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Lower Palaeolithic Transitions in the Northern Latitudesof Eurasia
Jan Michal Burdukiewicz
Abstract The northern latitudes of Eurasia were
inhabited temporarily, during favorable, warmer per-
iods since 1 Ma ago with a considerable transition
from Mode 1 (Oldowan) or Mode 2 (Acheulean) to
microlithic technology and introducing new andmore
effective composite tools. Until recent years, archae-
ologists believed that such technology occurred
almost exclusively during the Mesolithic and Late
Palaeolithic (Mode 5), in the form of small stone
inserts held by wooden or bone hafts, producing com-
posite tools. A significant spatial, chronological and
ecological variability ofLower Palaeolithicmicrolithic
assemblages suggests that they developed as a result of
the adaptation to local environment, possibly tempe-
rate and wooded, in different areas from North-East-
ern China to Northern Europe, parallel to Mode 1
and Mode 2 in southern and western part of Eurasia.
Keywords Acheulean � Lower Palaeolithic �
Microlithic technocomplex � Oldowan � Composite
tools
Introduction
The Lower Palaeolithic Period covers the longest time
span in human history, rich in numerous changes in
the widening of inhabited areas as well as in techno-
logical development. Archaeologists usually distin-
guish two main technologies of the Lower Palaeo-
lithic: Pebble Tool Technocomplex (called usually
Oldowan or Mode 1) and Technocomplex (called
Acheulean or Mode 2). Both these technocomplexes
originated inAfrica and dispersed to southern parts of
Asia and Europe. The climate of Africa was warm
enough for early hominins to survive. Southern Asia
and Europe offered more or less similar environmen-
tal conditions. This way technological equipment in
these areas was sufficient to exist and important tran-
sitions are not visible in the archaeological data.
Northern latitudes of Eurasia had very changeable
climatic conditions during the Pleistocene period.
According to recent research, the first inhabitants
in Northern Eurasia appeared c. 1 Ma ago. The
earliest colonization of northern latitudes (around
the 60th parallel) is connected with a considerable
transition fromMode 1 or 2 tomicrolithic technology.
Such sites are known in Eurasia, Central Europe,
and China, and span from c. 1 Ma to 300 ka BP, far
to the north from the Movius Line indicating the
northern border of the presence of Mode 2.
It is supposed that forested areas, especially the
northern zone of Eurasia, were settled because early
hominin groups carried out transitions to microlithic
technology, which meant introducing new and more
effective composite tools.Northern latitudes of Eurasia
were inhabited temporarily, during favorable, warmer
periods. These inventories are characterized by the
domination of microlithic technology in lithic produc-
tion (average length of artifacts c. 15–30 mm), which
were most probably hafted. Such invention in human
history is usually seen as very sophisticated and char-
acteristic forMode 5.Until recent years, archaeologists
believed that such technology occurred almost exclu-
sively during the Mesolithic and Late Palaeolithic
(Mode 5), in the form of small stone inserts held by
J.M. Burdukiewicz (*)Institute of Archaeology, University of Wroclaw, Wroclaw,Poland
M. Camps, P. Chauhan (eds.), Sourcebook of Paleolithic Transitions, DOI 10.1007/978-0-387-76487-0_11,� Springer ScienceþBusiness Media, LLC 2009
195
wooden or bone hafts, producing what is known as
composite tools. Finally they came to realize, however,
that small lithic tools, often no larger than a fingernail
and hard to hold using only fingers, started to be
produced much earlier, by hominin groups inhabiting
northern latitudes of Eurasia in the Lower Palaeolithic.
The Earliest Settling of Eurasia—Modes1 and 2
The earliest colonization of northern Eurasia is now
connected with the oldest archaeological site of
Dmanisi in Georgia, dated to c. 1.8 Ma (Vekua
et al. 2002). The artifacts of Dmanisi as well as
other older sites in Eurasia are connected with the
Pebble Tool Technocomplex. It should be men-
tioned further that sites with pebble tool technology
have similar tool-making techniques regardless of
raw material differences. The Oldowan appeared c.
2.6 Ma ago (Semaw et al. 2003; Stout et al. 2005). The
most important feature of Oldowan artifacts is the low
degree of standardization (Wynn and Tierson 1990).
The most diagnostic forms for such assemblages are
core tools and flakes with one or more sharp edges or
points. In other words, Mode 1 (Oldowan) can be
characterized by simple lithic processing techniques
with one, two, or more flaking directions (Plummer
et al. 2001; De la Torre and Mora 2005). Sometimes
centripetal flaking can also be seen. These flakes were
modified by the application of simple retouch, and
were frequently notched or denticulated. At archaeo-
logical sites from this period simple unmodified manu-
ports and hammerstones are common.
Further expansion to the north is connected with
the Acheulean technology, Mode 2, or more neutral
handaxe technology, which appeared c. 1.6Ma ago in
Africa (Dominguez-Rodrigo et al. 2001). Such tech-
nology was more sophisticated and standardized
(Mc Pherron 2000; Mc Nabb et al. 2004) compared
to the Oldowan. Handaxes, cleavers, and picks are
frequently seen as ‘‘preferred products’’ (Bordes 1968,
64; Debenath and Dibble 1994, 130, etc.), ‘‘cores for
flake extraction’’ (Davidson and Noble 1993), or
‘‘individualized memic constructs’’ (McNabb et al.
2004). Anyway, the handaxes are alike all through
their spatial and temporal allocation, although they
show a variety of shapes as well. Bifaces from Europe
and Asia are almost the same as African ones. How-
ever, cleavers in Europe are much rarer than in Africa
or South Asia.
From an archaeological point of view bifaces are
interpreted as a wide taxonomic unit, with recurring
‘‘mental templates’’ held by the knappers sharing simi-
lar cultural principles (Wynn and Tierson 1990). In
another explanation, technological and morphologi-
cal similarity did not reflect a taxonomic unit, in
opposition to the idea that it was the convergent result
of knapping techniques and the use of the artifacts
(Wynn 2004, 674). This way bifacial technology was
convergently invented in different local traditions, and
handaxes were made in various ways by individual
manufacturers, depending mostly on raw material.
Majority of archaeologists prefer to see the Acheu-
lean as a general taxonomic unit with several spatial
and temporal variations. The northern border of early
handaxe distribution is indicated by the Movius Line,
with some recent modifications like Bose Basin in
China (Hou et al. 2000). The oldest sites with handaxes
in Western Asia appeared c. 1.4 Ma ago in Ubeidiya
(Bar-Yosef 1987; 1998) and Gesher Benot Ya’aqov in
Israel aged 0.8 Ma (Goren-Inbar and Saragusti 1996).
