings of the National Academy of Science 101:15664-15669.

Wright, D. H. 1983. Species-energy theory: An extension of species-area theory. Oikos 41:496-506.

Wright, S. Joseph. 1999. Plant Diversity in Tropical Forests. Pp. 449-471 in Francisco I. Pugnaire & Fernando Valladares, eds. Handbook of Functional Plant Ecology. Marcel Dekker, Inc. New York, NY.

Wright, S. Joseph. 2002. Plant diversity in tropical forests: a review of mechanisms of species coexistence. Oecologia 130:1-14.

Yenni, Glenda, Peter B. Adler & S. K. Morgan Ernest. 2012. Strong self-limitation promotes the persistence of rare species. Ecology 93:456-461.

40

11. Origins and maintenance of species diversity and diversity gradients. II. Phylogenetic perspectives, community-level convergence, and etc.

Additional references: See: Integrating Ecology and Phylogenetics (2012) Ecology v. 93, no. 8 Supplement. See: Phylogenetic Approaches to Community Ecology (2006) Ecology v. 87 no. 7

Supplement. Ackerly, D. D. & W. K. Cornwell. 2007. A trait-based approach to community assembly:

partitioning of species trait values into within- and among-community components. Ecology Letters 10:135-145.

Anderson, T. Michael, Joey Shaw & Han Olff. 2011. Ecology’s cruel dilemma, phylogenetic trait evolution and the assembly of Serengeti plant communities. J. Ecol. 99:797-806.

Avise, J. C. 2000. Phylogeography: The History and Formation of Species. Harvard University Press, Cambridge, MA.

Barot, Sébastien & Jacques Gignoux. 2004. Mechanisms promoting plant coexistence: can all the proposed processes be reconciled? Oikos 106:185-192.

Belya, L. R. & J. Lancaster. 1999. Assembly rules within a contingent ecology. Oikos 86:402-416.

Blackburn, T. M. & K. J. Gaston. 2001. Local avian assemblages as random draws from regional pools. Ecography 24:50-58.

Brooks, Daniel R. & Deborah A. McLennan. 1991. Phylogeny, Ecology, and Behavior. Chicago U. Press, Chicago, IL, USA.

Brooks, Daniel R. & Deborah A. McLennan. 1993. Pp. 267-280 in Robert E. Ricklefs & Dolph Schulter, eds., Species Diversity in Ecological Communities: Historical and Geographical Perspectives. The University of Chicago Press, Chicago, IL.

Brown, J. M., M. A. McPeek & M. L. May. 2000. A phylogenetic perspective on habitat shifts and diversity in the North American Enallagma damselflies. Systematic Biology 49:697-712.

Cadle, John E. & Harry W. Green. 1993. Phylogenetic patterns, biogeography, and the ecological structure of Neotropical snake assemblages. Pp. 281-293 in Robert E. Ricklefs & Dolph Schulter, eds., Species Diversity in Ecological Communities: Historical and Geographical Perspectives. The University of Chicago Press, Chicago, IL.

Cavender-Bares, J., D. D. Ackerly, D. A. Baum & F. A. Bazzaz. 2004. Phylogenetic overdispersion in Floridian oak communities. Am. Nat. 163:823-843.

Cavender-Bares, Jeannine, David D. Ackerly & Kenneth H. Kozak. 2012. Integrating ecology and phylogenetics: the footprint of history in modern-day communities 93 (Supplement):S1-S3.

Cavender-Bares Jeanine, Kenneth H. Kozak, Paul V. A. Fine & Steven W. Kembel. 2009. The merging of community ecology and phylogenetic biology. Ecology Letters 12:693-715.

Cavender-Bares, Jeanine & Amity Wilczek. 2003. Integrating micro- and macroevolutionary processes in community ecology. Ecology 84:592-597.

Chazdon, Robin L., Sonia Careaga, Campbell Webb & Orlando Vargas. 2003. Community

41

and phylogenetic structure of reproductive traits of woody species in wet tropical forests. Ecological Monographs 73:331-348.

Cody, M. L. & H. A. Mooney. 1978. Convergence versus non-convergence in Mediterranean-climate ecosystems. Annual Review of Ecology and Systematics 9:265-321.

Cornell, H. V. & J. H. Lawton. 1992. Species interactions, local and regional processes, and limits to the richness of ecological communities: A theoretical perspective. Journal of Animal Ecology 61:1-12.

Emerson, Brent C. & Rosemary G. Gillespie. 2008. Phylogenetic analysis of community assembly and structure over space and time. Trends in Ecology & Evolution 23:619-630.

Esselstyn, Jacob A., Sean P. Maher & Rafe M. Brown. 2011. Species interactions during diversification and community assembly in an island radiation of shrews. PLoS ONE 6(7): e21885

Fordyce, James A. 2010. Host shifts and evolutionary radiations of butterflies. Proceedings of the Royal Society of London, B. Online…

Fox, B. J. 1999. The genesis and development of guild assembly rules. Pp. 23-57 in E. Weiher & P. Keddy, eds., Ecological Assembly Rules: Perspectives, Advances, Retreats. Cambridge University Press, Cambridge, UK.

Futuyma, D. J. & C. Mitter. 1996. Insect-plant interactions: The evolution of component communities. Philosophical Transactions of the Royal Society of London, Series B 351:1361-1366.

Gillespie, Rosemary. 2004. Community assembly through adaptive radiation in Hawaiian spiders. Science 303:356-359.

Godoy, Oscar, Nathan J. B. Kraft & Jonathan M. Levine. 2014. Phylogenetic relatedness and the determinants of competitive outcomes. Ecology Letters 17:836-844.

Graham, C. H. & P. V. A. Fine. 2008. Phylogenetic beta diversity: linking ecological and evolutionary processes across space and time. Ecology Letters 11:1265-1277.

Grandcolas, P. 1998. Phylogenetic analysis and the study of community structure. Oikos 82:397-400.

Grant, P. R. & B. R. Grant. 2002. Adaptive radiation in Darwin’s finches. American Scientist 90:130-139.

Hardy, O. J. 2008. Testing the spatial phylogenetic structure of local communities: statistical performances of different null models and test statistics on a locally neutral community. Journal of Ecology 96:914-926.

Hardy, O. J. & B. Senterre. 2007. Characterizing the phylogenetic structure of communities by an additive partitioning of phylogenetic diversity. Journal of Ecology 95:493-506.

Harvey, Paul H. & Mark D. Pagel. 1991. The Comparative Method in Evolutionary Biology. Oxford U. Press, Oxford, UK.

Hickerson, M. J., B. C. Carstens, J. Cavender-Bares, K. A. Crandall, C. H. Graham, J. B. Johnson, L. Rissler, P. F. Victoriano & A. D. Yoder. 2010. Phylogeography’s past, present, and future: 10 years after Avise, 2000. Molecular Phylogenetics and Evolution 54:291-301.

Kartzinel, Tyler R., Patricia A. Chen, Tyler C. Coverdale, David L. Erickson, W. John Kress, Maria L. Kuzmina, Daniel I. Rubenstein, Wei Wang & Robert M. Pringle. 2015. DNA metabarcoding illuminates dietary niche partitioning by African large herbivores. Proceedings of the National Academy of Sciences 112:8019-8024. [See also commentary: Tilman & Borer (2015) PNAS 112:7890-7891]

42

Keddy, P. A. 1992. Assembly and response rules: Two goals for predictive community ecology. Journal of Vegetation Science 3:157-164. Might be good for discussion!

Kraft, Nathan J. B., William K. Cornwell, Campbell O. Webb & David D. Ackerly. 2007. Trait evolution, community assembly, and the phylogenetic structure of ecological communities. Am. Nat. 170:271-283.

Losos, Jonathan B. 1990. A phylogenetic analysis of character displacement in Caribbean Anolis lizards. Evolution 44:558-569.

Losos, Jonathan B. 1992. The evolution of convergent structure in Caribbean Anolis communities. Systematic Biology 41:403-420.

Losos, Jonathan B. 1996. Phylogenetic perspectives on community ecology. Ecology 77:1344-1354.

Losos, Jonathan B. 2009. Lizards in an Evolutionary Tree: Ecology and Adaptive Radiation of Anoles. University of California Press, Berkeley, CA.

Losos, J. B., T. R. Jackman, A. Larson, K. de Queiroz & L. Rodriguez-Schettino. 1998. Contingency and determinism in replicated adaptive radiations of island lizards. Science 279:2115-2118.

