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XXXII.—Otoliths offishes from the lowertertiary formations ofSouthern England.—II.PercomorphiG. Allan Frost F.L.S. F.G.S.Published online: 18 Aug 2009.

To cite this article: G. Allan Frost F.L.S. F.G.S. (1934) XXXII.—Otolithsof fishes from the lower tertiary formations of Southern England.—II.Percomorphi , Annals and Magazine of Natural History: Series 10, 13:75,380-386, DOI: 10.1080/00222933408654825

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JOHANSSO~, K . E . 1927. " Bei t rago zur K e n n t n i s s der Polyeh~eton- Fami l i en Hermellid~e, Sabellidm trod Serpul idm." Zool. B id rag , Uppsa l a , xL 184 pp. , 5 pls. ,15 text-f igs.

M'INTOS•, W. C. 1885. " R e p o r t on t h e A n n e l i d a P o l y c h m t a col- lec ted by H.M.S. ' C h a l l e n g e r ' d u r i n g t h e Yea r s 1873-1876." ' Chal lenger ' Rep . , Zool. vol. xii., 554 pp. , 94 pls.

M~ENZELLER, E. VOW. 1879. " S t i d j a p a n i s c h e A n ne l i den . " Denks . K . Akad . Wiss . Vienna , vol. xli. pp. 109-154, pls. i . -v i . ; a n d 1885, op. cit. vol. xl lx. pp. 197-224, pls. L- iv .

M o ~ o , C. C. A. 1930. " Po lychmte W o r m s . " ' D i s c o v e r y ' Reps . vol. ii., 222 pp. , 91 text-f igs .

Moo~E, J . P- 1909. " Polych~etous Anne l ids f rom Mon te r ey B a y a n d San Diego, Cal i fornia ." Prec . Acad. Na t . Sci. Ph i l ade lph ia , vol. lxi. pp . 235-294, pls. v i i . - ix .

SEIDL~R, H . J . 1924. " Bei t rgge zur K e n n t n i s s der P o l y n o i d e n . - - I . " Arch . N a t u r g e s . lxxx ix . Abt . A, H e f t xi. , pp . 217, 22 text-f igs .

XXXII.--Otolith8 of Fishes from the Lower Tertiary For- mations of Southern England.--II. Percomorphi. By G. ALLAH FROST, F.L.S., F.G.S.

[Pla te XIV. ]

Order PERCOMORPHI .

Suborder P E ~ C O I D E A .

Otolithus (Dentex) pulcher, sp. n. (P1. XIV. fig. ~.)

Shape ovate, outer side concave, inner side convex; dorsal rim irregular, pointed; ventral rim curved, serrated, forming an angle with the oblique serrated posterior r im; anterior rim obtuse, small ; rostrum present in the specimen described, other examples without rostrum.

Sulcus horizontal ; upper angle rounded and placed anteriorly to the sharp lower angle; ostium distended, opening on the anterior r im; cauda long, with sharply curved end. A crista and a narrow depression are present above the cauda, and below the sulcus a fine groove extends from the rostrum to the end of the cauda, with furrows from it to the ventral rim.

Resembles that of the Recent species Dentex macro- phthalmus *, but differs in the dorsal rim and in the forwardly placed upper angle of the sulcus.

* J . Sanz Echever r ia , " Inves t igac iones sobre otol i tos de Espa f i a , " Bo le t in de la Rea l Soe iedad Espaf io la de Hi s to r i a .Na tu ra l , t. x x x . pp. 17,5-178, pl. i. fig. 3 (Madrid, 1930).

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Otolith8 of Fishes. 381

Six specimens, 5×3½ to 9 × 6 m m . Upper Eocene, Barton, Hampshire.

Otolithus (Dentex) ovatus, sp. n, (P1. XIV. fig. 2.)

Shape ovate, thick, symmetrical, resembling O. (Dentex) pulcher generally, but differing in its stouter build, in the upper distension of the ostium, in the position of the upper angle of the sulcus, and in the straighter termination of the cauda.

Two specimens, 8×6 and 7 × 5 ram. Upper Eocene, Barton, Hampshire.

Otolithus (Cantharus) denticulatus, sp. n. (P1. XIV. fig. 3.)

