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Occurrence Of Fishes At 38 Sites in the Mississippi Embayment Study Unit, 1996-98
By B.G. Justus and B.J. Caskey
U.S. Geological Survey
Open-File Report 99-605
National Water-Quality Assessment Program
Pearl, Mississippi
2000
U.S DEPARTMENT OF THE INTERIOR
BRUCE BABBITT, Secretary
U.S. GEOLOGICAL SURVEY
Charles G. Groat, Director
Any use of trade, product, or firm names is for identification purposes only and does not constitute endorsement by the U.S. Geological Survey.
For additional information write to:
District Chief U.S. Geological Survey 308 South Airport Road Pearl, MS 39208-6649
Copies of this report can be purchased from:
U.S. Geological Survey Branch of Information Services Box 25286 Denver, CO 80225-0286
Information regarding the National Water-Quality Assessment (NAWQA) Program is available on the Internet via the World Wide Web. You may connect to the NAWQA Home Page using the Universal Resource Locator (URL) at:
http://wwwrvares.er.usgs. gov/nawqa/nawqa_home.html
FOREWORD
The mission of the U.S. Geological Survey (USGS) is to assess the quantity and quality of the earth resources of the Nation and to provide informa tion that will assist resource managers and policymak- ers at Federal, State, and local levels in making sound decisions. Assessment of water-quality conditions and trends is an important part of this overall mission.
One of the greatest challenges faced by water- resources scientists is acquiring reliable information that will guide the use and protection of the Nation's water resources. That challenge is being addressed by Federal, State, interstate and local water-resource agencies and by many academic institutions. These organizations are collecting water-quality data for a host of purposes that include: compliance with permits and water-supply standards; development of remedia tion plans for a specific contamination problem; opera tional decisions on industrial, wastewater, or water- supply facilities; and research on factors that affect water quality. An additional need for water-quality infor mation is to provide a basis on which regional and national-level policy decisions can be based. Wise decisions must be based on sound information. As a society we need to know whether certain types of water-quality problems are isolated or ubiquitous, whether there are significant differences in conditions among regions, whether the conditions are changing over time, and why these conditions change from place to place and over time. The information can be used to help determine the efficacy of existing water-quality policies and to help analysts determine the need for and likely consequences of new policies.
To address these needs, the Congress appropri ated funds in 1986 for the USGS to begin a pilot pro gram in seven project areas to develop and refine the National Water-Quality Assessment (NAWQA) Pro gram. The NAWQA Program builds upon an existing base of water-quality studies of the USGS, as well as those of other Federal, State, and local agencies. The objectives of the NAWQA Program are to:
Describe current water-quality conditions for a large part of the Nation's freshwater streams, rivers, and aquifers.
Describe how water quality is changing over time.
Improve understanding of the primary natural and human factors that affect water-quality conditions.
This information will help support the development and evaluation of management, regulatory, and monitoring decisions by other Federal, State, and local agencies to protect, use and enhance water resources.
The goals of the NAWQA Program are being achieved through ongoing and proposed investigations of more than 50 of the Nation's most important river basins and aquifer systems, which are referred to as study units. These study units are distributed through out the Nation and cover a diversity of hydro-geologic settings. More than two-thirds of the Nation's freshwa ter use occurs within the more than 50 study units, and more than two-thirds of the people served by public water-supply systems live within their boundaries.
National synthesis of data analysis, based on aggregations of comparable information obtained from the study units, is a major component of the program. This effort focuses on selected water-quality topics using nationally consistent information. Comparative studies will explain difference and similarities in observed water-quality conditions among study areas and will identify changes and trends and their causes. The first topics addressed by the national synthesis are pesticides, nutrients, volatile organic compounds, and aquatic biology. Discussions on these and other water- quality topics will be published in periodic summaries of the quality of the Nation's ground and surface water as the information becomes available.
This report is an element of the comprehensive body of information developed as part of the NAWQA Program. The program depends heavily on the advice, cooperation, and information from many Federal, State, interstate, Tribal, and local agencies and the public. The assistance and suggestions of all are greatly appreciated.
Robert M. Hirsch Chief Hydrologist
iii
CONTENTSPage
Abstract.................................................................................................................................................... 1
Introduction............................................................................................................................................. 1
Purpose and Scope................................................................................................................................... 1
Acknowledgments................................................................................................................................... 2
Site selection criteria and site information.............................................................................................. 2
Methods of sampling and processing ...................................................................................................... 2
Results..................................................................................................................................................... 3
References............................................................................................................................................... 4
ILLUSTRATION
Figure 1. Map showing locations of 38 sites sampled in the Mississippi Embayment during 1996-98 5
TABLES
Table 1. Information for 38 fish sampling sites in the Mississippi Embayment Study Unit............. 6
2. Scientific and common names of fishes collected from 38 sites in the Mississippi
Embayment Study Unit, 1996-98 ..................................................................................... 7
3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment
Study Unit, 1996-98.......................................................................................................... 9
4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment
Study Unit, 1996-98.......................................................................................................... 17
CONVERSION FACTORS AND ABBREVIATIONS
Divide By To obtain
cubic meter per second (m3/s)
centimeter (cm)
meter (m)
meter per second (m/s)
f\
square kilometer (km)
0.02832
0.4
3.28
3.28
2.59
cubic foot per second (ft3/s)
inch (in.)
foot (ft)
foot per second (ft/s)
square mile (mi2)
The following acronyms are used in this report:
MAP Mississippi Alluvial PlainMISE Mississippi Embayment Study UnitNAWQA National Water-Quality AssessmentUSGS U.S. Geological Survey
VI
Occurrence of Fishes at 38 Sites in the Mississippi Embayment Study Unit, 1996-98
By B.C. Justus and B.J. Caskey
Abstract
Fishes were sampled at 38 sites (58 sampling reaches) in the Mississippi Embayment Study Unit in parts of Arkansas, Kentucky, Louisiana, Mississippi, Missouri, and Tennessee from 1996 through 1998 as part of the National Water-Quality Assessment Program. Fishes were collected by seining and electrofishing during low- flow periods from late spring to late summer. All fishes collected were identified to the lowest possible taxon (usually species), weighed, measured, and examined for anomalies. A total of 95 taxa, representing 94 species and 18 families, were collected during the study. All 94 species were known to occur in the study area. The number of species collected at each of the sampling reaches ranged from 13 to 37.
INTRODUCTION
In 1994, the U.S. Geological Survey (USGS) began a study of the Mississippi Embayment Study Unit (MISE) as part of the National Water-Quality Assessment (NAWQA) Program (fig. 1). The NAWQA Program is designed to assess status and trends in the quality of the Nation's water resources and to determine the natural and human factors affecting these resources (Hirsch and others, 1988). The Program will eventually integrate physical, chemical, and biological data from more than 50 study units across the Nation.
Fish community structure is one aspect of the biological data collected as part of the NAWQA Program. Most of the MISE overlies the Mississippi Alluvial Plain (MAP) Ecoregion, an area where fish communities have not been intensively sampled.
The MAP Ecoregion has been extensively altered by human activity; about 75 percent (or about 16 million acres) of the original forested wetlands has been cleared and drained (Nature Conservancy, 1992) and most streams in the MAP Ecoregion have some degree of channel and hydrological modifications. Land use is dominated by agriculture (85.1 percent) and is used for growing row crops, such as cotton, corn, and soybeans, and small grains, such as rice and wheat.
The climate of the MISE is characterized as humid, with southern parts being subtropical and northern parts being temperate. Mean annual temperatures range from about 64 °F in the south to about 57 °F in the north. Annual precipitation ranges from about 56 inches in the south to about 48 inches in the north (U.S. Department of Commerce, 1995).
Purpose and Scope
This report documents the occurrence of fishes sampled at 38 sites in the MISE (36 of which are in the MAP Ecoregion) from 1996 to 1998. These and other ecological data collected by NAWQA can be used to help describe water-quality conditions and improve understanding of the factors that affect water quality.
INTRODUCTION 1
Acknowledgments
The authors would like to acknowledge three museum curators for their willingness to verify identification of specimens: Dr. Neil Douglas at the University of Louisiana at Monroe; Dr. Todd Slack at the Mississippi Museum of Natural Sciences, and Dr. Steve Ross at the University of Southern Mississippi. Additional thanks are extended to Dr. Neil Douglas and Dr. Todd Slack for archiving museum specimens.
SITE SELECTION CRITERIA AND SITE INFORMATION
In 1995, eight sites were selected for which fish community sampling was planned from 1996 to 1998. The eight sites (which were located on seven streams in Arkansas, Kentucky, Louisiana, Mississippi, Missouri, and Tennessee, and within the MISE boundary) were selected because they either integrated most types of land use in the MISE or were representative of a specific land use in the MISE, collectively provided spatial coverage of the MISE, and had long-term gage records. Six of the eight sites and five of the seven streams are located in the MAP Ecoregion; of the two remaining sites, one site is located in the Mississippi Valley Loess Plains Ecoregion, and the other is located in the Southeastern Plains Ecoregion. As planned, fish communities were sampled annually from 1996 to 1998 at the eight sites.
In 1998, fish communities also were sampled at 30 sites that had not been previously sampled by NAWQA (table 1). These 30 sites were added to increase spatial coverage of the MAP Ecoregion. Two criteria were used for selection of these 30 additional sites: first, ten sites were chosen to represent a gradient of crop intensity for each of three major crops grown in the MAP Ecoregion~corn, rice, and cotton. Secondly, as with the first eight sites sampled, the remaining sites were selected that provided the best spatial coverage of the MAP Ecoregion. County-level land use information for 1995 and 1996 was used to
determine crop intensities. Photographs and maps showing the sampling locations at each of the 38 sampling sites can be viewed at
<http://ms.water.usgs.gov/misenawqa/>.
METHODS OF SAMPLING AND PROCESSING
Stream reaches, or lengths of the stream where sampling was to be done, were designated at each of the 38 sites before sampling. To designate reaches, a visual assessment of the stream was made, and lengths physically representative of the stream were measured and marked for sampling. A reach of stream 500-m long was selected for sampling at 36 sites; samples were collected from a 250-m reach at each of two small streams (LaGrue Bayou near Dewitt, Arkansas, and Second Creek near Palestine, Arkansas) because fallen trees and beaver dams restricted access.
Three separate 500-m reaches were sampled at two of the eight sites sampled in 1996 (the Bogue Phalia at Leland, Mississippi, and the Cache River at Cotton Plant, Arkansas) to assess sampling variability; consequently, fish were sampled from 12 stream reaches that year. Data from these samples are included in table 3 and 4.
