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This article was downloaded by: [Cornell University Library] On: 12 November 2014, At: 17:04 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Annals and Magazine of Natural History: Series 10 Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ tnah16 XXXVII.—Otoliths of Fishes from the lower tertiary formations of Southern England.—III. Percomorphi, Scleroparei G. Allan Frost F.L.S. F.G.S. Published online: 18 Aug 2009. To cite this article: G. Allan Frost F.L.S. F.G.S. (1934) XXXVII.—Otoliths of Fishes from the lower tertiary formations of Southern England.—III. Percomorphi, Scleroparei , Annals and Magazine of Natural History: Series 10, 13:76, 426-433, DOI: 10.1080/00222933408654834 To link to this article: http://dx.doi.org/10.1080/00222933408654834 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our

XXXVII.— Otoliths of Fishes from the lower tertiary formations of Southern England .—III. Percomorphi, Scleroparei

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Page 1: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

This article was downloaded by: [Cornell University Library]On: 12 November 2014, At: 17:04Publisher: Taylor & FrancisInforma Ltd Registered in England and Wales Registered Number:1072954 Registered office: Mortimer House, 37-41 MortimerStreet, London W1T 3JH, UK

Annals and Magazine ofNatural History: Series10Publication details, includinginstructions for authors andsubscription information:http://www.tandfonline.com/loi/tnah16

XXXVII.—Otoliths ofFishes from the lowertertiary formations ofSouthern England.—III.Percomorphi,ScleropareiG. Allan Frost F.L.S. F.G.S.Published online: 18 Aug 2009.

To cite this article: G. Allan Frost F.L.S. F.G.S. (1934) XXXVII.—Otolithsof Fishes from the lower tertiary formations of Southern England.—III.Percomorphi, Scleroparei , Annals and Magazine of Natural History: Series10, 13:76, 426-433, DOI: 10.1080/00222933408654834

To link to this article: http://dx.doi.org/10.1080/00222933408654834

PLEASE SCROLL DOWN FOR ARTICLE

Taylor & Francis makes every effort to ensure the accuracy ofall the information (the “Content”) contained in the publicationson our platform. However, Taylor & Francis, our agents, and our

Page 2: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

licensors make no representations or warranties whatsoever asto the accuracy, completeness, or suitability for any purpose ofthe Content. Any opinions and views expressed in this publicationare the opinions and views of the authors, and are not the viewsof or endorsed by Taylor & Francis. The accuracy of the Contentshould not be relied upon and should be independently verifiedwith primary sources of information. Taylor and Francis shall notbe liable for any losses, actions, claims, proceedings, demands,costs, expenses, damages, and other liabilities whatsoever orhowsoever caused arising directly or indirectly in connection with,in relation to or arising out of the use of the Content.

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426 Mr. G. Allan Frost on

XXXVII.--Otoliths of Fishes from the Lower Tertiary Formations of Southern Eng/and.--III . Percomorphi, Scleroparei. By G. A~a~a~ FROST, F.L.S., F.G..S.

[ P l a t XV. ]

Order PERC01~IOI~ PHI.

Suborder P ~. g c o i D ~ A. Otolithus (Percidarum) eoncavus Priom *.

(P1. XV. fig. 1.) Shape ovate, outer side smooth, concave, inner side

convex, thin and curved in its length ; the posterior part narrower than the anterior; above the sulcus a shallow depression is present. The sulcns with upper and lower angles opens widely above the ros t rum; cauda curved and narrow.

Described by Priem from the Lutdtien of Lo Bois- Goufit and Upper Ypr6sien of H6rouval.

Two specimens, 7 × 4 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Percidarum) bartonensis Priom t. (P1. XV. fig. 2.)

Shape ovate, thick, outer side with upper part hollowed, umbo in centre and lower part raised; inner side convex with crista and wide depression above sulcus; dorsal rim raised, sloping posteriorly and forming angle with the irregular posterior r im; ventral rim curved; anterior rim obtuse or pointed. Sulcus with rounded upper angle placed anteriorly to the sharp lower anglo ; upper margin of ostium shorter than the lower margin ; cauda curved and pointed.

