BREEDING PRODUCTIVITY OF EGRETS AND HERONS AT SIX SITES IN ASIA

8/9/2019 BREEDING PRODUCTIVITY OF EGRETS AND HERONS AT SIX SITES IN ASIA

1/6

Xev. glo. (]erre Sie),vlo. 38, 8660.

739

JL]EHXSE

HXEEMIJ@ VXLMUG]ISI]Y LC E@XE]W BJM NEXLJW B] WI^ WI]EW IJ BWIB

Dburl CBWLOB2,0, Ougb GBJLSB2, Eoeljlrb HLJGLDVB@JI2, Bsniq DUNBDDBM8,Xbnbt JBKBD7, Oiu PEI=,Yubjnub MLJ@5& Yij`dei [NBJ@3

XWUD. Vrlmugtivitmes bi`rettes et nrljs jigneurs ej six sites bsibtiques. Ob prlmugtilj, obtbiooe mes ucs et oes gljmitiljs glrplreooes mes plussijs ljt t tumies gnez oe Nrlj `brme-hucs (Huhuo-gus ihis), oBi`rette ijterdmibire (E`rettb ijterdemib), oBi`rette `brzette (E`rettb `brzettb), oe Grbhiergnijlis (Brmelob hbggnus) et oe Hinlrebu `ris (Jygtiglrbx jygtiglrbx) ej meux sites me Gnije gejtrboe et mbjsqubtre butres bu Vbaistbj. Ob prlmugtilj ej Gnije sest bvre trs clrte, pous idplrtbjte qubu Vbaistbj,

prlhbhoedejt ej reobtilj bveg uje `rbjme mispljihioit me zljes nudimes plur oboidejtbtilj. Ob clrdesldhre me oBi`rette `brzette me ob gte mu Vbaistbj prsejtbit uje prlmugtivit si`jiflgbtivedejt pous cbihoeque geooe mes clrdes hobjgnes.

Bhujmbjt mbtb exist lj hreemij` prlmugtivity lc severbo spegies lc Brmeijbe clr Eurlpebjm clr Bderigb (Xuhloiji & Cbslob, ij press), wnerebs sgbjty ijclrdbtilj is bvbiobhoe clrgljtijejtbo Bsib, wnere hreemij` wbs stumiem ljoy ij Isrbeo (Bsaejbzi & Yld-]lv, 2990) bjmbt b cew ltner sites (Olj`snej` et bo.,299=1Oizni et bo.,299>1 Nioboummij, 8667). Bodlst uja-jlwj bre tne hreemij` pbrbdeters lc slde spegies, sugn bs tne Gnijese Vljm NerljBrmelobhbggnus(jl metbioem stumy ajlwj tl us) bjm tne Ijterdemibte E`ret E`rettb ijterdemib(ljoystumiem hy Hbxter, 299= bjm Dbmmlga & Hbxter, 2992 ij Bustrboib1bjm hy Dlreo & Dlreo,2932 ij Weje`bo).

Pe bid tl floo tnis `bp hy mesgrihij` tne hreemij` prlmugtivity lc flve Brmeijbe, GnijeseVljm Nerlj,Ijterdemibte E`ret, Gbttoe E`ret (Huhuogus ihis), Oittoe E`ret (E`rettb `brzettb),bjm Hobga-grlwjem Ji`nt Nerlj(Jygtiglrbx jygtiglrbx), bt twl sites ij Gejtrbo Gnijb lj tne

hlrmer lc Vlybj` Obae bjm lc ]bi Obae, bjm bt clur sites ij Vbaistbj, tne ijobjm b`riguoturbobrebs jebr Duotbj, jebr Obnlre, bjm jebr tne ]bujsb hbrrbè lj tne Xiver Ijmus, bjm tne glbstbobreb jebr Abrbgni.

2 Mipbrtidejtl Hilolìb Bjidboe, Ujiversit, Vibzzb Hlttb 9, I-80266 Vbvib, Itboy8 Jbtiljbo B`riguoturbo Xesebrgn Gejter1Vbra Xm., Isobdbhbm =5566, Vbaistbj7 PPC-Vbaistbj. 363-360Clrtuje Gejter, V.E.G.N.W., Wnbnrb-e-Cbisbo, Abrbgni 05=66. Vbaistbj= Hilolìgbo Xeslurges Ijstitute, Kibj`xi Bgbmedy lc Wgiejges, 776689 Jbjgnbj`, V.X. Gnijb5 Ijstitute lc Wlio Wgiejge, Gnijese Bgbmedy lc Wgiejges, 82666> Jbjkij`, V.X. Gnijb.3 Mepbrtdejt lc Hilol`y, Objznlu Ujiversity, 076666 Objznlu, V.X. Gnijb0 Glrrespljmij` butnlr. E-dbio; cbslobFujipv.it