Shape ovate, outer side concave, inner side convex; dorsal rim low, regular; ventral rim keeled, deepest anteriorly ; posterior rim rounded, notched ; anterior rim oblique. Sulcus opens widely on the anterior r im; upper angle slight, lower angle well defined; ostium distended, cauda gently curved.

Resembles that of a young example of the Recent species Cantharus lineatus *.

Two specimens, 2× 1½ ram. Upper Eocene, Barton, Hampshire.

Otolithus (Pagrus) bartonensis, sp. n. (P1. XIV. fig. 4.)

Shape pentagonal, outer side concave, inner side convex, upper part furrowed ; dorsal rim oblique, crenu- la tcd; ventral rim keeled; posterior rim concave, dentated ; anterior rim irregular. Sulcus oblique, opening widely on the anterior rim ; no upper angle, but large lower angle present ; upper margin of ostium descending, lower margin distended ; cauda distended, lower margin straight.

Resembles generally that of the Recent species Pagrus cardinalis, but differs in the distended cauda, which is similar to that of the Recent species Sargus vulgaris.

Three specimens, 5½ × 5 to 8 × 7 ram. Upper Eocene, Barton, Hampshire.

* G. Allan Frost., " A Compara t ive S tudy of t he Otol i ths of t he Neopte ryg ian F ishes , " Ann. & Mag. Nat . Hist . vol. xx. p. 301, pl. v. fig. 17 (1927).

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Otolithus (Pagellus) symmetricus, sp. n. (el. X l V . fig. 5.)

Shape high, symmetrical, outer side concave, inner side convex ; crista and depression above the sulcus ; dorsal rim high, pointed ; ventral rim deep ; anterior and poste- rior rims oblique; rostrum wide and pointed. S u l c u s opens on anterior rim, upper and lower angles present ; upper margin of ostium distended, lower margin straight ; cauda curved, wide and pointed.

Resembles that of the Recent species Pagellus erythrinus, but differs in the height of the dorsal rim and in the general symmetry of the otolith.

One specimen, 81x6 mm. Upper Eocene, Barton, Hampshire.

Otolithus ( Sparidarum ) s611ingensis Koken * (P1. XIV. fig. 6.)

Described by Koken from the Oligocene of Sollingen, North Germany.

Two specimens, 1 -1 7~× o.~ mm. Upper Eocene, Barton, Hampshire.

Otolithus (Sargus) eocenicus, sp. n. (P1. XIV. fig. 7.)

Shape ovate, outer side concave, inner side convex, crista and depression above the sulcus ; dorsal rim high, irregular, forming a sharp angle with the oblique, slightly concave posterior rim ; ventral rim curved and serrated ; anterior rim sloping, forming a stout rostrum. Sulcus oblique, with lower angle; ostium distended; cauda distended as in O. (Sparidarum) sSllingensis.

Resembles that of the Recent species Sargus rondeletti, but differs in its greater height.

One specimen, 5½ × 4 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Spariclarum) densus, sp. n. (P1. XIV. fig. 8.)

Shape ovate, very thick, outer side fiat, inner side convex; dorsal rim highest posteriorly; ventral rim curved; posterior rim oblique; anterior rim irregular.

• E. Koken, Zeitschrift der deut. geol. Gesell. xxxvi, pp. 500-565, Taf. ix.-xii. (Berlin, 1884).

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Otoliths of Fishes. 383

Sulcus wide, deeply excavated, upper and lower angles present ; ostium distended and hollowed ; cauda oblique with downturned rounded end. This otolith is noticeable among the Barton forms for its heavy coarse appearance.

One specimen, 9 × 6mm. Upper Eocene, Barton Beds, Highcliffe, Hampshire.

Otolithus (Sparidarum) oligocenicus, sp. n. (P1. XIV. fig. 9.)

Shape circular, outer side fiat, inner side slightly convex; dorsal rim indented, forming angle with the oblique posterior r im; anterior rim consisting of wide bhmt rostrum. Sulcus with upper and lower angles; ostium distended; cauda narrow, straight, with downtarned pointed end.

Resembles that of Sargus eocenicus, but differs in the narrow cauda and in the height of the postero-dorsal angle.

One specimen, 6 ×41 ram. Lower Oligocene, White- cliffe Bay, Isle of Wight.

Otolithus (Ditrema) sheppeyensis, sp. n. (P1. XIV. fig. 10.)