Mayfield, Margaret M. & Jonathan M. Levine. 2010. Opposing effects of competitive exclusion on the phylogenetic structure of communities. Ecology Letters 13:1085-1093.

McGee, Matthew D., Samuel R. Borstein, Russell Y. Neches, Heinz H. Buescher, Ole Seehausen & Peter C. Wainwright. 2015. A pharyngeal jaw evolutionary innovation facilitated extinction in Lake Victoria cichlids. Science 350:1077-1079. [See also commentary by Geerat Vermeij (2015) Science 350:1038]

McPeek, Mark A. 1995. Morphological evolution mediated by behavior in the damselflies of two communities. Evolution 49:749-769.

McPeek, Mark A. & T. E. Miller. 1996. Evolutionary biology and community ecology. Ecology 77:1319-1320.

Merckx,Vincent S. F. T. et al. 2015. Evolution of endemism on a young tropical mountain. Nature 524:347-350. [“… most of [Mt. Kinabalu’s] unique biodiversity is younger than the mountain itself (6 million years), and comprises a mix of immigrant pre-adapted lineages and descendants from local lowland ancestors, although substantial shifts from lower to higher vegetation zones in this latter group were rare.”]

Milne, B. T. & R. T. T. Forman. 1986. Peninsulas in Maine: Woody plant diversity, distance, and environmental patterns. Ecology 67:967-974.

Morlon, Hélène. 2014. Phylogenetic approaches for studying diversification. Ecology Letters…

Pearse, Ian S. & Andrew L. Hipp. 2009. Phylogenetic and trait similarity to a native species predict herbivory on non-native oaks. PNAS 106:18097-18102.

Pigot, Alex L. & Rampal S. Etienne. 2014. A new dynamic null model for phylogenetic community structure. Ecology Letters…

Rabosky et al. 2018. An inverse latitudinal gradient in speciation rate for marine fishes. Nature 559:392-395.

RibasCC_etal_2011_ProcRoySocB_PalaeobiogeographicModelForAmazonianDiversification.pdf

Ricklefs, Robert E. & Dolph Schluter. 1993. Species diversity: Regional and historical influences. Pp. 350-363 in R. E. Ricklefs & D. Schluter, eds., Species Diversity in Ecological Communities. Chicago University Press, Chicago, IL.

43

Ricklefs, Robert E. & Dolph Schulter, eds. 1993. Species Diversity in Ecological Communities: Historical and Geographical Perspectives. The University of Chicago Press, Chicago, IL.

Schluter, Dolph. 2000. The Ecology of Adaptive Radiation. Oxford University Press, Oxford, UK.

Schweitzer, Jennifer A., Joseph K. Bailey, Brian J. Rehill, Gregory D. Martinsen, Stephen C. Hart, Richard L. Lindroth, Paul Keim & Thomas G. Whitham. 2004. Genetically based trait in a dominant tree affects ecosystem processes. Ecology Letters 7:127-134.

Stevens, Richard D., Maria M. Gavilanez, Juan S. Tello & David A. Ray. 2011. Phylogenetic structure illuminates the mechanistic role of environmental heterogeneity in community organization. Journal of Animal Ecology x:xx-xx.

Swenson, N. G. et al. 2007. The influence of spatial and size scale on phylogenetic relatedness in tropical forest communities. Ecology 88:1770-1780.

Swenson, N. G. et al. 2006. The problem and promise of scale dependency in community phylogenetics. Ecology 87:2418-2424.

ter Braak, Caro J. F., Anouk Cormont & Stéphane Dray. 2012. Improved testing of species traits-environment relationships in the fourth-corner problem. Ecology 93:1525-1526.

Tofts, R. & J. Silvertown. 2000. A phylogenetic approach to community assembly from a local species pool. Proceedings of the Royal Society of London, Series B 267:363-369.

Urban, Mark C., Mathew A. Leibold, Priyanga Amarasekare, Luc De Meester, Richard Gomulkiewicz, Michael E. Hochberg, Christopher A. Klausmeier, Nicolas Loeuille, Claire de Mazancourt, Jon Norberg, Jelena H. Pantel, Sharon Y. Strauss, Mark Vellend & Michael J. Wade. 2008. The evolutionary ecology of metacommunities. TREE 23:311-317.

Webb, Campbell O. 2000. Exploring phylogenetic structure of ecological communities: An example for rain forest trees. The American Naturalist 156:145-155.

Webb, Campbell O., David D. Ackerly, Mark A. McPeek & Micheal J. Donoghue. 2002. Phylogenies and community ecology. Annual Review of Ecology and Systematics 33:475-505.

Weiher, E. & P. A. Keddy. 1995. The assembly of experimental wetland plant communities. Oikos 73:323-335.

Whitham, Thomas G., William P. Young, Gregory D. Martinsen, Catherine A. Gehring, Jennifer A. Schweitzer, Stephen M. Shuster, Gina M. Wimp, Dylan G. Fischer, Joseph K. Bailey, Richard L. Lindroth, Scott Woolbright & Cheryl R. Kuske. 2003. Community and ecosystem genetics: a consequence of the extended phenotype. Ecology 84:559-573.

Wiens, John J. & Catherine H. Graham. 2005. Niche conservatism: integrating evolution, ecology, and conservation biology. Annual Reviews of Ecology, Evolution & Systematics 36:519-539.

Wiens, John J., Catherine H. Graham, Daniel S. Moen, Sarah A. Smith & Tod W. Reeder. 2006. Evolutionary and ecological causes of the latitudinal diversity gradient in hylid frogs: treefrog trees unearth the roots of high tropical diversity. Am. Nat. 168:579-596.

Willis, J. C. 1922. Area and Age: A Study of Geographical Distribution and Origin in Species. Cambridge University Press, Cambridge, UK.

Wilson, J. B., T. C. E. Wells, I. C. Trueman, G. Jones, M. D. Atkinson, M. J. Crawley, M. E. Dodd & J. Silvertown. 1996. Are there assembly rules for plant species abundance: An investigation in relation to soil resources and successional trends. Journal of Ecology 84:527-538.

44

12. Diversity-productivity, diversity-stability, diversity-invasibility, and richness-function relationships

Additional references: Abrams, Peter A. 1995. Monotonic or unimodal diversity-productivity gradients: What does

competition theory predict? Ecology 76:2019-2027. Adler, Peter B. et al. 2011. Science. Al-Mufti et al. 1977. J. Ecol. 65:759- [Good for “hump-shaped” figure] Benedetti-Cecchi, L. 2004. Increasing accuracy of causal inference in experimental analyses

of biodiversity. Functional Ecology 18:761-768. Cardinale, Bradley J., Margaret A. Palmer & Scott L. Collins. 2002. Species diversity

enhances ecosystem functioning through interspecific facilitation. Nature 415:426-429. Connell, J. H. & W. P. Sousa. 1983. On the evidence needed to judge ecological stability or

persistence. The American Naturalist 121:789-824. Coomes, David A., Georges Kunstler, Charles D. Canham & Elaine Wright. 2009. A greater

range of shade-tolerance niches in nutrient-rich forests: an explanation for positive richness-productivity relationships? Journal of Ecology 97:705-717.

Crutsinger, Gregory M., Michael D. Collins, James A. Fordyce, Zachariah Gompert, Chris C. Nice, Nathan J. Sanders. 2006. Plant genotypic diversity predicts community structure and governs an ecosystem process. Science 313:966-968.

Davis, M. B. 1986. Climatic instability, time lags, and community disequilibrium. Pp. 269-284 in J. Diamond & T. J. Case, eds., Community Ecology. Harper & Row, NY, NY.

Duffy, J. Emmett. 2009. Why biodiversity is important to the functioning of real-world ecosystems. Front. Ecol. Environ. 7:437-444.

DuffyJE_etal_2017_Nature_BiodivKeyDriverProdInTheWild.pdf Elton, C. S. 1958. The Ecology of Invasions by Animals and Plants. Methuen & Co.,

London, UK. Frank & McNaughton. 1991. Stability increases with diversity in plant communities:

Empirical evidence from the 1988 Yellowstone drought. Oikos 62:360-362. Fraser, Lauchlan H. et al. 2015. Worldwide evidence of a unimodal relationship between

productivity and plant species richness. Science 349:302-305. Fridley, J. D., J. J. Stachowicz, S. Naeem, D. F. Sax, E. W. Seabloom, M. D. Smith, T. J.

