54
MANU EXPEDITIONS BIRDING TOURS [email protected] www.Birding-In-Peru.com A BIRDING TRIP REPORT FOR A PRIVATE TRIP TO THE MANU BIOSPHERE RESERVE October 8 th -22 nd 2016 Tour Leader: Silverio Duri

MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

  • Upload
    others

  • View
    0

  • Download
    0

Embed Size (px)

Citation preview

Page 1: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

MANU EXPEDITIONS BIRDING TOURS

[email protected]

ABIRDINGTRIPREPORTFORAPRIVATETRIPTOTHEMANUBIOSPHERERESERVE

October8th-22nd2016

TourLeader:SilverioDuri

Page 2: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Bananarama-RosemaryCheshire

CochaCamungoinManu–CarolineSimpson

Page 3: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

TheManu Biosphere Reserve has the highest diversity of life on Earthandisoneofthemostimportantconservationunitsintheworld.Ifyouwere to choose one of THE best birding trips in the world, this mustsurelybea candidate. Thebeautyof this trip is thevarietyofhabitatsvisited ranging from orchid laden cloud forestwhere Spectacled Bearsand Cock-of-the-Rocks still live unmolested, to untouched Amazonrainforest where 13 species of Monkey abound and Giant Otters stillexist intheox-bowlakes.AtriptoManuisatriptooneoftheworld'sgreat wilderness areas where wildlife is still plentiful and over 1000speciesofbirdshavebeenrecorded.Imaginebirdingonforesttrailswithmixed species flocks that might contain over 50 species and bumpinginto a troop of Black Spider Monkeys around the corner, or quietlypaddling on a catamaran on an Ox-bow Lake with Giant Otters andteeminglakesidebirdlife!

BIRDINGLOCALITIESALONGTHEMANUROAD,CUSCO,MACHU-PICCHUANDPUERTOMALDONADO

HuacarpayLakesS1337462W07143109(3100mts)

AcjanacoPass:S1311929W07137057(TresCruces)GuardStation(3360mts)

WayqechaBiologicalStation:S1310478W07135478(2910mts)

Pillahuata(bellowtheTunnels):S1309725W07135670(2600mts)

Rocotal:S1306081W07134145(2010mts)

CockoftheRockLodge:(CORL)S1303863W07132377(1300mts)

QuitaCalzones:S1301550W07129979(1000m)

Pillcopata:S1254518W07124196(600m)

Mirador(AboveAtalaya):S1253730W07121751(751m)

Atalaya:S1253368W07121547(500mts)

AmazoniaLodge:S1252232W07122527(AL)(500mts)

ManuWildlifeCenter:(MWC)12°21'19.99"S70°42'18.06"W(250mts)

PuertoMaldonado:12°36'10.09"S69°11'28.54"W(220mts)

Page 4: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

DAY-BY-DAYACTIVITIESOct08th:AfulldayatHuacarpayLakesandnightintheIncaCapitalCusco

Oct09/10th:AtwodaytriptoMachu-Picchu(birdsnotincludedonlist)

Oct11th:CuscotoCockoftheRockLodgewithbirdingstopsalongtheway

Oct12rd:FulldayatCORL.Ambirdingupanddowntheroad;andintheafternoonup the road to the Mirador (lookout), ending in the evening at the Lyre-tailedNightjarstakeout.NightatCORL.

Oct 13th: Full day at Cock of the Rock Lodge. Morning to Rocotal and in theafternoontoQuitaCalzones(1000mts).NightCORL.

Oct14th:CockoftheRockLodgetoPantiacollaLodgebirdingalongtheway–boatridefromAtalayacommunitytoPantiacollaLodge.NightatPantiacolla.

Oct15th:FulldayatPantiacollaLodge.AmtoOropendolaandAracari trailand intheafternoontoTinamoutrail.NightatPantiacolla.

Oct16th:FulldayatPantiacollaLodge.AmtoCapybaratrailandintheafternoontotheMirador,MonkSakiandOropendolatrail.NightatPantiacollaLodge.

Oct17th:BoatridefromPantiacollaLodgetoManuWildlifeCenterontheMadredeDiosRiveraftermorningbirdingonTinamoutrail.NightatMWC.

Oct18th:FulldayatMWC.AmtoBlanquilloMacawclaylickandintheafternoontothecanopyplatform.NightatMWC.

Oct 19th: Full day atMWC. Am to Cocha Camungo ox-bow lake and the canopyplatformandintheafternoontocollpatrail.NightMWC.

Oct 20th: Full day at MWC. Am to Cocha Blanco ox-bow lake and Antthrush(bamboo)trailandintheafternoontotheGRIDtrailsystem.

Oct21th:FulldayatMWC.Alldayalongthecollpatrail,earlyafternoonbacktotheLodge.NightMWC.

Oct 22th: Travel day to Puerto Maldonado via Boca Colorado and onto PuertoPunkirichicobycarandthenshortboatridetocrosstheInambariRivertogettoPuerto Carlos and then to Santa Rosa on the high way that took as to PuertoMaldonadostraighttotheAirportfortheafternoonflghttoLima–Endofthetour.

Page 5: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

TheKeytotheBirdlist*Speciesheardonly

RED–IUCNREDLISTCATEGORY

SACC=SouthAmericanClassificationCommittee

IOC=InternationalOrnithologicalCongress

RR=NearEndemicorrangerestrictedspecies

E=PeruvianEndemic

THEBIRDLISTTINAMOUSGreatTinamou-Tinamusmajor–3intotalseeninonedayatPantiacolla.BrownTinamou-CrypturellusobsoletusWesaw2inthegardenononedayatCockoftheRocklodge.Greatviews!

HornedScreamer–CarolineSimpson

Page 6: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SCREAMERSHornedScreamer-AnhimacornutaWesawthisseveraltimesattheMadreDeDiosRiverandalsoatCamungoandBlancoOxbowLakemanyindividualsDUCKSOrinocoGoose-OressochenjubataWesaw10individualsontheRiverbankoftheMadredeDiosonourwaydowntoBocaColorado.Greatviews!MuscovyDuck-CairinamoschataSeen in threedifferentdaysontheMadredeDiosRiver.FormerlycalledMuskDuckbecausethedrakehasamuskysmellYellow-billedTeal-AnasflavirostrisMostCommonatHuacarpayLakes;TheSACCsays“Hellmayr&Conover(1948a)andmanyearlierclassificationstreatedandium(AndeanTeal)asaseparatespeciesfromA.flavirostris.Manyauthors,fromMeyerdeSchauensee(1970)toDickinson(2003),havetreated andium as a subspecies of A. flavirostris. Anas andium was considered aseparatespeciesfromAnasflavirostrisbyRidgelyetal.(2001),andthiswasfollowedby Hilty (2003). SACC proposal passed to recognize andium as separate species.Jaramillo (2003) further suggested that the subspecies oxyptera may also deserverecognitionasaseparatespeciesfromA.flavirostris.Yellow-billedPintail-AnasgeorgicaAlsoseenatHuacarpayLakes.PunaTeal-AnaspunaCommonatHuacarpayLakesCinnamonTeal-AnascyanopteraSeenatHuacarpayLakes.VerycommonCHACHALACAS,GUANS&CURASSOWSAndeanGuan-PenelopemontagniiOneseenaboveCockoftheRockLodge.InGreekmythologyPenelopewasdaughterofIcarusandwifeofUlysses,KingofIthaca.Spix’sGuan-PenelopeJacquacuOneseenatPantiacollaLodgeandanotheroneatManuWildlifeCenter.NamedaftertheGermannaturalistandcollectorinBrazil(1781-1826)JohannBaptistRittervonSpixBlue-throatedPipingGuan-PipilecumanensisSeenatPantiacollaandManuWildlifeCenter