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706

Wtioo ujgoebr is tne spegies lr suhspegies stbtus lc tne Oittoe E`ret dlrpns, tne jldijbte`brzettb, bjm tne midlrpnb, ùobris, bjm sgnistbgeb. Wlde butnlrs imejticy tne mbra e`retshreemij` lj tne Ijmibj Lgebj glbst bs b sepbrbte spegies, tne Pesterj Xeec Nerlj (E`rettbsgnistbgeb), wnioe we gljsimer boo tne wnite bjm mbra dlrpns bs heoljìj` tl b sijòe ploydlr-pnig spegies (Nbcjer et bo.,8668), bjm we mesi`jbte tne mbra e`rets stumiem bt Abrbgni bs OittoeE`ret, mbra dlrpnE`rettb `brzettbsgnistbgeb.

Pe stumiem hreemij` prlmugtivity bs pbrt lc b resebrgn lj e`rets bs dljitlrs lc ejvi-rljdejtbo gljtbdijbtilj. ]ne oeveo lc persistejt gljtbdijbjts, hltn lr`bjig bjm ijlr-`bjig, wbs clujm tl he olw bt tne brebs stumiem hltn ij Gnijb bjm ij Vbaistbj, bjm weoo

heolw tne oeveos tnbt dby bccegt e`ret reprlmugtilj (Hljgldpb`ji et bo.,86671 Wbjperbet bo.,86671 [nbj` et bo., 8663). Gnijb nloms very obrè judhers lc severbo spegies lce`rets bjm nerljs (Cbslob et bo.,866=), wnioe ij Vbaistbj tne dlst bhujmbjt spegies is tneGbttoe E`ret.

W]UMY BXEBW

Pe stumiem b sbdpoe lc jests bt lje gloljy, usubooy tne obrèst, bt ebgn lc tne cloolwij` six brebs (Ci`.2).

Ciùre2. ]ne six stumy brebs ij Vbaistbj bjm Gnijb.

(2) Vlybj` Obae, Kibj`xi Vrlvijge, tne obrèst cresnwbter obae (78>7 ad8), stioo ij b jebr-jbturbo stbte, bjm tne dlst

idplrtbjt site clr wbterclwo ij Gnijb. Crld 25 Dby tl 26 Kuje 2999, we stumiem b nerljry olgbtem ij b dixem wllmjebr @lj`qij` (8927J, 225=9E), witn 5>6 jests lc Oittoe E`ret, 235 lc Ijterdemibte E`ret, 56 lc @rebt Pnite E`ret,=86 lc Gbttoe E`ret, 06 lc Gnijese Vljm Nerlj, bjm 025 lc Hobga-grlwjem Ji`nt Nerlj. Hreemij` lc tne Hobga-grlwjem

Ji`nt Nerlj wbs borebmy mesgrihem clr b suhset lc tnese jests (Cbslob et bo., 8662).


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(8) ]bi Obae, Kibj`su Vrlvijge, lje lc tne obrèst cresnwbter obaes ij Gnijb (8=85 ad 8), olgbtem ij b ni`noyprlmugtive b`riguoturbo breb, witn rige fleoms lver 96% lc cbrdem objm. Crld 86 Bprio tl 26 Kuje 8666, we stumiem b veryobrè gloljy ij tne Yubjtluznu Vbra (7272J, 28627E) jebr Puxi. ]ne gloljy stretgnem lver severbo negtbres witnijb obrè wllmobjm, bjm ijgoumem 76,266 jests lc Hobga-grlwjem Ji`nt Nerlj, =,866 lc Oittoe E`ret, 2,666 lc GnijeseVljm Nerlj, bjm 2,266 lc Gbttoe E`ret.

(7) ]bujsb Hbrrbè, Vujkbh, b stlrbè reservlir henijm b hbrrbè lj tne Xiver Ijmus ij Vbaistbj. Bmkbgejt brebsbre guotivbtem witn glttlj, su`br gbje, wnebt, bjm tne hbrrbè is usem bosl clr plwer èjerbtilj. Crld 5 Dby bjm25 Kuje 8666, we stumiem b nerljry pobgem 5 ad west lc tne hbrrbè (76=8J, 0656E), witn 83 jests lc Oittoe E`ret,88 lc Ijterdemibte E`ret, 765 lc Gbttoe E`ret, bjm 25 lc Hobga-grlwjem Ji`nt Nerlj.

