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Evolution in Extant PopulationsVariation in Form
Differentiation of Parts: different songs in individualsOntogeny: similar beginning embryo which diverges through development
Ambystoma tigrinum: different diet, different behavior (carnivorous & herbivorous), identical
genotypeGrowthAccretion: shells and trees add new layers
Addition: Foraminifera and echinoderms add partsMolting: (crabs, trilobites) shedding outer layer for larger bodice underneath; outgrowing your
clothesModification: hands are born with all bones which lengthen and changes over time
Isometric growth: proportions stay the same, size changesAllometric growth: shape and features changes; apes and humans
Sexual dimorphismPopulation variation (colors in plants, finger lengths between gender)
geographic variation in deer mice leads to population variation of camouflaging colorpatterns
variations of monarch flycatchers across islandsPhenotypic plasticity: nutritional variation in beetles and Daphnia longiremis (plankton)
Phenotypic plasticity vs rapid evolution (Northern pike)
Cline: gradual change in phenotype across landscape (elevation)Ring species: variation across landscape until speciation amongst adjacent populations
Modes of SpeciationAllopatric: physical barrier divides population
Peripatric: small founding population enters isolated nicheParapatric: new niche found adjacent to original one
Sympatric: speciation without physical separation
Origin of LifeWhat is life
process energy separate from environment store and process information (DNA) replicate these instructions crown-group: life stems from MRCA of all extant living organisms
Domains of life: Eukarya, bacteria, archaea
Lateral gene transfer (esp bacteria) complicates tree of life to create web of lifePrimodial soup: amino acids synthesized from methane, ammonia, hydrogen and boiling water
using electric spark for over a week (Miller)Banded iron formation: indication of photosynthesis by cyanobacteria producing oxygen to
support life on Earth; also creating stromatoliteSnowball Earth
reduction of methane and CO2 due to lack of silicate weathering in warmer conditions; heat nolonger trapped within atmosphere (opposite of global warming today)
temperature cools, snow develops reflecting sunlight creating positive feedback for more coolingand snow
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volcanism puts CO2 back into atmosphere, rescue
Readings
Palumbi - Humans as the Worlds Greatest Evolutionary Force
Use of antibiotics have quickly evolved many micro-organisms in our world today, creating a
cascading effects on other species. Bacteria will develop resistance to treatment as quickly as a
year after first use, forcing us to constantly develop new vaccines and pesticides. Methods for
controlling this rapid evolution: withholding most powerful drugs, screening for resistance
before therapy, integrated pest manage where multiple methods are used simultaneously, refuge
planting, engineering evolution to make resistant individuals less fit when chemicals are
removed.
Trinkaus - Late Pleistocene adult mortality patterns and modern human establishment
High immature-adult ratios within fossil samples suggest generally low adult life expectancy.
However, they may be a bias in who was buried, and older adult skeletons are considered to bemore fragile and less likely to become fossilized. It is concluded that mobility was necessary for
survival, and individuals who are unable to keep up are left behind to die. Changes in human
evolution during the Late Pleistocene must have been the result of increase fertility or reduction
of immature mortality, not adult longevity.
Losos & Ricklefs - Adaptation and diversification on islands
Darwin mentioned that isolated lineages can evolve independently, ultimately forming new
species, so islands are excellent examples of this. Stephen Gould proposes that, if history were
repeated, the outcome of evolution would be entirely different. Islands are good models for
evolutionary studies because the first colonists are often unchallenged and free to diversify innovel ways. Unbalanced colonization and strong dispersal abilities sometimes make islands
more diversified than continents, and old/isolated islands contain species not found anywhere
else; Morris makes an argument that organisms would evolve to fill the same ecological niches
(woodpeckers replaced by lemur that can also extract grubs from trees), so not entirely novel.
Adaptive radiation: natural selection and sympatric speciation eventually leads to a clade of
coexistent taxa, adapted and specialized to use a broad spectrum of ecological space. Larger
islands have more complex geography and more opportunities for allopatric isolation (more
niches). Older islands are more prone to extinction of species, though the potential role of
extinction as a select agent and potentially creative force has not yet been resolved.
Lazcano - Historical Development of Origins Research
For a long time, the theory of spontaneous life remained unchanged due to
political/ideological pressures, until the birth of biochemistry suggested that organic molecules
can be synthesized using organics. Most of the early experiments did not consider Earths
primitive conditions, however. By considering the first life forms as a collection of single
molecules, researchers (Miller) hypothesized a primordial soup that can generate amino acids
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given Earths primitive weather of thunderstorm and heat. Model for single unit cell organisms
came about with theories such as Coacervates (charged microscopic colloidal droplets that
concentrate organic materials, formed spontaneously when two solutions of macromolecules
with opposite charges mix) - conflicts with Millers DNA model. Now, the gap between
primordial soup and the RNA world is discouragingly enormous, but continued research are
leading to a refined soup recipe, and self-assembly is found to be not unique to biology, seemingly
following general principles that might be equivalent to universal laws of nature.