Page 7: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SpeckledChachalaca-OrtalisguttataAverycommonbirdofAmazonianlowlands.CommoninmorelightlywoodedandforestedgeareasinthelowlandsNEWWORLDQUAILRufous-breastedWood-Quail-Odontophorusspeciosus*HeardbelowCockoftheRockLodge.GREBESLeastGrebe-TachybaptusdominicusOneseenatCochaBlancoSTORKSWoodStork-MycteriaAmericanaOneseenonourboatridefromAtalayatoPantiacollaLodgeontheMadredeDiosRiver.CORMORANTSNeotropicCormorant-PhalacrocoraxbrasilianusVerycommonbirdontheMadredeDiosRiverDARTERSAnhinga-anhingaanhingaCommonontheox-bowlakesintheLowlands

HERONS&BITTERNSFasciatedTigerHeron-TigrisomafasciatumCommonontheAltoMadredeDiosRiver–3seenonourboatridetoPantiacollaStriatedHeron-ButoridesstriatusoTwoseenatCochaBlancoox-bowLakeCattleEgret-BubulcusIbisCommonCocoiHeron-ArdeaCocoiCommonbirdonMadredeDiosRiverGreatEgret-ArdeaalbusCommonbirdonAmazonRiversCappedHeron-PilherodiuspileatusVeryprettybirdseenon4differentoccasionsontheAltoMadredeDiosRiverandBlancoLake

Page 8: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SnowyEgret-EgrettathulaAverycommonbirdontheMadredeDiosRiverIBIS&SPOONBILLSPunaIbis–plegadisridgwayiWerecordedthisatHuacarpayLakesandontheroadsideonourwaytoPuertoMaldonadoRoseateSpoonbill-AjaiaajajaSeenontheMadredeDiosRiverinourwaytoBocaColoradofromMWC.Greatviews.AMERICANVULTURESTurkeyVulture-CathartesauraVery common bird. Jaramillo (2003) suggested that the resident tropical subspeciesruficollis and the southern subspecies group (jota and "falklandica") might meritrecognitionasseparatespeciesfromthenorthernCathartesauragroup.GreaterYellow-headedVulture-CathartesmelambrotusCommoninthelowlandAmazonBlackVulture-CoragypsatratusMostcommonintheAmazonrainforestKingVulture-SarcoramphuspapaSeentwiceintheLowlandsOSPREYSOsprey-PandionhaliaetusOneseenatBlancoOxbowLakeinflightandanotheroneontheMadredeDiosRiveronourwaydowntoBocaColoradoKITES,HAWKSANDEAGLESSwallowtailedKite-ElanoidesforficatusGreatviewsatPantiacollaLodgeandManuWildlifeCenterBlackHawk-Eagle-SpizaetustyrannusOneindividualseenaroundManuWildlifeCenterBlackandchestnutEagle-SpizaetusisidoriA male seen above Cock of the Rock Lodge. Named after Isidore Geoffrey Saint –Hilliaire1805-1861Frenchzoologist.VULNERABLEDouble-toothedKite-HarpagusbidentatusSeenatManuWildlifeCenter.

Page 9: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

PlumbeousKite-IctiniaplumbeaVerycommononAmazonlowlandrainforestsSlate-coloredHawk-ButeogallusschistaceaSeentwicearoundManuWildlifeCenterandonemoreonourboatridetoBocaColorado.GreatBlackHawk-ButeogallusurubitingaCommonontheMadredeDiosRiverMontaneSolitaryEagle-ButeogallussolitariusAnimmatureseenonitsnestaboveCockoftheRockLodge.NEARTHREATENEDRoadsideHawk-RupornismagnirostrisVerycommonbirdinAmazonlowlandforestBlack-chestedBuzzard-Eagle-GeranoaetusmelanoleucusOnebirdatHuacarpayLakesLIMPKINSLimpkin-AramusguaraunaSeenatCochaBlancoox-bowLakeandonemoreintheagriculturefieldonourwaytoPuertoMaldonado.

Limpkin–CarolineSimpson

Page 10: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

TRUMPETERSPale-wingedTrumpeter–Psophialeucoptera2seenwellinaweirdway(uponthetreeafterbeenchasebyadomesticdoginthemiddleofthejunglewithoutitsowner)intheGRIDatManuWildlifeCenterRAILS&CRAKESGray-cowledWoodRail-AramidescajaneaSeencrossingtheroadinourwaytoPuertoMaldonado.TheoldGray-neckedWood-Rail has been split two ways – Russet-napped Wood-Rail Aramides albiventris ofMexicoandtheCaribbeanslopeofCostaRicaandtheonewesawofthePacificslopeofCostaRicatoArgentinaRufous-sidedCrake-Laterallusmelanophaius*Gray-breastedCrake–LaterallusexilisOneseenatCochaBlancoox-bowLakePlumbeousRail-Pardirallussanguinolentus3seenatHuacarpayLakesCommonGallinule-GallinulagaleataVerycommonatHuaxarpayLake.NotethattheNewWorldformhasbeensplitfromtheCommonMoorhenofEurasia.CommonGallinuleofWesternHemisphere is splitfromCommonMoorhenonthebasisofmorphological,genetic,andvocaldifferences(Groenenbergetal2008)Slate-coloredCoot-FulicaardesiacaCommonatHuacarpaylakes;Called"AndeanCoot"inFjeldså&Krabbe(1990),Taylor(1996andRidgelyetal.(2001)butotherauthorsuseSlate-coloredCoot.SUNGREBESSungrebe–Heliornisfulica5intotalseenatCochaCamungoandCochaBlancoox-bowLakesSUNBITTERNSSunbittern-EurypygaheliasOneseenontheMadredeDiosRiveraroundManuWildlifeCenterPLOVERSCollaredPlover-CharandriuscollarisSeentwiceontheMadredeDiosRiverbankintheManulowlandPiedLapwing-Vanellus(Haploxypterus)cayanusCommonontheMadredeDiosRiver–seenaroundMWC

Page 11: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SouthernLapwing–Vanelluschilensis10+seenontheroadsideofthehighwayinourwaytoPuertoMaldonadoAndeanLapwing.VanellusresplendensVerycommonatHuacarpayLakesandintheAndesSTILTSBlack-necked(White-backed)Stilt–Himantopus(melanurus)mexicanusOneseenonthefreshplowedfieldsontheroadsideofthehighwayinourwaytoPuertoMaldonadoSANDPIPERSANDSNIPESPunaSnipe-Gallinagoandina2seennearColquepatainourwaytoCockoftheRockLodgeSpottedSandpiper-ActitismaculariaAtHuacarpaylakeandintheManulowlandLesserYellowlegs-Tringaflavipes2seeninourwaytoPuertoMaldonadoPectoralSandpiper-CalidrismelanotosAfewseenontheMadredeDiosRiveraroundManuWildlifeCenterand3moreinthefreshagriculturefieldonourdrivetoPuertoMaldonadoonthehighwayJACANASWattledJacana-JacanajacanaVerycommonontwoOxbowLakesCamungoandCochaBlancoandfewmoreonourdrivetoPuertoMaldonadoGULLS&TERNSAndeanGull-ChroicocephlusserranusVerycommonatHuacarpayLakesLarge-billedTern-PhaetusasimplexCommoninlowerMadredeDiosRiverYellow-billedTern-SternasuperciliarisVerycommoninMadredeDiosRiverSKIMMERSBlackSkimmer-RynchopsnigerCommonontheMadredeDiosRiver