Stohlgren, D. Tilman & B. Von Holle. 2007. The invasion paradox: Reconciling pattern and process in species invasions. Ecology 88:3-17. [Add figure from Metha Klock’s presentation!]

Gough, Laura, Craig W. Osenberg, Katherine L. Gross & Scott L. Collins. 2000. Fertilization effects on species density and primary productivity in herbaceous plant communities. Oikos 89:428-439.

GraceJB_etal_2007_EcolLetters_DoesSpeciesDivLimitProdInGrasslands.pdf [SEM that simultaneously evaluates influence of abiotic factors on productivity and richness, influence of richness on prod., and influence of prod. on richness…; see Adler et al. 2011. Science]

45

Grace, James B., T. Michael Anderson, Eric W. Seabloom, Elizabeth T. Borer, Peter B. Adler, W. Stanley Harpole, Yann Hautier, Helmut Hillebrand, Eric M. Lind, Meelis Pärtel, Jonathan D. Bakker, Yvonne M. Buckley, Michael J. Crawley, Ellen I. Damschen, Kendi F. Davies, Philip A. Fay, Jennifer Firn, Daniel S. Gruner, Andy Hector, Johannes M. H. Knops, Andrew S. MacDougall, Brett A. Melbourne, John W. Morgan, John L. Orrock, Suzanne M. Prober & Melinda D. Smith. 2016. Integrative modelling reveals mechanisms linking productivity and plant species richness. Nature 529:xx-xx…

Grime, J. P. 1998. Benefits of plant diversity to ecosystems: immediate, filter and founder effects. Journal of Ecology 86:902-910. [Grime’s Mass Ratio Theory]

Grimm, V. 1996. A down-to-earth assessment of stability concepts in ecology: dreams, demands, and the real problems. Senckenbergiana maritima 27:215-226.

Grimm, V., and C. Wissel. 1997. Babel, or the ecological stability discussions: an inventory and analysis of terminology and a guide for avoiding confusion. Oecologia 109:323-334. [163 definitions of stability categorized into: persistence, constancy, resilience, resistance, elasticity, domain of attraction]

Gross, Katherine & B. J. Cardinale. 2007. Does species richness drive community production or vice versa? Reconciling historical and contemporary paradigms in competitive communities. American Naturalist 170:207-220.

Harpole, W. Stanley et al. 2016. Addition of multiple limiting resources reduces grassland diversity. Nature 537:93-96.

HarpoleWS_etal_2017_FunctionalEcology_MultNutrientLimitationsDriveBiomassProdRelationship.pdf

Hautier, Yann, David Tilman, Forest Isbell, Eric W. Seabloom, Elizabeth T. Borer & Peter B. Reich. 2015. Anthropogenic environmental changes affect ecosystem stability via biodiversity. Science 348:336-340.

He, Qiang & Mark D. Bertness. 2014. Extreme stresses, niches, and positive species interactions along stress gradients. Ecology 95:1437-1443. [An assessment of the Stress-Gradient Hypothesis]

Hector, A., B. Schmid, C. Beierkuhnlein, M. C. Caldeira, M. Diemer, P. G. Dimitrakopoulos, J. Finn, H. Freitas, P. S. Giller, J. Good, R. Harris, P. Högberg, K. Huss-Danell, J. Joshi, A. Jumpponen, C. Körner, P. W. Leadley, M. Loreau, A. Minns, C. P. H. Mulder, G. O'Donovan, S. J. Otway, J. S. Pereira, A. Prinz, D. J. Read, M. Scherer-Lorenzen, E.-D. Schulze, A.-S.D. Siamantziouras, E. M. Spehn, A. C. Terry, A. Y. Troumbis, F. I. Woodward, S. Yachi, S. & J. H. Lawton. 1999. Plant diversity and productivity experiments in European grasslands. Science 286:1123-1127.

Hocking, Morgan D. & John D. Reynolds. 2011. Impacts of salmon on riparian plant diversity. Science 331:1609-1612.

Hodgson, J. G., K. Thompson, P. J. Wilson & A. Bogaard. 1998. Does biodiversity determine ecosystem function? The Ecotron experiment reconsidered. Functional Ecology 12:843-848.

Huston, M. A. 1997. Hidden treatments in ecological experiments: Re-evaluating the ecosystem function of biodiversity. Oecologia 110:449-460.

Huston, Michael A. & Steve Wolverton. 2009. The global distribution of net primary production: resolving the paradox. Ecological Monographs 79:343-377.

Isbell, Forest et al. 2015. Biodiversity increases the resistance of ecosystem productivity to climate extremes. Nature 526:574-577.

46

Ives, Anthony R. & Stephen R. Carpenter. 2007. Stability and diversity of ecosystems. Science 317:58-62.

Jiang, Lin, Shiqiang Wan & Linghao Li. 2009. Species diversity and productivity: why do results of diversity-manipulation experiments differ from natural patterns? Journal of Ecology 97:603-608.

Karlson, Ronald H. 1999. Dynamics of Coral Communities. Kluwer Academic Pubs., Dordrecht, The Netherlands.

Larsen, Trond H., Neal M. Williams & Claire Kremen. 2005. Extinction order and altered community structure rapidly disrupt ecosystem functioning. Ecology Letters 8:538-547.

Lawton, J. H., S. Naeem, L. J. Thompson, A. Hector, and M. J. Crawley. 1998. Biodiversity and ecosystem function: Getting the Ecotron experiment in its correct context. Functional Ecology 12:848-852.

Leigh, Egbert Giles, Jr. & Geerat Jacobus Vermeij. 2002. Does natural selection organize ecosystems for the maintenance of high productivity and diversity? Philosophical Transactions of the Royal Society of London, Series B 357:709-718.

Levin, S. A. 1970. Community equilibria and stability, and an extension of the competitive exclusion principle. American Naturalist 104:413-423. [See McPeek (2014)]

Levine, Jonathan M. 2000. Species diversity and biological invasions: Relating local process to community pattern. Science 288:852-854.

Levine, Jonathan M. & C. M. D’Antonio. 1999. Elton revisited: a review of evidence linking diversity and invasibility. Oikos 87:15-26.

Lewontin, R. C. 1969. The meanings of stability. Brookhaven Symp. Biology 22:13-24. [On alternative stable states]

Liang, Jingjing et al. 2016. Positive biodiversity-productivity relationship predominant in global forests. Science 354:196. [“777,126 global forest biodiversity permanent sample plots… consistent positive and concave-down biodiversity-productivity relationship…”]

Loreau, M., S. Naeem, P. Inchausti, J. Bengtsson, J. P. Grime, A. Hector, D. U. Hooper, M. A. Huston, D. Raffaelli, B. Schmid, D. Tilman & D. A. Wardle. 2001. Biodiversity and ecosystem functioning: Current knowledge and future challenges. Science 294:804-808.

Loucks, O. L. 1970. Evolution of diversity, efficiency, and community stability. American Zoologist 10:17-25.

MacDougall, Andrew S., Benjamin Gilbert & Jonathan M. Levine. 2009. Plant invasions and the niche. Journal of Ecology 97:609-615.

Maron, John L., Marilyn Marler, John N. Kilronomos & Cory C. Cleveland. 2011. Soil fungal pathogens and the relationship between plant diversity and productivity. Ecology Letters 14:36-41.

May, Robert M. 2000. Relation between diversity and stability, in the real world. Science 290:51.

McCann, Kevin Shear. 2000. The diversity-stability debate. Nature 405:228-233. McNaughton, S. J. 1977. Diversity and stability of ecological communities: A comment on

the role of empiricism in ecology. The American Naturalist 111:515-525. Mittelbach, Gary G., Christopher F. Steiner, Samuel M. Scheiner, Katherine L. Gross, Heather

L. Reynolds, Robert B. Waide, Michael R. Willig, Stanley I. Dodson & Laura Gough. 2001. What is the observed relationship between species richness and productivity? Ecology 82:2381-2396.

47

Naeem, S. & J. P. Wright. 2003. Disentangling biodiversity effects on ecosystem functioning: Deriving solutions to a seemingly insurmountable problem. Ecology Letters 6:567-579.

Nowlin, Weston H., María J. González, Michael J. Vanni, M. Henry H. Stevens, Matthew W. Fields & Jonathan J. Valente. 2007. Allochthonous subsidy of periodical cicadas affects the dynamics and stability of pond communities. Ecology 88:2174-2186.

Oksanen, J. 1996. Is the humped relationship between species richness and biomass an artifact due to plot size? Journal of Ecology 84:293-295.

