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Scale Insects Hemiptera: Many families

Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

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Page 1: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Scale Insects

Hemiptera: Many families

Page 2: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

‘Soft’ Scales Armored Scales

Page 3: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Some Important Armored (Hard)

Scales in Colorado

• Oystershell scale

• Pine needle scale

• Scurfy scale

• Walnut scale

• San Jose scale

• Poplar scale

Page 4: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Armored Scales Family Diaspididae

Page 5: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Oystershell Scale

Page 6: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 7: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Oystershell scale can

develop thick colonies

on the bark of

susceptible hosts

Page 8: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Bark cracking is a

common symptom

of current or

previous oystershell

scale infestation of

aspen

Page 9: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Oystershell scale infested trees. Note bark cracking.

Page 10: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Oystershell scale flipped over to exposure lower surface.

Some eggs are visible at right where the ‘test’ has torn.

Page 11: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Oystershell scale eggs

(left) and recently

hatched eggs,

producing the Instar I

“crawler” stage

Eggs, under the cover

of the mother, are the

overwintering stage.

Page 12: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

The crawler stage usually

occurs in late May and early

June. It lasts for about a

week.

It is the only mobile stage of the

oystershell scale – and all other

armored scales.

Page 13: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Within about a week the

crawlers have either ‘settled’

or died. They remain in place

where they settled for the

rest of their life.

Page 14: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Pine Needle Scale

Page 15: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 16: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 17: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Mother scale and recently laid eggs. Eggs are the

primary overwintering stage.

Page 18: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 19: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Crawlers usually occur in early to mid May (ca. lilac

full bloom)

Page 20: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 21: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 22: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Natural Enemies of Pine Needle Scale

Lady beetles that

specialize in armored

scales (Coccidophilus

spp., Chilochorus spp.)

Page 23: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Natural Enemies of Pine Needle Scale

Parasitic Wasps

Page 24: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Poplar Scale on Aspen

Page 25: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Bark cracking

(right) and bubbling

(below) associated

with poplar scale

Page 26: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Overwintering stage of poplar scale in Salida,

March 7, 2007

Page 27: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Lady beetle predators of poplar scale

Page 28: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Some Important Soft Scales in

Colorado

• European elm scale

• Cottony maple scale

• Striped pine scale

• Pine tortoise scale

• European fruit lecanium

• Soft brown scale

Page 29: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scales Families Coccidae, Eriococcidae and others

Page 30: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scale Example – Cottony Maple Scale

Page 31: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Overwintering stage –

Adult females that have

not matured eggs.

Page 32: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Females swell with

eggs in late spring

Page 33: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 34: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Crawlers move to the leaves in

summer

Page 35: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Like aphids, soft

scales suck sap from

the phloem and

excrete honeydew.

Page 36: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Overwintering stage –

Adult females that have

returned to the twigs in

late summer.

Page 37: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

European Elm Scale

Page 38: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 39: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 40: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 41: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Honeydew is excreted.

Where it lands and

persists, sooty molds

grow.

Page 42: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Branch ‘flagging’ in late summer can be due to stress

associated with high scale populations.

Page 43: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Crawlers return to twigs in late summer for overwintering.

Page 44: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Brown Soft Scale

Coccus hesperidum

Page 45: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 46: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Hemispherical Scale, Saisettia coffae

Page 47: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 48: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 49: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Crawler Treatments

Page 50: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scales vs. Armored Scales

Soft scales produce

honeydew

Armored scales do not

Page 51: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scales vs. Armored Scales

Soft scales typically

produce several hundred

eggs

Armored scales

typically produce a

couple of dozen eggs

Page 52: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scales vs. Armored Scales

Soft scales retain mobility

through their lifetime,

moving from foliage to

twigs

Armored scales are only

active during the

crawler period

Page 53: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Soft Scales vs. Armored Scales

Soft scales typically

produce crawlers for

several weeks in late spring

and early summer

Armored scales typically

produce crawlers for a

week or two in spring

Page 54: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Principles of Scale Control

• Insecticides timed to coincide with crawlers

• Oils

–Dormant season treatment

–Post-crawler treatment

• Systemic insecticides (limited)

• Hand removal

Page 55: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 56: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Crawler Treatments

Page 57: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Horticultural Oils

• Primarily act by smothering insects

• Derived from various sources

– Petroleum/Mineral oils

– Vegetable oils (cottonseed, soybean)

– Neem seed extracts

– Fish oils

Page 58: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Petroleum Derived Horticultural Oils (Mineral oils, paraffinic oils)

Page 59: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

‘Soft’ Scales Armored Scales

Dormant season (dormant oil) applications

of oils tend to work more consistently on

soft scales

Page 60: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Many current

horticultural oils can

be used on trees

with foliage.

Crawlers and young

settled scales are

targets.

Page 61: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

‘Soft’ Scales Armored Scales

Systemic insecticides applied to the soil

for root uptake are primarily effective

against phloem-feeding soft scales

Page 62: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Scraping

oystershell

scale

Page 63: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated
Page 64: Scale Insects - pdfs.semanticscholar.org€¦ · Honeydew is excreted. Where it lands and persists, sooty molds grow. Branch ‘flagging’ in late summer can be due to stress associated

Principles of Scale Control

• Insecticides timed to coincide with crawlers

• Oils

–Dormant season treatment

–Post-crawler treatment

• Systemic insecticides (limited)

• Hand removal