Page 12: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

PIGEONSANDDOVESRockPigeon-ColumbaliviaCommonnearhumanhabitationSpotWingedPigeon–PatagioenasmaculosaSeenatHuacarpayLakescommon.Band-tailedPigeon-PatagioenasfasciataSeenatRocotalPale-ventedPigeon-PatagioenascayennensisCommonnearwaterinthelowlands

Pale-ventedPigeon–CarolineSimpson

Page 13: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

PlumbeousPigeon-PatagioenasplumbeaQuietcommonaroundCockoftheRockLodgeEaredDove–ZenaidaauriculataCommoninthehighlands.NumbersincreasingrapidlyWhite-tippedDove-Leptotilaverreauxidecipiens*GrayfrontedDove-LeptotilarufaxillaSeen very common at MWC in the garden. This and the previous species havecomplicateddistributionwithplaceswhere theyoccur sideby sideandotherwhereonly oneoccurs or ismore common. In this part of Peru the two species appear tosegregateoneanotherbyelevation.White-tipped is found in the foothillsandGray-frontedinthelowlands.At leastoneisfarmorecommonthantheotheratdifferentelevations.RuddyQuailDove-Geotrygonmontana2seenatPantiacollaLodgeRuddyGround-Dove–GeotrygonMontanaOneseenatBlanquillomacawclaylickBare-facedGround-Dove-MetriopeliaceciliaeOneseenatHuacarpayLakes

Hotazin–CarolineSimpson

Page 14: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

HOATZINHoatzin-OpisthocomushoazinCommononox-bowlakes–seenatBlancoandCamungoox-bowLakeCUCKOOSLittleCuckoo-CoccycuaminutaSeenatCochaCamungoandCochaBlanco.NiceviewsSquirrelCuckoo-PiayacayanaThecommonestCuckoointhefoothillandlowlandrainforestBlack-belliedCuckoo-Piayamelanogaster3individualsseenonthecollpatrailatManuWildlifeCenterGreaterAni–CrotophagamajorSeenatBlanquillomacawclaylickandalsoatCamungoandBlancoox-bowlakesSmoothbilledAni-CrotophagaaniVerycommoninamazonlowlandforestonsecondaryforestTYPICALOWLSRufescentScreech-Owl-MegascopsingensSeentwiceatCockoftheRockLodgeBurrowingOwl–AthenecuniculariaOneseenontheriverbankonourboatridefromPantiacollatoMWCPOTOOSGreatPotoo-NyctibiusgrandesOneindividualseenroostingbetweenPillcopataandAtalayacommunityLong-tailedPotoo–NyctibiusaethereusGreatviewsatPantiacollaLodge,neartheLodgeAndeanPotoo–NyctibiusmaculosusOneseenroostingbelowRocotalontheManuroadNIGHTHAWKS&NIGHTJARSSand-coloredNighthawk-ChordeilesrupestrisSeenroostingontheMadredeDiosRiverbankonourboat ride fromPantiacolla toManuWildlifeCenterCommonPauraque-Nyctidromusalbicollis*

Page 15: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

LyretailedNightjar-UropsalislyraWonderfuldisplayofamale(alittlebitbelowtheusualsite)apartfromtheoneattheusualspotaboveCockoftheRockLodge.

MaleLyre-tailedNightjar–CarolineSimpson

Page 16: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

OcellatedPoorwill-NyctiphrynusocellatusOneseenverywellatPantiacollaLodgeSWIFTSChestnut-collaredSwift-StreptoprocnerutilusSeenatCockoftheRockLodgeWhite-collaredSwift-StreptoprocnezonarisCommonGray-rumpedSwift-ChaeturacinereiventrisSeenatPantiacollaLodgeNeotropicalPalmSwift-TachornissquamataCommonatPantiacollaLodge–commonintheLowlandHUMMINGBIRDSWhite-neckedJacobin-FlorisugamellivoraCommonatPantiacollaLodgeRufous-breastedHermit-GlaucishirsutusSeenatBlanquillomacawclaylickandCochaCamungoGreenHermit-PhaethornisguySeenatCockoftheRockLodgeatthefeedersReddishHermit-PhaethornisruberAtPantiacollaLodgeWhite-browedHermit-PhaethornisstuartiOneseenbelowCockoftheRockLodgeinourwaydowntoAtalayaonourtraveldaytoPantiacollaLodgeLesserVioletear-ColibricyanotusSeenatRocotal,aboveCockoftheRockLodge.TheCentralandMiddleAmericanformofGreenVioletarhavebeensplit intoMexicanVioletearbytheAOUourformwhichoccursfromCostaRicatoBoliviaisnowthisspeciesSparklingVioletear-ColibricoruscansSeenseveraltimes

Page 17: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SparklingVioletear–CarolineSimpson

Black-earedFairy-Heliothryxaurita2atPantiacollaLodgeAmethyst-throatedSunangel-HeliangelusamethysticollisCommoninthehighercloudforestaroundWayqechaBiologicalStation.Feedingatflowers.Wire-crestedThorntail-DiscosuralangsdorffiSeenintwoconsecutivedaysontheVerbenaflowersatCockoftherocklodgeinthegarden–NEARTHREATNEDBlack-belliedThorntail–DiscosuralangsdorffiAfemaleseenatthegardenofPantiacollaLodgeFestiveCoquette-LophornischalybeusSeenatPantiacollaLodgeandManuWildlifeCenterEPeruvianPiedtail-PhlogophilushartertiThisendemicwasseenbellowCockoftheRockLodgeonourtraveldaytoPantiacollaLodge–NEARTHREATNED

Page 18: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SpeckledHummingbird-AdelomyiamelanogenysSeenatCockoftheRockLodgeinthegardenfeedingonflowersGreen-tailedTrainbearer–Lesbianuna2seenatHuacarpayLakes.Lesbia=AwomanofLesbosEBeardedMountaineer–Oreonymphanobilis2seenverywellatHuacarpayLakesTyrianMetaltail–MetalluratyrianthinasmaragdinicollisNamedafterthecolorTyrianpurple.VariouslyknownasRoyalpurple,Tyrianpurple,purpleoftheancients,thisancientdyestuff,mentionedintextsdatingabout1600BC,wasproducedfromthemucusofthehypobranchialglandofvariousspeciesofmarinemollusks,notablyMurex.AlthoughoriginatinginoldportofTireinmoderndaySyria(hence the name), man's first large scale chemical industry spread throughout theworld.With thedeclineof theRomanEmpire, theuseof thedye also declined andlarge-scaleproductionceasedwiththefallofConstantinoplein1453.ItwasreplacedbyothercheaperdyeslikelichenpurpleandmadderRRBuff-thighedPuffleg-HaplophaediaassimilisOneseenatRocotal–aboveCockoftheRockLodge. ShiningSunbeam–AglaeactiscupripenniscaumatonotusSeeninthecloudforestaroundWayqechaBiologicalStationBronzyInca-CoeligenacoeligenaAlsoseenatRocotalCollaredInca–CoeligenatorquataQuietcommonatRocotalAnna’s(Booted)Racket-tail-Ocreatus(underwoodii)annaeCommonatCockoftheRockLodge.The“Racquet-tailedPufflegwasunknownin lifebut specimens existed in various London cabinets, whence a drawing was sent toLesson (1832) byMr.Underwoodonbehalf of Charles Stokes a London stockbrokerandcollector.A new paper in Zootaxa 4200 (1): 083–108 by KARL-L. SCHUCHMANN1, ANDRÉ-A.WELLER1 & DIETMAR JÜRGENS, suggests that the Booted Raquet-tail is in fact 4species.IfcorrectthisarticlewouldprovideBoliviawithanendemicspeciesandgivesPeru2species -one an endemic! Which ones have you seen?Here is a resume by theauthorsOurtaxonomicanalysesshowthatthedivergenceinindependentdiagnosablemorphologicalcharactersandinbehavioraltraitssufficientlymeetsthecriteriasetbyHelbig et al. (2002) for assigning species rank to four populations: O. underwoodii(including polystictus, discifer, underwoodii, incommodus, melanantherus), White-bootedRacket-tail:RANGEVENEZUELATOECUADORO.peruanus,PeruvianRacket-tailRANGE - SE ECUADORTOHUANUCOPERU. O. addae,Adda’s Racket-tail LA PAZ TO

Page 19: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SANTACRUZBOLIVIAO.annae.Anna’sRacket-tailRANGE:PASCOTOCUSCO

Violet-frontedBrilliant-HeliodoxaleadbeateriVerycommonatCockoftherockLodgeGiantHummingbird-PatagonagigasSeenatHuacarpayLakesandalsoontheManuroad-theworld’slargestHummerFork-tailedWoodnymph-ThaluraniafurcataSeenatPantiacollaLodgeandManuWildlifeCenterGolden-tailedSapphire-ChrysuroniaoenoneQuietcommonatPantiacollaLodgeWhite-chinnedSapphire-HylochariscyanusSeenatManuWildlifeCenter.