In 1997 and 1998, only one reach was sampled at each of the two sites where three reaches were sampled in 1996. Because the three reaches at each of the two sites were physically similar, accessibility was the main consideration for selecting which reach would be sampled for both years. At the Bogue Phalia at Leland, Mississippi, the most upstream reach (reach A) was sampled. At the Cache River at Cotton Plant, Arkansas, the most downstream reach (reach C) was sampled. Only one reach was sampled at each of the 36 remaining sites in 1998.
Fishes were collected by seining and electrofishing during low-flow periods from late spring to late summer. In general, methods were consistent with NAWQA fish
2 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
sampling protocols (Meador and others, 1993).
Seining consisted of sampling available habitats with a 4-m x 2-m seine having a mesh size of 0.5 cm. In streams having some areas that were wadeable, seining was done by two people wading and pulling the seine through the water. Areas that were unwadeable but near-shore were seined by two people standing facing each other about 2 m apart at the edge of the water. Each person held the seine by the top of one of the seine-poles with the bottom of the poles resting on the bank about 1.5 m from the edge of water. On a signal, the seine was swung overhead, with the lead-line on the outer edge of the arc, and into the water as far away from the bank as possible. After the lead-line sank to the stream bottom, the seine was retrieved. The time spent seining at each site was approximately 45 minutes, and the number of seining efforts (hauls) ranged from 6 to 9. After each seine haul, specimens were placed in 10 percent formalin. All fishes collected by seining were taken to a lab at the USGS office in Pearl, Mississippi, where they were identified to the lowest possible taxon (usually species), weighed, measured, and examined for anomalies.
At 57 of the 58 reaches sampled, both banks of the sampling reach were sampled by electrofishing (one bank at the Skuna River was not sampled in 1996 because of equipment problems). Electrofishing was done using a 4.7-m X 1-m aluminum boat with a commercially manufactured electrofishing unit consisting of a 2,500-watt generator and a pulsator.
An electrofishing team consisted of a boat driver and a person positioned on the bow of the boat to net fish. For sites where the water was clear enough that samplers could see fish habitat (submerged woody debris was the dominant habitat type), the boat was maneuvered downstream and along the bank until the electrodes were positioned near habitat, at which time electrofishing began. At sites that lacked fish habitat or where fish habitat was not visible due to turbid water, the boat was maneuvered in the
same manner as above but with the electrofishing gear in continual use.
Samples from each side of the stream were collected and processed independently of each other. Electrofishing time averaged about 21 minutes per bank sample. Fish large enough were identified to the lowest taxon possible (usually species), and were weighed, measured, examined for anomalies, and then released. At sites where there were large numbers of fish, the time that the fish were out of water (and fish mortality) was reduced by subdividing the sampling reach and processing fish from each subreach immediately after they were collected. Fish that were too small for positive identification, or too small to be weighed and measured in the field, were preserved in 10 percent formalin and taken to the USGS office in Pearl, Mississippi for processing.
Standard quality-assurance and quality- control procedures were taken to ensure that the fish data were of high quality (Walsh and Meador, 1998). Common and scientific names reported were those established by the American Fisheries Society's Committee on Names of Fishes (Robins and others, 1991). Although most identifications were made by the study unit biologist; some individuals were of a size or species that made them challenging to identify. To ensure data quality, those specimens were also identified by curators of fish museums in Louisiana and Mississippi. Voucher specimens are stored in fish collections at two museums: the Museum of Natural Sciences in Jackson, Mississippi; and the Museum of Natural History (Zoology) at the University of Louisiana at Monroe in Monroe, Louisiana.
RESULTS
A list of the fishes collected from 1996 to 1998 is provided in table 2. A total of 95 fish taxa representing 94 species and 18 families were collected at the 38 sites (table 3). The literature indicates that all 94 species were known to occur in the study area (Douglas, 1974; Pfleiger, 1975;
RESULTS 3
Robison and Buchanan, 1988; Etnier and Starnes, 1993). The number of taxa collected at the sites for all sampling dates ranged from 13 at Bogue Phalia near Leland, Mississippi in 1996 to 37 at Little River Ditch near Morehouse, Missouri in 1996. The number of individuals collected on all sampling dates is provided; however, it should be noted that immature or damaged individuals were not identified beyond the family or genus level (table 3). Data in table 3 were used to calculate percent relative abundance of fishes at all taxonomic levels on all sampling dates (table 4).
REFERENCES
Douglas, N.H., 1974, Freshwater fishes of Louisiana: Claitor's Publishing Division, Baton Rouge, La., 443 p.
Etnier, D.A., and Starnes, W.C., 1993, The fishes of Tennessee: The University of Tennessee Press, Knoxville, Tenn., 681 p.
Hirsch, R.M., Alley, W.M., and Wilbur, W.G., 1988, Concepts for a National Water-Quality Assessment Program: U.S. Geological Survey Circular 1021, 42 p.
Meador, M.R., Cuffney, T.F., and Gurtz, M.E., 1993, Methods for sampling fish communities as part of the National Water-Quality Assessment Program: U.S. Geological Survey Open-File Report 93-104,40 p.
Nature Conservancy, 1992, The forested wetlands of the Mississippi River: An ecosystem in crisis, The Nature Conservancy, Baton Rouge, LA, 25 p.
Pflieger, W.L., 1975, The fishes of Missouri: Missouri Department of Conservation, 343 p.
Robins, C.R., Bailey, R.M., Bond, C.E., Brooker, J.R. Lachner, E.A., Lea, R.N., and Scott W.B., 1991, Common and scientific names of fishes from the United States and Canada: Bethesda, Md., American Fisheries Society, Special Publication 20,183 p.
Robison, H.W., and Buchanan, T.M., 1988, Fishes of Arkansas (5th ed.): The University of Arkansas Press, Fayetteville, Ark., 536 p.
U.S. Department of Commerce, 1995,Monthly station normals of temperature, precipitation, and heating and cooling degree days 1961-1990: Climatography of the United States No. 81, National Oceanic and Atmospheric Administration, National Climatic Data Center, Asheville, North Carolina.
U.S. Geological Survey, 1999,http://ms.watcr.usgs.gov/misenawqa. Homepage to the Mississippi Embayment National Water-Quality Assessment Program.
Walsh, S.J., and Meador, Michael R., 1998, Guidelines for quality assurance and quality control of fish taxonomic data collected as part of the National Water- Quality Assessment Program: U.S. Geological Survey Open-File Report 98-4239, 33 p.
4 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
ILLINOIS
KENTUCKY
TENNESSEE
ARKANSAS MISSISSIPPI
LOUISIANA
EXPLANATIONMississippi Embayment Study Unit
Mississippi Alluvial Plain Ecoregion
Streams
Base fromU.S. Geological Survey digital data, 1:2,000,000 Geographic projection
(3J>) Sampling site and number
0 40 MILES
0 40 KILOMETERS
Figure 1. Locations of 38 sites sampled in the Mississippi Embayment Study Unit during 1996-98.
Tabl
e 1.
Inf
orm
atio
n (li
sted
by
site
num
ber)
for
38 fi
sh s
ampl
ing
site
s in
the
Mis
siss
ippi
Em
baym
ent
Stu
dy U
nit [
km2,
squa
re k
ilom
eter
s; m
3/s, c
ubic
met
ers
per s
econ
d; m
, met
ers;
m
/s, m
eter
s pe
r sec
ond]
.en 8 o c 3 3 rn z o
rn Tl
C/3 rn C/3
U O)
C/3 1 Z ^ rn c/5 C/3 w
w TJ 2 m > m H C/3
O C H CO i
No.
of
Site
nam
e S
ite
time
s S
tatio
n nu
mbe
r sa
mpl
ed
num
ber
St.
John
s D
itch
near
Sik
esto
n, M
OLi
ttle
Riv
er D
itch
no.
1 ne
ar M
oreh
ouse
, M
OS
pillw
ay D
itch
at H
wy
102
near
Eas
t Pra
irie,
MO
Littl
e R
iver
Ditc
h no
. 25
1 ne
ar L
ilbou
m,
MO
Obi
on C
reek
nea
r H
ickm
an,
KY
Mai
n D
itch
at H
wy
153
near
Whi
te O
ak,
MO
Run
ning
Ree
lfoot
Bay
ou a
t H
wy
103,
TN
Elk
Chu
te n
ear G
oble
r, M
OC
ockl
e B
urr
Slo
ugh
Ditc
h ne
ar M
onet
te,
AR
St.
Fran
cis
Riv
er a
t La
ke C
ity, A
R
Cac
he R
iver
at E
gypt
, AR
Vill
age
Cre
ek n
ear
Sw
ifton
, A
RT
yron
za R
iver
nea
r Tw
ist,
AR
St.
Fran
cis
Riv
er n
ear
Col
dwat
er, A
RB
ayou
DeV
iew
at
Mor
ton,
AR
Sec
ond
Cre
ek n
ear
Pal
estin
e, A
RW
olf R
iver
at
LaG
rang
e, T
NL'
Ang
uille
Riv
er n
ear
Pal
estin
e, A
RC
ache
Riv
er n
ear
Cot
ton
Pla
nt, A
RBi
g C
reek
at
Pop
lar
Gro
ve,
AR
_
LaG
rue
Bay
ou n
ear
Dew
itt,
AR
Col
dwat
er R
iver
at
Mar
ks,
MS
Bay
ou M
eto
near
Bay
ou M
eto,
AR
Cas
sidy
Bay
ou a
t Web
b, M
SS
kuna
Riv
er a
t Bru
ce,
MS
Qui
ver
Riv
er n
ear
Dod
dsvi
lle,
MS
Big
Sun
flow
er R
iver
at
Sun
flow
er,
MS
Bog
ue P
halia
nea
r Le
land
, M
SB
ayou
Mac
on n
ear
Hal
ley,
AR
Dee
r C
reek
nea
r H
olan
dale
, M
S
Boe
uf R
iver
nea
r Ark
ansa
s/LA
Sta
te L
ine,
LA
Big
Sun
flow
er R
iver
nea
r Ang
uilla
, M
SS
teel
e B
ayou
Eas
t P
rong
nea
r R
ollin
g Fo
rk,
MS
Silv
er C
reek
nea
r B
ayla
nd,
MS
Yaz
oo R
iver
bel
ow S
teel
e B
ayou
nea
r Lo
ng L
ake,
MS
Ten
sas
Riv
er a
t Ten
dal,
LAB
ayou
Mac
on n
ear
Del
hi,
LABi
g C
reek
nea
r S
ligo,
LA
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38
1 3 1 1 1 1 1 1 1_
_ 1 3 1 1 1 1 1 3 1 5 1 1 1 1 1 3 1 1 5 1 1 1 1 1 1 3 3 1 1
0704
3300
0704
3500
0702
4160
0704
2500
0702
3800
0704
1120
0702
7050
0704
6515
0704
0496
0704
0450
0707
7380
0707
4660
0704
7700
0704
7520
0707
7700
0704
7947
0703
0392
0704
7950
0707
7555
0707
7950
0707
8040
0727
9950
0726
5099
0728
0900
0728
3000
0728
8570
0728
8500
0728
8650
0736
7659
507
2887
70
0736
7700
0728
8700
0728
8870
0728
8720
08_
0728
8955
0736
9500
0737
0000
0736
8580
Latit
ude
3656
0836
5003
3644
5436
3320
3638
58
3619
2736
0944
3610
1835
5139
3549
16_
3551
2835
4910
3522
2935
2152
3515
07
3502
2135
0157
3458
2035
0207
3433
20
3419
0034
1522
3412
0533
5659
3358
25
3338
2533
3250
3323
4733
3216
3308
59
3258
2532
5818
3254
4132
5208
3226
40
3225
5532
2725
3212
20
Longitu
de
8933
0289
4348
8921
1989
4012
.890
721
9000
2089
3036
8957
3490
1949
_902
556
9056
0091
0505
9028
0590
3436
9106
37
9054
4089
1448
9053
1091
1919
9050
44
9116
5790
1557
9131
4590
2028
8920
50
9024
0590
3235
9050
4791
1736
9050
47
9126
2590
4640
9057
1090
4145
9054
00
9122
0091
2830
9149
11
Dra
inag
e ba
sin
area
(k
m2)
101
1,14
418
662
778
4
356
751
218
146
6,15
0
1,81
641
01,
367
13,7
741,
081
111
543
1,98
32,
996
1,16
0
594
4,93
72,
078
536
668
651
2,01
01,
301
376
231
1,82
26,
675
1,12
247
.934
,850
721
2,14
11,
311
Per
cent
o
f ba
sin
in
agricu
lture
*79 61 81 87 32 88 37 95 96 28 78 92 92 59 73 65 14 77 79 77 71 43 55 85 19 81 81 80 85 81 83 78 81 56 41 74 78 76
Dis
char
ge
at g
age
(m3/
s)2.
