N Jb Geol Palaont Abh 2602165-171published online March 2011

Article

Synopsis of fossil decapod crustaceans from Slovakia(Western Carpathians)

Matus HyznyWith 1 figure and 1 table

HYZNY M (2011) Synopsis of fossil decapod crustaceans from Slovakia (Western Carpathians) - Nlb Geol Paliiont Abh 260 165-171 Stuttgart

Abstract In the past only very few works dealt with fossil decapod crustaceans from Slovakia Thesystematic research of this group has never been fully conducted there This contribution provides ashort review of all known Mesozoic and Cenozoic fossil decapod occurrences So far most of the taxawere described from the Eocene and Miocene strata

Key words Decapod crustaceans Slovakia Mesozoic Eocene Miocene

1 Introduction

The presence of fossil decapod crustaceans from thearea which is today Slovakia is known since 19thcentury (REuss 1859) However until very recentlyonly little attention has been paid to the systematicresearch of this arthropod group in this area Thus ithas virtually no history in Slovakia except the scatte-red mentions of the occurrences of fossil decapods inthe literature which have rather informative character(eg PAPsovA1970 1975 1977 1978 PAPsovAin GROSSet al 1973 FILO amp SIRANOvA1996 KREMPASKA1998)The only systematic works dealing partly with fossildecapod crustaceans from todays Slovakia are thoseby REuss (1859) DORNYAY(1913) and LORENTHEYampBEURLEN(1929) Several occurrences are also menti-oned in a palaeontological textbook by HousA in SPI-NARet al (1966)

The aim of this contribution is to provide a list ofall occurrences of fossil decapod crustaceans fromSlovakia known to date However it should be con-sidered as the preliminary one as the research is stillongomg

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2 Studied materialMaterial shortly reviewed here is deposited in several insti-tutions and private collections in Slovakia namely Prfrodo-vedne muzeum SNM (Bratislava) Muzeum Spisa (SpisskliNova Yes) Liptovske muzeum - Cierny Orol (LiptovskyMikulas) and Department of Geology and PalaeontologyComenius University (Bratislava)

The material can be divided in three groups 1) Taxacollected and described personally by myself 2) taxa depo-sited in the museums or other institutions and examined bymyself and 3) taxa mentioned in the literature only and stillwaiting for the re-examination Fossil decapod crustaceansare known from several formations mainly of Cenozoic ageUp to date over 20 genera in 17 families including 6 newspecies have been identified (see Table 1)

Still there is yet undescribed material from several 10-calities of the middle Miocene age known to me and pre-vious studies on many geological units were simply notfocused on fossil decapods Although a great collectingbias should be considered when making further conclusionsfrom the presented results in some cases the studied mate-rial provides enough data for considering palaeoecologicaland palaeobiogeographical implications (HYZNY amp SCHLOGLin press)

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166 M Hyzny

Mesozoic From Slovakia there are very few Mesozo-ic occurrences of decapods known This may in partbe attributed to collection failure

The oldest decapod remain from Slovakia is oneleft propodus of a presumed paguroid anomuran fromthe Pliensbachian of the Bela Unit (MAHEr 1959) If itsaffinity was confirmed it would be one of the oldestfossil representatives of that group

From the middle Oxfordian of the Bohunice For-mation (MISiK et ai 1994) several specimens assign-able to decapod crustaceans come from (HYZNY ampSCHLOGL2009) All material originates from a sin-gle layer of white ammonite coquina (Gregorycerastransversarium Zone) at the locality Stepnicka skala(AUBRECHTet ai 2002 AUBRECHTamp JAMRICHovA2009)It comprises one almost complete internal mould ofa dorsal carapace of Tanidromites insignis (v MEYER1857) three propodi of indeterminate anomurans orbrachyurans and further unidentifiable crustaceanfragments The material of Tanidromites SCHWEITZERamp FELDMANN2008 is the only occurrence of a Jurassiccrab reported from the Pieniny Klippen Belt and fromSlovakia as well

Paleogene The Borove Formation (GROSSet ai 1984)of the Subtatric Group is known of several brychyurangenera originating from several localities of middle andupper Eocene age

Decapods from middle Eocene strata of the BoroveFm of Liptovska kotlina Depression have been knownsince the work of DORNYAY(1913) who reported thepresence of Harpactoxanthopsis quadrilobata (DEs-MAREST1822) at the locality Mnich near the town ofRuzomberok (Rozsahegy) The presence of the specieswas confirmed by LORENTHEYamp BEURLEN(1929) andPAPsovA (1970) LORENTHEYamp BEURLEN(1929) men-tioned from the same area also Raninoides fabianii(LORENTHEYin LORENTHEYamp BEURLEN1929) and Har-pactocarcinus punctulatus A MILNE-EDWARDS1862

From the upper Eocene strata of the Ruzomberokarea DORNYAY(1913) reported Lophoranina bittneri(LORENTHEY1902) and L reussi (WOODWARD1866)However the re-examination of the material depositedin the Liptovske Muzeum in Liptovsky Mikulas revea-led the presence of L reussi only (HYZNY2010)

Several brachyuran genera have been identifiedin the upper Eocene Tomasovce Member (FILO ampSIRANovA1996) of the Borove Fm The first mentionis that by REUSS(1859) who described Ranina hazslin-skyi from Radacs (todays Radatice eastern Slovakia)on the basis of a rather poorly preserved specimen (see

REUSS1859 pI 4 figs 4-5 compare with the commentsby LORENTHEYamp BEURLEN1929 12 109) The place ofrepository of the specimen is unknown LORENTHEYampBEURLEN(1929 33) noted the late Oligocene age of thespecimen however the more recent geological surveyof the area identified the surrounding of the locality asa part of the exposed Tomasovce Mb and yielded thePriabonian - early Oligocene age (predominantly latestPriabonian) (FILO amp SIRANovA1996) Remains of fos-sil crabs of Tomasovce Mb exposed in the same area(locality Sedlice) are mentioned by PAPsovA in GROSSet ai (1973)

From Durkovec quarry (Hornadska kotlina Dep-ression northeastern Slovakia) the stratotype loca-lity of the Tomasovce Mb quantitatively rich deca-pod fauna has been collected (HYZNY2007) Severalspecies have been identified Ranina sp (presumablya new species) Calappilia tridentata (BEURLEN1939)(Fig ID) and Coeloma vigil A MILNE-EDWARDS1865(Fig IB) At the same locality also heavily bioturba-ted horizons occur with very abundant Thalassinoidesburrows These horizons usually yield very few deca-pod body fossils (pers obs)

The presence of indeterminate decapod remainsat several other localities of presumed middle to lateEocene age of Liptovska kotlina Depression were re-ported by PAPsovA (1970 1975 1978) Similarly de-capods were reported from borehole cores NB-l andS2-NB of the Paleogene basement of the Danube Ba-sin and the regions of Banska Bystrica and Brezno(PAPsovA 1977)

Neogene In Slovakia there are several depositionalbasins filled with sediments of Miocene age formingonce parts of Central Paratethys

The lower Miocene (upper Karpatian) sediments ofLaksarska Nova Ves Fm of the Vienna Basin yieldedup to now six species (HYZNYamp SCHLOGLin press) Cal-lianopsis sp nov Crosniera sp nov Agononida spnov Munidopsis sp nov Mursia sp nov and Styri-oplax exiguus (GLAESSNER1928) The representativesof Crosniera KENSLEYamp HEARD1991 and AgononidaBABAamp DESAINTLAURENT1996 are the first membersof these genera known from the fossil record The wholedecapod association is dominated by ctenochelid Cal-lianopsis sp nov representing more than 90 of morethan 120 studied specimens The association indicatesdeep-water conditions (HYZNYamp SCHLOGLin press)