Orians, Gordon H. 1975. Diversity, stability and maturity in natural ecosystems. In W. H. Van Dobben & R. H. Lowe-McConnell, eds., Unifying Concepts in Ecology. Dr. W. Junk Publishers, The Netherlands.

Paine, Robert T. 2002. Trophic control of production in a rocky intertidal community. Science 296:736-739.

Peterson, G., C. R. Allen & C. S. Holling. 1998. Ecological resilience, biodiversity, and scale. Ecosystems 1:6-8.

Pierik, Marleen, Jasper van Ruijven, T. Martijn Bezemer, Rob H. E. M. Geerts & Frank Berendse. 2011. Recovery of plant species richness during long-term fertilization of a species-rich grassland.

Pimm, Stuart L. 1984. The complexity and stability of ecosystems. Nature 307:321-326. Rosenzweig, Michael L. & Zvika Abramsky. 1993. How are diversity and productivity

related? Pp. 52-65 in Robert E. Ricklefs & Dolph Schulter, eds., Species Diversity in Ecological Communities: Historical and Geographical Perspectives. The University of Chicago Press, Chicago, IL.

Sankaran, M. & S. M. McNaughton. 1999. Determinants of biodiversity regulate compositional stability of communities. Nature 401:691-693.

Scherber, Christopher, Alexandru Milcu, Stephan Partsch, Stefan Scheu & Wolfgang W. Weisser. 2006. The effects of plant diversity and insect herbivory on performance of individual plant species in experimental grassland. Journal of Ecology 94:922-931.

Stevens, M. Henry H. & Walter Carson. 1999. The significance of assemblage-level thinning for species richness. Journal of Ecology 87:490-502.

Storkey, J., A. J. Macdonald, P. R. Poulton, T. Scott, I. H. Köhler, H. Schnyder, K. W. T. Goulding & M. J. Crawley. 2015. Grassland biodiversity bounces back from long-term nitrogen addition. Nature 528:401-404. [See also commentary: David Tilman & Forest Isbell. 2015. Recovery as nitrogen declines. Nature 528:336-337.]

Thompson, K. & J. G. Hodgson. 1998. Response to Lawton et al. Functional Ecology 12:852-853.

Tilman, David. 1997. Community invasibility, recruitment limitation, and grassland biodiversity. Ecology 78:81-92.

Tilman, David & J. A. Downing. 1994. Biodiversity and stability in grasslands. Nature 367:363-365.

Tilman, David, J. Knops, D. Wedin, P. Reich, M. Ritchie & E. Siemann. 1997. The influence of functional diversity and composition on ecosystem processes. Science 277:1300-1302.

Tilman, David, Clarence L. Lehman & Kendall T. Thomson. 1997. Plant diversity and ecosystem productivity: Theoretical considerations. Proceedings of the National Acadeny of Sciences 94:1857-1861.

Tilman, David, D. Wedin & J. Knops. 1996. Productivity and sustainability influences by biodiversity in grassland ecosystems. Nature 379:718-720.

48

Ushio, Masayuki, Chih-hao Hsieh, Reiji Masuda, Ethan R. Deyle, Hao Ye, Chun-Wei Chang, George Sugihara & Michio Kondoh. 2018. Fluctuating interaction network and time-varying stability of a natural fish community. Nature 554:360-363. [“interspecific interactions, community network structure and community stability are dynamic properties…”]

Vogelsang, K. M., H. L. Reynolds & J. D. Bever. 2006. Plant species diversity and productivity increase with mycorrhizal fungal diversity in a tallgrass prairie system. New Phytologist 172:554–562.

13. Disturbance and ecological succession; alternative stable states; priority effects; regime shifts, etc.

Additional references: Bazzaz, F. A. 1979. The physiological ecology of plant succession. Annual Review of

Ecology and Systematics 10:351-371. Bond, W. J. 1993. Keystone species. Pg. 237-253 in E. D. Schulze and H. A. Mooney, eds.

Ecosystem function and biodiversity. Springer-Verlag, Berlin, Germany. Bond, W. J. & J. E. Keeley. 2005. Fire as global ‘herbivore’: the ecology and evolution of

flammable ecosystems. Trends in Ecology and Evolution 20:387-394. Bond, W. J., F. I. Woodward & G. F. Midgley. 2005. The global distribution of ecosystems in

a world without fire. New Phytologist 165:525-538. Bongers, Frans, Lourens Poorter, William D. Hawthorne & Douglas Sheil. 2009. The

intermediate disturbance hypothesis applies to tropical forests, but disturbance contributes little to tree diversity. Ecology Letters 12:798-805.

Bormann, F. H. & G. E. Likens. 1979. Catastrophic disturbance and the steady state in northern hardwood forests. American Scientist 67:660-669.

Bormann, F. H. & G. E. Likens. 1979. Pattern and Process in a Forested Ecosystem… Bowman, D. Fire Ref. Brewer, J. Stephen. 2011. Disturbance-mediated competition between perennial plants along

a resource supply gradient. Journal of Ecology 99:1219-1228. Brokaw, N. V. L. 1987. Gap-phase regeneration of three pioneer tree species in a tropical

forest. Journal of Ecology 75:9-19. Brokaw, Nicholas & Richard T. Busing. 2000. Niche versus chance and tree diversity in

forest gaps. Trends in Ecology and Evolution 15:183-188. Brokaw, N., S. Fraver, J. S. Grear, J. Thompson, J. K. Zimmerman, R. B. Waide, E. M.

Everham III, S. P. Hubbell, and R. B. Foster. 2004. Disturbance and canopy structure in two tropical forests. Pages 177-194 in E. Losos and E. G. Leigh, Jr. (eds.). Tropical Forest Diversity and Dynamism: Results from a Long-Term Tropical Forest Network. University of Chicago Press, Chicago, USA.

Bruelheide et al. 2011. Community assembly during secondary forest succession in a Chinese subtropical forest. Ecological Monographs 81:25-41.

Bruno, John F. 2000. Facilitation of cobble beach plant communities through habitat modification by Spartina alterniflora. Ecology 81:1179-1192.

49

Bruno, John F., John J. Stachowicz & Mark D. Bertness. 2003. Inclusion of facilitation into ecological theory. Trends in Ecology & Evolution 18:119-125.

Buma, Brian, Sarah Bisbing, John Krapek & Glenn Wright. 2017. A foundation of ecology rediscovered: 100 years of succession on the William S. Cooper plots in Glacier Bay, Alaska. Ecology 98:1513-1523.

Bush, M. B. & R. J. Whittaker. 1991. Krakatau: Colonization patterns and hierarchies. Journal of Biogeography 18:341-356.

Caswell, H. & J. E. Cohen. 1991. Disturbance, interspecific interactions, and diversity in metapopulations. Biological Journal of the Linnean Society 42:193-218.

Chapin et al. 1994. Chazdon, R. L. 2008. Chance and determinism in tropical forest succession. In: Carson, W.

and S. Schnitzer, Eds. Tropical Forest Community Ecology. Wiley-Blackwell Publishing, pp. 384-408.

Clements, F. E. 1916. ??? Clements, F. E. 1928. Plant succession and indicators. A definitive edition of plant succession

and indicators. H.W. Wilson, New York, NY. Connell, J. H. & R. O. Slatyer. 1977. Mechanisms of succession in natural communities and

their role in community stability and organization. The American Naturalist 111:1119-1144.

Cooper, W. S. 1926. The fundamentals of vegetational change. Ecology 7:391-413. Cowles, H. C. 1899. The ecological relations of the vegetation on the sand dunes of Lake

Michigan. I. Geographical relations of the dune floras. Botanical Gazette 27:95-117,167-202,281-308,361-391.

Cowles, H. C. 1901. The physiographic ecology of Chicago and vicinity: A study of the origin, development, and classification of plant communities. Botanical Gazette 31:73-108,145-182.

Curran, L. M., I. Caniago, G. D. Paoli, D. Astianti, M. Kusneti, M. Leighton, C. E. Nirarita & H. Haeruman. 1999. Impact of El Niño and logging on canopy tree recruitment in Borneo. Science 286:2184-2188.

del Moral, R. & L. C. Bliss. 1993. Mechanisms of primary succession: Insights resulting from the eruption of Mount St. Helens. Advances in Ecological Research 24:1-66.

Deutschman, D. H., S. A. Levin & S. W. Pacala. 1999. Error propagation in a forest succession model: The role of fine-scale heterogeneity in light. Ecology 80:1927-1943.