Shape ovate, thick, outer side flat, inner side strongly convex; dorsal rim low; ventral rim curved, regular; posterior rim oblique, corrugated; anterior rim with rostrum, slight antirostrum, and small excisura. Sulcus opening on anterior rim, with upper and lower angles; ostium distended, deep ; cauda with curved rounded end.

Resembles that of the Recent species Ditrema temminckii, but differs in its greater height and thickness.

One specimen, 7 × 5 ram. Lower Eocene, London Clay, Warden Point, Isle of Sheppey.

Otolithus (Perca) similis, sp. n. (P1. XIV. fig. 11.)

Shape ovate, outer side concave, inner side convex, with crista above the sulcus ; dorsal and posterior rims irregular, separated by a deep indentation ; ventral rim curved; anterior rim with broad, pointed rostrum. Sulcus nearly straight with a lower angle only ; ostium opening on rostrum ; cauda straight with rounded end.

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Resembles that of the Recent species Perca fluviatilis, but differs in the deeper ventral rim and in the absence of an excisura.

One specimen, 4×21 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Dentex) regularis, sp. n. (P1. XIV. fig. 12.)

Shape ovate ; dorsal, ventral, and posterior rims regular ; upper part of anterior rim oblique, forming angle with the lower part. Sulcus straight, with lower angle ; upper margin of ostium descending; cauda waved and pointed, resembling the sulcus of the Recent species Dentex macrophthalmus. The otolith resembles generally Otolithus (Dentex) latior described by Schubert from the Miocene of Voslau *, but differs in the shortened dorsal area and oblique anterior rim.

One specimen, 3 × 2 mm. Upper Eocene, Barton, Hampshire.

Otolithus (Serranidarum) minsterensis, sp. n. (P1. XIV. fig. 13.)

Shape ovate, outer side concave, inner side convex; dorsal rim low ; ventral rim deep ; posterior rim concave ; anterior rim consisting of a broad rostrum. Sulcus horizontal, with lower angle; ostium with descending upper margin, opening on rostrum ; cauda elevated with rounded end.

Resembles that of the Recent species Mcena vulgaris (Frost, 1927, t. c. p. 300, pl. v. fig. 12), but differs in the wide elevated caud~. I t also resembles very closely a fossil form O. (Percidarum) plebejus t described by Koken from the Middle Oligocene of Waldbfickelheim, again differing in the cauda, which in the latter species has a turned-down end.

One specimen, 3½× 2½ mm. Lower Eocene, London Bay, Minster, Isle of Sheppey.

* R. J . Schuber t , "Die F isehoto l l then des 6ster-ung. T e r t i a r s . - - I I I . , " J a h r b . der k.-k. geol. Reichsanta l t , B, lvi. pp . 623-706, Tab. xv i iL-xx . (Wien, 1906).

~f E. Koken , " Neue Unte rsuchungen an te r t ia ren l~isch-Otol i then," Zeitsch. d. deut . geol. Gesellschaft , Band xtiii, p. 77, pl. x. fig. 1 (Berlin, 1891).

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Ololith8 of Fishes. 385

Otolithus (Perca) striatu, s, sp. n. (P1. XIV. fig. 14.)

Shape ovate, outer side concave, inner side convex; dorsal rim low, serrated ; ventral rim curved ; posterior rim rounded ; anterior rim with broad pointed rostrum ; upper part of otolith furrowed. Sulcus straight, with lower angle; upper margin of ostium not distended; cauda with distended end.

Resembles Otolithus (Percidarum) frequens *, described by Koken from the Upper Oligocene of Sternberger Gestein, but differs in the distended cauda and in the pointed front of the dorsal area.

One specimen, 4½× 2½ mm. Lower Eocene, London Clay, Minster, Isle of Sheppey.

Otolithus ( Lutianidarum) gracilis, sp. n. (P1. XIV. fig. 15.)

Shape ovate, outer side concave, inner side convex; dorsal rim high, curved, finely serrated, passing into the rounded posterior r im; ventral rim curved ; anterior rim angular, truncated. Sulcus with upper and lower angles; ostium wide, upper margin dis tended; cau4~ narrow, curved, and pointed.

Resembles that of the Recent species Apsilus microdon, ram. Lutianid~e, but differs in the blunt anterior rim. I t also resembles the fossil species Otolithus (Serranidarum) curvidorsalis "~ Frost from the Blackheath Beds of Abbey Wood, but differs in the unusually fine cauda, in the irregular front of the otolith, and in the deiieato striations of the dorsal area.