Page 20: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

White-chinnedSapphire–CarolineSimpson

TROGONSANDQUETZALSCrestedQuetzal–Pharomachrusantisianus3seenatCockoftheRockLodgeGolden-headedQuetzal–PharomachrusauricepsSeentwiceatCockoftheRockLodgeCollaredTrogon-TrogoncollarisAtManuWildlifeCenterMaskedTrogon-Trogonpersonatus2seenatRocotalBlue-crownedTrogon-TrogoncurucuiAmaleoneseenatPantiacollaLodgeAmazonianTrogon-TrogonramonianusSeenatManuWildlifeCenter.Thesubspeciesramonianusandcaligatuswereformerly(e.g., Cory 1919, Pinto 1937) considered separate species from Trogon violaceus(Violaceous Trogon) but Peters (1945) considered them all conspecific. Ridgely &

Page 21: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Greenfield (2001) considered caligatus (Gartered Trogon) of Middle America andnorthwesternSouthAmericatobeaseparatespeciesfromTrogonviolaceus,andthiswas followed by Hilty (2003); Genetic data (DaCosta & Klicka 2008) indicate thatcaligatus is basal toa group that includesAmazonianT. violaceus,T. curucui, andT.surrucura (and that Amazonian violaceusmay be paraphyletic with respect to thelattertwospecies).SACCproposalwaspassedtorecognizecaligatusasaspecies.SACCproposalpassedtorecognizeramonianusasaseparatespeciesfromT.violaceus.Soinshortyouhave3speciesthatcameoutofViolaceousTrogon.

GarteredTrogonTrogoncaligatus–S.MexicotoNWPeru

AmazonianTrogon-Trogonramonianus–theAmazonBasin

GuiananTrogon–Trogonviridis–TrinidadandtheGuiananShield

KINGFISHERSRingedKingfisher-MegaceryletorquataSeenatBlancoox-bowLakeAmazonKingfisher-ChloroceryleamazonaAlsoseenatBlancoox-bowLakeMOTMOTSBroad-billedMotmot-ElectronplatyrhynchumSeenatPantiacollaLodgeRufousMotmot-BaryphthengusmartiiSeentwiceatManuWildlifeCenterAmazonianMotmot-MomotusmomotaSeenatPantiacollaLodgeAndeanMotmot-MomotusaequatorialisOneoftheveryattractivebirdsseenatCockoftheRockLodgeinthegarden.Sotherearenow6recognizedspeciesthatcameoutoftheoldBlue-crownedMotmotMomotuscoerulicepsBlue-crownedMotmot–NEandCentralMexico

MomotuslessoniBlue-diademedMotmot–SouthMexicotoCentralPanama

MomotussubrufescensWhoopingMotmot-EPanamatoNCVenezuelaandtheMagdalenaValleyofColombia;SEEcuadorandextremeNWPeru.

MomotusbahamensisTrinidadMotmot–Trindad&Tobago

MomotusmomotaAmazonianMotmot-Venezuela(SoftheOrinoco)andtheGuianasSthroughtheentireAmazonbasintoextremeNArgentinaandParaguay.

Page 22: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

MomotusaequatorialisAndeanMotmot–TheAndesfromNCColombiatoNEBolivia.

AndeanMotmot–CarolineSimpson

Page 23: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

JACAMARSPurusJacamar-GalbalcyrhynchuspurusianusSeveralatCamungoandBlancoox-bowLakeinfamilygroup.NamedforthePurusRiverthatrunsfromeasternPeruintoBrazilBluish-frontedJacamar-GalbulacyanescensCommoninAmazonialowlandforest–seenbellowCockoftheRockLodgeandManuWildlifeCenter.ParadiseJacamar–GalbuladeaOneseenoncollpatrailatManuWildlifeCenterPUFFBIRDSANDNUNBIRDSChestnutcappedPuffbird-BuccomacrodactylusSeenatAntthrushtrailbellowManuWildlifeCenterSpottedPuffbird–BuccotamatiaOneseenoncollpatrailatManuWildlifeCenterWesternStriolatedPuffbird–NystalusobamaiOneseenatPantiacollaLodge.NamedfortheoutgoingUSPresidentBarackObamaBlack-streakedPuffbird-Malacoptilafulvogularis2seenatRocotalaboveCockoftheRockLodgeLanceolatedMonklet-MicromonachalanceolataOneseenbelowCockoftheRockLodgeRufous-cappedNunlet-Nonnularuficapilla2seenwellatPantiacollaLodgeBlack-frontedNunbird-MonasanigrifronsCommon in amazon lowland rainforest. Monasa is Greek for solitary or a monk areferencetotheplainplumageandquietbehavioroftheNunbirds.White-frontedNunbird-MonasamorphoeusA bird of terra firma forest seen at Pantiacolla Lodge and Manu Wildlife Center.MorpheouswasthesonofsleepandgodofdreamsareferraltothelethargicbehaviorofthisNunbird

Swallow-Wing-ChelidopteratenebrosaCommoninManuamazonlowlandforestandinriverislandsNEWWORLDBARBETSGildedBarbet-Capitoauratus*

Page 24: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Lemon-throatedBarbet-EubuccorichardsoniSeentwiceatPantiacollaLodgeScarlet-hoodedBarbet–EubuccoversicolorGreatviewsatPantiacollaLodgeVersicoloredBarbet-EubuccoversicolorSeenintwoconsecutivedaysatCockoftheRockLodgeTOUCANSChannel-billedToucan-RamphastosvitellinusSeenatManuWildlifeCentreWhite-throatedToucan-RamphastostucanusSeenatPantiacollaLodgeandManuWildlifeCenterEmerald(Black-throated)Toucanet-Aulacorhynchus(atrigularis)prasinusSeenbelowCockoftheRockLodgeChestnut-tippedToucanet-Aulacorhynchusderbianus1individualbelowCockoftheRockLodgeRRBluebandedToucanet-AulacorhynchuscoeruleicinctisQuietcommonatRocotalGray-breastedMountain-Toucan–AndigenahypoglaucaTwoseeninthecloudforestinourtraveldayfromCuscotoCockoftheRockLodge–NEARTREATNEDGolden-collaredToucanet-Seleniderareinwardtii2seenatPantiacollaLodgeIvory-billed(Brown-mandibled)Aracari–Pteroglossus(mariae)azaraeSeenintwoconsecutivedaysatManuWildlifeCenterChestnut-earedAracari-PteroglossuscastanotisSeen from the Blanquillo macaw clay lick. In Greek Pteroglossus means“feathertongued”areferencetotheslimfeather-liketonguesoftoucansandaracaris

