Sponge Porifera

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    Sponges are similar to other animals in that they are multicellular,heterotrophic,

    lack cell wallsand produce sperm cells. Unlike other animals, they lack true

    tissuesand organs, and have nobody symmetry. The shapes of their bodies are

    adapted for maximal efficiency of water flow through the central cavity, where it

    deposits nutrients, and leaves through a hole called the osculum.Many sponges

    have internal skeletons of sponginand/or spicules of calcium carbonateor silicon

    dioxide. ll sponges are sessilea!uatic animals. lthough there are freshwater

    species, the great ma"ority are marine #salt water$ species, ranging from tidal

    %ones to depths exceeding &,&'' m #(.( mi$.

    )hile most of the approximately (,'''*+',''' known species feed onbacteria

    and other food particles in the water, some hostphotosynthesi%ingmicroorganisms asendosymbiontsand these alliances often produce more food and

    oxygen than they consume. few species of sponge that live in foodpoor

    environments have become carnivoresthat prey mainly on small crustaceans.-+

    Most species use sexual reproduction, releasing spermcells into the water to

    fertili%e ovathat in some species are released and in others are retained by the

    mother. The fertili%ed eggs form larvaewhich swim off in search of places to

    settle.-0Sponges are known for regenerating from fragments that are broken off,

    although this only works if the fragments include the right types of cells. few

    species reproduce by budding. )hen conditions deteriorate, for example as

    temperatures drop, many freshwater species and a few marine ones produce

    gemmules, survival pods of unspeciali%ed cells that remain dormant until

    conditions improve and then either form completely new sponges or recoloni%e

    the skeletons of their parents.-1

    The mesohyl functions as an endoskeletonin most sponges, and is the only

    skeleton in soft sponges that encrust hard surfaces such as rocks. More commonly,

    the mesohyl is stiffened bymineralspicules, by sponginfibers or both.

    2emospongesuse spongin, and in many species, silicaspicules and in somespecies, calcium carbonateexoskeletons. 2emosponges constitute about 3'4 of

    all known sponge species, including all freshwater ones, and have the widest

    range of habitats. 5alcareous sponges,which have calcium carbonate spicules

    and, in some species, calcium carbonate exoskeletons, are restricted to relatively

    shallow marine waters where production of calcium carbonate is easiest.-6The

    fragile glass sponges, with scaffolding of silica spicules, are restricted to polar

    regions and the ocean depths where predators are rare. 7ossils of all of these types

    have been found in rocks dated from (&'million years ago. 8n addition

    rchaeocyathids, whose fossils are common in rocks from (1' to 63'million

    years ago, are now regarded as a type of sponge.

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