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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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PRLS =maximum lifespan − age at last birth

maximum lifespan − age of female reproductive maturity

Offspring Ratio =offspring weight at birth

adult weightOR

offspring length3

adult length3

Litter Size Adjusted Offspring Ratio = (Offspring Ratio)* (Average Litter Size)

Reproductive Window =1 −

Age at Last Birth − Age of Reproductive Maturity

Maximum Age

peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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PRLS =maximum lifespan − age at last birth

maximum lifespan − age of female reproductive maturity

Offspring Ratio =offspring weight at birth

adult weightOR

offspring length3

adult length3

peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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ReproductiveMaturity

AdulthoodDevelopment

Reproductive Senescence

DEATH

IZATION

Gestation Weaning

Birth

Maximum Documented Lifespan

Pro

po

rtio

n R

epro

du

ctiv

e o

r A

live

Fertilization

Reproductive Period PRLS

A B

CCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCC

peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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A B

C D

*

E F

frac

tion

repr

oduc

tive

frac

tion

aliv

efr

actio

n re

prod

uctiv

e

mat

ricid

e fr

actio

n

% re

prod

uctiv

e/al

ive

% re

prod

uctiv

e/al

ive

time (days of adulthood) time (days of adulthood)

time (days of adulthood) time (days of adulthood)

time (days of adulthood) time (days of adulthood)

peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;

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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;


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