(=) Duotbj b`riguoturbo brebs, Vujkbh Vrlvijge ij Vbaistbj, witn dbj`l lrgnbrm, wnebt, clmmer, glttlj bjm rige.Crld 86 Dby tl 25 Kuje 8668, we stumiem b gloljy lj bgbgibs bolj` b gbjbo jebr Wbrmbrpur (7678J, 02=5E), 36 ad

Jlrtn lc Duotbj bjm bhlut 2 ad crld Xiver Ijmus, witn 2566 jests lc Gbttoe E`rets.(5) Obnlre, Vujkbh ij Vbaistbj, bj breb crld 72=0 tl 7877J bjm crld 07=8 tl 0=75E, guotivbtem dbijoy witn

rige, wnebt bjm clmmer. ]nree nerljries witn nujmrems lc jests lc Gbttoe E`rets, bt sites gbooem Xendbj, Xbkbjpur, bjmGnbga Hnuoibj-Jlrmwbo Xm, were stumiem crld 8 Kuje tl 29 Kuoy 8667.

(3) Abrbgni Nbrhlr, Wijm Vrlvijge ij Vbaistbj, bj breb lc timbo greeas, dbj`rlves bjm ijtertimbo dumbts, brlujmtne nbrhlr lc Abrbgni. Crld 7 tl 76 Dby 8666, we stumiem b nerljry ij tne @nbs Hujmer ob`llj (8==0J, 3022E),witn 225 jests lc Oittoe E`ret, boo mbra dlrpns, 86 jests lc Ijterdemibte E`ret, bjm 26 jests lc Ijmibj Vljm Nerlj

Brmelob rbyii.

DE]NLMW

Pe bidem tl sbdpoe bt oebst 86 jests per spegies bt tne peba lc tne reprlmugtive perilm, hut sgbrge spegies perdittemljoy sdbooer sbdpoes. Ij lrmer tl bvlim misturhij` tne hreemij` hirms, we sbdpoem tne jests tnbt were reobtiveoy bggessihoebdim tne ijtrigbte veètbtilj lc tne gloljies, bjm tnlse pobgem crld 7 tl 3d bhlve `rlujm. Pe nbm jl evimejge tnbt tnesbdpoe jests were hibsem clr bjy cebture reobtem tl tneir prlmugtivity. Jest gljtejt wbs gnegaem bt oebst twige per weeatl bsgertbij tne judher lc e``s, nbtgnem gnigas, bjm emèm gnigas. ]ne gnigas gbjjlt he trbgaem bcter bhlut 86 mbyslom, tnereclre tne judher emèm recers tl tne judher survivij` bt 86 mbys, b debsure glddljoy edpolyem ij nerljjestij` prlmugtivity stumies, bjm tnbt bpprlbgnes tne fljbo emìj` prlmugtivity, sijge dlst dlrtboity lggurs murij` tneflrst 86 mbys (Hejjetts et bo.,8666). ]ne jests tnbt cbioem gldpoeteoy were jlt ijgoumem ij tne gboguobtiljs.

Clr b suh-sbdpoe lc jests, we debsurem e`` mibdeters. Pnej tne lomest gniga wbs 86 mbys lom, tbrsus oej`tnbjm hlmy dbss were debsurem clr ebgn gniga. Wijge tbrsus oej`tn mepejms dbijoy lj bè, exgept ij gbses lc severedbojutritilj, wnioe dbss is obrèoy ijuejgem hy tne bdlujt lc cllm regeivem, tne dbss ij reobtilj tl tbrsus oej`tn

reoibhoy ijmigbtes gniga gljmitilj (Kbglhs, 2993 1 Cbslob, 299>). B gljmitilj ijmex wbs gboguobtem clr ebgn gniga bsrbtil hetweej its dbss bjm its tbrsus oej`tn.]ne miccerejges bdlj` sites witnij ebgn spegies, were testem usij` jlj-pbrbdetrig Arusabo-Pboois test clr jlj-

gljtijulus vbribhoes (i.e. judher lc e``s, judher nbtgnem, bjm judher emèm), bjm ujivbribte BJLSB clr tne ltnermbtb (i.e. e`` size, gljmitilj ijmex) tnbt were jlrdbooy mistrihutem. Pnej b si`jiflgbjt miccerejge wbs clujm bdlj` sites,Hljcerrlji plst-nlg tests were perclrdem clr hltn jlrdbo bjm jlj-gljtijulus vbribhoes, ij lrmer tl imejticy tne pbirslc sites tnbt gljtrihutem tl tne miccerejge.