In South Asia, the early Acheulean site of Isampur
(India) is tentatively dated to 1.2Ma (Paddayya et al.
2002; Chauhan 2004). In Eastern Asia the oldest han-
daxes are lastly known from Bose Basin in Southern
China, and are dated to 0.8 Ma (Hou et al. 2000).
In EuropeMode 2 appeared later, probably 600 ka
ago, as indicated by the oldest sites with handaxes, like
Venosa Notarchirco in Southern Italy, Carriere Car-
pentier inNorthernFrance, andBoxgrove in southern
England. Much more numerous are slightly younger
lithic concentrations from La Galeria in Atapuerca
(Carbonell et al. 2001, 267–271).
Lower Palaeolithic Microlithic Sitesin the Levant
In the northern latitudes of Eurasia there are lithic
assemblages which are quite unlike the two men-
tioned above. They are characterized by core/flake
technology and by the very small size of the arti-
facts. These artifacts are frequently calledmicroliths
because of their small dimensions, like typicalMode
5 tools (Figs. 1, 2, 3, 4, 5, 6, 7, 8, and 9).
196 J.M. Burdukiewicz
Fig. 1 Mean length ofartifacts from the mainLower Palaeolithicmicrolithic sites in Eurasia
Fig. 2 Lower Palaeolithicmicrolithic sites in CentralEurope: Scatter plot of corelength and width
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 197
Nevertheless, they do not have frequent regular
geometric shapes like Mesolithic ones. Sometimes
they are associated with larger tools, like choppers,
which make them somewhat similar to the Pebble
Tool complex.
The oldest Lower Palaeolithic microlithic
assemblages known at present appeared around
1 Ma BP in the Levant. A unique microlithic site
is Ruhama near Ashqelon, in the border zone
between the coastal plain and the Negev Desert
(Ronen et al. 1998; Burdukiewicz and Ronen
2000; Zaidner et al. 2003). At Ruhama, flint arti-
facts are numerous and homogenous in size. Mean
lengths of cores, flakes, and tools are smaller than
3 cm (Fig. 1). The most numerous are single plat-
form cores, but double platform cores and items
Fig. 4 Lower Palaeolithicmicrolithic sites in CentralEurope: Frequency of maingroups of artifacts
Fig. 3 Lower Palaeolithicmicrolithic sites in CentralEurope: Mean length offlakes and tools
198 J.M. Burdukiewicz
with changed orientation are also present. The
retouch techniques are usually scalar, denticulated,
or notched. Bifacial retouch is also present but rare,
and some points are reminiscent of very small han-
daxes, which are also microlithic. It should be an
indication of a transition from Mode 2 technology.
The presence of warm-region fauna (bovids,
horses, and hippopotamuses) broadly indicates
the climatic conditions.
Another site, Evron-Quarry, is located in the wes-
tern Galilee coastal plain c. 2.5 km from the Medi-
terranean shore and 20 m above sea level, slightly
east of the major sandstone ridge along this coast.
The inventory includes three components: (1) 20
handaxes, (2) cores, flakes, and tools of ‘‘ordinary’’
size, and (3) small size cores and debitage (Ronen
2003). The presence of such small artifacts is excep-
tional for the Lower Palaeolithic assemblages in the
Levant. At Ruhama, only small artifacts are present,
but at Evron-Quarry, there are two separate
components: a large one with handaxes and a small
one showing a complete technological process, from
small core processing to small retouched tools
(Ronen 2003). In the Evron-Quarry case, it is not
clear if the ‘‘small-tool component’’ was fashioned by
the same hominin group or another one, which used
the same place in different periods.
Lower Palaeolithic Microlithic Sitesin Central and Eastern Asia
Other sites with microlithic artifacts, dated roughly
to 1 Ma ago, are recorded at Donggutuo and
Xiaochangliang in northeast China (Keates 2003,
149). At the present stage of research it is thought
that deposits with microlithic artifacts date from a
period somewhat before the Jarmillo event, i.e.,
Fig. 5 Bilzingsleben in Thuringia (Germany): Microlithiccores Fig. 6 Bilzingsleben in Thuringia (Germany): Noncortical
flakes
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 199
their dating is somewhat before 990 ka BP.
Xiaochangliang in the Nihewan Basin is rich in
microlithic artifacts with warm-region fauna (forest
elephants, hippopotamuses, horses, red deer,
gazelles, and small mammals) and plant remains of
birch and elm (Zhou et al. 2000; Keates 2003).
Similar artifacts were found in Donggutuo, located
on loess upland 1.5 km fromXiaochangliang. There
were several archaeological horizons with over 10
thousand artifacts and numerous bone fragments of
warm-region fauna. Lithic artifacts, like cores,
flakes, and retouched tools, at both sites are very
small (Fig. 1). Keates (2003, 149) suggests that these
dimensions resulted from the poor quality of local
raw material like chert, vein quartz, quartzite,
basalt, or sandstone.
The best known site with microlithic artifacts in
China is Zhoukoudian locality 1—a large cave 140
m long and up to 40 m broad. There were 17 layers
connected with 5 climatic cycles (Zhou et al. 2000).
Cycle 1 (layers 14–13) was cold and should be
linked with OIS 18–19. Cycle 2 (layers 12–9) was
rather warm; and layer 10, with a lower archaeolo-
gical horizon, dense ash, and remains of animals
from forest and steppe environments, was recently
dated c. 670 ka BP (Zhou et al. 2000, 105). Cycle 3
(layers 8–6) originated in a warm climate and was
closed by the collapse of the cave roof; it is corre-
lated with OIS 13–12. The warmest was cycle 4
(layers 5–3), when the travertine originated. From
a layer dated by the TMS method to c. 400 ka BP,
come traces of fireplaces and the youngest hominin
remains (Zhou et al. 2000, 108). The top layers 2
and 1 come from cycle 5, which originated during
the cold climate of OIS 11–10. In the cave were
found over 100,000 lithic artifacts, numerous ani-
mal remains, as well as bone fragments of 45 indi-
viduals of Homo erectus pekinensis. Forty-four var-
ious types of rocks had been transported from a
distance of up to 5 km (Zhou et al. 2000; Keates
2003, 151). The main raw material (over 88%) was
Fig. 8 Trzebnica 2LH in Silesia (Poland). Side scrapersFig. 7 Bilzingsleben in Thuringia (Germany): Bifacialpoints: 1–6, elongated points: 7, burins: 7–8
200 J.M. Burdukiewicz
vein quartz represented by cores, flakes, and tools
in the range of 1.9–7.3 cm, but over 75% of the
tools were smaller than 4 cm. There were numerous
pebble tools as well.