Curl-crestedAracari-PteroglossusbeauharnaesiiSeenintwodifferentdaysatManuWildlifeCenter.Thecoolafrocrestfeelslikeplastictothetouch!(Onmuseumspecimens)

Page 25: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Blue-bandedToucanet–CarolineSimpson

WOODPECKERS&PICULETSYellow-tuftedWoodpecker-MelanerpescruentatusSeenatPantiacollaLodgeRed-stainedWoodpecker–VeniliornisaffinisOneseenatManuWildlifeCenterWhite-throatedWoodpecker–Piculusleucolaemus*Golden-OliveWoodpecker-PiculusrubiginosusSeenaroundtheCockoftheRockLodgeCrimson-mantledWoodpecker-PiculusrivoliiSeenneartheWayqechaBiologicalStationSpot-breastedWoodpecker-ColaptespunctigulaOneonthetrailaswewalkbacktotheRiverfromtheBlanquillomacawclaylick

Page 26: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

AndeanFlicker–ColaptesrupícolaSeenatHuacarpayLakesandontheManuroadScaly-breastedWoodpecker-CeleusgrammicuslatifasciatusOneseenattheCamungocanopyplatformChestnutWoodpecker–CeleuselegansSeenonlybyRobinatPantiacollaLodgeCream-coloredWoodpecker–CeleusflavusSeenfromthecanopyplatformatCamungoRufous-headedWoodpecker–CeleusspectabilisOneseenatAntthrushtrailinthebambooforestLineatedWoodpecker-DryocopuslineatusSeenatManuwildlifeCenterRed-neckedWoodpecker-CampephilusrubricollisAtManuWildlifeCenterCrimson-crestedWoodpecker-CampephilusmelanoleucosSeenatPantiacollaLodgeCARACARAS&FALCONSLaughingFalcon-HerpetotherescachinnasOneseenatPantiacollaLodgeBlackCaracara-DaptriusaterCommoninthelowlandrainforestMountainCaracara–PhalcoboenusmegalopterusSeenwellatHuacarpaylakesandontheManuroadAmericanKestrel–FalcosparveriusCommonbirdoftheAndesBatFalcon-FalcorufigularisSeentwicearoundManuWildlifeCenterPARROTSAmazonianParrotlet-Nannopsittacadachilleae6seenattheBlanquillomacawclaylick

Page 27: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

TuiParakeet-BrotogerissanctithomaeAlsoseenatthemacawclaylickandelsewherearoundManuWildlifeCenterOrangeCheekedParrot-PyriliabarrabandiSeen at the Macaw clay lick very well. Named after Jaques Barrand (1767-1809),Frenchbirdandflowerillustrator

Blue-headedParrot-PionusmenstruusVerycommoninthelowlandsYellow-crownedAmazon-AmazonaochrocephalaAttheBlanquilloMacawclaylickinManuWildlifecenterScaly-nappedAmazon-AmazonamercenariaSeeninthecloudforestonourtraveldayfromCuscotoCockoftheRockLodgeMealyAmazon-AmazonafarinosaCommonattheBlanquillomacawclaylickandaroundMWC.FarinosaisLatinfor“SprinkledwithFlour”referringtothe“dusted”appearanceofthisAmazon.White-belliedParrot-PionitesleucogasterSeenatPantiacollaLodgeRose-frontedParakeet-Pyrrhuraroseifrons10+seenattheTapirclaylickatManuWildlifeCenterBlack-cappedParakeet–Pyrrhurarupicola3seenalsoattheTapirclaylickDusky-headedParakeet-AratingaweddelliiSeenatManuWildlifeCenterRed-billedMacaw-OrthopsittacamanilataSeenintwoconsecutivedaysatPantiacollaLodgeBlue-headedMacaw–Primoliuscouloni2seenatPantiacollaLodge-VULNERABLEBlueandYellowMacaw-AraararaunaQuietcommonintheManulowlandrainforestScarletMacaw-AramacaoSeenverycommonlyintheManulowlandRedandGreenMacaw-ArachloropterusVerycommonattheBlanquilloMacawClaylick

Page 28: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Red&GreenMacawsatthelick–CarolineSimpson

Chestnut-frontedMacaw-AraseveraOneoftheverycommonMacawsintheamazonlowlandforestWhite-eyedParakeet-PsitticaraleucophthalmusAnotherverycommonintheLowlandANTBIRDSRRYellow-rumpedAntwren-EuchrepomissharpieSeenatCockoftheRockLodge-ENDANGEREDFasciatedAntshrike-Cymbilaimuslineatus*GreatAntshrike-TarabamajorSeenatManuWildlifeCenterontheAntthrushtrail.BarredAntshrike-ThamnophilusdoliatusSeenattheBlanquilloMacawClaylickPlain-wingedAntshrike-ThamnophilusschistaceusAmaleoneseenatPantiacollaLodgeWhite-shoulderedAntshrike–Thamnophilusaethiops*Dusky-throatedAntshrike-ThamnomanesardesiacusSeenatPantiacollaLodgeBluish-slateAntshrike-ThamnomanesschistogynusAlsoseenatPantiacollaLodge–theunderstoryflockleader

Page 29: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Plain-throatedAntwren–IsleriahauxwelliSeenatPantiacollaLodgeandManuWildlifeCenter.ThegenusisnamedforMortandPhyllisIslerwhoworkedextensivelyonAntbirdsSpot-wingedAntshrike-PygiptilastellarisSeenatPantiacollaLodgeandManuWildlifeCenterWhite-eyedAntwren-EpinecrophyllaleucophthalmaGoodviewsof3atPantiacollaLodgePygmyAntwren-MyrmotherulabrachyuraAlsoseenatPantiacollaLodgeSclater´sAntwren-MyrmotherulasclateriOneseeninthemix-specieflockatPantiacollaLodgeAmazonianStreakedAntwren-MyrmotherulamultostriataWesawthisatBlancoOxbowlakeveryniceviewsofmaleandfemaleWhite-flankedAntwren-MyrmotherulaaxillarisSeen twiceatPantiacolla Lodge. Willis (1984),Ridgely&Tudor (1994),Hilty (2003),and Zimmer & Isler (2003) noted that vocal differences among several subspeciesofMyrmotherulaaxillarissuggestthatmorethanonespeciesisinvolvedLong-wingedAntwren-MyrmotherulalongipennisgarbeiVerygoodviewsatPantiacollaLodgeGrayAntwren-MyrmotherulamenetriesiiCommoninlowland–seenatPantiacollaLodgeandManuWildlifeCenterYellow-breastedAntwren-HerpsilochmusaxillarisSeentwicebellowCockoftheRockLodge.VULNERABLEStriatedAntbird-DrymophiladevilleiTwoseennearAtalayainourtraveldayfromCORLtoPantiacollaLodgeGrayAntbird-CercomacracinerascensOneseenatPantiacollaLodgeRRManuAntbird-CercomacraManuSeenatManuWildlifecenterontheAntthrushtrail–abamboospecialist!BlackAntbird-Cercomacroidesserva*White-browedAntbird-Myrmoborusleucophrys3intotalatPantiacollaLodge