Deutschman, D. H., S. A. Levin, C. Devine & L.A. Buttel. 1997. Scaling from trees to forests: Analysis of a complex simulation model. Science 227 [online] http://www.sciencemag.org/feature/data/deutschman/index.htm

Doyle, T. W. 1981. The role of disturbance in the gap dynamics of a montane rain forest: an application of a tropical forest succession model. Pages 56-73 in D. C. West, H. H. Shugart, and D. B. Botkin (eds.). Forest Succession: Concepts and Application. Springer-Verlag, New York.

Drayton, P. K. 1972. Toward an understanding of community resilience and the potential effects of enrichment to the benthos at McMurdo Sound, Antarctica. Pp. 81-95 in B. C. Parker, ed. Proceedings of the Colloquium on Conservation Problems in Antarctica. Allen Press, Lawrence, KS.

Drury, W. H. & I. C. T. Nisbet. 1973. Succession. Journal of the Arnold Arboretum 54:331-368.

50

Egler, F. E. 1954. Vegetation science concepts. I. Initial floristic composition – a factor in old-field vegetation development. Vegetatio 4:412-417.

Fagan, William F. & John G. Bishop. 2000. Trophic interactions during primary succession: herbivores slow a plant reinvasion at Mount St. Helens. Am. Nat. 155:238-251.

Fastie, Christopher L. 1995. Causes and ecosystem consequences of multiple pathways of primary succession at Glacier Bay, Alaska. Ecology 76:1899-1916.

Gleason, H. A. 1926. ??? Gleason, H. A. 1939. The individualistic concept of the plant association. American Midland

Naturalist 21:92-108. Grime, J. P. 1979. Plant strategies and Vegetation Processes. John Wiley & Sons,

Chichester, UK. Grubb, Peter J. 1985. Plant populations and vegetation in relation to habitat, disturbance and

competition: problems of generalization. Pp. 595-621 in J. White, ed. The Population Structure of Vegetation. Junk, Dordrecht.

Halpern, Charles B. 1989. Early successional patterns of forest species: interactions of life history traits and disturbance. Ecology 70:704-720.

Harper, J. L. 1967. A Darwinian approach to plant ecology. Journal of Ecology 55:247-270. Horn, H. S. 1975. Markovian properties of forest succession. Pp. 196-211 in M. L. Cody & J.

M. Diamond, Ecology and Evolution of Communities. Belknap Press, Cambridge, UK. Hubbell, S. P., R. B. Foster, S. T. O’Brien, K. E. Harms, R. Condit, B. Wechsler, S. J. Wright

& S. Loo de Lao. 1999. Light-gap disturbances, recruitment limitation, and tree diversity in a neotropical forest. Science 283:554-557.

Hughes, T. P. 1994. Catastrophes, phase shifts, and large-scale degradation of a Caribbean coral reef. Science 265:1547-1551.

Huston, M. & T. Smith. 1987. Plant succession: life history and competition. American Naturalist 130:168-198.

Levin, S. A. & R. T. Paine. 1974. Disturbance, patch formation, and community structure. PNAS 71:2744-2747.

Lorimer, C. G. 1989. Relative effects of small and large disturbances on temperate hardwood forest structure. Ecology 70:565-567.

McCook, L. J. 1994. Understanding ecological community succession: Causal models and theories, a review. Vegetatio 110:115-147.

Michaud, Jean-Philippe, Kenneth G. Schoenly & Gaétan Moreau. 2015. Rewriting ecological succession history: did carrion ecologists get there first? The Quarterly Review of Biology 90:45-66.

Oliver, C. D. 1980. Forest development in North America following major disturbances. Forest Ecology and Management 3:153-168.

Oliver, C. D. & B. C. Larson. 1996. Forest Stand Dynamics. John Wiley & Sons, New York, NY.

Pacala, Stephen W., Charles D. Canham, John Saponara, John A. Silander, Jr., Richard K. Kobe & Eric Ribbens. 1996. Forest models defined by field measurements: Estimation, error analysis and dynamics. Ecological Monographs 66:1-43.

Paine, R. T. & S. A. Levin. 1981. Intertidal landscapes: Disturbance and the dynamics of pattern. Ecological Monographs 51:145-178.

Paine, Robert T., Mia J. Tegner & Edward A. Johnson. 1998. Compounded perturbations yield ecological surprises. Ecosystems 1:535-545.

51

Peet, R. K. 1992. Community structure and ecosystem function. Pp. 103-151 in D. C. Glenn-Lewin, R. K. Peet & T. T. Veblen, eds. Plant Succession: Theory and Prediction. Chapman & Hall, London, UK.

Peet, R. K. & N. L. Christensen. 1980. Succession: A population process. Vegetatio 43:131-140.

Peltzer, Duane A. et al. 2010. Understanding ecosystem retrogression. Ecological Monographs 80:509-529.

Pickett, S. T. A. 1976. Succession: An evolutionary interpretation. The American Naturalist 110:107-119.

Pickett, S. T. A., S. L. Collins & J. J. Armesto. 1987. Models, mechanisms, and pathways of succession. Botanical Review 53:335-371.

Pickett, S. T. A. & M. J. McDonnell. 1989. Changing perspectives in community dynamics: A theory of successional forces. Trends in Ecology and Evolution 4:241-245.

Pickett, S. T. A. & P. S. White, eds. 1985. The Ecology of Natural Disturbance and Patch Dynamics. Academic Press, New York, NY.

Platt, William J. 1975. The colonization and formation of equilibrium plant species associations on badger disturbances in a tall-grass prairie. Ecological Monographs 45:285-305.

Platt, William J., Brian Beckage, R. F. Doren & H. H. Slater. 2002. Interactions of large-scale disturbances: prior fire regimes and hurricane mortality of savanna pines. Ecology 83:1566-1572.

Platt, William J. & Joseph H. Connell. 2003. Natural disturbances and directional replacement of species. Ecological Monographs 73:507-522.

Roxburgh, S. H., K. Shea & J. B. Wilson. 2004. The intermediate disturbance hypothesis: patch dynamics and mechanisms of species coexistence. Ecology 85:359-371.

Runkle, J. R. 1981. Gap regeneration in some old-growth forests of the eastern United States. Ecology 62:1041-1051.

Shea, K., S. H. Roxburgh & E. S. J. Raushert. 2004. Moving from pattern to process: coexistence mechanisms under intermediate disturbance regimes. Ecology Letters 7:491-508.

Sheil, D. & D. F. R. P. Burslem. 2003. Disturbing hypotheses in tropical forests. TREE 18:18-26.

Shugart, H. H. & D. C. West. 1977. Development of an Appalachian deciduous forest model and its application to assessment of the impact of the chestnut blight. Journal of Environmental Management 5:161-179.

Smith, T. & M. Huston. 1989. A theory of the spatial and temporal dynamics of plant communities. Vegetatio 83:49-69.

Sousa, Wayne P. 1979. Disturbance in marine intertidal boulder fields: the nonequilibrium maintenance of species diversity. Ecology 60:1225-1239.

Sousa, Wayne P. 1984. The role of disturbance in natural communities. Annual Review of Ecology and Systematics 15:353-391.

Stevens, Jens T. & Brian Beckage. 2009. Fire feedbacks facilitate invasion of pine savannas by Brazilian pepper (Schinus terebinthifolius). New Phytologist.

Suding, K.N., K.L. Gross, & G.R. Houseman. 2004. Alternative states and positive feedbacks in restoration ecology. Trends Ecol. Evol. 19:46-53.

52

Suding, K.N., & R.J. Hobbs. 2009a. Threshold models in restoration and conservation: a developing framework. Trends Ecol. Evol 24:271-279.

Tansley, A. G. 1920. The classification of vegetation and the concept of development. Journal of Ecology 8:118-149.

Thoreau, H. D. 1860. The succession of forest trees. Read to the Middlesex Agricultural Society, Concorde, MA, September 1860. Reprinted in 1863 in Excursions, pages 135-160. Ticknor and Fields, Boston, MA.

Turner, Monica G. 2010. Disturbance and landscape dynamics in a changing world. Ecology 91:2833-2849.

Uriarte, María, Charles D. Canham, Jill Thompson, Jess K. Zimmerman, Lora Murphy, Alberto M. Sabat, Ned Fetcher & Bruce L. Haines. 2009. Natural disturbance and human land use as determinants of tropical forest dynamics: results from a forest simulator. Ecological Monographs 79:423-443.