In China these sites are included in the ‘‘small
tool tradition,’’ also called the ‘‘Donggutuo-
Zhoukoudian-Xujiayao-Salawusu’’ tradition (in
chronological sequence), with slightly larger tools
(mean length c. 27–30 mm) made of a variety of
lithic raw material, such as chert, quartz, basalt,
and others. They are juxtaposed with the macro-
tool tradition ‘‘Kehe-Lantian-Sanmenxia-Dingcun,’’
with choppers, picks, and spheroids (Keates 2003).
However, the most complicated aspect of such a
division is the inseparability of several collections.
Several researchers believe that the usage of wood
and bamboo was very important (Keates 2003, 156).
A similar number of artifacts is characteristic of
Kuldara in Tajikistan, Central Asia. This site is
considered at present as the oldest in the region. Its
age, based on soil stratigraphy, is correlated with
OIS 23 or 25, i.e., c. 900 kaBP (Ranov andDodonov
2003). In Central Asia, microlithic assemblages are
known from recent research in Kazakhstan, at sites
like Koshguran and Shoktas, which are dated to
c. 500 ka BP (Derevianko et al. 2000).
Lower Palaeolithic Microlithic Sitesin Europe
In Europe microlithic sites are known from Italy,
Hungary, Germany, and Poland. The oldest are
Italian sites, which are dated from 0.7 to 0.3 Ma
BP. Isernia La Pineta in Central Italy was excavated
in 1978. Three archaeological layers were recog-
nized (Peretto 1994). The richest was horizon 3a,
which yielded over 10,000 artifacts and rich faunal
and floral collections. This horizon showed a K/Ar
date of c. 730 ka and a similar age estimation by
paleomagnetic method (Cremaschi and Peretto
1988). However, age estimation according to Arvi-
cola chronology by T. van Kolfschoten is 200 ka
younger (Roebroeks and van Kolfschoten 1995).
Organogenic data enable reconstruction of the eco-
logical setting as an open landscape with relatively
warm climatic conditions.
The small lithic artifacts were made of local flint
with small crystalline intrusions, and much less
numerous macro-tools were made of dolomite
chalk. For example, in sector II of the site 4589
artifacts were collected: 2.2% cores, 70% flakes,
and almost 28% retouched tools. The mean length
of the flakes was 23 mm and the tools were slightly
longer—31 mm (Crovetto 1991). The main goal of
flint processing in Isernia La Pineta was the produc-
tion of flakes with sharp edges, which were used as
functional tools; the retouch seems to have been
accidental damage rather than intentional reshap-
ing (Peretto 1994, 460).
Similar small artifacts like those at Isernia La
Pineta were also found at other Italian sites:
Venosa-Loreto (Basilicata), Quarto delle Cinfonare
(Latina), and Visogliano near Trieste. In Venosa-
Loreto the most important is horizon A, dated to
Late Cromer Complex and previously interpreted as
Tayacian (Crovetto 1991); however, it is much older
Fig. 9 Rusko 42 in Silesia (Poland). Side scrapers: 1–10,point fragments: 11–12
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 201
than traditional Tayacian in France. Nearby in
Venosa the important Acheulean site Notarchirico
has several archaeological horizons, and is dated to
the Middle Pleistocene (Piperno 1999). Quarto delle
Cinfonare yielded only microlithic artifacts. The
team which excavated the site believes that it should
be compared with Fontana Ranuccio nearby, which
is also characterized by the presence of small artifacts
associated with bone handaxes (Peretto et al. 1997,
613). In the Visogliano cave, a small collection of
microlithic artifacts was found to be associated with
a few pebble tools and forest/steppe fauna dated to
OIS 13 or 11 (Cattani et al. 1991).
The next region which has produced early micro-
lithic assemblages is Central Europe. The oldest are
Mauer in Southern Germany and Vertesszolos in
northern Hungary, which originated probably
0.6 Ma BP. Vertesszolos was inhabited several
times, probably up to 0.3 Ma BP. Additional sites,
Bilzingsleben and Schoningen in Central Germany,
were inhabited several times in the period
0.45–0.3 Ma . These sites yielded unique organo-
genic finds made of wood, bone, and antler. Trzeb-
nica and Rusko in southwestern Poland are the
most northern, and, like Schoningen in Germany,
they were covered by Scandinavian inland ice at one
time.
The lower deposits at Mauer in Baden-Wurttem-
berg include redeposited microlithic artifacts and a
jaw ofHomo heidelbergensis. These small lithic arti-
facts and remains of temperate fauna were for a
long time overlooked because of their post-deposi-
tional replacement (Beinhauer et al. 1992, 46). Pol-
len analysis indicates that Homo heidelbergensis
here lived in a forested environment (Urban 1992).
Occupation at Vertesszolos is documented in
nine layers with lithic artifacts, a skull fragment,
and two teeth of Homo erectus seu sapiens palaeo-
hungaricus, spanning from around 600 to 300 ka BP
in changeable climatic conditions, from Mediterra-
nean to cold forests of Pinus montana (Kretzoi and
Dobosi 1990). Over this long duration its archaeol-
ogy witnessed little transformation, despite evident
changes in the natural environment. At
Vertesszolos, almost 8,900 lithic artifacts in several
horizons were documented, and are characterized
by the presence of numerous small pebble and flake
tools made of flint (66%), quartzite (31%), and
limestone (3%).
Other microlithic sites are recorded in Lower
Silesia in Poland. Several years ago the brickyard
at Winna Gora in Trzebnica and the open cast
kaolinite mine at Rusko near Strzegom produced
four microlithic assemblages, relatively dated on
geological grounds to OIS 13 and 11. The most
interesting is the lower horizon from Trzebnica
(OIS 13), which included almost 1,500 lithic arti-
facts (Fig. 8) and several remains of forest and
steppe fauna. Other assemblages: Trzebnica upper
horizon, Rusko 33, and Rusko 42 from OIS 11
contained almost exclusively small lithic artifacts
(Fig. 9). The richest inventory was from Rusko 42,
with almost 3,800 lithic artifacts (Burdukiewicz
2003).
Another very rich site is Bilzingsleben, excavated
over 30 years by D. Mania (Mania and Mania
2003), which delivered 140,000 artifacts (Figs. 5, 6,
and 7). Seventy-five per cent of these artifacts were
made from Nordic flint. Among these 120,000 flint
artifacts there are about 30,000 retouched tools.