222.
960.
933.
601.
22
2.20
0.68
0.70
3.31
11.0
8.84
4.71
4.59
36.1
4.61
2.52
5.04
8.69
14.6
6.33
0.00
90.7
0.00
2.41
0.49
5.77
15.0
5.35
9.80
0.00
6.17
46.0
2.88
0.00
405
2.35
6.80
1.56
Mea
n ch
anne
l w
idth
(m
)10
.933
.710
.522
.412
.5
19.9
13.9
11.7
53.1
24.6
21.7
115.
819
.058
.928
.8
15.1
15.6
24.6
35.0
17.8
8.1
37.5
25.5
62.2
37.4
16.3
23.2
37.7
14.6
19.2
45.0
89.8
49.6
20.4
91.4
19.3
55.4
48.4
Mea
n ch
anne
l de
pth
(m)
0.52
0.35
0.40
0.73
0.84
0.33
0.16
0.20
1.06
2.90
2.34
0.74
0.81
1.27
2.62
0.73
1.29
2.09
2.00
2.21
0.61
4.60
3.54
0.85
1.18
1.65
2.19
1.34
1.20
1.71
2.32
3.98
2.02
0.13
.5.9
2
1.26
2.02
1.75
Mea
n in
stan
tane
ous
velo
city
(m
/s)
0.25
0.13
0.15
0.27
0.17
0.36
0.26
0.14
0.08
0.45
0.18
0.11
0.21
0.46
0.06
0.28
0.30
0.18
0.22
0.23
0.00
0.50
0.00
0.04
_0.2
6__
0.24
0.56
0.09
0.45
0.00
0.07
0.15
0.08
0.00
0.49
0.26
0.08
0.02
* In
clud
es a
ll ar
eas
used
for
the
prod
uctio
n of
cro
ps s
uch
as c
orn,
soy
bean
s, v
eget
able
s, t
obac
co,
and
cotto
n, a
s w
ell a
s sm
all g
rain
s su
ch a
s w
heat
and
ric
e.
Exc
lude
s ar
eas
used
for
the
prod
uctio
n of
hay
and
pas
ture
.
Table 2. Scientific and common names of fishes collected from 38 sites in the Mississippi EmbaymentStudy Unit, 1996-98, listed in phylogenetic order [Rare, occurs at 2 or less (approximately 5 percent) of the sites sampled; common, occurs at 19 or more (50 percent) of the sites sampled; 1, indicates species was rare or common; 0, indicates the species was not rare or common; (34), indicates the number of sites where the species was collected]
Scientific namePetromyzontidae - lampreys
Petromyzontidae speciesLepisosteidae - gars
Lepisosteus oculatus (Winchell, 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820
Amiidae - bowflnsAmiacalva Linnaeus, 1766
Hiodontidae - mooneyesHiodon alosoides (Rafinesque, 1819)
Anguillidae - freshwater eelsAnguilla rostrata (Lesueur, 1817)
Clupeidae - herringsDorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)
Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venusta Girard, 1856Cyprinus carpio Linnaeus, 1758Hybognathus hayi Jordan, 1885Hybognathus nuchalis Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbratilis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storer/ana (Kirtland, 1847)Notemigonus crysoleucas (Mitchill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856)Notropis volucellus (Cope, 1865)Opsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque, 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchill, 1818)
Catostomidae - suckersCarp/odes carp/o (Rafinesque, 1820)Carp/odes cyprinus Lesueur, 1817Erimyzon obtongus (Mitchill, 1814)Hypentelium nigricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cyprinellus (Valenciennes, 1 844)Ictiobus niger (Rafinesque, 1819)Minytrema melanops (Rafinesque, 1820)Moxostoma macrolepidotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)
Ictaluridae - bullhead cattrshesAmeiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819)
Common name
lamprey
spotted garlongnose garshortnose gar
bowfin
goldeye
american eel
gizzard shadthreadfin shad
grass carpbluntface shinerred shinerspotfin shinerblacktail shinercommon carpcypress minnowmississippi silvery minnowribbon shinerredfin shinerspeckled chubsilver chubgolden shinerorangefin shinerpallid shineremerald shinerghost shinerironcolor shinertaillight shinerweed shinermimic shinerpugnose minnowsuckermouth minnowbluntnose minnowfathead minnowbullhead minnowcreek chub
river carpsuckerquillbackcreek chubsuckernorthern hogsuckersmallmouth buffalobigmouth buffaloblack buffalospotted suckershorthead redhorseblacktail redhorse
black bullheadyellow bullhead
Rare
1(1)
00
0
0
1(1)1(1)
00
000100000
1(1)0
1(2)0
1(1)1(2)
00
1(1)
1(2)1(2)
00
1(1)0
1(1)0
1(2)
00
1(2)1(1)
0000
1(2)1(1)
00
Common
0
1(35)0
1(30)
0
0
0
1(35)0
0000
1(24)
1(35)0000
00000000000
0000
1(25)0
0000
1(32)1(19)1(24)
000
00
Introduced
0
000
0
0
0
00
100001000000000000000000100
0000
000000
00
Table 2. Scientific and common names of fishes collected from 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued.
Scientific nameIctaluridae - bullhead catfishes - - continued
Ictalurus furcatus (Lesueur, 1840)Ictalurus punctatus (Rafinesque, 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846
Aphredoderidae - pirate perchesAphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - kllllfishesFundulus chrysotus (Gunther, 1866)Fundulus notatus (Rafinesque, 1820)Fundulus olivaceus (Storer, 1845)
Poeciliidae - livebearersGambusia affinis (Baird and Girard, 1853)
Atherinidae - silversidesLabidesthes sicculus (Cope, 1865)Menidia beryllina (Cope, 1866)
Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensis Jordan and Eigenmann, 1887Morone saxatilis (Walbaum, 1792)
Centrarchidae - sunfishesAmbloplites ariommus Viosca, 1936Elassoma zonatum Jordan, 1877Lepomis cyanellus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829)Lepomis humilis (Girard, 1858)Lepomis macrochirus Rafinesque, 1819Lepomis marginatus (Holbrook, 1 855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus Evermann, 1899Lepomis hybridMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802)Pomoxis annularis Rafinesque, 1818Pomoxis nigromaculatus (Lesueur, 1829)
Percidae - perchesAmmocrypta beani Jordan, 1877Ammocrypta Clara Jordan and Meek, 1885Ammocrypta vivax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1 881 )Etheostoma fusiforme (Girard, 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare (Hay, 1881)Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Percina maculate (Girard, 1859)Percina sciera (Swain, 1883)
Sciaenidae - drumsAplodinotus grunniens Rafinesque, 1819
Common name
blue catfishchannel catfishtadpole madtombrindled madtomfreckled madtombrown madtomflathead catfish
grass pickerel
pirate perch
golden topminnowblackstripe topminnowblackspotted topminnow
western mosquitofish
brook silversideinland silverside
white bassyellow bassstriped bass
shadow bassbanded pygmy sunfishgreen sunfishwarmouthorangespotted sunfishbluegilldollar sunfishlongear sunfishredear sunfishredspotted sunfishsunfish hybridspotted basslargemouth basswhite crappieblack crappie
naked sand darterwestern sand darterscaly sand dartermud darterbluntnose darterswamp darterslough darterharlequin dartercypress darterspecked darterlogperchblackside darterdusky darter
freshwater drum
Rare
000
1(1)1(2)1(1)
0
0
0
1(2)00
0
01(2)
000
01(2)
00000000
1(2)0000
1(1)1(1)
000
1(1)0
1(2)0
1(1)1(1)
00
0
Common
01(30)
0000
1(19)
0
0
000
1(37)
00
000
00
1(27)1(30)1(30)1(34)
01(32)
0000
1(35)1(32)1(24)
0000000000000
1(37)
Introduced
0000000
0
0
000
0
00
000
000000000000000
0000000000000
0
8 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 [96,1996; A, one of three reaches sampled; , no individuals collected]
II
SCIENTIFIC NAMEPetromyzontidae - lampreys
Petromyzontidae species Lepisosteidae - gars _~^_^,
Lepisosteus oculatus {Winchell, 1864) Lepisosteus osseus {Linnaeus, 1758) ^^^~ Lepisosteus platostomus Rafinesque, 1820 Lepisosteus species
Amiidae - bowfins Amia calva Linnaeus, 1766 flHH^
Hiodontidae - mooneyes Hiodon alosoides {Rafinesque, 1819)
Anguiiiidae - freshwater eels Anguilla rostrate (Lesueur, 1817) ^Hflffc
Clupeidae - herrings Dorosoma cepedianum {Lesueur, 1818) ~ ~ Dorosoma petenense {Gunther, 1867) Dorosoma species
Cyprinidae - carps and minnows Cteriopharyngodon idella (Valenciennes, 1844) Cyprinella camura {Jordan and Meek, 1884) Cyprinella lutrensis {Baird and Girard, 1853) Cyprinella spitoptera {Cope, 1868) Cyprinella venusta Girard, 1856 _^|HB Cyprinus carpio Linnaeus, 1758
Hybognathus hayi Jordan, 1885 Hybognathus nuchalis Agassiz, 1855 *^^BIB Lythrurus fumeus {Evermann, 1892) Lythrurus umbratilis {Girard, 1856) Macrhybopsis aesSvalis {Girard, 1856) Macrhybopsis storeriana {Kirtland, 1847) Notemigonus ciysoleucas {Mitchill, 1814) Notropis ammophilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896
Notropis maculatus (Hay, 1881) Notropis texanus (Girard, 1856) ! Notropis volucellus :Co^e, 1865 ^^^^^^
Opsopoeodus emiliae hay, 1881 Phenacobius mlrabllis (Girard, 1856) Pimephales notatus {Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vigilax {Baird and Girard, 1856! St.', "jr.;, , acu/afus (Mitchill, 1818)
Catostomidae - suckers CarpKXtes carpio (Rafinesque, 1820) __ Carptodes cyprinus Lesueur, 1817