Described by Priem from Barton. Numerous specimens. Upper Eocene, Barton, Hamp-

shire. Otolithns (Centropomns) superpendens, sp. n.

(PI. XV. fig. 3.) Shape ovate, outer side concave, inner side convex;

dorsal rim low, curved; ventral rim keeled, deepest

* F . Pr iem, " Sur des otol i thes des Poissons 6oc~nos du Bass in Par i s ien ," Bull. Soc. g~ol. de France , s~r. 4, vi. p. 269, figs. 11-18(1906).

t F. Pr iem, " Sar des otoli$hes 6oc~nes de F rance et d 'Ang le te r re , " Bull. Soc. g6ol. de France , s~r. 4, xii. p. 248, figs. 7-8 (1912).

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Otoliths of Fishes. 427

anteriorly ; posterior rim oblique, the lower part rounded and overhanging; anterior rim with broad rostrum. Sulcus with upper and lower angles; upper margin of ostium distended ; cauda curved and pointed, constricted at junction with ostium.

Resembles that of the Recent species Centropomus undecimalis *.

One specimen, 7 × 4 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Scicenidarum) sheppeyensis, sp. n. (P1. XV. fig. 4.)

Shape lanceolate, longitudinally curved, outer side concave, inner side highly convex ; dorsal rim low, with median angle; ventral rim curved; posterior rim oblique, with an acute projection at the base ; anterior rim consisting of a broad pointed rostrum. Sulcus straight, with upper and lower angles; ostium flush with surrounding surfaces ; cauda straight, narrow, with small down-turned rounded end.

Resembles the otoliths of the Recent American species Cynoscion nebulosus ~f in the flush ostium, in the presence of a second lower angle in the cauda, and in the down- turned rounded end of the sulcus, but differs in the shape of the ostlum and in the angularity of the dorsal, anterior, and posterior rims.

One specimen, 12 × 5½ ram. Lower Eocene, London Clay, Warden Point, Isle of Sheppey.

Otolithus (Serranidarum ) rectangulus, sp. n. (P1. XV. fig. 5.)

Shape elongate, outer side concave, inner side convex ; dorsal rim low, forming angle with the oblique posterior rim ; ventral rim curved, regular, forming rounded anglo with the posterior rim ; anterior rim consisting of rounded rostrum. Sulcus horizontal, with an upper and two lower angles; ostium shallow, rectangular; cauda deep with down-turned end.

* G. Al lan Fros t , " A C o m p a r a t i v e S t u d y of t h e Oto l i th s of t h e N e o p t e r y g i a n Fishes , " Ann . & M a g . Nat . His t . ser. 9, vol. xx . p. 299. pl. v. fig. 7 (1927).

t T. c. p. 303, pl. v. fig. 28.

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428 Mr. G. Allan Frost on

Resembles generally that of the Recent North American species Perca flavescens, but differs in the squared and angular ostium.

One specimen, 8½ × 4 mm. Lower Eocene, London Clay, Warden Point, Isle of Sheppey.

Otolithus ( Smaris) eocenicus, sp. n. (P1. XV. fig. 6.)

Shape elliptical, outer side slightly concave, inner side convex; dorsal rim straight, forming anglo with the oblique posterior r im; ventral rim regular, curved; anterior rim irregular, concave. Suleus wide, undivided, curved, with rounded end.

Resembles the otoliths of the Recent species Smaris australis (Frost, t. c. p. 300, pl. v. fig. 11).

One specimen, 9½ × 4½ ram. Lower Eocene, London Clay, Worcester Park, Surrey.

Otolithus (Serranidarum) ostiolatus, sp. n. (P1. XV. fig. 7.)

Shape oblong, very thin, curved in its length ; dorsal rim low, straight ; ventral rim curved with slight median angle; posterior rim pointed, notched and furrowed; anterior rim upright, irregular ; cauda upwardly inclined with distended down-turned end.

Resembles tha t of the Recent American species Morone lineatus in the thin curved form of the otolith, in the relative positions of the angles of the sulcus, and in the upward inclination of the cauda, but differs in the vertical front rim, in the height of the ostium, and in the width and distension of the cauda.