Several pebble tools were present, made of large
pieces of crystalline rocks, and rare bifacial tools
made from flint similar to miniature Acheulean
handaxes (Fig. 7; Bruhl 2003, 51). Exceptional are
the rare bone handaxes of normal size (Bruhl 2003,
52). These artifacts were associated with 3 or 4
individuals of Homo erectus bilzingslebenensis and
a very rich warm ecological context represented by
thousands of floral and faunal remains (Mania and
Mania 2003).
An extremely interesting discovery was made at
Schoningen in Lower Saxony, where H. Thieme
identified seven assemblages, of which only three
were published in detail. Schoningen produced a
number of truly exceptional wooden objects in asso-
ciation with lithic artifacts: namely, several spears, a
throwing stick, and small handles—hafts for micro-
lithic stone tools (Thieme 2003). The findings from
Schoningen have shed new light on the more poorly
preserved wooden artifacts discovered at
Bilzingsleben.
The microlithic assemblages recently recorded in
Central Europe cited above apparently jointly
represent a distinct taxonomic complex, sharing a
number of common attributes, which are analyzed
in the coming section. Most probably, the complex
in question developed as a result of adaptation to
the conditions of the moderate climate in a
202 J.M. Burdukiewicz
woodland zone, with some elements of aMediterra-
nean climate in the southern region. In the north,
certain boreal elements are observed, as evidenced
by finds of spruce and fir wood objects recorded at
Schoningen (Fig. 10).
Lithic Artifacts
Analysis of lithic artifacts is usually limited to gen-
eral descriptions of cores and flakes and more
detailed classification of retouched tools. Therefore,
the size of the artifacts and the peculiarities of the
microlithic assemblages were not perceptible. The
author prefers a morphometric approach, showing
a technological and three-dimensional analysis.
Experience was gained from the study of other
Palaeolithic assemblages (Schild 1980) and formu-
lated as a dynamic technological analysis. A list of
all artifact categories and attributes was designed in
the form of a hierarchical sequence of production of
all artifacts (including waste and broken pieces)
classified with several levels from each category
into main groups, keeping with the technological
sequences and statistics: I—raw material procure-
ment, II—preparation and early core exploitation,
III—advanced core exploitation, IV—final core
exploitation, and V—tool production (Table 1).
Intentional selection of raw stone material is the
first important factor, which indicates the users’
familiarity with the properties of the rocks. For
example, at Bilzingsleben lithic tools were made
mainly from flint nodules and chunks. Chert and
hornstone were used less often. Only in areas with
limited flint deposits, such as Vertesszolos, were the
dominant materials quartz and quartzite. Flint
accounted for more than 90% of the inventories.
The early stage of core exploitation is represented
by initially struck cores and by cortical or cortical-
natural flakes. A comparison of average flake
length in Lower Palaeolithic microlithic inventories
shows them to have an astounding similarity of
between 16 and 23 mm (Fig. 1).
The more advanced stage of working is repre-
sented by cores with more than three removals and
Fig. 10 Possiblereconstruction of the LowerPalaeolithic composite tool(c. 20 cm-long fir stick withdiagonal groove and insertedpointed flint) according tofinds more than 400 ka oldfrom Schoningen 12(Germany)
Table 1 Lower Palaeolithic microlithic sites in Central Europe: Dynamic technological sequences of lithic artifacts
Rusko 33 Rusko 42 Trzebnica 2LH Trzebnica 2UH BilzingslebenTechnological sequences N % N % N % N % N %
I. Raw material procurement 0 0.00 0 0.00 376 25.67 57 26.76 1388 22.21
II. Preparation and early coreexploitation
31 8.93 133 3.58 132 9.01 10 4.69 306 4.90
III. Advanced core exploitation 88 25.36 411 11.05 158 10.78 15 7.04 1174 18.78
IV. Final core exploitation 193 55.62 2822 75.86 601 41.02 102 47.89 1808 28.93
V. Tool production 35 10.09 354 9.52 198 13.52 29 13.62 1574 25.18
Total 347 100.00 3720 100.00 1465 100.00 213 100.00 6250 100.00
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 203
flakes partially covered by cortex on the dorsal face
or noncortical flakes (Fig. 5; 6). Altogether they
account for 7–25% of all products (Fig. 4). Some
difficulty for classification is posed by flake frag-
ments, which form one of the largest categories of
flints in all assemblages. The set of cores analyzed
with the DTAmethod shows interesting tendencies.
Slightly over half of the pieces were produced dur-
ing the sequence of core exploitation. Preparation
of core platforms played an important role inmicro-
lithic assemblages of the Lower Palaeolithic. Other
procedures, including preparation of distal end and
side edges, are less frequent. The techniques of core
exploitation show an evident and recurrent ten-
dency. In Lower Palaeolithic microlithic assem-
blages, early exploitation started generally from
single- or double-platform cores (Fig. 5). Change
of flaking direction was the main technique of
adjusting the angle of core exploitation—used fre-
quently, as indicated by the 65% of cores showing
changed orientation in the group of advanced cores,
and their equally high percentage in the group of
residual cores.
Another indication that the process of core
exploitation involved core platform preparation or
change of direction are flake butts. There are no
evident differences in the percentage proportion of
flake butts from sequences II and III in all the
analyzed inventories. In Sequence III there was a
slightly decreased ratio of corticated to natural
butts and a decrease in the percentage share of
punctiform butts. At the same time, paradoxically,
sequence II showed a higher share of facetted and
dihedral butts than in sequence III (Fig. 6).
Microlithic assemblages from the Lower Palaeo-
lithic show a greater similarity in the percentage parti-
cipation of butts. Next to percussion cones, scars and
wavy rings are considered a diagnostic feature of core
exploitation technique. Other features taken into
account in determining flake shape include transverse
and longitudinal cross-section. Once the flake shape is
defined, the next step is to assess the degree of mod-
ification during tool production (Fig. 3).
The main attribute used in defining flakes is their
shape; along with flake proportions, it is an impor-
tant feature helping distinguish blades among the
flakes. The basic categories are represented by irre-
gular flakes, considered as the most characteristic
for early lithic industries, followed by parallel,
diverging, converging, oval, and segmented flakes.
Contrary to recurring views, the flaking technique
during the Lower Palaeolithic was not random.
Final core exploitation is documented by a group of
residual cores, core fragments, and fragments of
rejected flakes, partly cortical or entirely without cor-
tex.Cores in this sequence are typically small in size and
show a marked degree of exploitation. The high per-
centage of fragmented cores and diverse residual forms
is most probably the effect of using the direct technique
of percussion with a hard hammer, and the bipolar
technique, which do not allow for proper control,
resulting in a substantial quantity of waste (Table 1).