Erimyzon species Hypentelium nigncans (Lesueur, 181?) ^^^HK Ictiobus bubalus {Rafinesque, 1818) ^^^^^ Ictiobus cypnnellus (Valenciennes, 184tt^m^p. Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820) Moxostoma macrolepidotum (Leuseur, 1817) Moxostoma poecilurum (Jordan, 1877)
Catostomidae species
96 A
8
-
30 32
-
-
._
-
-
-
-
-
-
6
-
-
96 B
7
8
33 17
-
-
I
-
-
-
-
-
-
5
1
96 C
15
4
5 14
-
-
-
-
-
~
-
10
4
-
4
97
14
3
1 1
1
43 4
-
17
-
-
-
-
-
10
-
2
96
6
6
38
164
5 25 5
-
7
-
-
~-
24 1 4
-
-
96 A
4 2 3
1
8 14
9
-
_
-
-
6
-
-
5
-
-
96 B
6
2
-
26 11
-
-
-
-
8
-
-
21
-
-
96 C
7
5
3
1
2 12 2 2
-
1
-
-
7
-
-
2
-
97
4
4
3
41 15
10
1
178
1
1
16
_
-
7 1
-
-
98
3 1 1
3
17
21 17 1
72
5
I
-
-
5
1
-
14 3 6
-
-
96
42 7
77 36
30
-
_
-
-
-
-
-
62 5 1
-
97
3 22
35
21 14
-
3
-
-
-
_
-
48 1 4
-
-
98
3
5
4
2 13
-
-
-
-
6
-
-
36 10 22
-
-
96
2
10
193 9
3 1
38
21
1
751
g
-
6
2
I
-
97
4
1
70 5
1
26
17
53
2 3 1
5
-
98
3
3
122 6
5
15
1
18
29
13 _
1 5
2
-
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 Continued
SCIENTIFIC NAMEIctaluridae - bullhead catflshes
Ameiurus me/as (Rafinesque, 1820)
Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Rafinesque, IB 18) ^^HBV Noturus gyrinus (Mitchill, 1817)
Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocldae - pikes Esox americanus vermiculatus Lesueur, 1846
Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - kiliif ishes jgffE*.
Fundulus notatus (Rafinesque, 1820) jjffgf Fundulus olivaceus (Storer, 1845)
Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853) ^ff^^
Atherinldae - silversides Labidesthes sicculus (Cope, 1865) ^^^Bi
Menidia beryllina (Cope, 1B66) Percichthyidae - temperate basses
Morons chrysops (Rafinesque, 1820)
Morons saxatilis (Walbaum, 1792) Centrarchldae - sunfishes
Amb/oplites ariommus Viosca, 1936
Lepomis cyanellus Rafinesque, 1819 Lepomisgulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotls (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis mlniatus Evermann, 1899
Lepomis species Micropterus punctulatus (Rafinesque, 1819) Mlcroptarus salmoides iLaca^ede, 1802) Micropterus species Pomoxls annularis Rafinesque, 1818 ^g^^^p Pomoxis nigromaculatus (Lesueur, 1829)
Percidae - perches
Ammocrypta clara Jordan and Meek, 1885
Etheostoma asprigene (Forties, 1878) Etheostoma chlorosomum (Hay, 1881)
Etheostoma gracile (Girard, 1859)
Etheostoma proeliare (Hay, 1881) ^^^^ Etheostoma stigmaeum (Jordan, 1877)
Percina maculata (Girard, 1859)
Sciaenidae - drumsAplodinotus gwnniens Rafinesque, 1819
Unclassified fishes Unknown fry
96 A
1
8
2
-
1
~
:1 1
34
3
3
»
-
-
-
7
96 B
-
4
1
-
-
::4 12 3
1
-
-
-
-
-
-
4
<D96 C
--
1
3
-
3
_
1 4 25 9
1
7
-
-
-
-
-
19
97
-
4
1
-
31
~
1 102 19
1
1
6
5
-
-
-
-
-
17
98
-
1
-
-
53
~
:5 2 19 13
1
526
1 1
37 4
-
-
-
1
-
7
196 A
-
2 2
1
3
1
5
:7 7
2
1
3 2
8 4
-
2
-
-
-
10
96 B
-
2
4
1
1
:2 3 9
1
1
9
-
-
-
-
-
8
o96 C
-
67
2
-
-
__
4 3 1 4 1 4
8 2
161
-
2
-
-
8
97
-
3 2
2
1
5
34
~~
::3 2
4
2
2
11
-
41
-
~
1
tt
2
98
-
9 2
4
11
7
56
::10 10 15
26
3
5
27
-
1
3
-
-
26
1
I96
-
3
8
-
~
9 1
7
1 2
9 1
-
-
-
-
-
11
0"
97
-
2
1
-
35
~~
'_
5 17 9
2
2
15 2
-
6
-
-
5
12
98
-
7
5
-
136
_
:2
14
2
108
6
-
8
-
-
4
4
96
-
8
2
1
4 2
2
_
13
14 5 3 16
35
12
1 9 67
1
1 1
-
4
1
10
w97
-
6
1
3
3
~
1
10
11 3 1 5
35
6
1 8 5
1
-
-
-
-
-
5
98
-
6
3
1
m? --T
12
6 -^m.
-
>
23 2
19
64
2
5 16 10
1
-
-
-
-
1
11
10 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAME'" Petromyzontidae - lampreys
Petromyzontidae species Leplsosteidae - gars
Lepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820
Amildae - bowfins Amutcalva Linnaeus, 1766
Hiodontidae - mooneyesHiodon alosoides (Rafinesque, 1819)
Anguillidae - freshwater eelsAnguilla rostrate (Lesueur, 1817)
Clupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) ^ ""soma species
Cyprinidae - carps and minnows
Cyprinella camura (Jordan and Meek, 1884) Cyprinella lutransis (Baird and Girard, 1853)
Cyprinella veriusta Girard, 1856 Cyprinus carpio Linnaeus, 1 758 Cyprinidae speciesHybognathus hayi Jordan, 1885
Lythrurus fumeus (Evermann, 1892) Lythrurus umbratilis (Girard, 1856)
Macrhybopsis storariana (Kirtland, 1847) Notemigonus crysoleucas (Mitchiil, 1814)
Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856) Notropis volucellus (Cope, 1865)
Opsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856) Pimephales notatus (Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchiil, 1818)
Catostomidae - suckersCarp/odes carpio (Rafinesque, 1820)
Erimyzon oblongus (Mitchiil, 1814)
Hypentelium nigricans (Lesueur, 1817) ictiobus bubalus (Rafinesque. 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820)
Moxostoma poecilurum (Jordan, 1877)
96
2 1
-
8
-
251 1
-
-
10
-
-
28
-
1 1
-
c?97
9 1 1
-
2
6
1007
-
1
23 1
-
-
18
1
1 1
-
98
7 1 1
-
11
9
541 5 1
3
1
9
-
1
1
26 1
1
22
-
96
7
8
-
14
1
5 2
-
-
1
-
-
-
-
15 1
5
-
1
197
8
3
-
2
-
8 21
-
-
18
-
-
-
-
30
1
-
98
4
2
-
5
2
2
-
-
18
-
-
7
-
29
3
--
/96
3
1
15
57
1
223 3
-
1 11
-
1
2
r
2
4
197
7
-
3
60
1
3
-
3
-
-
1
-
1
1
5
98
1
-
6
23
91
-
1
-
-
:
-
4 1
1
6
3
96
2 1 14
31
5584
-
_
-
1
12
-
-
-
-
2
-
I97
1 2 2
-
19 60
-
4
-
-
14 3
1
1
1
-
2
-
98
2 2
-
171 836
2
3
-
22
161 1
5
1
-
2
-
/I98
15 1 1
-
12
12
-
27 2
16
-
1
13
-
3
-
29 9 3
-
/I / 03
98
1
-
8
25
2 3
-
-
-
--
-
30
-
-
-
/I / °>
98
3
4
-
23 15 15
1
302
18 11
-
-
1 1
--
-
29
-
41 1 8
-
/I / °>
98
11 4 3
-
28
195
2
18
9
-
22
-
-
2
2
-
29 6
2
-
/I98
7
2
1
21
-
16 11
2
-
2
-
-
15
1
-
27
4
-
11
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEIctalurldae - bullhead catfishes
Ameiurus me/as (Rafinesque, 1820) Amelurus natalls (Lesueur. 1819) Ictalurus furcatus (Lesueur. 1840) Ictalurus punctatus (Rafinesque. 1818) Noturus gyrinus (Mitchill, 1817) Noturus miurus Jordan, 1877 Noturus nocturnus Jordan and Gilbert. 1886 Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikes Esox americanus vermiculatus Lesueur. 1 846 j|} Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)
Cyprinodontldae - kllllflshes Fundulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845) Fundulus species
Poeclllldae - livebearers Gambusia affinis (Baird and Girard, 1853 1
Atherinidae - sllversldes Labidesthes slcculus (Cope, 18651 Menidia beryllina (Cope, 1866)
Percichthyidae - temperate basses Morone chrysops (Ra*inesque, 1820) Morone mississippiensis Jordan and Morone saxatilis (Walbaum, 1792)
Centrarchldae - sunfishes Ambloplites ariommus Viosca, 1936 Elassoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalobs (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis mmiatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomoxis species
Percldae - perches Ammocrypta beam Jordan, 1877 Ammocrypta tiara Jordan and Meek, 1885 Ammocrypta vivax Hay. 1882 Etheostoma asprigene (Fortes, 1878) Etheostoma chlorosomum (Hay, 1881) Etheostoma fusiforme (Girard, 1854) Etheostoma gracile (Girard, 1859) Etheostoma histrio Jordan and Gilbert, 1887 Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877) Percina caprodes (Rafinesque, 1818) Percina maculata (Girard, 1859) Percina sctera (Swain, 1883)
Sciaenidae - drums Apkxtnolusgrunniens Rafinesque 1H1Q
Unclassified fishes
/ £96
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6 21 9
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34 18
128
147
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12 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98~Continued
SCIENTIFIC NAMEPetromyzontidae - lampreys
Lepisosteidae - garsLepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820
Amiidae - bowfins /Am/a calva Linnaeus, 1766
Hiodontidae - mooneyes
Anguiiiidae - freshwater eeis
Ciupeidae - herringsDorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867)
Cyprinidae - carps and minnows
Cyprinella lutrensis (Baird and Girard, 1853)
Cyprinella venusta Girard, 1856 Cyprinus carpio Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885
Lythrurusfumeus (Evermann, 1892)
Macrhybopsis aestivalis (Girard, 1856)
Notemigonus crysoleucas (Mitchill, 1814)
Notropis atherinoides Rafinesque, 1818 Notropis buchanani Meek, 1896 Notropis chalybaeus (Cope, 1869)
Notropis volucellus (Cope, 1865)
Opsopoeodus emiliae Hay, 1881
Pimephales notatus (Rafinesque, 1820)
Pimephales vigilax (Baird and Girard, 1856)Semotilus atromaculatus (Mitchill, 1818)
Catostomidae - suckers Carp/odes carpio (Rafinesque, 1820) Carpiodes cyprinus Lesueur, 1817
Ictiobus bubalus (Rafinesque, 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820)
1 S1
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19 13 10
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18 5