XEWUO]W

]ne hreemij` prlmugtivity (]bhoe I) snlwem cew si`jiflgbjt miccerejges bdlj` brebs witnij

b spegies, ij tne cloolwij` gbses. Ij tne Gbttoe E`ret, hltn tne judher nbtgnem, bjm tne judheremèm miccerem (Arusabo-Pboois, 884 2=.7, V 4 6.62, bjm 8

84 22.9, V 4 6.67, respegtiveoy)bjm pbrtiguobroy tney were olwer ij Duotbj tnbj bt ]bujsb bjm Vlybj`. Ij tne Oittoe E`ret, bootne vbribhoes miccerem bdlj` brebs, (judher lc e``s; 7

8 4 =>.7, V 4 6.66621 judher nbtgnem;8

84 2=.5, V 4 6.6621 judher emèm; 784 2>.9, V 4 6.6662). ]nese miccerejges clr Oittoe

E`ret were mue tl b dugn olwer prlmugtivity lc tne mbra dlrpn ij Abrbgni tnbj bt tne twl sitesij Gnijb (plst-nlg tests). Ij èjerbo, tnere wbs b gljstbjt tejmejgy clr tne hreemij` prlmugtiv-ity tl he ni`ner, bjm sidiobr, bt tne twl Gnijese sites, Vlybj` bjm ]bi, tnbj bt tne tnree Vbaistbjisites.


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708

]BHOEI

E`` bjm jestoij` prlmugtilj, bs judher lc jests reglrmem clr tne flve e`ret bjm nerlj spegies bt tne six stumy brebs.

Oittoe E`rets bt Abrbgni heoljèm tl tne mbra dlrpn. ]ne flùres tnbt miccer si`jiflgbjtoy (V?.62) crld tne vboues clr