In the middle Miocene (lower Badenian) depositsof the Plastovce Mb (VASS 1971) of the SebechlebyFm of the Novohrad Basin (southern Slovakia) various

Synopsis of fossil decapod crustaceans from Slovakia 167

Fig 1 Some Cenozoic decapod crustaceans from Slovakia A - Szaboa inermis (BROCCHI1883) inner mould of rightcheliped consisting of carpus propodus and dactylus Studienka Fm Sandberg B - Coeloma vigil A MILNE-EDWARDS1865dorsal carapace preserved in the sandstone nodule Borove Fm Hlinisko C - Tasadia carniolica (BITINER1884) imprintof dorsal carapace Sebechleby Fm PhiStovce whitened with ammonium chloride D - Calappilia tridentata (BEURLEN1939) dorsal carapace Borove Fm Durkovec E - Calappa heberti BROCCHI1883 isolated left propodus StudienkaFm Sandberg F - Retropluma borealis FRAAlJEHANSENamp HANSEN2005 imprint of dorsal carapace Sebechleby FmPhiStovce- Scale bars equal 1 em

decapod species were identified coming from severallocalities around the PhiStovce village (SUKATCHEVAetal 2006) The most abundant are commonly completeindividuals of Tasadia carniolica (BITTNER1884) (FigIe) Less frequent are Jaxea kuemeli BACHMAYER1954(see HYZNY20ll this volume) and Retropluma borealisFRAAuE HANSENamp HANSEN2005 (Fig IF) The lat-ter has been previously only reported from the upperMiocene of Gram Denmark (FRAAuEet al 2005)

The middle Miocene (upper Badenian) strata ofthe Studienka Fm of the Vienna Basin in the vicin-ity of Bratislava represent several different facies withdifferent decapod associations The near-shore faciesexposed at Dubravska hlavica locality are represent-ed by hundreds of claws of Callianassa brocchiLORENTHEY1898 and Callianassa pseudorakosensisLORENTHEYin LORENTHEYamp BEuRLEN1929 Togetherwith them many burrow structures are preserved inseveral occasions also containing the callianassid

body fossils (HYZNYin press) Both species are cur-rently under detailed revision (HYZNYin prep) Anoth-er facies exposed at the localities of Devln-Merice andSandberg contain remains of Calappa heberti BRoc-CHI 1883 (Fig IE) Szaboa inermis (BROCCHI1883)(Fig lA) Tasadia carniolica (BITTNER1884) and yetindeterminate brachyuran claws (presumably xanthidsand portunids) The presence of C heberti at Sand-berg was already mentioned by LORENTHEYamp BEuRLEN(1929) Finally a more basinal lithology exposed in theformer claypit in Devlnska Nova Ves (the locality isno more accessible) yielded abundant specimens of Tcarniolica and much less frequent Mursia sp and un-described galatheids The Tasadia claws were alreadymentioned and figured by TouLA (1900) and the pres-ence of crabs was later noted by TOMASOVYCH(1998)None of these works however provided a systematictreatment ofthis material The material from the local-ity is currently under description

168 M Hyzny

Decapods from the localities where upper Badeni-an strata are exposed show similarities to coeval verywell known decapod associations described from theBudapest area (MOLLER 1984)

The upper Miocene strata yielded up to now onlyfew decapod remains The strata of the Stretava Fmof the early Sarmatian (Serravallian) age of Zdana andSlanska Huta (eastern Slovakia) are known of some re-mains of presumably freshwater crabs (KOROSSY 1940ZLINSKA amp FORDINAL 1995) They were described asPotamon hungaricum KOROSSY 1940 however KLAUSamp GROSS (2010) recently cast doubts on their potamidaffinities

True potamids were identified in upper Pannon-ian (Tortonian) strata of the Beladice Fm in boreholePiD-l in the vicinity of the Ordany village (FORDINAL1994 FORDINAL amp NAGY 1997) and recently assignedto Potamon hegauense KLAUS amp GROSS 2010 (KLAUSamp GROSS 2010)

From middle Pleistocene strata of the Mogyorosquarry which is today in Slovakia (Muzla southernSlovakia) KLAUS amp GROSS (2010 13) reported thepresence of Potamon antiquum SZOMBATHY1916

Acknowledgements

Material presented here was donated by KLEMENTFORDlNAL(Statny Geologicky Ustav Dionyza Stura Bratislava Slova-kia) MARIANGOLEJ(Geological Institute of Slovak Academyof Sciences Bratislava Slovakia) ADRIANHARNICAR(SvitSlovakia) PETER LEDVAK(Geological Institute of SlovakAcademy of Sciences Bratislava Slovakia) JAN SCHLOGL(Comenius University Bratislava Slovakia) STEFANSZALMA(PhiStovce Slovakia) and PETERVRSANSKY(Geological Ins-titute of Slovak Academy of Sciences Bratislava Slovakia)RENEH B FRAAlJE(Oertijdmuseum De Groene Poort Box-tel the Netherlands) ADRIANHARNICARNATALIAHLAVATA-HUDAcKovA(Comenius University Bratislava Slovakia) andJAN SCHLOGLhelped with the fieldworks Access to musealcollections was provided by BARBARACHALUPovA(Prfrodo-vedne muzeum SNM Bratislava Slovakia) PAVOLHRIADEL(Liptovske muzeum - Cierny Orol Liptovsky Mikulas Slo-vakia) PETER KLEPSATEL(formerly Prfrodovedne muzeumSNM Bratislava Slovakia) and ZUZANAKREMPASKA(Muze-um Spisa Spisska Nova Yes Slovakia) ANTONIODE ANGELI(Museo Civico G Zannato Montechio Maggiore Italy)RODNEYM FELDMANN(Kent State University Ohio USA)RENE H B FRAAlJEALESSANDROGARASSINO(Mus eo Civicodi Storian Naturale Milano Italy) and CARRIEE SCHWEIT-ZER (Kent State University Ohio USA) discussed the affi-nities of fossil material GONTERSCHWEIGERT(Natural His-tory Museum Stuttgart Germany) acted as a referee Sin-cere thanks to all of these individuals The work has beensupported by research grants APVV 0280-07 to DANIELAREHMovA (Comenius University Bratislava Slovakia) andgrant PRIF UK 192010 to MATOSHYZNY

References

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BROCCHIP (1883) Notes sur les Crustaces fossiles desterres tertiaires de la Hongrie - Annales des SciencesGeologiques 14 1-8

DESMARESTAG (1822) Les Crustaces Proprement Dits -In BRONGNIARTA amp DESMARESTA-G (Eds) HistoireNaturelle des Crustaces Fossiles sous les Rapports Zoo-logiques et Geologiques 67-154 Paris (Levrault)

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FORDlNALK (1994) Vrchny panon (zona H) vychodnehookraja Povazskeho Inovca - Geologicke prace Spravy99 67-75

FORDlNALK amp NAGYA (1997) Hlavinske vrstvy - okra-jove vrchnopanonske sedimenty risiiovskej priehlbiny- Mineralia Slovaca 29 401-406

FRAAlJER HANSEN1 amp HANSENT (2005) Late Miocenedecapod faunas from Gram Denmark - Palaeontos 751-61

GLAESSNERME (1928) Die Dekapodenfauna des oster-reichischen Jungtertiiirs - Jahrbuch der GeologischenBundesanstalt78 161-219

GROSS P KOHLER E amp SAMUEL0 (1984) Litostrati-graficka klasifikacia vnUtrokarpatskeho paleogennehosedimentacneho cyklu - Geologicke prace Spravy 81103-107