Page 30: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Black-facedAntbird-MyrmoborusmyotherinusSeenatPantiacollaLodgeandManuWildlifeCenterRRYellow-breastedWarblingAntbird-HypocnemissubflavaSeenatPantiacollaLodgePeruvianWarblingAntbird–Hypocnemisperuviana2seenatManuWildlifeCenterBand-tailedAntbird-Hypocnemoidesmaculicauda2seenatBlancoLakeSilveredAntbird–SclaterianaeviaAtPantiacollaLodgeandManuWildlifeCenterWhite-linedAntbird-PercnostolalophotesSeenatPantiacollaLodgeandManuWildlifeCenter.Anotherbamboospecialist!RangejustcreepsovertheborderintoNWBoliviaChestnut-tailedAntbird-MyrmecizahemimelaenaSeenatPantiacollaLodgePlumbeousAntbird-MyrmecizahyperythraOneseenatManuWildlifeCenterHairy-crestedAntbird-Rhegmatorhinamelanosticta*AtManuWildlifeCenterANTPITTASUndulatedAntpitta–Grallariasquamigera*Thrush-likeAntpitta-MyrmotheracampanisonaVerygoodviewsoftwoatPantiacollaLodge

TAPACULOSRusty-beltedTapaculo-Lioscelesthoracicus2seenatManuWildlifeCenterDiademedTapaculo-ScytolopusschulenbergiGreatviewofonenearAcjanacoontheManuroadonourtraveldaytoCockoftheRockLodgeANTHRUSHESRufous-cappedAntthrush-FormicariuscolmaSeenatManuWildlifeCenter

Page 31: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Black-facedAntthrush-FormicariusanalisGoodviewsatPantiacollaLodgeRRRufous-frontedAntthrush–FormicariusrufifronsOneseenbrieflyatAntthrushtrailbellowManuWildlifeCenterOVENBIRDSBlack-tailedLeaftosser-Scleruruscaudacutus*Slender-billedMiner-GeosittatenuirostrisTwoseenontheManuroadonourtraveldaytoCockoftheRockLodgefromCuscoOlivaceousWoodcreeper-SittasomusgriseicapillusamazonusSeen at Pantiacolla Lodge. Certainly consists of multiple species (Hardy et al. 1991,Ridgely&Tudor1994,Parkeretal.1995,Ridgely&Greenfield2001,Hilty2003),withatleastfivesubspeciesgroupspossiblydeservingseparatespeciesstatus(Marantzetal. 2003). The subspecies viridis and amazonus are elevational replacements insouthernPeruwithdifferentsongtypes(Robbinsetal.2013).Wedge-billedWoodcreeper-GlyphorynchusspirurusCommonInthelowland–seenatPantiacollaLodgeandManuWildlifeCenterCinnamon-throatedWoodcreeper-DendrexetastesrufigulaAlsoseenatPantiacollaLodgeandManuWildlifeCenterLong-billedWoodcreeper-NasicalongirostrisSeenatManuWildlifeCenterAmazonianBarredWoodcreeper-DendrocolaptescerthiaSeenatManuWildlifeCenteronthecollpatrailBlack-bandedWoodcreeper-DendrocolaptespicumnusAlsoseenoncollpatrailatManuWildlifeCenterElegantWoodcreeper-XiphorhynchuselegansjuruanusSeenatPantiacollaLodgeandManuWildlifeCenter.NotethatZimmer(1934d),Pinto(1937), Ridgely & Tudor (1994), and Ridgely & Greenfield (2001) consideredXiphorhynchuselegansandX. spixii (Spix’sWoodcreeper) conspecific, but seeHaffer(1997) for rationale for treating them as separate species, as in Peters (1951) andMeyerdeSchauensee(1970).Aleixo(2002)alsofoundmolecularsupportfortreatingnominatespixiiasaseparatespeciesfromallothertaxainthegroup.Cory&Hellmayr(1925)treatedthesubspecies juruanusand insignisasseparatespeciesfromX.spixii(JuruaWoodcreeper),andPinto(1947)alsomaintainedjuruanusasaseparatespecies;but theywereconsideredconspecificbyZimmer (1934d)andPeters (1951).<incorp.Aleixo2004>

Page 32: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Red-billedScythebill-CampylorhamphustrochilirostrisSeenatPantiacollaLodgeMontaneWoodcreeper-LepidocolapteslacrymigerSeenatRocotalaboveCockoftheRockLodgeInambariWoodcreeper-LepidocolaptesfatimalimaeSeentwiceatManuWildlifeCenter.SACCsaysTheAmazonianfuscicapillussubspeciesgroup(withmadeiraeand layardi)wasformerly(e.g.,Cory&Hellmayr1925)treatedas a separate species from Lepidocolaptes albolineatus –LineatedWoodcreeper butrecent authors have followed Zimmer (1934c) in treating them as conspecific;Hilty(2003)suspectedthatthis treatmentwillbeshowntobecorrect,andMarantzetal.(2003) suspected thatmore than one specieswas involved. Rodrigues et al. (2013)treatedalldiagnosabletaxainthecomplexasseparatespecies;theyalsodescribedanew species, Lepidocolaptes fatimalimae, from southwestern Amazonia. SACCproposal passed to recognize fatimalimae as a species and also elevate subspecieslayardiandfuscicapillustospeciesrank.Sotherearenow5speciesA.Lepidocolaptesalbolineatus=GuiananWoodcreeperB.Lepidocolaptesduidae=DuidaWoodcreeperC.Lepidocolaptesfatimalimae=InambariWoodcreeperD.Lepidocolaptesfuscicapillus=RondoniaWoodcreeperE.Lepidocolapteslayardi=Layard’sWoodcreeper

PlainXenops-XenopsminutusSeenatManuWildlifeCenterRufous-tailedXenops–MicroxinopsmilleriOneseenoncollpatrailatManuWildlifeCenterPale-leggedHornero–Furnariusleucopus2seenatCamungoox-bowLakeWren-likeRushbird-PhleocryptesmelanopsSeenatHuacarpayLake–onlyfoundinreedbedsWhite-wingedCinclodes–Cinclodesatacamensis2seennearColquepatainourtraveldaytoCockoftheRockLodgeDusky-cheekedFoliage-gleaner-AnabazenopsdorsalisSeeninbambooatAntthrushtrailRufous-rumpedFoliage-gleaner-PhilydorerythrocercumOneseenatManuWildlifeCenter

Page 33: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Chestnut-wingedFoliage-gleaner-PhilydorerythropterumSeenatPantiacollaLodgeBuff-frontedFoliageGleaner-PhilydorrufusSeenatCockoftheRockLodgeMontaneFoliage-gleaner-AnabacerthiastriaticollisSeenbellowCockoftheRockLodgeChestnut-wingedHookbill-AncistropsstrigilatusAtPantiacollaLodgeandManuWildlifeCenterOlive-backedFoliage-gleaner-AutomolusinfuscatusAtManuWildlifeCenterChestnut-crownedFoliage-gleaner-AutomolusrufipileatusSeenatPantiacollaLodgeStreak-frontedThornbird-Phacellodomusstriaticeps3seenatHuacarpayLakeERusty-frontedCanatero-AsthenesottonisSeenatHuacarpayLakesAsh-browedSpinetail-CranioleucacurtataOneseenatCockoftheRockLodge.VULNERABLEECreamy-crestedSpinetail-CranioleucagutturataSeenbetweenPaucartamboanAjanaco1individualSpeckledSpinetail–CranioleucagutturataOneseenatPantiacollaLodgeDark-breastedSpinetail–Synallaxisalbigulares1seenatCamungoox-bowLakeCabani´sSpinetail-Synallaxiscabanisi*Plain-crownedSpinetail-Synallaxisgujanensis*TYRANTFLYCATCHERS.Yellow-crownedTyrannnulet–TyrannuluselatusOneseenatPantiacollaLodgeMottle-backedElaenia-ElaeniagigasSeenbelowCockoftheRockLodge