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196
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118
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19 9
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1 24
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20 30
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8
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177
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I ^98
3
1
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2061
10
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11
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598
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6
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-
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49
-
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11
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1 31 2
-
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-
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52 9 6
/£98
5 1 1
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91 12E1
-
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56
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73
24 1
14 1 4
2
*98
4 1
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-
5
79 4
9
1
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43
3
9
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27
-
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6
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41
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73
14 10
-
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127
-
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18
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°98
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6
14
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3 37
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1
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1 12
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92 10 6
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Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEIctalurldae - bullhead catfishes
Ameiurus me/as (Rafinesque, 1820) Ameiurus natatis (Lesueur, 1819) Ictalurus furcatus (Lesueur, 1840) ICtalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817)
Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocldae - pikes Esox americanus vermiculatus Lesueur, 1846
Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - killifishM Funuulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845)
Poeciliidae - llvebearersGambus/a affinis (Baird and Girard, 1853)
Atherinldae - silversides Labidesthes sicculus (Cope, 1865)
Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)
Morone saxatilis (Walbaum, 1792) Centrarchidae - sunfishes
Ambloplites ariommus Viosca, 1936
Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819
Lepomis megalotis (Rafinesque, 1820) Lepomis microlophus (Gunther, 1859) Lepomis miniatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802)
Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829)
Percldae - perches
Ammocrypta Clara Jordan and Meek, 1 885 Ammocrypta vivax Hay, 1882 Etheostoma asprigene (Forbes, 1878) Etheostoma chlorosomum (Hay, 1881)
Etheostoma gracile (Girard, 1859)
Etheostoma proeliare (Hay, 1881)
Percina maculata (Girard, 1859)
Sclaenidae - drums /(psxftwlu* grunniens Rafinesque, 1819
Unclassified fishes
98
2
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1
217
1
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2 3 6
12
4 9 15
161
-
-
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2
1
98
2 1
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2
124
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59
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51
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15
98
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6 6 16 16
2
66
1
111
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13
98
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215
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6 3 5 10
13
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8
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14
98
1 1 4 1
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6
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98
1 6
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72 3
134 32 6 7 13
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9
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14 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEPetromyzontidae - Lamprey*
Petromyzontidae speciesLepisosteidae - gars
Lepisosteus oculatus (Winchell, 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820Lepisosteus species
Amiidae - bowfinsAm&calva Linnaeus, 1766
Hiodontidae - mooneyes
98
6
3
98
-
-
2
1
Hiodon alosoides (Rafinesque, 1819) ^ miBI -Anguillidae - freshwater eels
Anguilla rostrata (Lesueur, 1817)Clupeidae - herrings
Dorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)Dorosoma species
Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venuste Girard, 1856 -: ,;Cyprinus carpio Linnaeus, 1758Cyprinidae species ~^^^IKHybognathus hayi Jordan, 1885Hybognathus nuchalls Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbrahlis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storeriana (Kirtland, 1847)Notemigonus crysoleucas (Mitchill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869 iNotropis maculatus (Hay, 1881)Notropis texanus (Girard, 1856)Notropis volucellus (Cope, 1865)Notropis speciesOpsopoeodus em///ae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque, 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)S»'V.T,,'!,"= .,tromaculatus (Mitchill, 1818)
Catostomidae - suckersCarp/odes carp/o (Rafinesque, 1820)Carp/odes cyphnus Lesueur, 1817Erimyzon oblongus (Mitchill, 1814)Erimyzon speciesHypentelium nigricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cypnnellus (Valenciennes 1844Ictiobus niger (Rafinesque. 1819)Ictiobus speciesMinytrema melanops (Rafinesque, 1820)Moxostoma macrolep'dotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)Moxostoma speciesCatostomidae species
19
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8 - - - 91 - -
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5221
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12 8 - ---
98
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7
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654 - - - - -
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98
-
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8186 14 t
1 5
93
- - -
1112
1 - --
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1 ~
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1920 20 - -_ ~
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4 13 --
98
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3 - -
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98
4
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98
14
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179643
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1999
4 - -
93 -- - 1 -
1
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-- -
- -_- -- - 2 - - -- -
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15
Table 3. Number of individuals for fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98~Continued
SCIENTIFIC NAMEIctaluridae - buiihead catfishM .
Ameiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819) ^jiSifIctalurus furcatus (Lesueur, 1840)Ictalurus punctatus {Rafinesque. 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846T, Aphredoderidae - pirate perchesAphreooderus sayanus (Gilliams, 1824)<T Cyprinodontidae - kiliifishesFunoulus chrysotus (Gunther, 1866)Fundulus notatus {Rafinesque, 1820)Fundulus divaceus (Storer, 1845) ^^^^^Fundulus species ifffff
Poeciiiidae - livebearers Gambusia affims (Baird and Girard, 1853)
Atherinidae - siiversidesLabidesthes sicculus (Cope, 1865) flHHfrMenidia beryllina (Cope, 1866)
Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensis Jordan and Eigenmann,Morone saxatilis (Walbaum, 1792)