HYZNYM (2007) Paleogene crab fauna of Borove Forma-tion (localities Durkovec and Hlinisko) Western Car-pathians Slovakia - In GARASSINOA FELDMANNRM amp TERUZZIG (Eds) 3rd Symposium on Mesozoicand Cenozoic Decapod Crustaceans Museo di StoriaNaturale di Milano - Memorie della Societa Italiana diScienze Naturali e del Museo Civico di Storia Naturaledi Milano 35 (2) 59-61

HYZNYM (in press) In situ mud shrimps (Decapoda Axi-idea Callianassidae) preserved within their burrowsfrom the middle Miocene of the Central Paratethys -Bulletin of the Mizunami Fossil Museum 37

HYZNYM (2010) Revision of the Eocene decapod crusta-ceans deposited in the Liptovske muzeum Cierny Orolin Liptovsky Mikulas (Slovakia) - Acta Geologica Slo-vaca 2 (2) 117-122

HYZNYM (2011) Revision of Jaxea kuemeli BACHMAYER1954 (Decapoda Gebiidea Laomediidae) from the Mi-ocene of Europe with remarks on the palaeobiogeog-raphy of the genus Jaxea NARDO1847 - Neues Jahr-buch fUr Geologie und Paliiontologie - Abhandlungenthis volume

HYZNYM amp SCHLOGL1 (2009) The first occurence ofProsopidae sl (Crustacea Decapoda Brachyura)and other findings of crustaceans in the Upper Jurassiclimestones of the Pieniny Klippen Belt - In KROBlC-KI M (Ed) Jurassica VIII Vrsatec 09-11102009 -Geologia 35 (31) 61-63

HYZNYM amp SCHLOGL1 (in press) An Early Miocene de-ep-water decapod crustacean faunule from the ViennaBasin (Western Carpathians Slovakia) - Palaeontology

KENSLEYB amp HEARDRW (1991) An examination of theshrimp family Callianideidae (Crustacea DecapodaThalassinidea) - Proceedings of the Biological Societyof Washington 104 493-537

KLAUSS amp GROSSM (2010) Synopsis of the fossil fresh-water crabs of Europe (Brachyura Potamoidea Potami-dae) - Neues Jahrbuch fUrGeologie und PaliiontologieAbhandlungen 256 39-59

KOROSSYL (1940) Kurze geologische Beschreibung derUmgebung von Hernadzsadany - F6ldtani K6z16ny7083-109 143-144

KREMPASKAZ (1998) Niektore vysledky geologickeho apaleontologickeho vyskumu v lome Durkovec v Sloven-skom raji - Natura Carpatica 39 27-32

LORENTHEYE (1898)Beitriige zur Decapodenfauna der un-garischen Tertiiirs - Termeszetrajzi Ftizetek 21 1-133

LORENTHEYE (1902) Neuere Beitriige zur Tertiiiren Deka-podenfauna Ungarns - Mathematische und Naturwis-senschaftliche Berichte aus Ungarn 18 98-120

LORENTHEYE amp BEURLENK (1929) Die fossilen Deka-poden der Liinder der Ungarischen Krone - GeologicaHungarica Series Palaeontologica 3 1-421

MAHEJM (1959) Nove Clenenie a pohlad na historicko-geologicky vyvoj mezozoika centralnych Karpat -Geologicke Prace 55 61-81

MEYERHv (1857) [Briefiiche Mittheilungen] - JahrbuchfUrMineralogie Geologie Geognosie und Petrefakten-Kunde 1857 556

Synopsis of fossil decapod crustaceans from Slovakia 169

MILNE-EDWARDSA (1862) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (4) 18 31-85

MILNE-EDWARDSA (1865) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (5) 3 297-351

MISIKM SIBLlKM SYKORAM amp AUBRECHTR (1994)Jurassic brachiopods and sedimentological study of theBabina Klippe near Bohunice (Czorsztyn Unit PieninyKlippen Belt) - Mineralia Slovaca 26 255-266

MOLLERP (1984) Decapod Crustacea of the Badenian -Geologica Hungarica Series Palaeontologica 42 3-317

GROSSP PAPsovAJ amp KOHLERE (1973) Sedimentol6giaa stratigrafia vrchnej casti bazalnej litofacie centralno-karpatskeho paleogenu od Sarisskej vrchoviny po Lip-tovsku kotlinu Dielcia zaverecna sprava - 57 pp Bra-tislava (GlmS)

PAPsovAJ (1970) Zaverecna sprava 0 makrofaune paleo-genu Liptovskej kotliny na liste Ruzomberok 1 50000 Ciastkova zaverecna sprava - 30 pp Bratislava(GlmS)

PAPsovAJ (1975) Makrofauna strednej casti Liptovskejkotliny na listoch Huty Demiinova Smrecany Lip-tovsky Hradok Ciastkova sprava - 41 pp Bratislava(GlmS)

PAPsovAJ (1977) Celkove zhodnotenie makrospolocens-tiev Zapadnych Karpat Ciastkova zaverecna sprava -111pp Bratislava (GlmS)

PAPsovA1 (1978) Makrofauna paleogenu vychodnej castiLiptovskej kotliny Ciastkova sprava - 56 pp Bratisla-va (GlmS)

REuss AE (1859)Zur Kenntnis fossiler Krabben - Denk-schriften der kaiserlichen Akademie der Wissenschaf-ten Wien 17 1-90

SCHWEITZERCE amp FELDMANNRM (2008 [imprint2007]) A new classification for some Jurassic Brachy-ura (Crustacea Decapoda Brachyura Homolodromio-idea) Families Goniodromitidae BEURLEN1932 andTanidromitidae new family - Senckenbergiana lethaea87 (2) 119-156

SUKATCHEVAID SZALMAS VRSANSKYP CHALUpovAB GOLEJM LABAJOvAE amp SMRECKovAM (2006)Caddis-fly (Insecta Trichoptera) from the Badenianvolcano-sedimentary succession (Western CarpathiansSlovakia) - Geologica Carpathica 57 (6) 531-534

SZOMBATHYK (1916) Die tertiiiren Formen der GattungPotamon (Telphusa) und ihre paliiarktischen Nachkom-men - Annales historico-naturales Musei NationalisHungarici 14 281-472SPlNARZ CEPEKP BOUCEKB EuAsovA-FRAJovAH HAVLlCEKV HORNYRHOUSAV KALABISV KUKALovA1 MAREKL NE-KVASILovA0 POKORNYV PROKOPR PRIBYLA Ru-ZICKAB SNAJDRM STEMPROKovAD amp ZIKMUNDovA1 (1966) Systematicka paleontologie bezobratlYch -1052 pp Praha (Academia)

TOMASOVYCHA (1998) Badenske miikkyse z tehelne De-vfnska Nova Yes (Bratislava Slovensko) - MineraliaSlovaca 30 357-386

Synopsis of fossil decapod crustaceans from Slovakia 167

Fig 1 Some Cenozoic decapod crustaceans from Slovakia A - Szaboa inermis (BROCCHI1883) inner mould of rightcheliped consisting of carpus propodus and dactylus Studienka Fm Sandberg B - Coeloma vigil A MILNE-EDWARDS1865dorsal carapace preserved in the sandstone nodule Borove Fm Hlinisko C - Tasadia carniolica (BITINER1884) imprintof dorsal carapace Sebechleby Fm PhiStovce whitened with ammonium chloride D - Calappilia tridentata (BEURLEN1939) dorsal carapace Borove Fm Durkovec E - Calappa heberti BROCCHI1883 isolated left propodus StudienkaFm Sandberg F - Retropluma borealis FRAAlJEHANSENamp HANSEN2005 imprint of dorsal carapace Sebechleby FmPhiStovce- Scale bars equal 1 em

decapod species were identified coming from severallocalities around the PhiStovce village (SUKATCHEVAetal 2006) The most abundant are commonly completeindividuals of Tasadia carniolica (BITTNER1884) (FigIe) Less frequent are Jaxea kuemeli BACHMAYER1954(see HYZNY20ll this volume) and Retropluma borealisFRAAuE HANSENamp HANSEN2005 (Fig IF) The lat-ter has been previously only reported from the upperMiocene of Gram Denmark (FRAAuEet al 2005)