All tools and production wastes are taken into
consideration in this sequence. We defined as tools
all specimens showing evidence of retouch, i.e.,
flakes, cores, and chunks. Many tools were only
partly retouched, repaired, and damaged. In keep-
ing with the principles of DTA, the degree of mod-
ification of the debitage was determined together
with the statistical extent of this modification.
In comparison to the preceding stages, sequence
V is represented by only a small number of artifacts.
In the study of Palaeolithic assemblages, it is com-
mon to calculate the ratios between tools, cores, and
flakes (Fig. 4). In the examined Lower Palaeolithic
microlithic assemblages, tools made up no less than
15% all lithic artifacts at Bilzingsleben, nearly 13%
at Trzebnica 2LH, 10% at Rusko 33, 8% at Trzeb-
nica 2 UH and Quarto delle Cinfonare, and a mere
4% at Rusko 42 (Burdukiewicz 2003a, 2003b,
2006).
The criteria of flake selection for tool production
are less easily understood. Analysis was made of the
relationship between core, flake, and tool size and
of the change in the frequency of flakes and tools in
terms of their form and their transverse and long-
itudinal cross-sections. Statistical analysis deter-
mined important differences in the size of cores,
flakes, and tools. In all of the analyzed microlithic
assemblages, length, width, and thickness medians
were higher for tools than for flakes. A particularly
great difference between tools and flakes could be
noted with regard to the thickness median.
Next to the criterion of size, which was evidently
taken into consideration when selecting flakes for
tool production, there may have been a preference
for flakes of a specific form. Certain differences were
observed in the frequency of specimen forms. Flake
204 J.M. Burdukiewicz
form modification by means of retouch, or even the
very selection of flakes of a preferred form, testifies
to the deliberate selection of preforms (Fig. 3).
On the basis of the percentage of tool forms, inven-
tories could be divided into two broad groups. One
includes material from Bilzingsleben and Rusko 33,
dominated by tools with converging edges. Rusko 42,
and Trzebnica 2LH and 2UH, on the other hand,
registered an obvious domination of forms with diver-
ging edges. Participation of tools having parallel or
oval shape is quite different than in the case of flakes.
As follows from the above discussion, differences in
the amounts of flakes and tool forms in each assem-
blage are clearly visible. Therefore, retouch led to the
modification of shapes, and consequently the distribu-
tion of flakes and tool form participation is comple-
tely different. This shows that the view proposed byC.
Peretto and his team, that retouched tools generally
represent waste, is hardly justifiable (Peretto 1994).
The demonstrated intentionality of rawmaterial selec-
tion and methods of retouch is evidently inconsistent
with the supposedly ‘‘opportunistic’’ approach of early
hominins to stone working.
The study of microlithic assemblages using the
DTA and statistical methods has made it possible to
detect the earliest indication of standardization in
stone working during the Lower Palaeolithic.
Obviously it was less developed than in the Levallois
or blade production techniques, but all the same,
observable in larger statistical samples. Also evident
is the considerable similarity of technological
sequences from Bilzingsleben and both levels at
Trzebnica (Table 1). Inventories from Rusko 33 and
Rusko 42 are slightly different, but this may be due to
the influence of postdepositional processes which led
to the redeposition of artifacts and are responsible for
removing heavier forms from sequence I and for the
low representation of sequence II. In viewof the above
discussion, the Lower Palaeolithic microlithic assem-
blages described here may best be defined as techno-
complexes in the sense proposed by D. Clarke (1968).
Organic Artifacts
Organic artifacts during the Palaeolithic are rarely pre-
served anywhere in the world, chiefly because organic
material tends to perish shortly after having been dis-
carded by its users or later, as a result of the destructive
action of postdepositional processes. At Vertesszolos
almost all bone artifacts were discovered in level 1 of
Vertesszolos site I. Most were fashioned from the long
bones of large mammals like Bovidae, Cervidae, Equi-
dae, and Proboscidae. The hunters from Vertesszolos
apparently preferred herbivores, which accounted for
as much as 92.5% of animal remains, whereas in a
natural faunal spectrum nearly 1/3 were predators.
At Bilzingsleben themain source of rawmaterial for
bone tool production was definitely Palaeoloxodon
antiquus,which represented only 12%of general faunal
inventory. It is noteworthy that juvenile individuals
with milk teeth (60%) outnumbered mature and aged
individuals (40%). Substantial variability of forms,
numerous incomplete or damaged individuals, coupled
with the as yet incomplete publication of the site, all
make a more detailed classification quite difficult. D.
Mania uses functional terms, applied to stone artifacts,
such as side scrapers, points, cleavers, chisels, hammers,
etc. (Mania and Mania 2003). Another group of
organic objects featured in large number at Bilzingsle-
ben is comprised of deer antler. Based on careful ana-
lysis, Mania was able to identify the process of antler
working, distinguishing many antler tools.
Unusually favorable post-depositional conditions at
Schoningen and Bilzingsleben assisted the survival of a
great number of worked wooden objects. The oldest of
these are four wooden pieces discovered at Schoningen
12 on the fossil lake shore, in association with numer-
ous flint artifacts. All were fashioned from silver fir and
had lengths of 12, 17, 19.1, and 32.2 cm. The shortest of
these pieces had diagonal grooves at both ends, the
other three only a single such groove (Thieme 2003).
Most probably, its purpose was to hold flint inserts,
forming a tool combining two types of raw material
(Burdukiewicz and Ronen 2003).
In 1995 excavation of level 4 at Schoningen 13 II–
4 uncovered finely preserved wooden spears. The
objects rested within a level of organic mud, covered
by a layer of peat—the dried out littoral zone of a flat
channel lake. The same area furnished 30,000 faunal
remains, including 17 well preserved skulls of young
horses (Equus mosbachensis), some of them with cut
marks. Numerous plant remains and abundant mol-
lusk fauna indicate a boreal climate, continental in
character, with coniferous forests (Thieme 2003).
The spears were recovered on the whole from a 25
by 10m zone of the largest concentration of finds. The
first three were discovered by H. Thieme; by 2003 five
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 205
more had been recorded (Thieme 2003). The spears
weighed around 500–600 g. Their average length and
weight—about 230 cm and 600 g, respectively—are
similar to those of modern women’s Olympic equip-
ment. Experiments have shown that the maximum
distance achieved with this type of spear is as much
as 75 m, but an experienced spear thrower is able to
achieve an accurate throw over a distance of up to 35
m. A large number of diverse pieces of worked wood
were discovered at Schoningen 13 II–4, but they still
await publication. Although Bilzingsleben produced a
greater variety of wooden tools, their function is not
easy to establish because of substantial damage as a
result of pos-tdepositional processes.