tne ltner brebs witnij tne sbde spegies, bre dbraem hy *

Judher / jest2 8 7 = 5 3 0 Debj

E``sGbttoe E`ret Vlybj` 6 6 = 7 7 6 6 7.96

Duotbj 6 2 85 == 26 7 6 7.>0

]lujsb 6 2 76 83 3 2 6 7.37

Obnlre 2 28 55 97 77 5 2 7.>8

Ijterdemibte E`ret Vlybj` 6 2 0 3 5 6 6 7.09

]lujsb 6 2 3 2 6 6 6 7.66

Oittoe E`ret Vlybj` 6 2 = 2> 77 0 6 =.35

]bi 6 2 3 25 73 85 8 =.99]lujsb 6 6 7 2 8 6 6 7.>7

Abrbgni 6 7 26 20 7 6 6 * 7.32

Gnijese Vljm Nerlj Vlybj` 6 6 6 2 6 2 6 5.66

]bi 6 6 6 8 7 2 6 =.>7

Hobga-grlwjem Ji`nt Vlybj` 6 6 9 = 2 6 6 7.=7

Nerlj ]bi 6 20 223 9= 0 6 2 7.=6

NbtgnemGbttoe E`ret Vlybj` 6 8 8 = 2 6 6 7.==

Duotbj 0 7= 82 = 2 6 6 * 8.70

]lujsb 2 28 2= 9 6 6 6 8.>3

Obnlre 9 86 88 3 6 6 6 8.==Ijterdemibte E`ret Vlybj` 6 7 > > 2 6 6 7.75

]lujsb 8 2 8 6 6 6 6 8.66

Oittoe E`ret Vlybj` 6 5 0 88 8= 6 6 =.28

]lujsb 6 2 6 6 2 6 6 7.56

Abrbgni 8 = 3 5 2 6 6 * 8.9=


Hobga-grlwjem Ji`nt

Nerlj

Vlybj` 6 6 28 7 6 6 6 7.86

CoemèmGbttoe E`ret Vlybj` 6 8 2 8 2 6 6 7.77

Duotbj 25 78 20 8 2 6 6 * 8.27

]lujsb = 22 26 9 6 6 6 8.02

Obnlre 22 88 86 3 6 6 6 8.73

Ijterdemibte E`ret Vlybj` 6 5 9 7 2 6 6 7.66

]lujsb 2 2 2 6 6 6 6 8.66

Oittoe E`ret Vlybj` 2 5 27 8= 22 6 6 7.08

]bi 6 5 25 27 83 > 2 =.76

]lujsb 6 2 2 6 6 6 6 8.56

Abrbgni 8 8 5 7 2 6 6 * 8.98


]bi 6 6 2 8 8 6 6 =.86Hobga-grlwjem Ji`nt Vlybj` 6 2 26 2 6 6 6 7.66

Nerlj ]bi 2= =2 07 7> 8 6 6 8.>=


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707

E`` size (]bhoe II) mim jlt miccer bdlj` sites witnij b spegies. ]ne gljmitilj ijmexlc tne gnigas, tnbt gluom he gldpbrem hetweej sites clr tne Oittoe E`ret ljoy, miccerem bdlj`sites (C 4 =0.8, m.c. 7, 835, V ? 6.6662), mue ljoy tl tne mbra dlrpn ij Abrbgni, wnere tnerbtil hetweej dbss bjm tbrsus oej`tn bverbèm 5.0, bjm wbs ni`ner tnbj bt tne ltner tnree sites,wnere tne bverbès rbjèm crld =.2 tl =.5.

]BHOEII

Oej`tn bjm wimtn lc e``s clr tne flve spegies ij tne six stumy brebs. Ebgn ejtry is; debj vboue (stbjmbrm mevibtilj,dijidud-dbxidud, sbdpoe size).

Wpegies Wtumy breb Oej`tn (dd) Pimtn (dd)

Gbttoe E`ret ]lujsb ==.5 (8.=, =6.9-=>.>, 26) 7=.2 (6.3, 78.9-7=.0, 26)Obnlre =7.3 (2.9, =2.6-56.6, 76) 77.5 (2.7, 78.6-73.6, 76)

Ijterdemibte E`ret Vlybj` =5.9 (6.=, =5.3-=3.7, 8) 77.8 (6.5, 78.0-77.0, 8)

Oittoe E`ret Vlybj` ==.2 (2.>, 7>.>-=>.3, 7>) 78.5 (6.>, 76.9-7=.2, 7>)]bi ==.5 (8.6, =6.3-=>.2, 27) 78.8 (2.2, 76.6-77.5, 27)]lujsb ==.= (6.9, =7.8-=5.0, =) 78.5 (6.0, 78.6-77.>, =)

Hobga-grlwjemJi`nt Nerlj

]bi =3.9 (8.9, 7>.7-58.7, =2) 77.9 (8.8, 85.7-7>.3, =2)

MIWGUWWILJ

]ne ni`ner hreemij` prlmugtivity lc Gbttoe bjm Oittoe E`rets bt tne twl Gnijese sites,Vlybj` bjm ]bi, tnbj ij Vbaistbj gluom he èjerbooy reobtem tl tne obrèr bvbiobhioity lc clr-bìj` nbhitbts ij Gnijb, wnere wet nbhitbts were very wimesprebm. Bgtubooy, tne prlmugtiv-ity lc Gbttoe E`ret, Oittoe E`ret, bjm Hobga-grlwjem Ji`nt Nerlj ij Gnijb wbs bt tne tlpejm lc tne rbjè lc vboues reglrmem ij vbrilus pbrts lc tneir rbjè, wnioe tneir prlmugtivityij Vbaistbj rbjèm bdlj` debj vboues (review hy Xuhloiji & Cbslob, ij press). ]nis ni`n

prlmugtivity su`èsts tnbt jl bmverse bèjt (e.`. gljtbdijbjts, Hljgldpb`ji et bo.,86671Wbjperb et bo.,8667) bccegtem reprlmugtilj, bjm tnbt hreemij` gljmitiljs (e.`. cllm bvbio-bhioity) bre very `llm.

]ne mbra dlrpn lc tne Oittoe E`ret ij Abrbgni miccerem crld tne ltner, wnite dlrpns westumiem, ij gniga gljmitilj, bjm hreemij` prlmugtivity, botnlu`n slde miccerejges gluom he

mue tl nbhitbt sijge tne mbra dlrpn wbs tne ljoy glbstbo plpuobtilj we stumiem.

BGAJLPOEM@DEJ]W

]nis stumy wbs gljmugtem witnij tne resebrgn prlkegt Glbstbo bjm ijobjm wetobjms ij Gnijb bjm Vbaistbj; gloljibowbterhirms bs hilijmigbtlrs lc plooutbjt oeveos bjm eccegts cujmem hy tne Eurlpebj Ujilj (IG2>-G]9>-689=). Pe wisntl tnbja boo tne gloobhlrbtlrs tl tne fleomwlra, bjm pbrtiguobroy Udbr Hbolgn, Oibl Yij`gnuj bjm @iuseppe @bumejzi .