The middle Miocene (upper Badenian) strata ofthe Studienka Fm of the Vienna Basin in the vicin-ity of Bratislava represent several different facies withdifferent decapod associations The near-shore faciesexposed at Dubravska hlavica locality are represent-ed by hundreds of claws of Callianassa brocchiLORENTHEY1898 and Callianassa pseudorakosensisLORENTHEYin LORENTHEYamp BEuRLEN1929 Togetherwith them many burrow structures are preserved inseveral occasions also containing the callianassid

body fossils (HYZNYin press) Both species are cur-rently under detailed revision (HYZNYin prep) Anoth-er facies exposed at the localities of Devln-Merice andSandberg contain remains of Calappa heberti BRoc-CHI 1883 (Fig IE) Szaboa inermis (BROCCHI1883)(Fig lA) Tasadia carniolica (BITTNER1884) and yetindeterminate brachyuran claws (presumably xanthidsand portunids) The presence of C heberti at Sand-berg was already mentioned by LORENTHEYamp BEuRLEN(1929) Finally a more basinal lithology exposed in theformer claypit in Devlnska Nova Ves (the locality isno more accessible) yielded abundant specimens of Tcarniolica and much less frequent Mursia sp and un-described galatheids The Tasadia claws were alreadymentioned and figured by TouLA (1900) and the pres-ence of crabs was later noted by TOMASOVYCH(1998)None of these works however provided a systematictreatment ofthis material The material from the local-ity is currently under description

168 M Hyzny

Decapods from the localities where upper Badeni-an strata are exposed show similarities to coeval verywell known decapod associations described from theBudapest area (MOLLER 1984)

The upper Miocene strata yielded up to now onlyfew decapod remains The strata of the Stretava Fmof the early Sarmatian (Serravallian) age of Zdana andSlanska Huta (eastern Slovakia) are known of some re-mains of presumably freshwater crabs (KOROSSY 1940ZLINSKA amp FORDINAL 1995) They were described asPotamon hungaricum KOROSSY 1940 however KLAUSamp GROSS (2010) recently cast doubts on their potamidaffinities

True potamids were identified in upper Pannon-ian (Tortonian) strata of the Beladice Fm in boreholePiD-l in the vicinity of the Ordany village (FORDINAL1994 FORDINAL amp NAGY 1997) and recently assignedto Potamon hegauense KLAUS amp GROSS 2010 (KLAUSamp GROSS 2010)

From middle Pleistocene strata of the Mogyorosquarry which is today in Slovakia (Muzla southernSlovakia) KLAUS amp GROSS (2010 13) reported thepresence of Potamon antiquum SZOMBATHY1916

Acknowledgements

Material presented here was donated by KLEMENTFORDlNAL(Statny Geologicky Ustav Dionyza Stura Bratislava Slova-kia) MARIANGOLEJ(Geological Institute of Slovak Academyof Sciences Bratislava Slovakia) ADRIANHARNICAR(SvitSlovakia) PETER LEDVAK(Geological Institute of SlovakAcademy of Sciences Bratislava Slovakia) JAN SCHLOGL(Comenius University Bratislava Slovakia) STEFANSZALMA(PhiStovce Slovakia) and PETERVRSANSKY(Geological Ins-titute of Slovak Academy of Sciences Bratislava Slovakia)RENEH B FRAAlJE(Oertijdmuseum De Groene Poort Box-tel the Netherlands) ADRIANHARNICARNATALIAHLAVATA-HUDAcKovA(Comenius University Bratislava Slovakia) andJAN SCHLOGLhelped with the fieldworks Access to musealcollections was provided by BARBARACHALUPovA(Prfrodo-vedne muzeum SNM Bratislava Slovakia) PAVOLHRIADEL(Liptovske muzeum - Cierny Orol Liptovsky Mikulas Slo-vakia) PETER KLEPSATEL(formerly Prfrodovedne muzeumSNM Bratislava Slovakia) and ZUZANAKREMPASKA(Muze-um Spisa Spisska Nova Yes Slovakia) ANTONIODE ANGELI(Museo Civico G Zannato Montechio Maggiore Italy)RODNEYM FELDMANN(Kent State University Ohio USA)RENE H B FRAAlJEALESSANDROGARASSINO(Mus eo Civicodi Storian Naturale Milano Italy) and CARRIEE SCHWEIT-ZER (Kent State University Ohio USA) discussed the affi-nities of fossil material GONTERSCHWEIGERT(Natural His-tory Museum Stuttgart Germany) acted as a referee Sin-cere thanks to all of these individuals The work has beensupported by research grants APVV 0280-07 to DANIELAREHMovA (Comenius University Bratislava Slovakia) andgrant PRIF UK 192010 to MATOSHYZNY

References

AUBRECHTR amp JAMRICHovAM (2009) Stepnicka skalaKlippe - unique type of the Czorsztyn Succession (Pie-niny Klipped Belt Western Carpathians) - Acta Geolo-gica Slovaca 1 (2) 141-158

AUBRECHTR KROBICKIM MATYJABA WIERZBOWSKIA amp SCHLOGLJ (2002) Jurassic stromatactis mud-mounds in the Pieniny Klippen Belt (Western Carpat-hians) - Petrography and Stratigraphy - In BucuR 11amp FILIPESCUS (Eds) Research advances in calcareousalgae and microbial carbonates (Proceedings of the 4thIFAA Regional Meeting Cluj-Napoca August 29-Sep-tember 5 2001) 1-16 Cluj (Cluj University Press)

BABAK amp DESAINTLAURENTM (1996) Crustacea Deca-poda Revision of the genus Bathymunida BALss 1914and description of six new related genera (Galatheidae)- In CROSNIERA (Ed) Resultats des Campagnes MU-SORSTOM Volume 15 - Memoires du Museum Nati-onal dHistoire Naturelle Paris 168 433-502

BACHMAYERE (1954) Zwei bemerkenswerte Crustaceen-Funde aus dem Jungtertiiir des Wiener Beckens - Sit-zungsberichte der Osterreichischen Akademie der Wis-senschaften in Wien mathematisch-naturwissenschaft-liche Klasse I 163 63-70

BEURLENK (1939) Neue Dekapoden-Krebse aus dem un-garischen Tertiiir - Paliiontologische Zeitschrift 21135-161

BITTNERA (1884) Beitriige zur Kenntnis tertiiirer Brachy-urenfaunen - Denkschriften der Kaiserlichen Akade-mie der Wissenschaften mathematisch-naturwissen-schaftliche Classe 48 15-30

BROCCHIP (1883) Notes sur les Crustaces fossiles desterres tertiaires de la Hongrie - Annales des SciencesGeologiques 14 1-8

DESMARESTAG (1822) Les Crustaces Proprement Dits -In BRONGNIARTA amp DESMARESTA-G (Eds) HistoireNaturelle des Crustaces Fossiles sous les Rapports Zoo-logiques et Geologiques 67-154 Paris (Levrault)

DORNYAYB (1913) Rozsahegy kornyekenek foldtani viszo-nyairol - 51 pp Budapest (E A Konyvnyomdaja)

FILO I amp SIRANovAZ (1996) Tomasovske vrstvy - novalitostratigraficka jednotka podtatranskej skupiny -Geologicke prace Spravy 102 41-49

FORDlNALK (1994) Vrchny panon (zona H) vychodnehookraja Povazskeho Inovca - Geologicke prace Spravy99 67-75