Centrarcnidae - sunfishesAmbloplites ariommus Viosca, 1936Elassoma zonatum Jordan, 1877 ^^f^^fLepomis cyanelus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829) ^^f^fLepomis humiis (Girard, 1858)Lepomis macrochirus Rafinesque, 1819Lepomis marginatus (Holbrook, 1855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus Evermann, 1899Lepomis hybridLepomis speciesMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802) ____Micropterus speciesPomoxis annularis Rafinesque, 1818^^HBPomoxis nigromaculatus (Lesueur, 1829)Pomoxis species
Percidae - perchesAmmocrypta beam Jordan, 1877Ammocrypta Clara Jordan and Meek, 1885Ammocrypta vivax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1881)Etheostoma fusiforme (Girard. 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare {Hay, 1881)Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Percina maculata (Girard, 1859)Percina sciera (Swain, 1883) flBHB
Sciaenidae - drumsAplodinotus gr unniens Rafinesque, 1819
Unclassified fishesUnknown fry
/ co98
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16 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 [96, 1996; A, one of three reaches sampled; -, no individuals collected; 0, value less than .005]
SCIENTIFIC NAMEPetromyzontidae - lampreys
Leplsosteldae - garsLepisosteus oculatus (Winchell, 1864) Lepisosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820 Lepisosteus species
Amiidae - bowfins Amia calva Linnaeus, 1766
Hiodontidae - mooneyes
Anguillidae - freshwater eelsAnguilla rostrata (Lesueur, 1817)
Clupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther. 1867) Doroxmia species
Cyprinidae - carps and minnows
Cyprinella lutrensis (Baird and Girard, 1853) Cyprinella spiloptera (Cope, 1868) Cyprinella venusta Girard, 1856 Cyprinus carpio Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885 Hybognathus nuchahs Agassiz, 1855
Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814)Notropls ammopbilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropls atherinoides Rafinesque, 1818 Notropis buchanani Meek. 1896 Notropis chalybaeus (Cope, 1869} Notropis maculatus (Hay, 1881)
Notropis volucellus (Cope, 1865)
Opsopoeodus emillae Hay, 1881 Phenacobius mlrabilis (Girard, 1856)
Pimephales promelas Rafinesque, 1820 Plmephales vigilax (Baird and Girard, 1856) Semotilus atromaculatus (Mitchill, 1818)
Catostomldae - suckers Carp/odes carpio (Rafinesque, 1820)
Erimyzon oblongus (Mitchill, 1814)
Ictiobus bubalus (Rafinesque. 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819)
/96 A
5.84
-
21.90 24.62
-
-
-
-
-
4.38
96 B
6.93
7.92
32.67 16.83
-
0.99
-
-
-
4.95
nQQ
#96 C
12.00
3.20
4.00 11.20
-
-
-
-
-
8.00
3.20
*9n
97
4.93
1.06
0.35 0.35
0.35
15.14 1.41
-
5.99
-
-
-
3.52
n7n
98
0.63
0.63
4.00
17.25
0.53 2.63 0.53
-
0.11
0.11
-
-
2.52 0.11 0.42
/96 A
3.57 1.79 2.68
089
7.14 12.50
8.04
-
-
-
-
5.36
4.46
96 B
5.22
1.74
E
22.61 9.57
-
-
-
-
6.96
18.26
O96 C
5.98
4.27
2.56
0.85
1.71 10.26 1.71 1.71
-
0.85
-
-
5.98
3.42
1.71
97
1.27
1.27
0.95
12.97 4.75
3.16
0.32
0.32 24.68
0.32
0.32
5.06
2.22 0.32
98
0.78 0.26 0.26
0.78
4.39
5.43 4.39 0.26 18.60
1.29
0.26
-
-
1.29
0.26
3.62 0.78 1.55
/96
1.44 0.72 2.53
27.80 13.00
10.83
-
-
-
-
-
22.38 1.81 0.36
o"97
1.27 0.84 0.84
14.77
8.86 5.91
-
1.27
-
-
-
20.25 0.42 1.69
98
0.79
1.32
1.06
0.53 3.43
-
-
-
-
1.58
9.50 2.64 0.53 n CQ
/96
0.38
1.88
36.35 1.69
0.56
7.16
3.95
0.19
14.12 0.19
1.69
1.13
0.38
0.19
o97
1.37
0.34
23.89 1.71
0.34
8.87
5.80
-
18.09
0.68 1.02
1.71
98
0.70
0.70
-
28.50 1.40
-
1 17
3.50
0.23
4.21
6.78
3.04
0.23 1.17
0.47
17
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEIctaluridae - bullhead catfishes
Ameiurus natalis (Lesueur, 1819) Ictalurus furcat us (Lesueur, 1840) Ictalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817)
Noturus noctumus Jordan and Gilbert. 1886Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikes
Aphredoderidae - pirate perchesAphredoderus sayanus (Gilliams, 1824)
Cyprmodontidae - kiliifishes
Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845)
Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853)
Atherinidae - siiversides Labidesthes sicculus (Cope, 1865)
Percichthyidae - temperate basses
Centrarchidae - sunfishes
E/assoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819
Lepomis megalotis (Rafinesque, 1820)
Lepomis miniatus Evermann, 1899
Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomoxis species
Percidae - perches
Ammocrypta Clara Jordan and Meek, 1885
Etheostoma asprigene (Forbes, 1878) Etheostoma chtorosomum (Hay, 1881)
Etheostoma gracile (Girard, 1859)
Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877)
Percina sciera (Swain, 1883) Sciaenidae - drums
Aplodinotus grunniens Rafinesque, 1819 Unclassified fishes
Unknown fry
96 A
0.73
5.84
-
1.46
-
-
0.73
0.73 0.73 24.82
2.19
-
2.19
-
-
-
-
511
96 B
3.96
-
0.99
-
-
-
3.96 11.88 2.97
-
0.99
-
-
-
-
-
3.96
I
196 C
0.80
-
2.40
-
-
2.40
0.80 3.20 20.00 720
-
-
080
5.60
-
-
-
-
12.00
'
97
1.41
-
0.35
-
-
10.92
0.35 35.92 6.69
0.35
-
0.35
2.11
1.76
-
-
-
-
5.99
98
011
-
-
-
-
5.57
0.53 0.21 2.00 1.37
011
-
55.31
011 0.11 3.89 0.42
-
-
-
0.11
0.74
96 A
1.79 1.79
-
0.89
2.68
0.89
4.46
6.25 6.25
1.79
0.89
2.68 1.79
7.14 3.57
-
1.79
-
-
8.93
96 B
1.74
-
3.48
-
087
087
1.74 2.61 7.83
-
-
0.87
0.87
7.83
-
-
-
-
6S6
k$
C?96 C
5.13 5.98
-
-
1.71
-
-
3.42 2.56
3.42
3.42
6.84 1.71
13.68 0.85
-
1.71
-
-
6.84
-
97
0.95 0.63
-
0.63
0.32
1.58
10.76
0.95 0.63
1.27
0.63
0.63
3.48
-
12.97
-
-
032
5.70
0.63
98
2.33 0.52
-
1.03
2.84
1.81
14.47
2.58 2.58 3.88
6.72
0.78
1.29
6.96
-
0.26
0.78
672
0.26
/96
1.08
-
2.89
-
-
0.36
3.25 036
2.53
-
0.36 0.72
3.25 0.36
-
-
-
-
3.97
J
I97
0.84
-
0.42
-
-
14.77
2.11 0.42 2.95 3.80
0.84
-
0.84
6.33 0.84
-
2.53
-
-
2.11
5.06
ID
98
1.85
-
1.32
-
-
35.88
0.53
3.69
0.53
-
28.50
1.58
-
2.11
-
-
A f)f>
1.06
96
1.51 0.19
0.38
0.19
-
0.75 0.38
0.38
2.45
2.64 094 0.56 3.01
6.59
2.26
0.19 1.69 1.13 1.32
0.19
0.19 n *IQ
-0.75
0.19
1.88
1 if
.1 -j97
2.05
-
0.34
-
1.02
t.02
0.34
3.41
3.75 1.02 0.34 1.71
11.95
2.05
0.34 2.73 1.71
0.34
-
-
-
-
1 71
98
1.40
-
0.70
0.23
3.04 0.93
2.80
1.40
5.37 0.47
4.44
14.95
0.47
1.17 3.74 2.34
0.23
-
-
-
-
0.23
2.57
18 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEPetromyzontidae - lampreys
Petromyzontidae species Lepisosteidae - gars
Lepisosteus oculatus (Wmchell. 1864) Leplsosteus osseus (Linnaeus, 1758) Lepisosteus platostomus Rafinesque, 1820
Amiidae - bowf ins Amia calva Linnaeus, 1766
Hiodontidae - mooneyes Hiodon alosoides (Rafinesque, 1819)
Anguiiiidae - freshwater eeis Anguilla rostrate (Lesueur, 1817)
Ciupeidae - herringsDorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) Dorosoma species
Cyprinidae - carps and minnows
Cyprinella camura (Jordan and Meek, 1884) Cypnnella lutrensis (Baird and Girard, 1853)
Cyprinella venusta Girard, 1858 Cyprinus carpio Linnaeus, 1758 Cyprinidae species
Lythrurus fumeus (Evermann, 1892) Lythrurus umbratilis (Girard, 1858)
Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814)
Notmpis athennoides Rafinesque, 1818 Notropis buchanam Meek. 1896
Votrop/s volucellus (Cope, 1865)
Opsopoeodus emiliae Hay, 1881
Pimephales vigilax (Baird and Girard, 1856)
Catostomidae - suckers
Carpiodes cypnnus Lesueur, 1817
Hypentelium nigricans (Lesueur, 1817) Wiobus bubalus (Rafinesque, 1818) 'ctiobus cyprinellus (Valenciennes. 1844) 'ctiobus niger (Rafinesque, 1819)
Minytrema metenops (Rafinesque, 1820)
Woxostoma poecilurum (Jordan, 1877)
96
0.60 0.30
-
2.40
-
75.15 0.30
-
-
2.99
-
-
8.38
-
0.30 0.30
-
-
J
197
3.61 0.40 0.40
-
0.80
2.41
40.16 2.81
-
0.40
9.24 0.40
-
-
7.23
0.40
0.40 0.40
-
-
i
98
1.00 0.14 0.14
-
1.56
1.28
76.96 0.71 0.14
0.43
0.14
1.28
-
0.14
0.14
3.70 0.14
-
0.14
3.13
--
--
96
7.14
8.16
-
14.29
1.02
5.10 2.04
-
-
1.02
-
-
-
-
15.31 1.02
5.10
-
1 n?
1 <t #
197
4.57
1.71
-
1.14
-
4.57 12.00
-
-
10.29
-
-
-
-
17.14
0.57
-
-
i>
98
1.60
0.80
-
2.00
0.80
0.80
-
-
7.20
-
-
2.80
-
11.60
1.20
-
-
96
0.76
0.25
3.82
14.50
0.25
56.74 0.76
-
0.252.80
-
0.25
0.51
0.25
051
-
1.02
ID
1
197
3.68
-
1.58
31.58
0.53
1.58
-
1.58
-
-
0.53
-
0.53
0.53
2.63
98
0.44
-
2.62
10.04
;;39.74
-
0.44
-
-
-
-
1.75 0.44
0.44
2.62
1.31
96
0.47 0.23 3.27
724
12.85 19.63
-
-
0.23
2.80
-
-
-
-
0.47
-
-
£s
I97
0.39 0.78 0.78
-
7.39 23.35
-
1.56
-
-
545 1.17
0.39
0.39
0.39
-
0.78
-
-
98
0.10 0.10
-
8.96 43.79
0.10
0.16
_
1.15
8.43 0.05
0.26
-
0.05
-
0.10
-
-
/J //
98
2.28 0.15 0.15
-
1.82
1.82
_
4.10 0.30
2.43
-
0.15
1.98
-
0.46
-
4.41 1.37 0.46
-
-
- / *£
198
0.32
2.57
8.04
0.64 0.96
-
-
-
-
-
9.65
-
-
-
-
7J I98
0.48
0.64
-
3.67 2.40 2.40
0.16
4824
2.88 1.76
-
-
0.16 0.16
--
--
4.63
1 OR
6.55 0.16 1.28
-
-
7J //
98
2.18 0.79 0.60
-
556
38.69
0.40
3.57
1.79
-
4.37
-
-
0.40
-
D.40
5.75 1.19
0.40
-
> / 9
It/<?
