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© 1996 Nature Publishing Group http://www.nature.com/naturegenetics

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© 1996 Nature Publishing Group http://www.nature.com/naturegenetics

© 1996 Nature Publishing Group http://www.nature.com/naturegenetics

© 1996 Nature Publishing Group http://www.nature.com/naturegenetics

© 1996 Nature Publishing Group http://www.nature.com/naturegenetics