FORDlNALK amp NAGYA (1997) Hlavinske vrstvy - okra-jove vrchnopanonske sedimenty risiiovskej priehlbiny- Mineralia Slovaca 29 401-406

FRAAlJER HANSEN1 amp HANSENT (2005) Late Miocenedecapod faunas from Gram Denmark - Palaeontos 751-61

GLAESSNERME (1928) Die Dekapodenfauna des oster-reichischen Jungtertiiirs - Jahrbuch der GeologischenBundesanstalt78 161-219

GROSS P KOHLER E amp SAMUEL0 (1984) Litostrati-graficka klasifikacia vnUtrokarpatskeho paleogennehosedimentacneho cyklu - Geologicke prace Spravy 81103-107

HYZNYM (2007) Paleogene crab fauna of Borove Forma-tion (localities Durkovec and Hlinisko) Western Car-pathians Slovakia - In GARASSINOA FELDMANNRM amp TERUZZIG (Eds) 3rd Symposium on Mesozoicand Cenozoic Decapod Crustaceans Museo di StoriaNaturale di Milano - Memorie della Societa Italiana diScienze Naturali e del Museo Civico di Storia Naturaledi Milano 35 (2) 59-61

HYZNYM (in press) In situ mud shrimps (Decapoda Axi-idea Callianassidae) preserved within their burrowsfrom the middle Miocene of the Central Paratethys -Bulletin of the Mizunami Fossil Museum 37

HYZNYM (2010) Revision of the Eocene decapod crusta-ceans deposited in the Liptovske muzeum Cierny Orolin Liptovsky Mikulas (Slovakia) - Acta Geologica Slo-vaca 2 (2) 117-122

HYZNYM (2011) Revision of Jaxea kuemeli BACHMAYER1954 (Decapoda Gebiidea Laomediidae) from the Mi-ocene of Europe with remarks on the palaeobiogeog-raphy of the genus Jaxea NARDO1847 - Neues Jahr-buch fUr Geologie und Paliiontologie - Abhandlungenthis volume

HYZNYM amp SCHLOGL1 (2009) The first occurence ofProsopidae sl (Crustacea Decapoda Brachyura)and other findings of crustaceans in the Upper Jurassiclimestones of the Pieniny Klippen Belt - In KROBlC-KI M (Ed) Jurassica VIII Vrsatec 09-11102009 -Geologia 35 (31) 61-63

HYZNYM amp SCHLOGL1 (in press) An Early Miocene de-ep-water decapod crustacean faunule from the ViennaBasin (Western Carpathians Slovakia) - Palaeontology

KENSLEYB amp HEARDRW (1991) An examination of theshrimp family Callianideidae (Crustacea DecapodaThalassinidea) - Proceedings of the Biological Societyof Washington 104 493-537

KLAUSS amp GROSSM (2010) Synopsis of the fossil fresh-water crabs of Europe (Brachyura Potamoidea Potami-dae) - Neues Jahrbuch fUrGeologie und PaliiontologieAbhandlungen 256 39-59

KOROSSYL (1940) Kurze geologische Beschreibung derUmgebung von Hernadzsadany - F6ldtani K6z16ny7083-109 143-144

KREMPASKAZ (1998) Niektore vysledky geologickeho apaleontologickeho vyskumu v lome Durkovec v Sloven-skom raji - Natura Carpatica 39 27-32

LORENTHEYE (1898)Beitriige zur Decapodenfauna der un-garischen Tertiiirs - Termeszetrajzi Ftizetek 21 1-133

LORENTHEYE (1902) Neuere Beitriige zur Tertiiiren Deka-podenfauna Ungarns - Mathematische und Naturwis-senschaftliche Berichte aus Ungarn 18 98-120

LORENTHEYE amp BEURLENK (1929) Die fossilen Deka-poden der Liinder der Ungarischen Krone - GeologicaHungarica Series Palaeontologica 3 1-421

MAHEJM (1959) Nove Clenenie a pohlad na historicko-geologicky vyvoj mezozoika centralnych Karpat -Geologicke Prace 55 61-81

MEYERHv (1857) [Briefiiche Mittheilungen] - JahrbuchfUrMineralogie Geologie Geognosie und Petrefakten-Kunde 1857 556

Synopsis of fossil decapod crustaceans from Slovakia 169

MILNE-EDWARDSA (1862) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (4) 18 31-85

MILNE-EDWARDSA (1865) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (5) 3 297-351

MISIKM SIBLlKM SYKORAM amp AUBRECHTR (1994)Jurassic brachiopods and sedimentological study of theBabina Klippe near Bohunice (Czorsztyn Unit PieninyKlippen Belt) - Mineralia Slovaca 26 255-266

MOLLERP (1984) Decapod Crustacea of the Badenian -Geologica Hungarica Series Palaeontologica 42 3-317

GROSSP PAPsovAJ amp KOHLERE (1973) Sedimentol6giaa stratigrafia vrchnej casti bazalnej litofacie centralno-karpatskeho paleogenu od Sarisskej vrchoviny po Lip-tovsku kotlinu Dielcia zaverecna sprava - 57 pp Bra-tislava (GlmS)

PAPsovAJ (1970) Zaverecna sprava 0 makrofaune paleo-genu Liptovskej kotliny na liste Ruzomberok 1 50000 Ciastkova zaverecna sprava - 30 pp Bratislava(GlmS)

PAPsovAJ (1975) Makrofauna strednej casti Liptovskejkotliny na listoch Huty Demiinova Smrecany Lip-tovsky Hradok Ciastkova sprava - 41 pp Bratislava(GlmS)

PAPsovAJ (1977) Celkove zhodnotenie makrospolocens-tiev Zapadnych Karpat Ciastkova zaverecna sprava -111pp Bratislava (GlmS)

PAPsovA1 (1978) Makrofauna paleogenu vychodnej castiLiptovskej kotliny Ciastkova sprava - 56 pp Bratisla-va (GlmS)

REuss AE (1859)Zur Kenntnis fossiler Krabben - Denk-schriften der kaiserlichen Akademie der Wissenschaf-ten Wien 17 1-90

SCHWEITZERCE amp FELDMANNRM (2008 [imprint2007]) A new classification for some Jurassic Brachy-ura (Crustacea Decapoda Brachyura Homolodromio-idea) Families Goniodromitidae BEURLEN1932 andTanidromitidae new family - Senckenbergiana lethaea87 (2) 119-156

SUKATCHEVAID SZALMAS VRSANSKYP CHALUpovAB GOLEJM LABAJOvAE amp SMRECKovAM (2006)Caddis-fly (Insecta Trichoptera) from the Badenianvolcano-sedimentary succession (Western CarpathiansSlovakia) - Geologica Carpathica 57 (6) 531-534

SZOMBATHYK (1916) Die tertiiiren Formen der GattungPotamon (Telphusa) und ihre paliiarktischen Nachkom-men - Annales historico-naturales Musei NationalisHungarici 14 281-472SPlNARZ CEPEKP BOUCEKB EuAsovA-FRAJovAH HAVLlCEKV HORNYRHOUSAV KALABISV KUKALovA1 MAREKL NE-KVASILovA0 POKORNYV PROKOPR PRIBYLA Ru-ZICKAB SNAJDRM STEMPROKovAD amp ZIKMUNDovA1 (1966) Systematicka paleontologie bezobratlYch -1052 pp Praha (Academia)

TOMASOVYCHA (1998) Badenske miikkyse z tehelne De-vfnska Nova Yes (Bratislava Slovensko) - MineraliaSlovaca 30 357-386

168 M Hyzny

Decapods from the localities where upper Badeni-an strata are exposed show similarities to coeval verywell known decapod associations described from theBudapest area (MOLLER 1984)