Veblen, T. T. 1992. Regeneration dynamics. Pp. 152-187 in D. C. Glenn-Lewin, R. K. Peet & T. T. Veblen, eds. Plant Succession: Theory and Prediction. Chapman & Hall, London, UK.

Veldman, Joe – old-growth grasslands… Vitousek, P. M. & L. R. Walker. 1987. Colonization, succession, and resource availability:

ecosystem-level interactions. Pp. 207-233 in A. J. Gray, M. J. Crawley & P. J. Edwards, eds. Colonization, Succession, and Stability. Blackwell, Boston, MA.

Vitousek, P. M & P. S. White. 1981. Process studies in succession. Pp. 267-276 in D. C. West, H. H. Shugart & D. B. Botkin, eds. Forest Succession. Springer-Verlag, New York, NY.

Walker, Lawrence R. 2012. The Biology of Disturbed Habitats. The Biology of Habitats Series. Oxford University Press, Oxford, UK.

Wardle, David A., Lawrence R. Walker & Richard D. Bardgett. 2004. Ecosystem properties and forest decline in contrasting long-term chronosequences. Science 305:509-513.

Werner, P. A. 1976. Ecology of plant populations in successional environments. Systematic Botany 1:246-268.

Whittaker, R. H. 1975. The design and stability of plant communities. Pp. 169-181 in W.H. Dobben & R.H. Lowe-McConnell, eds. Unifying Concepts in Ecology: Report of the Plenary Session of the 1st International Congress of Ecology. Junk, The Hague, The Netherlands.

Whittaker, R. J. 1994. ??? Whittaker, R. J., M. B. Bush & K. Richards. 1989. Plant recolonization and vegetation

succession on the Krakatau Islands, Indonesia. Ecological Monographs 59:59-123. Williamson – pine savanna. Young, Truman P., Jonathan M. Chase & Russell T. Huddleston. 2001. Community

succession and assembly: Comparing, contrasting and combining paradigms in the context of ecological restoration. Ecological Restoration 19:5-18. [Contrasts of assembly and succession]

14. Scaling relationships within communities; Metabolic Ecology Theory; etc. Additional references:

53

Bonner, John Tyler. 1985. The Evolution of Complexity. Princeton University Press,

Princeton, NJ. Boyer, Alison G., Jean-Luc E. Cartron & James H. Brown. 2010. Interspecific pairwise

relationships among body size, clutch size and latitude: deconstructing a macroecological triangle in birds. Journal of Biogeography 37:47-56.

Brown, J. H. et al. 2004. Constraints on negative relationships… In Taper et al., eds. The Nature of Scientific Evidence…

Brown, James H., James F. Gillooly, Andrew P. Allen, Van M. Savage & Geoffrey B. West. 2004. Toward a metabolic theory of ecology. Ecology 85:1771-1789. [See also the commentaries on this research framework, same issue of Ecology]

Enquist, Brian J. 2002. Universal scaling in tree and vascular plant allometry: toward a general quantitative theory linking plant form and function from cells to ecosystems. Tree Physiology 22:1045-1064.

Enquist, Brian, James Brown & G. West. 1998. Allometric scaling of plant energetics and population density. Nature 395:163-165.

Enquist, Brian J., John P. Haskell & Bruce H. Tiffney. 2002. General patterns of taxonomic and biomass partitioning in extant and fossil plant communities. Nature 419:610-613.

Enquist, Brian J. & Karl J. Niklas. 2001. Invariant scaling relations across tree-dominated communities. Nature 410:655-660.

Fridley, Jason D., Robert K. Peet, Thomas R. Wentworth & Peter S. White. 2005. Connecting fine- and broad-scale species-area relationships of southeastern U.S. flora. Ecology 86:1172-1177.

Gaines, S. & M. Denny. 1993. The largest, smallest, highest, lowest, longest, and shortest: extremes in ecology. Ecology 74:1677-1692.

Greenhill, A. G. 1881. Determination of the greatest height consistent with stability that a verticle pole or mast can be made, and the greatest height to which a tree of given proportions can grow. Proceedings of the Cambridge Philosophical Society 4:65-73.

Huxley, J. S. 1932. Problems of Relative Growth. Methuen & Co., London, UK. Kerkhoff & Enquist. 2009 (in press). Multiplicative by nature: why log-transformation is

necessary for allometry. J. Theor. Biol. Mandelbrot, B. B. 1983. The Fractal Geometry of Nature. Freeman, New York, NY. McMahon, Thomas A. 1973. Size and shape in biology. Science 179:1201-1204. McMahon, Thomas A. 1975. The mechanical design of trees. Scientific American 233:93-

102. McMahon, Thomas A. & John Tyler Bonner. 1983. Size and Life. Scientific American

Library, NY, NY. Morse, D. R., J. H. Lawton, M. M. Dodson & M. H. Williamson. 1985. Fractal dimension of

vegetation and the distribution of arthropod body lengths. Nature 314:731-733. Payne, J., Boyer, A. G., et al. 2009. Two-phase increase in the maximum size of life over 3.5

billion years reflects biological innovation and environmental opportunity. Proceedings of the National Academy of Sciences USA. 106:24-27.

SavageV_etal_2008_PlosCompBiol.pdf Steele, Mark A. & Graham E. Forrester. 2005. Small-scale field experiments accurately scale

up to predict density dependence in reef fish populations at large scales. PNAS 102:13513-13516.

54

Sugihara, G. & R.M. May. 1990. Applications of fractals in ecology. Trends in Ecology and Evolution 5:79-86.

Thompson, D’Arcy W. 1942. On Growth and Form. Cambridge University Press, Cambridge, UK.

Wang, Zhiheng, James H. Brown, Zhiyao Tang & Jingyun Fang. 2009. Temperature dependence, spatial scale, and tree species diversity in eastern Asian and North America. PNAS.

Westcott, David A. & Devon L. Graham. 2000. Patterns of movement and seed dispersal of a tropical frugivore. Oecologia 122:249-257.

West, G. B., J. H. Brown & B. J. Enquist. 1997. A general model for the origins of allometric scaling laws in biology. Science 276:122-126.

Whitfield, John. 2006. In the Beat of Heart: Life, Energy, and the Unity of Nature. Joseph Henry Press, Washington, D. C.

Wright, Ian J., Peter B. Reich, Mark Westoby, David D. Ackerly et al. 2004. The worldwide leaf economics spectrum. Nature 428:821-827.

Yoda, Kyoji, Tatuo Kira, Husato Ogawa & Kazuo Hozumi. 1963. Self-thinning and overcrowded pure stands under cultivated and natural conditions (intraspecific competition among higher plants XI). Journal of Biology, Osaka City University 14:107-129.

15. Community & ecosystem ecology plus conservation within human-dominated landscapes & seascapes; global change; etc.

Additional references: Aide, T. Mitchell & H. Ricardo Grau. 2004. Globalization, migration, and Latin American

ecosystems. Science 305:1915-1916. Balmford, Andrew, Aaron Buner, Philip Cooper, Robert Costanza, Stephen Farber, Rhys E.

Green, Martin Jenkins, Paul Jeffries, Valma Jessamy, Joah Madden, Kat Munro, Norman Myers, Shahid Naeem, Jouni Paavola, Matthew Rayment, Sergio Rosendo, Joan Roughgarden, Kate Trumper & R. Kerry Turner. 2002. Economic reasons for conserving wild nature. Science 297:950-953.

Baron, J. S., N. L. Poff, P. L. Angermeier et al. 2002. Meeting ecological and societal needs for freshwater. Ecol. Apps. 12:1247-1260.

Baum - sharks Blossey, Bernd & Rolf Notzold. 1995. Evolution of increased competitive ability in invasive

nonindigenous plants: a hypothesis. J. Ecol. 83:887-889. Bohlen, Patrick J., Stefan Scheu, Cindy M. Hale, Mary Ann McLean, Sonja Migge, Peter M.

Groffman & Dennis Parkinson. 2004. Non-native invasive earthworms as agents of change in northern temperate forests. Frontiers in Ecology and the Environment 2:427-435.

Bruner, Aaron G., Raymond E. Gullison, Richard E. Rice & Gustavo A. B. da Fonseca. 2001. Effectiveness of parks in protecting tropical biodiversity. Science 291:125-128.

Cardinale, Bradley J., Emmett Duffy, Andrew Gonzalez, David U. Hooper, Charles Perrings, Patrick Venail, Anita Narwani, Georgina M. Mace, David Tilman, David A. Wardle, Ann P. Kinzig, Gretchen C. Daily, Michel Loreau, James B. Grace, Anne Larigauderie, Diane S.