Page 34: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

White-throatedTyrannulet–MecocerculusleucophrysCommoninthecloudforest–seenatRocotalRingedAntpipit-CorythopistorquataSeenatPantiacollaLodgeandManuWildlifeCenterBolivianTyrannulet-ZimmeriusbolivianusQuietcommonaroundCockoftheRockLodgeMarble-facedBristleTyrant-PhylloscartesopthalmicusSeenatCockoftheRockLodgeCinnamon-facedTyrannulet-Phylloscartesparkeri2seenbellowCockoftheRockLodge.OneoftheseveralbirdsnamedforTheodoreAParker the III –pioneer US ornithologist tragically killed in plane crash in southernEcuadorin1993Streak-neckedFlycatcher-MionectesstriaticollisSeenatCockoftheRockLodgewhilstroadsidebirdingOlive-stripedFlycatcher–MionectesolivaceusOneseenatQuitaCalzonesbellowCockoftheRockLodgeOchre-belliedFlycatcher–MionectesoleagineusOneseenbellowQuitaCalzonesinourtraveldayfromCORLtoPantiacollaLodgeEIncaFlycatcher-LeptopogontaczanowskiiThisPeruvianendemicwasseenatRocotal.NamedforWladislawTaczanowskii,Polishornithologistandauthorof“OrnithologieduPerou”1884OrnateFlycatcher-MyiotriccusornatusSeenatQuebradaQuitacalzon2individualsofthisprettybirdMany-coloredRush-Tyrant-TachurisrubrigastraThisreedbedspecialistwasseenatHuacarpayLakes–astunningbirdShort-tailedPygmy-Tyrant-MyiornisecaudatusSeenatPantiacollaLodgeScale-crestedPygmy-Tyrant-LophotriccuspileatusSeenatCockoftheRockLodgeWhite-belliedTodyTyrant-HemitriccusgriseipectusOneseenoncollpatrailatManuWildlifeCenter

Page 35: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Rusty-FrontedTody-Flycatcher-Poecilotriccuslatirostre1seenatAntthrushtrailOchre-facedTody-Flycatcher-Poecilotriccusplumbeiceps2seenattheMirador,aboveCockoftheRockLodgeEBlack-backedTody-Flycatcher-Poecilotriccuspulchellum2seenafterChontachacaontheroadsideonourtraveldayfromCockoftheRockLodgetoPantiacollaLodgeSpottedTody-Flycatcher-TodirostrummaculatumOneseenattheMacawclaylickFulvous-breastedFlatbill-RhynchocyclusfulvipectusGoodviewofonebellowCockoftheRockLodgeYellow-oliveTolmomyias-TolmomyiassulphurescensSeenbelowCockoftheRockLodgeOlive-facedTolmomyias-TolmomyiasviridicepsAlsoseenbellowCockoftheRockLodgeGolden-crownedSpadebill-PlatyrinchuscoronatusOneseenatPantiacollaLodgeBran-coloredFlycatcher-MyiophobusfasciatusSeeninourtraveldaytoPantiacollafromCockoftheRockLodgeRuddy-tailedFlycatcher-TerenotriccuserythrurusSeenatPantiacollaLodgeCinnamonFlycatcher-PyrrhomyiascinnamomeaVerycommonbirdintheManuCloudForest–seenatRocotalSmoke-coloredPewee-ContopusfumigatusSeenatRocotalEasternWoodPewee–ContopusvirensAtCockoftheRockLodgeandPantiacollaLodgeBlackPhoebe-SayornisnigricansSeenatPaucartamboinbelowthebridgeandinManuAndeanNegrito-LessoniaoreasAnimmatureseenatHuacarpayLakes

Page 36: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

White-wingedBlack-Tyrant-KnipolegusaterrimusSeenabovePaucartambomaleandfemaleDrabWater-Tyrant-OchthornislittoralisVerycommonbirdonthebanksoftheMadredeDiosRiverLittleGround-Tyrant-MuscisaxicolafluviatilisSeenontheRiverbankbellowManuWildlifeCenterRufous-nappedGround-Tyrant-Muscisaxicolarufivertex2seenontheManuroadonourtraveldaytoCockoftheRockLodgetoPantiacollaBlack-billedShrike-Tyrant–AgriornisMontanaOneseenontheManuroadonourtraveldaytoCockoftheRockLodgeStreak-throatedBush-Tyrant-MyiotheretesstriaticollisSeenbelowthetwotunnelswhilstroadsidebirdingRRRufous-webbedBush-Tyrant–Polioxolmisrufipennis2seenontheManuroadinourtraveldaytoCockoftheRockLodgeRufous-breastedChat-Tyrant-OchthoecarufipectoralisSeenbellowAcjanacointhecloudforestonourtraveldaytoCockoftheRockLodgeBrown-backedChat-Tyrant-OchthoecafumicolorSeenatAcjanacopassandgoodlooksindeedWhite-browedChat-Tyrant-OchthoecaleucophrysSeenontheManuroadonourtraveldaytoCockoftheRockLodgeLong-tailedTyrant-Coloniacolonus2seennearChontachacaonourtraveldaytoPantiacollafromCockoftheRockLodgePiraticFlycatcher-LegatusleucophaiusSeenfromtheCamungocanopyplatform-goodviewSocialFlycatcher-MyiozatetessimilisCommoninAmazonlowlandforestGreatKiskadee-PitangussulphuratusSeenattheBlanquillomacawclaylickLesserKiskadee-PitanguslectorSeenatCamungoox-bowLake–alwaysnearwaterLemon-browedFlycatcher-ConopiascinchonetiSeenatCockoftheRockLodgeVULNERABLE

Page 37: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Golden-crownedFlycatcher-MyiodynasteschrysocephalusTwoseenbelowCockoftheRockLodgeStreakedFlycatcher-MyiodynastesmaculatusSeentwiceatCockoftheRockLodgeSulphur-belliedFlycatcher-MyiodynastesluteiventrisSeenintwodifferentdaysatPantiacollaLodgeSulphuryFlycatcher-TyrannopsissulphureaEverydayatPantiacollaLodgejustatthegardenCrownedSlatyFlycatcher-EmpidonomusaurantioatrocristatusSeenattheCamungocanopyplatform.AnAustralmigrantandthebirdwiththelongestscientificnameintheworldTropicalKingbird-TyrannusmelancholicusVerycommonbirdinamazonlowlandforestEasternKingbird–TyrannustyrannusCommonborealmigrant.SeenatManuWildlifeCenterGrayishMourner-RhytipternasimplexSeenaroundManuWildlifeCenterShort-crestedFlycatcher-MyiarchusferoxCommonintheLowland–seenaroundManuWildlifeCenterLarge-headedFlatbill–RamphotrigonmegacephalaOneseenatAntthrushtrail–abamboospecialist!Dusky-tailedFlatbill-Ramphotrigonfuscicauda*Rufous-tailedFlatbill-Ramphotrigonruficauda*Bright-rumpedAttila-Attilaspadiceus

COTINGASRed-crestedCotinga-AmpelionrubrocristataSeenontheManuroadonourtraveldaytoCockoftheRockLodge

Page 38: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

AndeanCockoftheRock-RupicolaperuvianaGreatviewsaroundCockoftheRockLodge,apartfromtheonesatseenattheleekaboveCORL

MaleCockoftheRock–CarolineSimpson

ScreamingPiha-LipaugusvociferansSeenatManuWildlifeCenterBare-neckedFruitcrow-Gymnoderusfoetidus3intotalseenatManuWildlifeCenter

MANAKINSDwarfTyrantManakin-TyranneutesstolzmanniSeenatManuWildlifeCenterBlue-backedManakin-ChiroxiphiapareolareginaAfemaleseenoncollpatrailatManuWildlifeCenterFiery-cappedManakin-MachaeropteruspyrocephalusAmaleoneseenatManuWildlifeCenterBand-tailedManakin-Ceratopiprafasciicauda2seenatthetapirclaylickatManuWildlifeCenter