The upper Miocene strata yielded up to now onlyfew decapod remains The strata of the Stretava Fmof the early Sarmatian (Serravallian) age of Zdana andSlanska Huta (eastern Slovakia) are known of some re-mains of presumably freshwater crabs (KOROSSY 1940ZLINSKA amp FORDINAL 1995) They were described asPotamon hungaricum KOROSSY 1940 however KLAUSamp GROSS (2010) recently cast doubts on their potamidaffinities

True potamids were identified in upper Pannon-ian (Tortonian) strata of the Beladice Fm in boreholePiD-l in the vicinity of the Ordany village (FORDINAL1994 FORDINAL amp NAGY 1997) and recently assignedto Potamon hegauense KLAUS amp GROSS 2010 (KLAUSamp GROSS 2010)

From middle Pleistocene strata of the Mogyorosquarry which is today in Slovakia (Muzla southernSlovakia) KLAUS amp GROSS (2010 13) reported thepresence of Potamon antiquum SZOMBATHY1916

Acknowledgements

Material presented here was donated by KLEMENTFORDlNAL(Statny Geologicky Ustav Dionyza Stura Bratislava Slova-kia) MARIANGOLEJ(Geological Institute of Slovak Academyof Sciences Bratislava Slovakia) ADRIANHARNICAR(SvitSlovakia) PETER LEDVAK(Geological Institute of SlovakAcademy of Sciences Bratislava Slovakia) JAN SCHLOGL(Comenius University Bratislava Slovakia) STEFANSZALMA(PhiStovce Slovakia) and PETERVRSANSKY(Geological Ins-titute of Slovak Academy of Sciences Bratislava Slovakia)RENEH B FRAAlJE(Oertijdmuseum De Groene Poort Box-tel the Netherlands) ADRIANHARNICARNATALIAHLAVATA-HUDAcKovA(Comenius University Bratislava Slovakia) andJAN SCHLOGLhelped with the fieldworks Access to musealcollections was provided by BARBARACHALUPovA(Prfrodo-vedne muzeum SNM Bratislava Slovakia) PAVOLHRIADEL(Liptovske muzeum - Cierny Orol Liptovsky Mikulas Slo-vakia) PETER KLEPSATEL(formerly Prfrodovedne muzeumSNM Bratislava Slovakia) and ZUZANAKREMPASKA(Muze-um Spisa Spisska Nova Yes Slovakia) ANTONIODE ANGELI(Museo Civico G Zannato Montechio Maggiore Italy)RODNEYM FELDMANN(Kent State University Ohio USA)RENE H B FRAAlJEALESSANDROGARASSINO(Mus eo Civicodi Storian Naturale Milano Italy) and CARRIEE SCHWEIT-ZER (Kent State University Ohio USA) discussed the affi-nities of fossil material GONTERSCHWEIGERT(Natural His-tory Museum Stuttgart Germany) acted as a referee Sin-cere thanks to all of these individuals The work has beensupported by research grants APVV 0280-07 to DANIELAREHMovA (Comenius University Bratislava Slovakia) andgrant PRIF UK 192010 to MATOSHYZNY

References

AUBRECHTR amp JAMRICHovAM (2009) Stepnicka skalaKlippe - unique type of the Czorsztyn Succession (Pie-niny Klipped Belt Western Carpathians) - Acta Geolo-gica Slovaca 1 (2) 141-158

AUBRECHTR KROBICKIM MATYJABA WIERZBOWSKIA amp SCHLOGLJ (2002) Jurassic stromatactis mud-mounds in the Pieniny Klippen Belt (Western Carpat-hians) - Petrography and Stratigraphy - In BucuR 11amp FILIPESCUS (Eds) Research advances in calcareousalgae and microbial carbonates (Proceedings of the 4thIFAA Regional Meeting Cluj-Napoca August 29-Sep-tember 5 2001) 1-16 Cluj (Cluj University Press)

BABAK amp DESAINTLAURENTM (1996) Crustacea Deca-poda Revision of the genus Bathymunida BALss 1914and description of six new related genera (Galatheidae)- In CROSNIERA (Ed) Resultats des Campagnes MU-SORSTOM Volume 15 - Memoires du Museum Nati-onal dHistoire Naturelle Paris 168 433-502

BACHMAYERE (1954) Zwei bemerkenswerte Crustaceen-Funde aus dem Jungtertiiir des Wiener Beckens - Sit-zungsberichte der Osterreichischen Akademie der Wis-senschaften in Wien mathematisch-naturwissenschaft-liche Klasse I 163 63-70

BEURLENK (1939) Neue Dekapoden-Krebse aus dem un-garischen Tertiiir - Paliiontologische Zeitschrift 21135-161

BITTNERA (1884) Beitriige zur Kenntnis tertiiirer Brachy-urenfaunen - Denkschriften der Kaiserlichen Akade-mie der Wissenschaften mathematisch-naturwissen-schaftliche Classe 48 15-30

BROCCHIP (1883) Notes sur les Crustaces fossiles desterres tertiaires de la Hongrie - Annales des SciencesGeologiques 14 1-8

DESMARESTAG (1822) Les Crustaces Proprement Dits -In BRONGNIARTA amp DESMARESTA-G (Eds) HistoireNaturelle des Crustaces Fossiles sous les Rapports Zoo-logiques et Geologiques 67-154 Paris (Levrault)

DORNYAYB (1913) Rozsahegy kornyekenek foldtani viszo-nyairol - 51 pp Budapest (E A Konyvnyomdaja)

FILO I amp SIRANovAZ (1996) Tomasovske vrstvy - novalitostratigraficka jednotka podtatranskej skupiny -Geologicke prace Spravy 102 41-49

FORDlNALK (1994) Vrchny panon (zona H) vychodnehookraja Povazskeho Inovca - Geologicke prace Spravy99 67-75

FORDlNALK amp NAGYA (1997) Hlavinske vrstvy - okra-jove vrchnopanonske sedimenty risiiovskej priehlbiny- Mineralia Slovaca 29 401-406

FRAAlJER HANSEN1 amp HANSENT (2005) Late Miocenedecapod faunas from Gram Denmark - Palaeontos 751-61

GLAESSNERME (1928) Die Dekapodenfauna des oster-reichischen Jungtertiiirs - Jahrbuch der GeologischenBundesanstalt78 161-219

GROSS P KOHLER E amp SAMUEL0 (1984) Litostrati-graficka klasifikacia vnUtrokarpatskeho paleogennehosedimentacneho cyklu - Geologicke prace Spravy 81103-107

HYZNYM (2007) Paleogene crab fauna of Borove Forma-tion (localities Durkovec and Hlinisko) Western Car-pathians Slovakia - In GARASSINOA FELDMANNRM amp TERUZZIG (Eds) 3rd Symposium on Mesozoicand Cenozoic Decapod Crustaceans Museo di StoriaNaturale di Milano - Memorie della Societa Italiana diScienze Naturali e del Museo Civico di Storia Naturaledi Milano 35 (2) 59-61

HYZNYM (in press) In situ mud shrimps (Decapoda Axi-idea Callianassidae) preserved within their burrowsfrom the middle Miocene of the Central Paratethys -Bulletin of the Mizunami Fossil Museum 37

HYZNYM (2010) Revision of the Eocene decapod crusta-ceans deposited in the Liptovske muzeum Cierny Orolin Liptovsky Mikulas (Slovakia) - Acta Geologica Slo-vaca 2 (2) 117-122

HYZNYM (2011) Revision of Jaxea kuemeli BACHMAYER1954 (Decapoda Gebiidea Laomediidae) from the Mi-ocene of Europe with remarks on the palaeobiogeog-raphy of the genus Jaxea NARDO1847 - Neues Jahr-buch fUr Geologie und Paliiontologie - Abhandlungenthis volume

