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Rhoeadales:Rhoeadales:Disbanded!Disbanded!
REBECCA S. FRANKLINREBECCA S. FRANKLIN
Laboratory of Tree-Ring Research, Department of Geosciences, University Laboratory of Tree-Ring Research, Department of Geosciences, University of Arizona, Tucson, AZ, 85721of Arizona, Tucson, AZ, 85721
[email protected]@ltrr.arizona.edu
Outline
• Location of families in Rhoeadales in Engler (1936) v. Cronquist (1988) v. APG II (2003)
• Description of families in Engler’s Rhoeadales, unifying characteristics. (many not covered in class)
• Regrouping in Cronquist, APG II
Engler & Diels 1936• Division Embryophyta• Sub-division Angiospermae• Class Dicotyledonae
• Order Rhoeadales• Families
– Papaveraceae– Fumariaceae– Capparidaceae (Capparaceae)– Brassicaceae– Tovariaceae– Resedaceae– Moringaceae– Bretschneideraceae
The disbanding and regrouping
Family Engler (1936) Cronquist (1988) APG II (2003)
Papaveraceae Rhoeadales Papaverales Ranunculales
Fumariaceae Rhoeadales Papaverales Ranunculales
Capparidaceae (Cappadaceae)
Rhoeadales Capparales Brassicales (in Brassicaceae)
Brassicaceae Rhoeadales Capparales Brassicales
Tovariaceae Rhoeadales Capparales Brassicales
Resedaceae Rhoeadales Capparales Brassicales
Moringaceae Rhoeadales Capparales Brassicales
Bretschneideraceae Rhoeadales Sapindales Brassicales (in Akaniaceae)
Engler’s morphological circumscription :
• Families have :– Cymose or panniculate inflorescences– Highly dissected simple leaves (compound)– Numerous stamens
Cronquist (1988)
• Papaverales– Papaveraceae
– Fumariaceae• 2 sepals• 3-aperature pollen • Absence of ethereal
oils• Recent origin• Isoquinoloine alkaloids
• Capparales– Capparaceae
– Brassicaceae
– Tovariaceae
– Resedaceae
– Moringaceae• Compound leaves• Parietal placentation• Presences of
glucosinolates• Mustard oil!• Hypogynous
• Sapindales– Bretschneideraceae
• Compound/cleft leaves• Haplostemonous/diplostemo-
nous androecium• Well developed nectary disk• Syncarpous ovary (1-2
ovules per)• (Char indiv. In Rosales but
not all together)
Angiosperm Phylogeny Group II (2003)• Ranunculales
– Papaveraceae
– Fumariaceae• (Berberidaceae)• (Ranunculaceae)
• Combined Ranunculales + Papaverales
• Herbaceous• Toothed/lobed/compound
leaves• Presence of alkaloids
(isoquinoline type)• Hypogynous, parts free and
distince• Numerous stamens• Seeds w/tiny embryos• Copious endosperm
• Brassicales– Capparaceae (in
Brassicaceae)
– Brassicaceae
– Tovariaceae
– Resedaceae
– Moringaceae– Bretschneideraceae
(in Akaniaceae)
• Monophyly of Brassicales and Capparales based on chloroplast sequence
• Mustard oils! (glucosinolates)• 4-merous flowers,
tetradynamous stamens• Seeds with curved/folded
embryos• Lacking/nearly lacking
endosperm– (5-merous straight embryos,
parietal plac, endosperm in Moringaceae & Akariaceae)