One specimen, 5 × 3 mm. Lower Eocene, London Clay, Warden Point, Isle of Sheppey.

E X P L A N A T I O N OF P L A T E X I V .

Fig. 1. Otolithus (Dentex) pulcher, sp. n. Fig. 2. - - - - ( . .) ovatus, sp. n. Fig. 3. - - (Cantharus) denticulatus, sp. n. Fig. 4. - - (Pagrus) bartonensis, sp. n. Fig. 5. - - (PageUu~¢) symmetricus, sp. n. Fig. 6. - - (Sparidarum) sOllingensis Koken.

* E. Koken, t. c. p. 77, pl. viii. figs. 4-4 a. t E. I . White, " The Ver tebra te F a u n a s of the Engl ish Eocene ,"

Brit . Mus. Nat . Hist . vol. i. p. 108, pl. fig. 17 (1931). Ann. & Mat. N. Hist. Ser. 10. VoL xiii. 25

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386 Monsieur M. Pic sur les

Fig. 7. Otolithus (Sargus) eocenicus, s p . n . JFig. 8. - - (Sparidarum) dcnsus, s p . n . .Fig. 9. - - ( ) oligocenicus, s p . n . Fig. 10. - - (Ditrema) shcppeyensis, s p . n . Fig. 11. - - (Perca) similis, sp. n. Fig. 12. - - (Dentex) regularis, sp. n. Fig. 13. - - (Serranidarum) minstercnsis, s p . n . Fig. 14. - - (Perca) striatus, s p . n . Fig. 15. - - (Lutianidarum) gracilis, s p . n .

X X X I I I . - - N ouveaux Coldopt~res d' A frique. Par M. PIC.

Eubrianax vittaticollis [Helodid~e], sp. n., ~ *

Oblongus, parum nitidus, breve griseo pubescens, niger, thorace pro parte, elytris, abdomine pedibusque ruffs, geniculis piceis. Capite medio sulcato ; antonnis sat gracilibus, articulo 2 ° breve, 3 ° et sequentibus longe flabeUatis ; thoraco brevi et lato, lateraliter subarcuato et explanato, rufo, antice flavo-notato, medio nigro-vittato, minute et diverse punctato ; scutollo grandi, apice arcuato, dense mguloso- punctato; elytris thorace paulo latioribus, postice paulo dilatatis, apice oblique attenuatis, diverse et irregulariter punctatis.

Long. 5 mm.

CONGO BELGE : Elisabethville, 11. ix. 1931 (J. Ogilvie). Voisin de E. africanus Pic, s'en distingue, & premiere

vue, par le prothorax orn6 d'une bande m6diane noire.

Silidius m~Iciei [Malacodermata], sp. n.

Elongatus, nitidus, griseo-pubescens, niger, thorace, scutello, pectore elytrisclue rufo-testaceis, his apice late nigris, pedibus nigris, femoribus ad basin late et coxis testaceis. Capite sat lato, parum punctato, oculis validis ; antennis nigris, longissimis, gracilibus; thorace parum brevi, non transverso, medio late impresso, postice arcuate sulcato, lateraliter post ice inciso et bilobato-dentato; elytris thorace paulo latioribus, elongatis, postice paulo dilatatis, apice subattenuatis, diverse, postice densiore, punctatis; pedibus gracilibus.

Long. 8 mm.

• L e s t y p e s d e s p r 6 s e n t e s n o u v e a u t ~ s s e t r o u v e n t a u B r i t i s h M u s e u m ; l e s c o - t y p e s d e s Sil idius n~ackiei, Hedibius particularicornis, Pseudo- colotes binotaticollis, Dasytes mackiei, Pagurodactylu, limbatus e t v a r . , a i n s i q u e Pagurodactylus congoensis f i g u r e ~ l t d a n s l a c o l l e c t i o n P i c .

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F R O S T . Ann. ~f .~Iag. Nat. Hist. S. I0. Vol. X~I. _Pl. XIK.

2 . I,

4 . 5. 6.

9. 7. 8.

I0. II, 12.

13. 14. IS.

G . A H a n Fro s"5 - de l ,

FOSSIL OTOLITHS OF THE ORDER PERCOMORPHi

SUBORDER PERCOIDELA.

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