Two specimens, 6×3½ mm. Lower Eocene, London Clay, Warden Point, Isle of Sheppey.

Otolithus ( Trachinus) falcatus, sp. n. (P1. XV. fig. 8.)

Shape lanceolate, biconvex; dorsal and ventral rims curved; posterior rim oblique, with pointed projection at base; anterior rim pointed. Sulcu~ undivided, sigmoidal ; cauda elevated.

Closely resembles that of the Recent species Trachinus draco (Frost, 1928, loc. cir. vol. i. p. 454, pl. xvii. fig. 13).

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Otoliths of Fishes. 429

One specimen, 4 × 2 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Trachinus) cequalis, sp. n. (P1. XV. fig. 9.)

Shape shuttle-like, symmetrical ; dorsal and ventral rims curved, meeting in the anterior and posterior pointed ends. Sulcus divided, ostium and cauda equal ; ostium rounded at junction with the slightly elevated cauda.

Resembles a fossil form erroneously described by Priem* from the Burdigalien of Martillac, S.W. France, as O. (Trachinus) miocenicus Bassoli & Schubert, to which it shows no resemblance, but differs from the otolith figured by Priem in the extra length of the cauda and in the absence of crenulations of the rims.

Two specimens, 2 × 1 and 4 × 2 mm. Upper Eocene, Barton, Hampshire.

Otolithus ( A pogon ) eocenicus Shepherd. (PL XV. fig. 10.)

Otolithus ( Monocentrls) bellovac~,nus Pr iem, 1911. Otolithus ( Monocentris ?) bellovacinus Schube r t , 1915.

This otoIith was first described by Priem t from the Lut6tien of the Paris Basin as belonging to the genus Monocentris. In 1915 Schubert:~, in retaining Priem's nomenclature for a number of specimens from Barton, but with the addition of a mark of interrogation, gives expression to his doubt as to the correctness of the description, and notes that the sulcus is nearer to that of Apogon; while in 1922 C. E. Shepherd §, comparing it with those of the Recent species Apogon rex mullorum, definitely places it as an otolith of Apogon, renaming it as Apogon eocenicus.

I t also resembles that of the Recent species Apogon melanota~nia (Frost, 1927, t. c. pl. v. fig. 25) in the sulcus,

* 1% Pr iem, " Sur des Oto l i thes de Po i s sons fossi les des t e r r a ins ter t ia i res sup6r ieurs du S u d - O u e s t de la F r a n c e , " Bull . See. g6ol. do F rance , s6r. 4, t . xi~. p. 275, fig. 74 (1914).

F. P r i em, " E t u d e des Po issons foss. du B a s s i n Pa r i s i en , " Anna l . de Pal . t . vi. p. 27, figs. 21-23 (Par is , 1911).

:~ R. J . Schuber t , " Obereoe~ne Oto l i then yon B a r t o n Cliff bei Chr i s t chu rch , t t a m p s h l r e , " J a h r b . der k .-k. geol. R e i c h s a n s t . Bd. lxv. p. 280, Taf . vii. figs. 1-5 (Wien, 1915).

§ C. E . Shepherd , " Sur que l ques e r reurs clans la d 6 t e r m i n a t i o n des o to l i thes fossi les ," Bull . Soc. g6ol. de F rance , s6r. 4, t. xxii. p. 139, pl. vii. fig. 3 (Par is , 1922).

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Page 7: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

430 Mr. G. Allan Frost on

but differs in the height of the otolith and in the sharply upturned rostrum.

The otoliths of the Recent species, however, while retaining a general resemblance to the Tertiary forms, appear to have undergone a process of degeneration and to have lost the bold outline and striking features of their fossil predecessors.

Eleven specimens, up to 5 × 3~ ram. Upper Eocene, Barton, Hampshire.

Otolithus ( A pogon ) tuberculatu8, sp. n. (P1. XV. fig. 11.)

Resembles O. (Apogon) eocenicus in shape and in the sulcus, but differs in the tubercle-covered furrowed outer side, which is in strong contrast to the smooth exterior of the latter-mentioned species.

Two specimens, 4× 3 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Cepola) prerube3cens Bassoli & Schubert. (P1. XV. figs. 12, 13, 14.)

