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SUPPLEMENTARY INFORMATION doi:10.1038/nature25453 WWW.NATURE.COM/NATURE | 1

SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

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Page 1: SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

SUPPLEMENTARY INFORMATIONdoi:10.1038/nature25453

WWW.NATURE.COM/NATURE | 1

Page 2: SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

SUPPLEMENTARY INFORMATIONRESEARCHdoi:10.1038/nature25453

WWW.NATURE.COM/NATURE | 2

Page 3: SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

SUPPLEMENTARY INFORMATIONRESEARCHdoi:10.1038/nature25453

WWW.NATURE.COM/NATURE | 3

Page 4: SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

SUPPLEMENTARY INFORMATIONRESEARCHdoi:10.1038/nature25453

WWW.NATURE.COM/NATURE | 4

Page 5: SU PPLEMENTAR Y INFORMATION - Nature10 .1038/nature research supplementary information ... rg pnpipnfi- gvnt rg pnpvpnfi- ... 1748 1748 2053 1983 1789 f

SUPPLEMENTARY INFORMATIONRESEARCHdoi:10.1038/nature25453

WWW.NATURE.COM/NATURE | 5

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