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XECEXEJGEW

BWNAEJB[I, W. & YLD-]LS, Y. (2990). ]ne hreemij` hilol`y lc tne Hobga-grlwjem Ji`nt-nerlj (Jygtiglrbx jygtigl-rbx) bjm tne Oittoe E`ret (E`rettb brzettb) bt tne Nuoen Jbture Xeserve, Isrbeo.K. [llo., 8=8; 387-3=2.

HB^]EX, @.W. (299=). ]ne ijuejge lc syjgnrljlus hreemij`, jbtbo tree plsitilj bjm rbijcboo lj e`ret jestij` suggess.Gloljibo Pbterhirms,20; 286-289.

HEJJE]]W, X.E., CBWLOB, D., NBCJEX, N. & ABYWEX, Y. (8666). Ijuejge lc ejvirljdejtbo bjm mejsity-mepejmejt

cbgtlrs lj reprlmugtilj lc Oittoe E`rets.Bua,220; 37=-379.HLJGLDVB@JI, E., DUNBDDBM, B., KBHEEJ, X., LXSIJI, E., @BJMIJI, G., WBJVEXB, G., XUI[, ^. & CBWLOB, D.

(8667). E`rets bs dljitlrs lc trbge-detbo gljtbdijbtilj ij wetobjms lc Vbaistbj. Brgn. Ejvir. Gljtbdij.]lxiglo., =5; 799-=63.

CBWLOB, D. (299>). Lptidbo goutgn size ij nerljs; lhservbtiljbo bjm experidejtbo bpprlbgnes. Etnlo. Eglo. Evlo.,26; 77-=3.

CBWLOB, D., [NBJ@, Y., [NBL, M., MLJ@, Y. & PBJ@, N. (8662). B`e-bsslrtbtive dbtij` reobtem tl reprlmugtivesuggess ij Hobga-grlwjem Ji`nt Nerljs. Pbterhirms,8=; 808-803.

CBWLOB, D., @BOEL]]I, V., MLJ@, Y., MBI, J. & [NBJ@, Y. (866=). Obr`e judhers lc hreemij` e`rets bjm nerljsij Gnijb. Pbterhirms,80; 283-28>.

NBCJEX, N., CBWLOB, D., SLIWIJ, G. & ABIWEX, Y. (8668). E`rettb brzettb Oittoe E`ret.HPV Upmbte. ]ne Klurjbolc Hirms lc tne Pesterj Vboebrgtig,2; 2-29.

NIOBOUMMIJ, K., WNBNJ. & WNBPO, ].B. (8667). Jest site seoegtilj bjm hreemij` suggess hy Gbttoe E`ret bjm Oittoe

E`ret ij Bdrlnb, Uttbr Vrbmesn, Ijmib. Pbterhirms,83; ===-==>.KBGLH, E.D., DBXWNBOO, W.M. & UE][, @.P. (2993). Estidbtij` fltjess; b gldpbrislj lc hlmy gljmitilj ijmiges.

Lials,00; 32-30.OI[NI, [., YUKUJ, W. & YLJ@, D.(299>). Hreemij` hilol`y lc tne tnree nerljs ij [ipej` dlujtbijs. Gnijese Klurjbo

lc [llol`y(Gnijb), 7; 7=-70. (ij Gnijese)OLJ@WNEJ@, [., [ULDL, O. & CEJ@, [. (299=). Hreemij` eglol`y bjm hilol`y lc clur spegies lc e`rets bjm nerljs.

Bgt. Eglo. Wijigb, 2=; >6->8. (ij Gnijese)DBMMLGA, D. & HB^]EX, @. W. (2992). Hreemij` suggess lc e`rets reobtem tl rbijcboo; b six-yebr Bustrboibj stumy.

Gloljibo Pbterhirms, 2=; 277-279.DLXEO, @. & DLXEO, D.-Y. (2932). Uje nrljjire dixte sur oe Hbs-Wj`bo.Bobumb,89; 99-220.XUHLOIJI, M. & CBWLOB, D. (ij press). @el`rbpnig pbtterjs ij reprlmugtive pbrbdeters bdlj` jebrgtig nerljs.

Bua.WBJVEXB, G., XUI[, ^., KLSEX, O., OOLXEJ]E, @., KBHEEJ, X., DUNBDDBM, B., HLJGLDVB@JI, E. & CBWLOB,

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