G. G. Bassoli, " Otili fossili terzi~ri dell ' E m i l i a . - - I V . , " Riv i s ta I t a l i ana eli Paloontologia, Anne xii, Fasc. I. p. 54, T~v. ii. fig. 43 (Perugia, 1906).

Described by Bassoli from the Middle Miocene of Monte Gibio, this otolith, differing in no perceptible way from those of the Recent species Cepola rubescens, is a striking example of the persistence of a form prevailing through- out the Tertiary epoch down to the present day. In P1. XV. fig. 14 the shape is slightly modified, the posterior rim becoming straightened and resembling examples described by Sulc * from the Bartonian of Biarritz, but otherwise conforming to type.

Six specimens, 3½ × 2½ to 4½ × 2½ ram. Upper Eocene, Barton, Hampshire.

Otolithus (Cepola) elegans, sp. n. (eL x v . fig. 15.)

Shape lanceolate, outer side fiat, sculptured, inner side convex; dorsal rim short, curved; ventral rim curved and long ; posterior rim concave, irregular, and distended ;

• Ja ros lav Sule, " L e s Otol i thes dut)al~ogSne des Env i rons deBian~ tz , " Rosp r a vy S ta tn iho geol. nst . Ceskoslovenske Republ icky , vii. pp. 70-71, tab. i. figs. 41 a-15 b (Prague, 1932).

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Page 8: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

Otoliths of Fishes. 431

anterior rim consisting of a large, level, pointed rostrum. Sulcus with strong lower angle; ostium longer and straighter than in O. (Cepola) prerubescens.

Two specimens, 5½x 3 ram. Upper Eocene, Barton, Hampshire.

Otolithus (Cepola) bartonensis Schubert. (P1. XV. fig. 16.)

R. Schuber t , 1915, t. c. p. 282, Tar . vii. fig. 15.

Shape lanceolate; dorsal and ventral rims curved; anterior and posterior rims narrow and pointed ; ventral area raised. Sulcus horizontal, angles slight; upper margin of ostium distended ; cauda merging into groove surrounding the ventral area.

Described by Schubert from the Upper Eocene of Barton. Two specimens, 4 x 2 mm. Upper Eocene, Barton,

Hampshire. Division LABRIFORMES.

Otolithus (Pseudoscarus) crenulatus, sp. n. (P1. XV. fig. 17.)

Shuttle-shaped, outer side nearly flat, with radial furrows, inner side convex; dorsal rim crenulated; ventral rim curved ; anterior and posterior rims bluntly pointed. Sulcus waved, upper margin only faintly indicated.

Resembles very closely that of the Recent species Pseudoscarus viridis (Frost, 1928, Ioc. cit. set. 10, vol. i. p. 453, fig. 16).

Two specimens, 3½ x 2 mm. Upper Eocene, Barton, Hampshire.

Suborder T R: c HI VR o 1D ~ A.

Otolithus (Sphyrvena) oblongus, sp. n. (P1. XV. fig. 18.)

Shape oblong, outer side concave, inner side convex; dorsal rim short, highest anteriorly ; ventral rim curved ; posterior rim oblique ; anterior.rim with pointed rostrum. Sulcus straight; ostium small; cauda straight with rounded end.

Resembles in shape generally the otoliths of the Recent species Sphyrcena guachancho, but differs in the short narrow ostium and in the straight end of the cauda;

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Page 9: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

432 On Otoliths of Fishes.

it also differs in the square posterior part of the otolith, which resembles that of the fossil species O. (Sphyr~ena) handsfuchsi described by Schubert * from the Miocene of Voslau.

One specimen only, 3 × 2 ram. Lower Eocene, London Clay, Minster, Isle of Sheppey.

Order SCLEROPAREI.

Otolithus ( Scorpcenidarum) acutus, sp. n. (P1. x v . ~g. 19.)

Shape lanceolate, outer side flat, inner side convex; dorsal and ventral rims curved; anterior and posterior rims pointed; dorsal rim short, and the upper part of rostrum concave. Sulcus horizontal, with an upper angle only.

Resembles generally that of the Recent species Scorpoena dactyloptera, the regularity of the outline being probably due to the immatur i ty of the example.