Pioneer Settlers of Northern Latitudes ofEurasia—Technological Transitions andAdaptations to Wooded Environment
From the time of the first investigations made at
Vertesszolos by Vertes (1965), microlithic assem-
blages have continued to pose a problem for archae-
ologists concerned with the Lower Palaeolithic,
because they did not fit easily into the traditional
culture scheme adopted at the time. Eventually, a
considerable number of similar assemblages were
recorded in Eurasia, ranging from Central Europe
to China, spanning the period from c. 1 Ma to c.
120 kaBP.According to the current state of research,
sites with microlithic artifacts appeared in Eurasia in
the following chronological order: the Levant, the
Middle and Far East, the Apennine Peninsul, and
Central Europe (cf. Derevianko et al. 2000; Burdu-
kiewicz andRonen 2003). A new site with small lithic
flakes was lastly found in Parkfield (Suffolk, United
Kingdom) in Cromerian layer dated 0.7 Ma ago
(Parfitt et al. 2005; Roebroeks 2005).
Given their substantial spatial, temporal, and eco-
logical variability, the most reasonable explanation is
that microlithic assemblages emerged from Pebble
Tool or Handaxe Technocomplexes, more or less
autonomous of each other, as a result of adaptation
to specific environmental conditions. In several sites
with microlithic artifacts, larger tools like choppers or
rare handaxes are more or less frequently present, in
particular in the southern zone, like Evron-Quarry, in
the vicinity of Quarto delle Cinfonare or Bilzingsleben
in the north. In Northern China, these associations
are still less clear.
The principal motivation for their increase was
possibly the abundance of organic material, wood
in particular, which came to be used as the main raw
material. Wooden and bone tools are easier to pro-
duce, but they are rather less effective. Innovation of
composite tools, made of wood or bone with lithic
inserts (Fig. 10), seems to be themost effective way of
tool production during the Stone Age. However,
until recently, wooden, bone, or resin hafts are
known in very limited numbers. New evidence from
Schoningen and Bilzingsleben offers sufficient proof
that composite tools were used during the Lower and
Middle Palaeolithic in Northern Eurasia around 608north latitude. The presence of several microlithic
assemblages in this zone supports such a concept. A
similar conceptwas developed entirely independently
in the Far East, where the functioning of microlithic
assemblages was probably associated with the wide-
spread use of bamboo (Keates 2003). It seems correct
to assume therefore that the Euroasiatic Microlithic
Technocomplexes developed in other environmental
conditions continuing a relationship—unspecified as
yet—with communities of Pebble Tool or Acheulean
Technocomplexes. Whatever may have been the
case, the assemblages in the two regions are partly
parallel to each other chronologically.
The situation of microlithic assemblages is remark-
able, in that their decline was multistage in character.
They are recorded during warmer and wetter periods
with prevailing woodland vegetation, and apparently
disappear during cooler periods, the time of the devel-
opment of steppe or tundra landscape. The first per-
iod of microlithic assemblage decline is noted at the
close of the Holsteinian interglacial (OIS 11), around
300 ka BP, a time of expansion in Europe of Acheu-
lean handaxes and flake tools of larger dimensions.
The second period of development dates to OIS 7, 5,
and 4, i.e., stages of the Middle Palaeolithic. Finally,
microlithic industries developed fully towards the end
of the Pleistocene and during the early Holocene.
Final Remarks
The study of the Lower Palaeolithic, the earliest
stage of human history, has made important pro-
gress during the latter half of the 20th century. The
206 J.M. Burdukiewicz
discovery of numerous hominin fossil remains and
new archaeological sites has furthered our knowl-
edge of palaeoecological conditions and climatostra-
tigraphic changes. Evidence newly recovered in Eur-
ope and northeastern China has made it necessary to
distinguish two or three taxonomic units, defined as
the Lower PalaeolithicMicrolithic Technocomplexes
in the Levant, Eastern Asia, and Europe. Their
assemblages are characterized by the domination of
microlithic technology in lithic production and the
use of microliths as inserts in composite tools. Such
invention in human history is usually seen as very
sophisticated and characteristic for Mode 5.
An important role was apparently played also by
tools from organic materials: wood, bone, and
antler in particular. Substantial spatial, chronologi-
cal, and ecological variability of microlithic assem-
blages suggests that they developed as a result of
adaptation to the conditions of the local environ-
ment, the adaptation processes presumably follow-
ing their individual courses in different areas.
Further research is needed on the mobility of homi-
nin groups who as early as around 1.8Ma appear to
have been able to travel over substantial distances in
a relatively short time, as indicated by new discov-
eries at Dmanisi (Georgia), several thousand kilo-
meters from southern and eastern Africa.
The Microlithic technocomplexes could have
developed in relative isolation from each other as
well as from other technocomplexes like Pebble
Tools (Mode 1) or Acheulean (Mode 2) in the south-
ern zone of Eurasia. In any case, these assemblages
are partly parallel to each other chronologically. In
Europe,microlithic assemblages occur in an environ-
ment with a climate ranging from theMediterranean
to the boreal, but always in association with wood-
land or woodland-steppe vegetation. No assem-
blages of similar type are recorded for colder periods.
Another interesting group of issues relates to the
beginnings of a cultural organization of the microen-
vironment, the domestication of fire, and the emer-
gence of hunting. Spears discovered at Schoningen
and surviving evidence of selective hunting now
make it possible to discard the hypothesis of the
long-lived persistence of scavenging. Exceptionally
favorable conditions for subsistence, offered by
springs in travertine areas (Bilzingsleben, Isernia),
enabled some hominin groups to occupy a single site
over a long period or to return frequently to the same
area (Vertesszolos; Schoningen 13–4). Travertine and
bog settlements, due to their exceptional properties
favoring preservation of organic materials, have con-
served traces of such occupation to this day.
The production of small artifacts, that were not
easy to hold and use with the fingers, is not easy to
explain. It is worth stressing the presence of some
bifacial tools bearing various links with Mode 2 tech-
nology. Recent excavations of worked wooden
objects in Schoningen (Lower Saxony, Germany)
and wooden sticks with diagonal grooves at the ends
suggest that they were handles to hold stone inserts,
forming tools combining two types of raw material.
As opposed to the early assemblages from Africa
and SouthernEurasia in northern latitudes, the usage
of microliths as inserts in composite tools seems to be
the most important feature. It is difficult to explain
why early forager groups changed their lithic tech-
nology. The most possible transition process can be
explained as an adaptation to new conditions of the
local environments, where technological innovations
provided survival and demographic success.