HYZNYM amp SCHLOGL1 (2009) The first occurence ofProsopidae sl (Crustacea Decapoda Brachyura)and other findings of crustaceans in the Upper Jurassiclimestones of the Pieniny Klippen Belt - In KROBlC-KI M (Ed) Jurassica VIII Vrsatec 09-11102009 -Geologia 35 (31) 61-63

HYZNYM amp SCHLOGL1 (in press) An Early Miocene de-ep-water decapod crustacean faunule from the ViennaBasin (Western Carpathians Slovakia) - Palaeontology

KENSLEYB amp HEARDRW (1991) An examination of theshrimp family Callianideidae (Crustacea DecapodaThalassinidea) - Proceedings of the Biological Societyof Washington 104 493-537

KLAUSS amp GROSSM (2010) Synopsis of the fossil fresh-water crabs of Europe (Brachyura Potamoidea Potami-dae) - Neues Jahrbuch fUrGeologie und PaliiontologieAbhandlungen 256 39-59

KOROSSYL (1940) Kurze geologische Beschreibung derUmgebung von Hernadzsadany - F6ldtani K6z16ny7083-109 143-144

KREMPASKAZ (1998) Niektore vysledky geologickeho apaleontologickeho vyskumu v lome Durkovec v Sloven-skom raji - Natura Carpatica 39 27-32

LORENTHEYE (1898)Beitriige zur Decapodenfauna der un-garischen Tertiiirs - Termeszetrajzi Ftizetek 21 1-133

LORENTHEYE (1902) Neuere Beitriige zur Tertiiiren Deka-podenfauna Ungarns - Mathematische und Naturwis-senschaftliche Berichte aus Ungarn 18 98-120

LORENTHEYE amp BEURLENK (1929) Die fossilen Deka-poden der Liinder der Ungarischen Krone - GeologicaHungarica Series Palaeontologica 3 1-421

MAHEJM (1959) Nove Clenenie a pohlad na historicko-geologicky vyvoj mezozoika centralnych Karpat -Geologicke Prace 55 61-81

MEYERHv (1857) [Briefiiche Mittheilungen] - JahrbuchfUrMineralogie Geologie Geognosie und Petrefakten-Kunde 1857 556

Synopsis of fossil decapod crustaceans from Slovakia 169

MILNE-EDWARDSA (1862) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (4) 18 31-85

MILNE-EDWARDSA (1865) Monographie des Crustaces dela famille Canceriens - Annales des Sciences Natu-relIes (Zoologie) (5) 3 297-351

MISIKM SIBLlKM SYKORAM amp AUBRECHTR (1994)Jurassic brachiopods and sedimentological study of theBabina Klippe near Bohunice (Czorsztyn Unit PieninyKlippen Belt) - Mineralia Slovaca 26 255-266

MOLLERP (1984) Decapod Crustacea of the Badenian -Geologica Hungarica Series Palaeontologica 42 3-317

GROSSP PAPsovAJ amp KOHLERE (1973) Sedimentol6giaa stratigrafia vrchnej casti bazalnej litofacie centralno-karpatskeho paleogenu od Sarisskej vrchoviny po Lip-tovsku kotlinu Dielcia zaverecna sprava - 57 pp Bra-tislava (GlmS)

PAPsovAJ (1970) Zaverecna sprava 0 makrofaune paleo-genu Liptovskej kotliny na liste Ruzomberok 1 50000 Ciastkova zaverecna sprava - 30 pp Bratislava(GlmS)

PAPsovAJ (1975) Makrofauna strednej casti Liptovskejkotliny na listoch Huty Demiinova Smrecany Lip-tovsky Hradok Ciastkova sprava - 41 pp Bratislava(GlmS)

PAPsovAJ (1977) Celkove zhodnotenie makrospolocens-tiev Zapadnych Karpat Ciastkova zaverecna sprava -111pp Bratislava (GlmS)

PAPsovA1 (1978) Makrofauna paleogenu vychodnej castiLiptovskej kotliny Ciastkova sprava - 56 pp Bratisla-va (GlmS)

REuss AE (1859)Zur Kenntnis fossiler Krabben - Denk-schriften der kaiserlichen Akademie der Wissenschaf-ten Wien 17 1-90

SCHWEITZERCE amp FELDMANNRM (2008 [imprint2007]) A new classification for some Jurassic Brachy-ura (Crustacea Decapoda Brachyura Homolodromio-idea) Families Goniodromitidae BEURLEN1932 andTanidromitidae new family - Senckenbergiana lethaea87 (2) 119-156

SUKATCHEVAID SZALMAS VRSANSKYP CHALUpovAB GOLEJM LABAJOvAE amp SMRECKovAM (2006)Caddis-fly (Insecta Trichoptera) from the Badenianvolcano-sedimentary succession (Western CarpathiansSlovakia) - Geologica Carpathica 57 (6) 531-534

SZOMBATHYK (1916) Die tertiiiren Formen der GattungPotamon (Telphusa) und ihre paliiarktischen Nachkom-men - Annales historico-naturales Musei NationalisHungarici 14 281-472SPlNARZ CEPEKP BOUCEKB EuAsovA-FRAJovAH HAVLlCEKV HORNYRHOUSAV KALABISV KUKALovA1 MAREKL NE-KVASILovA0 POKORNYV PROKOPR PRIBYLA Ru-ZICKAB SNAJDRM STEMPROKovAD amp ZIKMUNDovA1 (1966) Systematicka paleontologie bezobratlYch -1052 pp Praha (Academia)

TOMASOVYCHA (1998) Badenske miikkyse z tehelne De-vfnska Nova Yes (Bratislava Slovensko) - MineraliaSlovaca 30 357-386

HYZNYM (2007) Paleogene crab fauna of Borove Forma-tion (localities Durkovec and Hlinisko) Western Car-pathians Slovakia - In GARASSINOA FELDMANNRM amp TERUZZIG (Eds) 3rd Symposium on Mesozoicand Cenozoic Decapod Crustaceans Museo di StoriaNaturale di Milano - Memorie della Societa Italiana diScienze Naturali e del Museo Civico di Storia Naturaledi Milano 35 (2) 59-61

HYZNYM (in press) In situ mud shrimps (Decapoda Axi-idea Callianassidae) preserved within their burrowsfrom the middle Miocene of the Central Paratethys -Bulletin of the Mizunami Fossil Museum 37

HYZNYM (2010) Revision of the Eocene decapod crusta-ceans deposited in the Liptovske muzeum Cierny Orolin Liptovsky Mikulas (Slovakia) - Acta Geologica Slo-vaca 2 (2) 117-122

HYZNYM (2011) Revision of Jaxea kuemeli BACHMAYER1954 (Decapoda Gebiidea Laomediidae) from the Mi-ocene of Europe with remarks on the palaeobiogeog-raphy of the genus Jaxea NARDO1847 - Neues Jahr-buch fUr Geologie und Paliiontologie - Abhandlungenthis volume

HYZNYM amp SCHLOGL1 (2009) The first occurence ofProsopidae sl (Crustacea Decapoda Brachyura)and other findings of crustaceans in the Upper Jurassiclimestones of the Pieniny Klippen Belt - In KROBlC-KI M (Ed) Jurassica VIII Vrsatec 09-11102009 -Geologia 35 (31) 61-63

HYZNYM amp SCHLOGL1 (in press) An Early Miocene de-ep-water decapod crustacean faunule from the ViennaBasin (Western Carpathians Slovakia) - Palaeontology

KENSLEYB amp HEARDRW (1991) An examination of theshrimp family Callianideidae (Crustacea DecapodaThalassinidea) - Proceedings of the Biological Societyof Washington 104 493-537