55

Srivastava & Shahid Naeem. 2012. Biodiversity loss and its impact on humanity. Nature 486:59-67.

Chapin, III, F. Stuart, Pamela A. Matson & Harold A. Mooney. 2002. Principles of Terrestrial Ecosystem Ecology. Springer, New York, NY.

Chapin, III, F. Stuart, Mary E. Power, Steward T. A. Pickett, Amy Freitag, Julie A. Reynolds, Robert B. Jackson, David M. Lodge, Clifford Duke, Scott L. Collins, Alison G. Power & Ann Bartuska. 2011. Earth Stewardship: science for action to sustain the human-earth system. Ecosphere 2:1-20.

Chapin, III, F. Stuart, Erika S. Zavaleta, Valerie T. Eviner, Rosamond L. Naylor, Peter M. Vitousek, Heather L. Reynolds, David U. Hooper, Sandra Lavorel, Osvaldo E. Sala, Sarah E. Hobbie, Michelle C. Mack & Sandra Diaz. 2000. Consequences of changing biodiversity. Nature 405:234-242.

Cole, D. N. 2000. Paradox of the primeval: ecological restoration in wilderness. Ecological Restoration 18:77-86.

Costanza, Robert, Ralph d'Arge, Rudolf de Groot, Stephen Farber, Monica Grasso, Bruce Hannon, Karin Limburg, Shahid Naeem, Robert V. O'Neill, Jose Paruelo, Robert G. Raskin, Paul Sutton & Marjan van den Belt. 1997. The value of the world's ecosystem services and natural capital. Nature 387:253-260.

Cronin, James T., Ganesh P. Bhattarai, Warwick J. Allen & Laura A. Meyerson. 2015. Biogeography of a plant invasion: plant-herbivore interactions. Ecology 96:1115-1127.

D’Antonio, C. M. & P. M. Vitousek. 1992. Biological invasions by exotic grasses, the grass/fire cycle, and global change. Ann. Rev. Ecol. Syst. 23:63-87.

Daily, Gretchen C., Susan Alexander, Paul R. Ehrlich, Larry Goulder, Jane Lubchenco, Pamela A. Matson, Harold A. Mooney, Sandra Postel, Stephen H. Schneider, David Tilman, and George M. Woodwell. 1997. Ecosystem services: Benefits supplied to human societies by natural ecosystems. Issues in Ecology 2:1-16.

Daily, Gretchen, Tore Söderqvist, Sara Aniyar, Kenneth Arrow, Partha Dasgupta, Paul R. Ehrlich, Carl Folke, AnnMari Jansson, Bengt-Owe Jansson, Nils Kautsky, Simon Levin, Jane Lubchenco, Karl-Göran Mäler, David Simpson, David Starrett, David Tilman & Brian Walker. 2000. The value of nature and the nature of value. Science 289:395-396.

Damschen, Ellen I., Susan Harrison & James B. Grace. 2010. Climate change effects on an endemic-rich edaphic flora: resurveying Robert H. Whittaker’s Siskiyou sites (Oregon, USA). Ecology 91:3609-3619.

DeWalt, Saara J. 2004. Natural-enemy release facilitates habitat expansion of the invasive tropical shrub Clidemia hirta. Ecology 85:471-483.

Diamond, Jared. 2005. Collapse: How Societies Choose to Fail or Succeed. Penguin Books, London, UK.

Dorcas, Michael E., John D. Willson, Robert N. Reed, Ray W. Snow, Michael R. Rochford, Melissa A. Miller, Walter E. Meshaka, Jr., Paul T. Andreadis, Frank J. Mazzotti, Christina M. Romagosa & Kristen M. Hart. 2012. Severe mammal declines coincide with proliferation of invasive Burmese pythons in Everglades National Park. Proceedings of the National Academy of Science, USA…

Ehrlich, Paul R. & Edward O. Wilson. 1991. Biodiversity studies: Science and policy. Science 253:758-762. [Reasons to preserve biodiversity: ethics, esthetics, economics & ecosystem services]

56

Fahey, Timothy J. et al. 2015. The promise and peril of intensive-site-based ecological research: insights from the Hubbard Brook ecosystem study. Ecology 96:885-901.

Fayle, Tom M., Edgar C. Turner, Yves Basset, Robert M. Ewers, Glen Reynolds & Vojtech Novotny. 2015. Whole-ecosystem experimental manipulations of tropical forests TREE…

Fine, Paul V. A. 2002. The invasibility of tropical forests by exotic plants. Journal of Tropical Ecology 18:687-705.

Fleeger, John W., Kevin R. Carman & Roger M. Nisbet. 2003. Indirect effects of contaminants in aquatic ecosystems. The Science of the Total Environment 317:207-233.

Fleishman, Erica, David E. Blockstein, John A. Hall, Michael B. Mascia, Murray A. Rudd, J. Michael Scott, William J. Sutherland, Ann M. Bartuska, A. Gordon Brown, Catherine A. Christen, Joel P. Clement, Dominick De la Sala, Clifford S. Duke, Marietta Eaton, Shirley J. Fiske, Hannah Gosnell, J. Christopher Haney, Michael Hutchins, Mary L. Klein, Jeffrey Marqusee, Barry R. Noon, John R. Nordgren, Paul M. Orbuch, Jimmie Powell, Steven P. Quarles, Kathryn A. Saterson, Charles C. Savitt, Brcue A. Stein, Michael S. Webster & Amy Vedder. 2011. Top 40 priorities for science to inform US conservation and management policy. BioScience 61:290-300.

Freed, Leonard A. & Rebecca L. Cann. 2009. Negative effects of an introduced bird species on growth and survival in a native bird community. Current Biology 19:1736-1740.

Gonzalez, Andrew et al. 2016. Estimating local biodiversity change: a critique of papers claiming no net loss of local biodiversity. Ecology 97:1949-1960.

Graham, Nicholas A. J., Shaun K. Wilson, Peter Carr, Andrew S. Hoey, Simon Jennings & M. Aaron MacNeil. 2018. Seabirds enhance coral reef productivity and functioning in the absence of invasive rats. Nature 559:250-253.

Green, Peter T., Dennis J. O’Dowd, Kirsti L. Abbott, Mick Jeffery, Kent Retallick & Ralph Mac Nally. 2011. Invasional meltdown: Invader-invader mutualism facilitates a secondary invasion. Ecology 92:1758-1768.

Imhoff, M. L., L. Bounoua, T. Ricketts et al. 2004. Global patterns in human consumption of net primary production. Nature 429:870-873.

Janzen, D. 1998. Gardenification of wildland nature and the human footprint. Science 279:1312-1313.

Kleijn, David, Frank Berendse, Ruben Smit & Niels Gilissen. 2001. Agri-environment schemes do not effectively protect biodiversity in Dutch agricultural landscapes. Nature 413:723-725.

Kremen, C., J. O. Niles, M. G. Dalton, G. C. Daily, P. R. Ehrlich, J. P. Fay, D. Grewal & R. P. Guillery. 2000. Economic incentives for rain forest conservation across scales. Science 288:1828-1832.

LaffertyKD_etal_2018_BiolLtrs_RatEradicationPalmyraRidsIslandOfMosquitoes.pdf Larsen, Trond H., Alejandro Lopera & Adrian Forsyth. 2008. Understanding trait-dependent

community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22:1288-1298.

Levin, Simon A. 1999. Fragile Dominion: Complexity and the Commons. Perseus Publishing, Cambridge, MA.

Liu, Z. G. & X. M. Zou. 2002. Exotic earthworms accelerate plant litter decomposition in a Puerto Rican pasture and a wet forest. Ecological Applications 12:1406-1417.

Laurance

57

Lonsdale, W. M. 1999. Global patterns of plant invasions and the concept of invasibility. Ecology 80:1522-1536.

McKinney, M. L. & J. L. Lockwood. 1999. Biotic homogenization: a few winners replacing many losers in the next mass extinction. Trends Ecol. Evol. 14:450–453.

Moles, Angela T., Habacuc Flores-Moreno, Stephen P. Bonser, David I. Warton, Aveliina Helm, Laura Warman, David J. Edridge, Enrique Jurado, Frank A. Hemmings, Peter B. Reich, Jeannine Cavender-Bares, Eric W. Seabloom, Margaret M. Mayfield, Douglas Sheil, Jonathan C. Djietror, Pablo L. Peri, Lucas Enrico, Marcelo R. Cabido, Samantha A. Setterfield, Caroline E. R. Lehmann & Fiona J. Thomson. 2012. Invasions: the trail behind, the path ahead, and a test of a disturbing idea. Journal of Ecology 100:116-127.