The primary motivation for the transition to a
new lithic technology seems to be the abundance of
organic material, wood in particular, which came to
be used as the main raw material for the production
of composite tools. Until recently, our record
included only a very limited number of wooden
finds, but evidence offers proof that microlithic
assemblages indeed functioned during the Lower
and Middle Palaeolithic in Northern Europe. A
similar concept was developed independently in
northeastern China and Central Asia, where the
functioning of microlithic assemblages was prob-
ably associated with the widespread use of bamboo
and wood.
References
Bar-Yosef, O., 1987, Pleistocene connections between Africaand Southwest Asia: An archaeological perspective. TheAfrican Archaeological Review 5:29–38.
Bar-Yosef,O., 1998,Early colonizations and cultural continuitiesin the Lower Palaeolithic of western Asia. In Early HumanBehaviour in Global Context, edited byM.D. Petraglia andR.Korisettar, pp. 221–279. Routledge, London.
Beinhauer, K.W., Fiedler, L., and Wegner D., 1992,Hornstein-Artefakte von der Fundstelle desHomo erectusheidelbergensis aus Mauer. In Schichten von Mauer – 85
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 207
Jahre Homo erectus heidelbergensis, edited K.W. Bein-hauer and G.A. Wagner, pp. 46–73. Edition Braus,Meinheim.
Bordes, F., 1968, The Old Stone Age. Weidenfeld andNicolson, London.
Bruhl, E., 2003,The small flint tool industry fromBilzingsleben– Steinrinne. In Lower Palaeolithic Small Tools in Europeand the Levant, edited by J.M.Burdukiewicz andA. Ronen,pp. 65–92. BAR International Series 1115, Oxford.
Burdukiewicz, J.M., 2003a, Lower Palaeolithic sites withsmall artefacts in Poland. In Lower Palaeolithic SmallTools in Europe and the Levant, edited by J.M. Burdukie-wicz and A. Ronen, pp. 65–92. BAR International Series1115, Oxford.
Burdukiewicz, J.M., 2003b, Technokompleks mikrolityczny wpaleolicie dolnym srodkowej Europy. Instytut ArcheologiiUniwersytet Wrocawski, Wroclaw (in Polish).
Burdukiewicz, J.M., 2006, Lithic artifacts – Typology, tech-nology and morphometrics. In The Stone: Technique andTechnology, edited by A. Wisniewski, T. Plonka and J.M.Burdukiewicz, pp. 11–19. UniwersytetWroclawski, Insty-tut Archeologii, Wroclaw.
Burdukiewicz, J.M. and Ronen A., 2000, Ruhama in theNorthern Negev Desert. A new microlithic site of LowerPalaeolithic in Israel. Praehistoria Thuringica 5:32–46.
Burdukiewicz, J.M. and Ronen A., 2003, Research problemsof the Lower and Middle Palaeolithic small tool assem-blages. In Lower Palaeolithic Small Tools in Europe andthe Levant, edited by J.M. Burdukiewicz and A. Ronen,pp. 235–239. BAR International Series 1115, Oxford.
Carbonell, E., Mosquera, M., Olle, A., Rodriguez, X.P.,Sahnouni, M., Sala, R., and Verg es J.M., 2001, Structuremorphotechnique de l ’industrie lithique du Pl eistoc eneinf erieur et moyen d ’Atapuerca (Burgos, Espagne).L ’Anthropologie (Paris) 105:259–280.
Cattani, L., Cremaschi, M., Ferraris, M.R., Mallegni, F.,Masini, F., Scola, V., and Tozzi C., 1991, Le gisementdu Pleistocene moyen de Visogliano (Trieste): resteshumains, industries, environnement. L’Anthropologie(Paris) 91:9–36.
Chauhan, P.R., 2004, A review of the early Acheulian evi-dence from South Asia. Assemblage – The Sheffield Grad-uate Journal of Archaeology December 2004.
Clarke, D.L., 1968, Analytical Archaeology. Methuen & CoLtd., London.
Cremaschi,M. and Perreto C., 1988, Les sols d’habitat du sitepaleolithique d’Isernia La Pineta (Molise, Italie centrale).L’Anthropologie (Paris) 93:1017–1040.
Crovetto, C., 1991, Resultats preliminaires de la comparai-son des industries d’Isernia La Pineta et de Venosa-Loreto. In Isernia La Pineta. Nuovi contributi scientifici,edited by C. Peretto, pp. 79–95. Instituto Regionale per gliStudi Storici del Molise, V. Cuoco, Isernia.
Davidson, I. and Noble W., 1993, Tools and Language inHuman Evolution. In Tools, Language and Cognition,edited by K. Gibson and T. Ingold, pp. 363–388. Cam-bridge University Press, New York.
De la Torre, I. andMora R., 2005, Technological strategies inthe Lower Pleistocene at Olduvai Beds I & II. Etudes etRecherches Archeologiques de l’Universite de Liege 112,Liege.
Debenath, A. and Dibble H.L., 1994, Handbook of Paleo-lithic Typology: Lower and Middle Paleolithic of Europe.University of Pennsylvania Museum, Philadelphia.
Derevianko, A.P., Petrin V.T., and Taimagambetov J.K.,2000, The phenomenon of microindustrial complexes inEurasia. Archaeology, Ethnology & Anthropology of Eur-asia 4:2–18.
Dominguez-Rodrigo, M., Serrallonga, J., Juan-Tresserras,J., Alcala, L., and Luque L., 2001, Woodworking activ-ities by early humans: a plant residue analysis on Acheu-lian stone tools from Peninj (Tanzania). Journal of HumanEvolution 40:289–299.
Goren-Inbar, N. and Saragusti I., 1996, An Acheulian bifaceassemblage from the site of Gesher Benot Ya’aqov, Israel:indications of African affinities. Journal of Field Archae-ology 23:15–30.
Hou, Y.M., Potts, R., Yuan B.Y., Guo, Z.T., Deino, A.,Wang, W., Clark, J., Xie, G.M., and Huang W.W.,2000, Mid-Pleistocene Acheulean-like stone technologyof the Bose Basin, South China. Science 287:1622–1626.
Keates, S.G., 2003, The role of raw material in explainingtool assemblage variability in Palaeolithic China. InLower Palaeolithic Small Tools in Europe and the Levant,edited by J.M. Burdukiewicz and A. Ronen, pp. 149–168.BAR International Series 1115, Oxford.
Kretzoi, M. and Dobosi, V.T., (eds.) 1990, Vertesszo´ lo ´ s,Site, Man and Culture. Akademiai Kiado, Budapest.