98
2.60
0.74
0.37
7.81
-
595 4.09
0.74
-
0.74
-
-
5.58
0.37
10.04
1.49
-
-
19
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEIctaluridae - bullhead catfishes
Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Raflnesque, 1818) Noturus gyrinus (Mitchill, 1817) Noturus murus Jordan, 1877
Noturus phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikes Esox americanus vermiculatus Lesueur, 1846
Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - kiliifishes
Fundulus olivaceus (Storer, 1845)
Poeciiiidae - livebearersCambusia affmis (Baird and Girard, 1853)
Atherinidae - siiversides Labidesthes sicculus (Cope, 1865) Meriidia berylina (Cope, 1866)
Percichthyidae - temperate basses Morone chrysops (Rafinesque, 1820)
Morone saxatilis (Walbaum, 1792) Centrarchidae - sunfishes
Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotis (Rafinesque, 1820)
Lepomis species Micropterus punctulatus (Raflnesque, 1819) Micropterus salmoides (Lacepede, 1602) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829)
Percidae - perches
Etheostoma fusiforme (Girard, 1854)
Etheostoma stigmaeum (Jordan, 1877)
Percina maculata (Girard, 1859) Percina sciera (Swain, 1883)
Sciaenidae - drums Aplodinotus grunniens Raflnesque, 1819
Unclassified fishes Unknown fry
/96
-
;3.89
1.50
0.30
-
0.90
0.30 0.30
~
-
-
2.10
5 $
1CO97
2.01
;-
18.47
0.40
-
1.20 040 0.80 1.20
1.61 0.40
0.80
0.40
-
-
3.21
-
98
0.57 0.71
-
-
0.71
0.71
-
0.43
0.43 1.85
0.43
1.28
0.14
-
1.56
96
1.02
1.02
-
306
1.02
11.22 2-04
5.10
3.06 2.04
-
-
9.18
1 £p
/97
3.43
0.57
0.57
1.14
-
571
--
1.14 10.29
1.14
12.57
-
-
686
4.57
;
98
1.60 0.40
0.40
-
31.20
-
0.80 0.40 2.40
1.20
23.60
3.20 0.40
-
-
5.60
96
025
0.25
3.82
0.25
0.25
-
0.25
2.04 1.02 2.04
1.02
0.25 2.80 0.25
0.25
0.51
-
0.51
0.51
0.25
<D
1 1
97
-
0.53
0.53
11.05
1.05
-
1.58
11.58 1.05 2.11
O CO
5.26 1.05
1.05
6.84
3.68
0.53
98
-
0.44 0.44
3.49
-
-
0.44 0.44
5.68
13.10 n o."7
8.73 1.75 1.31
-
0.44
1.75
1.31
96
1.87 1.64
-
1.64
19.16
1.87
1.17
2.10 2.10
3.04
13.79 0.93
-
--
3.50
1 1
97
3.89 3.50
0.39
-
16.34
5.45
0.39
0.39
5.45
0.39
10.12
0.78
5.45
-
-
4.67
98
0.42 2.15
0.10
0.05
0.21
6.44
0.16 0.05
0.05
-
0.31 1.10 0.47 14.51
0.26
042
0.42
6.13 0.26
-
-
320
iIf
98
0.15
122
015
0.91
6.23
-
5.17 2.74 19.45
22.34
1.37
1033 0.61
3 34
030
-
380
7J /I
98
0.64
-
-
26.05
2.89
-
1.61 0.96 3.22 0.96
9.32
2605
1.61
2.89
-
-
1.61
yj /;
98
0.80
-
11.66
-
1.28 0.32 0.16
2.72
2.56
0.16
0.48
-
-
2.88
iI98
020
0.20
7.14
0.40
0.20 0.60 3.17 4.96
1.98
0.60
1.79
7.34 1.39
1.19
0.40
0.20
1.98
> / SF
/ 6
/«?98
0.37
0.74
0.37
1.12
27.51
-
0.37 3.35 0.37 5.95
5.20 0.37
1.86
4.83
9 en
1.12
0.37
2.97
20 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEPetromyzontidae - lampreys
Petromyzontidae species Lepisosteldae - gars
Lepisosteus oculatus (Winchell, 1864) Leplsosteus osseus (Linnaeus, 1758) Lepisosteus p/atostomus Rafinesque, 1820 Lephoit-u" svecies
Amiidae - bowfins Amiacalva Linnaeus, 1766
Hiodontidae - mooneyes Hiodon alosoides (Rafinesque, 1819)
Anguiiiidae - freshwater eels Angullla rostrata (Lesueur, 1817)
Ciupeidae - herrings Dorosoma cepedianum (Lesueur, 1818) Dorosoma petenense (Gunther, 1867) Dorosoma species
Cyprinidae - carps and minnows Ctenopharyngodon idella (Valenciennes, 1844) Cyprinella camura (Jordan and Meek, 1884) Cyprinella lutrensis (Baird and Girard, 1853) Cyprinella spiloptera (Cope, 1868) Cyprinella venusta Girard, 1856 Cypmus carp/'o Linnaeus, 1758 Cyprinidae species Hybognathus hayi Jordan, 1885 Hybognathus nuchalis Agassiz, 1855 Lythrurus fumeus (Evermann, 1892) Lythrurus umbratltis (Girard, 1856) Macrhybopsis aestivalis (Girard, 1856) Macrhybopsis storeriana (Kirtland, 1847) Notemigonus crysoleucas (Mitchill, 1814) Notropis ammophilus Suttkus and Boschung, 1990 Notropis amnis Hubbs and Greene, 1951 Notropis atherinoides Rafinesque 1818 Notropis buchanani Meek, 1896 Notropis chalybaeus (Cope, 1869) Notropis maculatus (Hay, 1881) Notropis texanus (Girard, 1856) Notropis volucellus (Cope, 1865) Notropis species Opsopoeodus emiliae Hay, 1881 Phenacobius mirabilis (Girard, 1856) Pimephales notatus (Rafinesque, 1820) Pimephales promelas Rafinesque, 1820 Pimephales vlgilax (Baird and Girard. 1856) Semotilus atromaculatus (Mitchill, 1818)
Catostomidae - suckers Carp/odes carpio (Rafinesque, 1820) Carp/odes cyprinus Lesueur, 1817 Erimyzon ob/ongus tMitchiil 1814) Enmyzon species Hypentelium nigricans (Lesueur, 1817) Ictiobus bubalus (Rafinesque, 1818) Ictiobus cyprinellus (Valenciennes, 1844) Ictiobus niger (Rafinesque, 1819) Ictiobus species Minytrema melanops (Rafinesque, 1820) Moxostoma macrolepidotum (Leuseur, 1817) Moxostoma poecilurum (Jordan, 1877) Moxostoma species Catostomidae species
/ «
/I s98
2.74
1.00
8.48
-
1.25
0.75 3.49
-
-
-
-
-
-
8.98
::-
0.25
0.25
-
i * isI $ $98
1.38
0.69
3.81
-
~
6.57 4.50 3.46
-
-
-
0.35
-
-
-
-
-
2.08 0.35
-
98
2.07
6.22
021
2.07
-
-
2.07
-
-
-
_
-
-
-
-
-
4.36 2.90 0.21 34.85
-
* i$ S
/ 0° / * / <8 / O
98
4.37
1.87 021
6.65
0.21
19.13
3.74 104
1.25
-
-
1.04
-
-
-
-
2.49 0.21
0.21
-
98
0.21
0.10
-
20.44
-
-
0.10 1.88
-
-
-
-
-
-
-
-
:;
2.50
-
/l/l/o198
0.370.37
0.19
1.85 0.19
-
3.52 1.67
0.19
-
1.67
2.04
20.74 0.74
-
3-33
0.74
-
0.19
0.37
0.56
019
98
0.330.66 1.32
~
0.33
-
1.32 46.86
6.60 9.90
-
0.33
0.33
0.66 2.97
-
-
-
2.64
-
0.66 0.33
-
98
1.56
-
46.09
-
2.08
-
-
0.52
-
-
-
-
-
-
? / c
^
$98
055
0.18
0.37
0.18
-
-
37.94 0.18 1.84
-
-
0.18
0.18
2.39
0.37
-
2.03
2.21
0.18
0.55
-
/ *
s 398
0.28
0.56
028
2.81
-
-
1.69
-
-
-
-
0.84
-
13.76
::
3.09 0.56 0.84
-
/ Cj98
3.700.34 0.67
-
3.70
-
0.34
0.34 10.44 0.67
-
-
0.34
1.68
-
-
-
0.34
0.67
17.51 3.03 2.02
-
//98
0.600.12 0.12
0.24
0.12
-
-
10.91 1.44 6.12
-
-
6.71
1.08
0.12
-
8.75
2.68 0.12
1.68 0.12 0.48
0.24
//98
1.370.29
-
-
-
1.46
23.03 1.17
2.62
0.29
-
12.54
0.87
2.62
0.58
7.87
-
0.58
0.29
//98
101
0.50
0.84
6.88
-
-
12.25
2.35 1.68
-
-
21.31
2.18
-
0.17
1.17
3.02
0.84
/<!98
4.19
2.99
-
3.59
8.38
-
1.80 22.16
-
060
-
-
-
0.60
5.39
0.60 7.19
//98
1.03
1.37
-
31.51 3.42 2.05
-
1.03 2.40
0.34
-
-
_
-
-
3.42
-
0.68
//98
i 1.37
0.19
0.59
-
0.20
-
-
-
-
-
-
~
21
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMtIctaluridae - bullhead catflshes
Ameiurus me/as (Rafinesque, 1820) Ameiurus natalis (Lesueur, 1819) Ictalurus furcatus (Lesueur, 1840) Ictalurus punctatus (Rafinesque, 1818) Noturus gyrinus (Mitchill, 1817) Noturus miurus Jordan, 1877 Noturus nocturnus Jordan and Gilbert, 1886 Notuws phaeus Taylor, 1969 Pylodictis olivaris (Rafinesque, 1818)
Esocidae - pikes Esox amencanus vermiculatus Lesueur. 1846
Aphredoderidae - pirate perches Aphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - kllilflshes Fundulus chrysotus (Gunther, 1866) Fundulus notatus (Rafinesque, 1820) Fundulus olivaceus (Storer, 1845) Fundulus species
Poeciiiidae - iivebearers Gambusia affinis (Baird and Girard, 1853)
Atherinidae - silversides Labidesthes sicculus (Cope, 1865) Menidia beryllma (Cope, 1866)
Percichthyidae -temperate tis-se« Morone chrysops (Rafinesque, 1820) Morone mississippiensls Jordan and Eigenmann, Morone saxatilis (Walbaum, 1792)
Centrarchidae - sunfishes AmbloplXes ariommus Viosca, 1936 Elassoma zonatum Jordan, 1877 Lepomis cyanellus Rafinesque, 1819 Lepomis gulosus (Cuvier, 1829) Lepomis humilis (Girard, 1858) Lepomis macrochirus Rafinesque, 1819 Lepomis marginatus (Holbrook, 1855) Lepomis megalotis (Rafinesque, 1820) Lepomis micro/ophus (Gunther, 1859) Lepomis minlatus Evermann, 1899 Lepomis hybrid Lepomis species Micropterus punctulatus (Rafinesque, 1819) Micropterus salmoides (Lacepede, 1802) Micropterus species Pomoxis annularis Rafinesque, 1818 Pomoxis nigromaculatus (Lesueur, 1829) Pomona species