One specimen, 2½× 1 mm. Lower Eocene, London Clay, Minster, Isle of Sheppey.

Otolithus (incert~e sedis) densus, sp. n. (P1. XV. fig. 20.)

Shape subovate, thick, biconvex ; dorsal rim irregular, highest anteriorly; ventral rim deep, curved, passing into the rounded posterior rim ; anterior r im--upper part vertical, lower part forming an upturned rostrum. Sulcus horizontal, divided, without angles ; ostium small, ovate ; eauda with distended end. A erista is present above the sulcus and a deep groove runs level with the ventral rim.

Remarkable for its unusually stout build and strongly marked features, this otolith resembles generally that of the Recent species Paracembraz robinsoni (Frost, 1929, loc. cir. ser. 10, vol. iv. p. 261, pl. iii. Rig. 19), but differs in the short ovate ostium and in the depth of the ventral rim.

In placing it among those of the order Scleroparei it is undesirable without further material to a t tempt to assign it to any particular family.

* R. J . Schuber t , " Die F i scho to l i thon des 6s te r r . - tmgar . T e r t i a r s . - - I I I . , " J a h r b . der k .-k. Geol. Re i chsans t . Bd. 56, p. 643 (Wien, 1906).

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Page 10: XXXVII.—               Otoliths of Fishes from the lower tertiary formations of Southern England               .—III.               Percomorphi, Scleroparei

On new Trichoptera in the French Pyrenees. 433

One specimen, 3½ × 2½ ram. Lower Eocene, London Clay, Minster, Isle of Sheppey.

Otolithus (Agonus) eocenicus, sp. n. (P1. XV. fig. 21.)

Shape lanceolate, outer side straight, inner side convex ; dorsal and ventral rims curved ; posterior rim bluntly pointed. Sulcus oblique, undivided, with rounded end.

Resembles that of the Recent species Agonus cataphractus (Frost, t. c. p. 262, pl. iii. fig. 26).

Two specimens, 5½ × 2½ ram. Upper Eocene, Barton, Hampshire.

Fig . Fig. Fig . F ig . F ig . F ig . F ig . F ig . F ig . F ig . Fi,q. F igs . F ig . F ig . F i g . FW. Fig . F ig . F ig .

E X P L A N A T I O N OF PLATE XV. 1. Otol i thus ( P e r c / d a r u m ) concawls Priem. 2. ~ ( ) bartonen~is Priem. 3. - - - - ( Gentropomu~) superpendens, sp. n. 4. - - (Scia~nidarum) sheppeyensis, sp. n . 5. ~ (Sdrranidarum) rectangu~us, sp . n . 6. ~ ( S m a r i s ) eocenicus, sp. n. 7. ..- (Serranidarum) ostiolatus, sp. n. 8. ( T r a c h i n u s ) f a l c a t ~ , sp . n . 9. ( ) ~qua l i s , sp. n .

10. (Apogon) eocenicu~ Shepnercl. 11. - - ( ) tuberc~datus, sp . n . 12-14 . . (Cepola) prcrubescens B a s s . & Sch . 15. - ( ) d e g a n s , sp. n . 16. ( ) bar$onensis S c h u b e r b . 17. ( Pseudoscarus ) crenulatus, sp. n. 18. ~ (Sphyr~ena) oblongus, sp . n . 19. .. ( S c o r p o ~ i d a r u m ) acut~.~, sp. n . 20. - - (incert~ sedis) den,~us, sp. n. 2 ] . (Agonus ) eocenicus, sp. n.

XXXVIII . - -New Trichoptera in the French Pyrenees. By lVIxRTr~¢ E. MOSELY, F.R.E.S.

DVRrtCG the summer of 1933, two months spent in the Haute-Garonne with a visit of ten days to the Hautes- 1)yr~n6es, in search mainly of Trichoptera, resulted in the discovery of four species new to science as well as a few new to the French fauna. As a full record of this trip is to be published at some future date, a record which will include as well the Plecoptera and Neuroptera taken during my visit, it will suffice here to give descriptions of the new species.

Ann. & Mag. N. Hist. Ser. 10. Vol. xiii. 28

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