KLAUSS amp GROSSM (2010) Synopsis of the fossil fresh-water crabs of Europe (Brachyura Potamoidea Potami-dae) - Neues Jahrbuch fUrGeologie und PaliiontologieAbhandlungen 256 39-59

KOROSSYL (1940) Kurze geologische Beschreibung derUmgebung von Hernadzsadany - F6ldtani K6z16ny7083-109 143-144

KREMPASKAZ (1998) Niektore vysledky geologickeho apaleontologickeho vyskumu v lome Durkovec v Sloven-skom raji - Natura Carpatica 39 27-32

LORENTHEYE (1898)Beitriige zur Decapodenfauna der un-garischen Tertiiirs - Termeszetrajzi Ftizetek 21 1-133

LORENTHEYE (1902) Neuere Beitriige zur Tertiiiren Deka-podenfauna Ungarns - Mathematische und Naturwis-senschaftliche Berichte aus Ungarn 18 98-120

LORENTHEYE amp BEURLENK (1929) Die fossilen Deka-poden der Liinder der Ungarischen Krone - GeologicaHungarica Series Palaeontologica 3 1-421

MAHEJM (1959) Nove Clenenie a pohlad na historicko-geologicky vyvoj mezozoika centralnych Karpat -Geologicke Prace 55 61-81

MEYERHv (1857) [Briefiiche Mittheilungen] - JahrbuchfUrMineralogie Geologie Geognosie und Petrefakten-Kunde 1857 556

Synopsis of fossil decapod crustaceans from Slovakia 169

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Manuscript received July 1st 2010Revised version accepted July 6th 2010

Address of the author

MATUSHYZNYDepartment of Geology and PalaeontologyFaculty of Natural Sciences Comenius University Mlynsklidolina Gl Bratislava 842 15 Slovakiae-mail hyznymatusgmailcom

Synopsis of fossil decapod crustaceans from Slovakia 171

Appendix

Table 1 List of known fossil decapod crustaceans from Slovakia Only taxa determined to at least genus level are listedFor all known occurrences see text

FORMATION (AGE)FAMILYBohunice Fm (middle Oxfordian)TanidromitidaeBorove Fm (middle Eocene)RaninidaeZanthopsidae

Zanthopsidae

Borove Fm (upper Eocene)RaninidaeBorove Fm - Tomasovce Mb(upper Eocene)RaninidaeCalappidaeMathildellidaeBorove Fm - Tomasovce Mb(upper Eocene)RaninidaeLaksarska Nova Yes Fm (lowerMiocene)CtenochelidaeThomassiniidaeGalatheidaeGalatheidaeCalappidaePilumnidaeSebechleby Fm (middle Miocene)LaomediidaeCancridaeRetroplumidaeStudienka Fm (middle Miocene)CallianassidaeCallianassidaeStudienka Fm (middle Miocene)CalappidaeMatutidaeCancridaeStudienka Fm (middle Miocene)CalappidaeCancridaeStretava Fm (middle Miocene)PotamidaeBeladice Fm (upper Miocene)Potamidaemiddle PleistocenePotamidae

LOCALITYTAXONStepnicka skalaTanidromites insignisRuzomberok - MnichRaninoides fahianiiHarpactocarcinus punctulatus

Harpactoxanthopsis quadrilohata

Ruzomberok - Mnich Kleinov lornLophoranina reussi

Durkovec Hlinisko

Ranina spCalappilia tridentataCoeloma vigil

Radatice

Ranina hazslinskyi

Cerova-Lieskove

Callianopsis sp novCrosniera sp novAgononida sp novMunidopsis sp novMursia sp novStyrioplax exiguusPlastovceJaxea kuemeliTasadia carniolicaRetropluma horealisDevinska Kobyla - Dubravska hlavicaCallianassa hrocchiCallianassa pseudorakosensisDevinska Kobyla - SandbergCalappa hehertiSzahoa inermisTasadia carniolicaDevinska Nova YesMursia spTasadia carniolicaZdana Slanska HutaPotamon hungaricumOresanyPotamon hegauenseMuzlaPotamon antiquum

REFERENCES

HYZNYamp SCHLOGL(2009)

LORENTHEYamp BEURLEN(1929)LORENTHEYamp BEURLEN(1929)DORNYAY(1913) LORENTHEYamp BEURLEN(1929) PAPsovA (1970) HYZNY(2010)

DORNYAY(1913) herein HYZNY(2010)

HYZNY(2007)HYZNY(2007) hereinHYZNY(2007)

REUSS(1859)

HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)

HYZNY(2011)

hereinherein

hereinHYZNY(in press) herein

LORENTHEYamp BEURLEN(1929) hereinhereinherein

hereinTOULA(1900) herein

KLAUSamp GROSS(2010)

FORDlNAL(1994) KLAUSamp GROSS(2010)

KLAUSamp GROSS(2010)

Synopsis of fossil decapod crustaceans from Slovakia 171

Appendix

Table 1 List of known fossil decapod crustaceans from Slovakia Only taxa determined to at least genus level are listedFor all known occurrences see text

FORMATION (AGE)FAMILYBohunice Fm (middle Oxfordian)TanidromitidaeBorove Fm (middle Eocene)RaninidaeZanthopsidae

Zanthopsidae

Borove Fm (upper Eocene)RaninidaeBorove Fm - Tomasovce Mb(upper Eocene)RaninidaeCalappidaeMathildellidaeBorove Fm - Tomasovce Mb(upper Eocene)RaninidaeLaksarska Nova Yes Fm (lowerMiocene)CtenochelidaeThomassiniidaeGalatheidaeGalatheidaeCalappidaePilumnidaeSebechleby Fm (middle Miocene)LaomediidaeCancridaeRetroplumidaeStudienka Fm (middle Miocene)CallianassidaeCallianassidaeStudienka Fm (middle Miocene)CalappidaeMatutidaeCancridaeStudienka Fm (middle Miocene)CalappidaeCancridaeStretava Fm (middle Miocene)PotamidaeBeladice Fm (upper Miocene)Potamidaemiddle PleistocenePotamidae

LOCALITYTAXONStepnicka skalaTanidromites insignisRuzomberok - MnichRaninoides fahianiiHarpactocarcinus punctulatus

Harpactoxanthopsis quadrilohata

Ruzomberok - Mnich Kleinov lornLophoranina reussi

Durkovec Hlinisko

Ranina spCalappilia tridentataCoeloma vigil

Radatice

Ranina hazslinskyi

Cerova-Lieskove

Callianopsis sp novCrosniera sp novAgononida sp novMunidopsis sp novMursia sp novStyrioplax exiguusPlastovceJaxea kuemeliTasadia carniolicaRetropluma horealisDevinska Kobyla - Dubravska hlavicaCallianassa hrocchiCallianassa pseudorakosensisDevinska Kobyla - SandbergCalappa hehertiSzahoa inermisTasadia carniolicaDevinska Nova YesMursia spTasadia carniolicaZdana Slanska HutaPotamon hungaricumOresanyPotamon hegauenseMuzlaPotamon antiquum

REFERENCES

HYZNYamp SCHLOGL(2009)

LORENTHEYamp BEURLEN(1929)LORENTHEYamp BEURLEN(1929)DORNYAY(1913) LORENTHEYamp BEURLEN(1929) PAPsovA (1970) HYZNY(2010)

DORNYAY(1913) herein HYZNY(2010)

HYZNY(2007)HYZNY(2007) hereinHYZNY(2007)

REUSS(1859)

HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)HYZNYamp SCHLOGL(in press)

HYZNY(2011)

hereinherein

hereinHYZNY(in press) herein

LORENTHEYamp BEURLEN(1929) hereinhereinherein

hereinTOULA(1900) herein

KLAUSamp GROSS(2010)

FORDlNAL(1994) KLAUSamp GROSS(2010)

KLAUSamp GROSS(2010)