Percidae - perches Ammocrypte beam Jordan, 1877 Ammocrypta clara Jordan and Meek. 1885 Ammocrypta vivax Hay, 1882 Etheostoma asprigene (Forbes, 1878) Etheostoma chlorosomum (Hay, 1881) Etheostoma fusiforme (Girard, 1854) Etheostoma gracile (Girard, 1859) Etheostoma histrto Jordan and Gilbert, 1887 Etheostoma proeliare (Hay, 1881) Etheostoma stigmaeum (Jordan, 1877) Percina caprodes (Rafinesque, 1818) Percina maculata (Girard, t859) Percina sciera (Swain, 1883)
Sciaenidae - drums Aplodinotus grunniens Rafinesque, 1819
Unciassified fishes Unknown fry
S"
98
0.50
0.25
-
54.11
025
-
0.50 0.75 150
2.99
1.00 2.24 3.74
3.99 0.25
-
0.50
025
/<§>
98
0.69 0.35
0.69
0.35
-
42.91
-
0.35
0.35 1.38 0.35
20.42
1.73
1.73 0.35
-
5.19
£*98
-
0.41
-
1598
-
:1.24 1.24 3.32 3.32
0.41
13.69
0.21
2.28 0.21
-
-
2.70
*98
0.21
-
44.70
0.21
-
;;1.25 0.62 1.04 2.08
2.70
0.21
1.66
:
-
2.91
/cf98
0.10 0.10 0.42 0.10
-
-
69.97
-
1.67 0.10
0.63
0.10
0.73 0.52
;;-
0.31
/<?98
0.19 1.11
0.56
0.56
0.37
-
0.74
1.85 0.37
3.89
13.33 0.56
24.81 5.93 1.11 1.30 2.41
0.19
-
0.19
1.67
/ <J98
0.33 0.66
0.33
-
2.31
7.26
;;0.66 0.33
4.95
0.33
429 0.33
;;-
2.97
*98
0.26
0.26 0.26
0.26
-
22.40
-
;;1.04 3.65 5.99
3.13
7.29
1.30
3.65
:
-
0.26
/<?98
0.18
1.84 0.37
-
0.18
25.23
-
4.42 0.18 4.24 0.18
9.76
1.66 0.55
0.55
0.18
-
1.10
-j98
1.40
0.56
0.84
0.56
1.69
-
1.69 6.18 0.84 23.88
10.39 0.56 0.84
3.93
13.76 2.25
028
-
5.62
1 598
2.69
1.01
3.70
13.80
-
;;0.34 2.36 7.41
1.35 0.34 0.34
3.03 1.01
4.04 0.34
6.06
1.35
5.05
*98
0.12
1.08 0.12
0.72
2.88
28.42
0.60
-
0.12
6.71 1.20
1.44
7.67 0.12 120
2.40 1.20 0.48
0.24 0.24
0.72
0.12
-
036
/I98
2.62 0.58
-
7.00
0.29
-
058
3.21
0.29
16.62
3.79 7.29 0.29
-
0.29
-
-
146
0.29
§98
1.68 1.34 0.50 5.87 1.01
-
1.34
503
0.34
1.34
0.50
3.02
4.19 0.17
3.02
2.68
3.52 0.34
0.84 0.34
-
-
8.72
98
1.20 1.80 0.60
4.19
-
14.37
-
;;0.60
0.60
8.38
0.60
7.78
-
-
-
2.40
Ij
98
0.34 0.34
1.03
0.34
0.34
-
13.36
0.34
2.05 1.71
1.37 0.34
2.74
0.68
19.86 0.34
3.42
__
-
-
4.11
*98
-
1.37
0.39
0.20
5.29
10.20
11.18 5.69 0.20 21.18
30.98 1.96 5.10 0.39 0.39
2.35 0.20
"-
-
0.20
020
22 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-98
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98-Continued
SCIENTIFIC NAMEPetromyzontidae - lampreys
Petromyzontidae speciesLepisosteidae - gars
Lepisosteus oculatus (Winchell. 1864)Lepisosteus osseus (Linnaeus, 1758)Lepisosteus platostomus Rafinesque, 1820Lepisosteus species
Amiidae - bowfinsAmia calva Linnaeus, 1766
Hiodontidae - mooneyesHiodon alosoides (Rafinesque, 1819)
Anguillidae - freshwater eelsAnguilla rostrata (Lesueur, 1817)
Clupeidae - herringsDorosoma cepedianum (Lesueur, 1818)Dorosoma petenense (Gunther, 1867)Dorosoma species
Cyprinidae - carps and minnowsCtenopharyngodon idella (Valenciennes, 1844)Cyprinella camura (Jordan and Meek, 1884)Cyprinella lutrensis (Baird and Girard, 1853)Cyprinella spiloptera (Cope, 1868)Cyprinella venusta Girard, 1856Cyprinus carplo Linnaeus, 1758Cyprinidae speciesHybognathus hayi Jordan, 1885Hybognathus nuchalis Agassiz, 1855Lythrurus fumeus (Evermann, 1892)Lythrurus umbratilis (Girard, 1856)Macrhybopsis aestivalis (Girard, 1856)Macrhybopsis storeriana (Kirtland, 1847)Notemigonus crysoleucas (Mitehill, 1814)Notropis ammophilus Suttkus and Boschung, 1990Notropis amnis Hubbs and Greene, 1951Notropis atherinoides Rafinesque, 1818Notropis buchanani Meek, 1896Notropis chalybaeus (Cope, 1869)Notropis maculatus (Hay, 1881)Notropis taxanus (Girard, 1856)Notropis volucellus (Cope, 1865)Notropis speciesOpsopoeodus emiliae Hay, 1881Phenacobius mirabilis (Girard, 1856)Pimephales notatus (Rafinesque. 1820)Pimephales promelas Rafinesque, 1820Pimephales vigilax (Baird and Girard, 1856)Se- LI romaculatus (Mitehill, 1818)
Catostomidae - suckersCarp/odes carpro (Rafinesque, 1820)Carp/odes cyprinus Lesueur, 1817Erimyzon oblongus (Mitchiil, 1814)Erimyzon speciesHypentelium nlgricans (Lesueur, 1817)Ictiobus bubalus (Rafinesque, 1818)Ictiobus cyprinellus (Valenciennes, 1844)Ictiobus niger (Rafinesque, 1819)Ictiobus speciesMinytrema melanops (Rafinesque, 1820)Moxostoma macrolepidotum (Leuseur, 1817)Moxostoma poecilurum (Jordan, 1877)Moxostoma speciesCatostomidae species
98
2.30
1.15
7.28
3.07
34.87
1.15 -
3.456.13
0.38 -
98
0.27
0.14
6.84
0.14
0.6830.230.68 -
1.6418.60
2.19
0.14
0.411.23
0.27
2.60
1.64
1.09 -
S8
--
5.34
3.05
1.533.82
49.623.05-
5.34 -
2.29
0.76
1.53
2.29 -
98
0.170.170.34
3.42-
1.373.081.03
2.39 --
0.17
0.17
0.85
15.90
1.882.050.17 - -
98
0.29
1.17
1.47
5.575.87
5.87
-
1.17
3.81 -
98
1.190.090.090.09
0.09
6.21
0.34 -
2.55
0.26-
1.190.510.090.77 -
98
0.79 -
3.3518.938.48
39.251.78
0.79 - -
1.780.59
0.20 --
0.20
0.20 - -
98
4.81
0.34
0.34
-
0.69 -
0.69 -
23
Table 4. Percent relative abundance of fishes collected at 38 sites in the Mississippi Embayment Study Unit, 1996-98 Continued
SCIENTIFIC NAMEIctaluridae - bullhead catflshes
Ameiurus me/as (Rafinesque, 1820)Ameiurus natalis (Lesueur, 1819)Icta/urus furcatus (Lesueur, 1840)Ictalurus punctatus (Rafinesque, 1818)Noturus gyrinus (Mitchill, 1817)Noturus miurus Jordan, 1877Noturus nocturnus Jordan and Gilbert, 1886Noturus phaeus Taylor, 1969Pylodiclis olivaris (Rafinesque, 1818)
Esocidae - pikesEsox americanus vermiculatus Lesueur, 1846
Aphredoderidae - pirate perchesAphredoderus sayanus (Gilliams, 1824)
Cyprinodontidae - kiilif ishesFundulus chrysotus (Gunther, 1866)Furidulus notatus (Rafinesque, 1820)Fundulus olivaceus (Storer, 1845)Fundulus species
Poeciiiidae - iivebearersGambusia affinis (Baird and Girard, 1853)
Atherinidae - siiversidesLabidesthes sicculus (Cope, 1865)Menidia beryllina (Cope, 1S66)
Percichthyidae - temperate bassesMorone chrysops (Rafinesque, 1820)Morone mississippiensls Jordan and Eigenmann,Morone saxatilis (Walbaum, 1792)
Centrarchidae - sunflshmAmbloplites ariommus viosca, 1936Elassoma zonatum Jordan, 1877Lepomis cyanellus Rafinesque, 1819Lepomis gulosus (Cuvier, 1829)Lepomis humilis (Girard, 1858)Lepomis macmchirus Rafinesque, 1819Lepomis marginatus (Hdbrook, 1855)Lepomis megalotis (Rafinesque, 1820)Lepomis microlophus (Gunther, 1859)Lepomis miniatus rmarm, 1899Lepomis hybridLepomis speciesMicropterus punctulatus (Rafinesque, 1819)Micropterus salmoides (Lacepede, 1802)Micropterus speciesPomoxis annularis Rafinesque, 1818Pomoxis nigromaculatus (Lesueur. 1829)Pomoxis species
Percidae - perchesAmmocrypta beam' Jordan, 1877Ammocrypta clara Jordan and Meek, 1885Ammocrypta vlvax Hay, 1882Etheostoma asprigene (Forbes, 1878)Etheostoma chlorosomum (Hay, 1881)Etheostoma fusiforme (Girard, 1854)Etheostoma gracile (Girard, 1859)Etheostoma histrio Jordan and Gilbert, 1887Etheostoma proeliare (Hay, 1 881 )Etheostoma stigmaeum (Jordan, 1877)Percina caprodes (Rafinesque, 1818)Perdna maculata (Girard, 1859)Percina sciera (Swain, 1883)
Sciaenidae - drumsAplodinotus grunniens Rafinesque, 1819
Unclassified fishesUnknown fry
98
--__,_-
~
0.38--
2.30
-
.-__-
0.381.5313.03
0.38 --
9.96
6.904.60-
-- ~ --
0.77
-
98
-~
1.92 __-
0.14
0.27
0.41
8.48
0.82
-
0.410.820.143.42
8.89
0.14
0.140.412.33
0.140.41
-- -~ --
0.14
1.78
-
98
-
3.051.53 _
0.76
0.76
--- -
3.82
-
---
-_- «-
0.76 - -
0.760.76
0.76--
- - -
-
8*0
-
98
- -
0.17 ___ -
0.51
3.59
3.76
0.17--
0.17__
2.052.390.179.57
17.26
2.055.980.85
0.510.17
--
0.680.1711.62
0.17
0.34-
3.59_
085
-
98
-
029-
0.59 __---
0.88
0.59
-
3.81-
40.76
-
---
0.290.290.291.76
6.74
9.38
1.76
1.172.64-~
1.47
- -
0.29 -
0.59
1.17
-
98
- -
0.17--_.---
- -
49.66
0.340.09-
.-__
1.45
4.082.04
1.19
22.110.090.60
2.550.51
-- -~ --
1.70
-
98
- -
0.59 __-
0.79
---~-
18.54
---
__
0.39
0.39-
1.58--
0.39
-- --- -- __
0.99
-
98
-0.34 __- -
1.37
-
1.03-
6.19
---
__
3.782.411.03
46.740.698.9310.654.12 -
412
0.69
-
1.03 -- ~
-
-
c en O§3 I-o33
24 OCCURRENCE OF FISHES AT 38 SITES IN THE MISSISSIPPI EMBAYMENT STUDY UNIT, 1996-96