Mania, D. and Mania U., 2003, Bilzingsleben – Homo erec-tus, his culture and his environment. The most importantresults of research. In Lower Palaeolithic Small Tools inEurope and the Levant, edited by J.M. Burdukiewicz andA. Ronen, pp. 65–92. BAR International Series 1115,Oxford.
McNabb, J., Binyon, F., and Hazelwood L., 2004, The largecutting tools from the South African Acheulean and thequestion of social traditions. Current Anthropology45:653–677.
McPherron, S., 2000, Handaxes as a measure of the mentalcapabilities of early hominids. Journal of ArchaeologicalScience 27:655–663.
Paddayya, K., Blackwell, B.A.B., Jhaldiyal, R., Petraglia,M.D., Fevrier, S., Chaderton, II D.A., Blickstein, J.I.B.,and Skinner A.R., 2002, Recent findings on the Acheulianof the Hunsgi and Baichabl valleys, Karnataka, withspecial reference to the Isampur excavation and its dating.Current Science (Bangalore), 83/5:641–647.
Parfitt, S.A., et al. 2005, The earliest record of human activityin northern Europe. Nature 438:1008–1012.
Peretto, C., (ed.), 1994, Le industrie litiche del giacimentopaleolitico di Isernia La Pineta: LA tipologia, le tracce diutilizzazione, la seprimentazione.Cosmo Iannone Editore,Isernia.
Peretto, C., La Rossa,M., Liboni, A., Milliken, S., Sozzi,M.,and Zaratini A., 1997, Le gisement de Quarto delle Cin-fonare dans le cadre du Paleolithique inferieur de l’ItalieOuest-Centrale. L’Anthropologie (Paris) 101:597–615.
Piperno, M., (ed.), 1999, Notarchirico, un sito del Pleistocenemedio antico nel bacino di Venosa. Osanna, Venosa.
Plummer, T., Ferraro, J., Ditchfield, P., Bishop, L., and PottsR., 2001. Late Pliocene Oldowan excavations at KanjeraSouth, Kenya. Antiquity 75:809–810.
208 J.M. Burdukiewicz
Ranov, V.A. and Dodonov A.E., 2003, Small instruments ofthe Lower Palaeolithic site Kuldara and their geoarchaeo-logical meaning. In Lower Palaeolithic Small Tools inEurope and the Levant, edited by J.M. Burdukiewicz andA. Ronen, pp. 133–147. BAR International Series 1115,Oxford.
Roche, H., Delagnes, A., Burgal, J.P., Feibel, C., Kibunjia,M., Mourre, V., and Texler P.J., 1999, Early hominidstone tool production and technical skill 2.34 Myr agoin West Turkana, Kenya. Science 399:57–60.
Roebroeks, W., 2005, Archaeology: Life on the Costa delCromer. Nature 428:921–922.
Roebroeks, W. and van Kolfschoten T., 1995, The earliestoccupation of Europe: A reappraisal of artefactual andchronological evidence. In The Earliest Occupation ofEurope. Proceedings of the European Science Foundation.Workshop at Tautavel (France), 1993, edited byW.Roeb-roeks and T. van Kolfshoten, pp. 297–315. University ofLeiden, Leiden.
Ronen, A., 2003, The small tools of Evron-Quarry, WesternGalilee, Israel. In Lower Palaeolithic Small Tools inEurope and the Levant, edited by J.M. Burdukiewicz andA. Ronen, pp. 113–120. BAR International Series 1115,Oxford.
Ronen, A., Burdukiewicz, J.M., Laukhin, S.A., Winter, Y.,Tsatskin, A., Dayan, T., Kulikov, O.A., Vlasov, V.K.,and Semenov V.V., 1998, The Lower Palaeolithic siteBizat Ruhama in the Northern Negev, Israel. Archaolo-gisches Korrespondenzblatt 28:163–173.
Schild, R., 1980, Introduction to dynamic technological ana-lysis of chipped stone assemblages. In UnconventionalArchaeology. New Approaches and Goals in Polish Archae-ology, edited by R. Schild, pp. 57–85. Ossolineum,Wroclaw.
Semaw, S., Rogers, M.J., Quade, J., Renne, P.R., Butler, R.F., Dominguez-Rodrigo, M., Stout, D., Hart, W.S.,Pickering, T., and Simpson S.W., 2003, 2.6-Million-year-old stone tools and associated bones from OGS-6
and OGS-7, Gona, Afar, Ethiopia. Journal of HumanEvolution 45:169–77.
Stout,D.,Quade, J., Semaw, S.,Rogers,M.J. andLevinN.E.,2005, Raw material selectivity of the earliest stonetoolmakers at Gona, Afar, Ethiopia. Journal of HumanEvolution 48:365–380.
Thieme, H., 2003, Lower Palaeolithic sites at Schoningen,Lower Saxony, Germany. In Lower Palaeolithic SmallTools in Europe and the Levant, edited by J.M. Burdukie-wicz and A. Ronen, pp. 101–111. BAR InternationalSeries 1115, Oxford.
Urban, B., 1992, Die Rolle der Quartarbotanik und ihreBedeutung fur die Fundstelle desHomo erectus heidelber-gensis von Mauer. In Schichten von Mauer – 85 JahreHomo erectus heidelbergensis, edited by K.W. Beinhauerand G.A. Wagner, pp. 111–120. Edition Braus,Meinheim.
Vekua, A., Lordkipanidze, D., Rightmire, G.P., ReidFerring, J.A., Maisuradze, G., Mouskhelishvili, A.,Nioradze, M., de Leon, M.P., Tappen, M., Tvalchrelidze,M., and Zollikofer C., 2002, A new skull of early Homofrom Dmanisi, Georgia. Science 297:85–89.
Vertes, L., 1965, Typology of the Buda Industry, a pebble-tool industry from Hungarian Lower Palaeolithic. Qua-ternaria 7:185–195
Wynn, T., 2004, Comment on ‘‘The large cutting tools fromthe South African Acheulean and the question of socialtraditions.’’ Current Anthropology 45:672–673.
Wynn, T. and Tierson F., 1990, Regional comparison of theshapes of later Acheulean handaxes.American Anthropol-ogist 92:73–84.
Zaidner, Y., Ronen, A., and Burdukiewicz J.M., 2003, Lamicroindustrie du Paleolithique ancien de Bizat Ruhama,Israel. L’Anthropologie (Paris) 105:203–222.
Zhou, C., Liu, Z., and Wang Y., 2000, Climatic cycles inves-tigated by sediment analysis in PekingMan’s Cave, Zhou-koudian, China. Journal of Archaeological Science27:101–109.
Lower Palaeolithic Transitions in the Northern Latitudes of Eurasia 209