Moorcroft Morton, S. R., O. Hoegh-Guldberg, D. B. Lindenmayer, M. Harriss Olson, L. Hughes, M. T.

McCulloch, S. McIntyre, H. A. Nix, S. M. Prober, D. A. Saunders, A. N. Andersen, M. A. Burgman, E. C. Lefroy, W. M. Lonsdale, I. Lowe, A. J. McMichael, J. S. Parslow, W. Steffen, J. E. Williams & J. C. Z. Woinarski. 2009. The big ecological questions inhibiting effective environmental management in Australia. Austral Ecology 34:1-9.

Myers, Norman, Russell A. Mittermeier, Cristina G. Mittermeier, Gustavo A. B. da Fonseca & Jennifer Kent. 2000. Biodiversity hotspots for conservation priorities. Nature 403:853-858.

Myers, R. A. & B. Worm. 2003. Rapid worldwide depletion of predatory fish communities. Nature 423:280-283.

Naeem, Shahid, Lindsey J. Thompson, Sharon P. Lawler, John H. Lawton & Richard M. Woodfin. 1994. Declining biodiversity can alter the performance of ecosystems. Nature 368:734-737.

Naeem, Shahid, F. S. Chapin III, Robert Costanza, Paul R. Ehrlich, Frank B. Golley, David U. Hooper, J. H. Lawton, Robert V. O'Neill, Harold A. Mooney, Osvaldo E. Sala, Amy J. Symstad and David Tilman. 1999. Biodiversity and ecosystem functioning: maintaining natural life support processes. Issues in Ecology 4:1-12.

Ngai & Srivastava. 2006. Predators accelerate nutrient cycling in a bromeliad ecosystem. Science 314:963. [Illustrates microcosm experiments to explore general principles…]

Odadi et al. 2011. Cattle facilitate & compete with wild ungulates – depends on season. Science. [Illustrates context dependencies…]

O’Dowd, Dennis J., Peter T. Green & P. S. Lake. 2003. Invasional ‘meltdown’ on an oceanic island. Ecology Letters 6:812-817.

Olden, J. D., N. L. Poff, M. R. Douglas, M. E. Douglas & K. D. Fausch. 2004. Ecological and evolutionary consequences of biotic homogenization. Trends in Ecology & Evolution 19:18-24.

Pacala, S. W., E. Bulte, J. A. List & S. A. Levin. 2003. False alarm over environmental false alarms. Science 301:1187-1188.

Palumbi, Stephen R. 2001. Humans as the world's greatest evolutionary force. Science 293:1786-1790.

PastorJohn_Naiman_1992_SelectiveForaging_by_Beaver_vs_Moose_AltersEcosystems.pdf Pearse, Ian S., David J. Harris, Richard Karban & Andrew Sih. 2013. Predicting novel

herbivore-plant interactions. Oikos 122:1554-1564. Pereira, Henrique M., Paul W. Leadley, Vania Proenca, Rob Alkemade, Jorn P. W.

Scharlemann, Juan F. Fernandez-Manjarres, Miguel B. Araujo, Patricia Balvanera, Reinette

58

Biggs, William W. L. Cheung, Louise Chini, H. David Cooper, Eric L. Gilman, Sylvie Guenette, George C. Hurtt, Henry P. Huntington, Georgina M. Mace, Thierry Oberdorff, Carmen Revenga, Patricia Rodrigues, Robert J. Scholes, Ussif Rashid Sumaila & Matt Walpole. 2010. Scenarios for global biodiversity in the 21st century. Science 330:1496-1501.

Rahel, F. J. 2000. Homogenization of fish faunas across the United States. Science 288:854-856.

Rahel, F. J. 2002. Homogenization of freshwater faunas. Annual Review of Ecology and Systematics. Volume 33:291-315.

Reider, Kelsey E., Walter P. Carson & Maureen A. Donnelly. 2013. Effects of collared peccary (Pecari tajacu) exclusion on leaf litter amphibians and reptiles in a Neotropical wet forest, Costa Rica. Biological Conservation…

Resasco, Julian, Nick M. Haddad, John L. Orrock, DeWayne Shoemaker, Lars A. Brudvig, Ellen I. Damschen, Joshua J. Tewsbury & Douglas J. Levey. 2014. Landscape corridors can increase invasion by an exotic species and reduce diversity of native species. Ecology 95:2033-2039. [Solenopsis invicta]

Robinson, Doug – birds on BCI; especially the experiments… Roemer, Gary W., C. Josh Donlan & Franck Courchamp. 2002. Golden eagles, feral pigs, and

insular carnivores: How exotic species turn native predators into prey. Proceedings of the National Academy of Science, USA 99:791-796.

Rompré, Ghislain, W. Douglas Robinson, André Desrochers & George Angehr. 2009. Predicting declines in avian species richness under nonrandom patterns of habitat loss in a Neotropical landscape. Ecological Applications 19:1614-1627.

Rosenzweig, Michael L. 2001. The four questions: What does the introduction of exotic species do to diversity? Evolutionary Ecology Research 3:361-367.

Sanders, Nathan J., Nichoas J. Gotelli, Nicole E. Heller & Deborah M. Gordon. 2003. Community disassembly by an invasive species. Proceedings of the National Academy of Science, USA 100:2474-2477.

Sanderson, E. W., M. Jaiteh, M. A. Levy et al. 2002. The human footprint and the last of the wild. BioScience 52:891-904.

Schlesinger, William H. 1997. Biogeochemistry: An Analysis of Global Change. Second ed. Academic Press, San Diego, CA.

Simberloff, D. et al. 2012. The natives are restless, but not often and mostly when disturbed. Ecology 93:598-607.

Smith, Kevin G., Karen R. Lips & Jonathan M. Chase. 2009. Selecting for extinction: nonrandom disease-associated extinction homogenizes amphibian biotas. Ecology Letters 12:1069-1078.

Smith, Scott A., Graham Bell & Eldredge Bermingham. 2004. Cross-cordillera exchange mediated by the Panama Canal increased the species richness of local freshwater fish assemblages. Proc. R. Soc. Lond. B 271:1889-1896.

Sterner, Robert W. & James J. Elser. Ecological Stoichiometry: The Biology of Elements from Molecules to the Biosphere. Princeton University Press, Princeton, NJ.

Stokstad, Erik. 2014. Death of the Stars. Science 344:464-467. [Report on the biggest known die-off of sea stars, some of which are keystone species.]

Stouffer, Philip C., Richard O. Bierregaard Jr., Cheryl Strong & Thomas E. Lovejoy. 2006. Long-term landscape change and bird abundance in Amazonian rainforest fragments.

59

Conservation Biology 20:1212-1223. Terborgh, John. 1989. Where Have All the Birds Gone? Princeton University Press,

Princeton, NJ. Terborgh, John. 1999. Requiem for Nature. Island Press, Washington, DC. Terborgh, John, Lawrence Lopez, Percy Nuñez V., Madhu Rao, Ghazala Shahabuddin,

Gabriela Orihuela, Mailen Riveros, Rafael Ascanio, Greg H. Adler, Thomas D. Lambert & Luis Balbas. 2001. Ecological meltdown in predator-free forest fragments. Science 294:1923-1926.

Torchin, Mark, Kevin D. Lafferty, Andrew P. Dobson, Valerie J. McKenzie & Armand M. Kuris. 2003. Introduced species and their missing parasites. Nature 42:628-630.

Vitousek, P. 1994. Beyond global warming: Ecology and global change. Ecology 75:1861-1876.

Vitousek, Peter M., Harold A. Mooney, Jane Lubchenco & Jerry M. Melillo. 1997. Human domination of Earth's ecosystems. Science 277: 494-499.

Wilcove, David S. 2008. No Way Home: The Decline of the World’s Great Animal Migrations. Island Press, Washington, DC.

Willis, K. J., L. Gillson & T. M. Brncic. 2004. How "virgin" is virgin rainforest? 304:402-403.

Wilson, Edward O. 2002. The Future of Life. Vintage Books, New York, NY. Western, D. 1992. The biodiversity crisis: a challenge for biology. Oikos 63:29-38. Western, D. 2001. Human-dominated ecosystems and future evolution. PNAS 98:5458-5465. Wright, S. Joseph. 2005. Tropical forests in a changing environment. TREE 20:553-560.