Page 39: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

White-beardedManakin–Manacusmanacus1seennearPatriaonourtraveldaytoPantiacollaLodgefromCROLRound-tailedManakin-PiprachloromerosSeenintwodifferentdaysatPantiacollaLodgeBlue-crownedManakin–Pipracoronataexquisite2atPantiacollaLodgeand2moreatManuWildlifeCenter

TITYRASANDBECARDSBlac-tailedTityra-TityracayanaSeenatManuWildlifeCenter.MaskedTityra-TityrasemifasciataSeenattheCamungocanopyplatformCinereousMourner-LaniocerahypopyrraGoodviewsatPantiacollaLodgeBarredBecard-Pachyramphusversicolor2seenatRocotalontheManuroadChestnut-crownedBecard-PachyramphuscastaneusSeenatCamungocanopyplatformWhite-wingedBecard-PachyramphuspolichopterusSeenatPantiacollaLodgePink-throatedBecard-PachyramphusminorSeenatManuWildlifeCenter

VIREOS&GREENLETSRed-eyedVireo–VireoolivaceuschiviSeenatPantiacollaLodge.Someclassifications(e.g.,Pinto1944)haveconsideredtheSouthAmericanchivigroupasaseparatespecies("ChiviVireo")fromV.olivaceus,oras conspecificwithV. flavoviridis (Hamilton 1962), but seeHellmayr (1935), Zimmer(1941d),Eisenmann1962a,Johnson&Zink(1985),andRidgely&Tudor(1989).Ridgely&Greenfield(2001)suggested,however,thatmorethanonespeciesmaybeinvolvedwithintheSouthAmericanchivigroup.Brown-cappedVireo–VireoleucophrysSeenatRocotalDusky-cappedGreenlet-PachysylviahypoxanthusSeenintwoconsecutivedaysatPantiacollaLodge

Page 40: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Tawny-crownedGreenlet-Pachysylviaochraceiceps2seenwithmix-specieflockatManuWildlifeCenter

JAYSPurplishJay-CyanocoraxcyanomelasSeenatPantiacollaLodgeandManuWildlifeCenterViolaceousJay-CyanocoraxviolaceusBellowCockoftheRockLodgeandPantiacollaLodgeGreenJay-Cyanocoraxluxuosus2seenatCockoftheRockLodge.

SWALLOWS&MARTINSBlueandWhiteSwallow-PygochelidoncyanoleucaCommonWhite-bandedSwallow-AtticorafasciataCommonbirdalongtheMadredeDiosRiverSouthernRough-wingedSwallow-StelgidopteryxruficollisAnothercommonBirdinthelowlandrainforestBrown-chestedMartin-PrognetaperaSeenperchedontheMadreDeDiosRiveronourtraveldaytoManuWildlifeCenterWhite-wingedSwallow-TachycinetaalbiventerMostcommonbirdinMadreDeDiosRiverandlakesBarnSwallow–HirundorusticaSeenatHuacarpayLakes

WRENSGray-mantledWren-OdontorchilusbranickiiOneseenatCockoftheRockLodgeHouseWren-Troglodytesaedon*MountainWren-TroglodytessolstitialisSeenatRocotalontheManuroadSedge(Puna)Wren-Cistothorus(minimus)platensisOneseenbelowAcjanacoontheManuroad.

Page 41: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

ArecentpaperCANADATOTIERRADELFUEGO:SPECIESLIMITSANDHISTORICALBIOGEOGRAPHYOFTHESEDGEWREN(CISTOTHORUSPLATENSIS)byMARKB.ROBBINSANDARPADS.NYARIsuggestthatthereare11speciesintheSedgeWrencomplex Cistothorus stellaris (Naumann 1823) Sedge Wren migratory; breeds in mesic grasslands from central Canada south to central United States; primarily winters in southeastern United States and northeastern Mexico. Cistothorus meridae Hellmayr 1907 Merida Wren Distribution: paramo in Venezuelan Andes Cistothorus apolinari Chapman 1914 Apolinar’s Wren Distribution: cattail and bulrush dominated marshes in northern Colombian Andes Cistothorus elegans Sclater and Salvin 1859 Grass Wren Distribution: upland grassland and marshes (at least formerly) in transvolcanic belt in centralMexico, lowlands in southeastern Mexico, patchily distributed in both lowlands and highlands through Central America south to western Panama. Cistothorus hornensis (Lesson 1834) Austral Wren Distribution: Coquimbo, Chile and Neuque´n, Argentina south through Tierra del Fuego, including Falkland Islands/Malvinas. Comments: includes falklandicus. Cistothorus tucumanus Hartert and Venturi 1909 Tucuman Wren Distribution: highlands of Santa Cruz, Bolivia south to Tucuma´n and at least the mountains in western Co´rdoba province, Argentina Cistothorus platensis (Latham 1790) Pampas Wren Distribution: lowland grasslands in southeastern Brazil (southern Goı´s, Parana´), southern Paraguay, northeastern Argentina, Rio Grande do Sul, Brazil, Uruguay south to central Argentina, and west to dptos Beni and Santa Cruz, Bolivia. Cistothorus minimus Carriker 1934 Puna Wren Distribution: puna grassland from dpto Ayacucho, Peru south to La Paz, Bolivia. Cistothorus graminicola Taczanowski 1874 Junın Wren Distribution: paramo south of the Rıo Maranon drainage to dpto Junın, Peru Cistothorus aequatorialis Lawrence 1871 Paramo Wren Distribution: paramo of Andean Venezuela and northern Colombia south to northern Peru Cistothorus alticola Salvin and Godman, 1883 Venezuelan Wren Distribution: high elevation grassland in Gran Sabana, coastal mountains of Venezuela from Carabobo east to Sucre/Monagas border Gray-breastedWood-Wren-HenicorhinaleucophrysSeennearRocotalandcommonlyheardChestnut-breastedWren-Cyphorhinusthoracicus*

Page 42: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Ferruginous(Musician)Wren-Cyphorhinus(aradus)modulatorOneseenatPantiacollaLodge.ArecentpaperAtaxonomicrevisionoftheMusicianWren,Cyphorhinusarada(Aves,Troglodytidae), reveals the existence of six valid species endemic to the AmazonbasinCyphorhinusarada(Hermann,1783)MusicianWrenTotheeastoftheRioNegro,onthenorthernbankoftheAmazonRiver,includingsoutheasternandeasternVenezuela,theGuianasandnortheasternBrazilCyphorhinustransfluvialis(Todd,1932)ImeriWrenFromEasternbaseoftheAndesisSEColombiaandnorthbankofputumayoRivertoeasttoAmazonaandRioNegroCyphorhinusmodulator-FerruginousWrenEasternPeruandWesternBraziltoNWBoliviaCyphorhinussalviniSharpe,1881-Grey-earedWrenBetweenRioMaranonandRioCaquetainSWColombia,EasternEcuadorandNEPeruCyphorhinusinterpositus(Todd,1932)-RondoniaWrenSouthbankofAmazonfromeastbankofRioMadeiratowestbankofRioTapajosCyphorhinusgriseolateralisRidgway,1888-Gray-flankedWrenSouthbankofAmazonfromeastbankofTapajosRivertowestbankofXinguRiver

Black-cappedDonacobius–CarolineSimpson

Page 43: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

DONACOBIUSBlack-cappedDonacobius-DonacobiusatricapillusSeenatCamungoandBlancoox-bowLakes-fantasticviews

DIPPERSWhite-cappedDipper–CinclusleucocephalusOneseenaroundCockoftheRockLodge

THRUSHSANDSOLITAIRESAndeanSolitaire-Myadestesralloides*Slaty-backedNightingale-Thrush–CatharusfuscaterGoodviewofoneindividualabovePaucartamboinourtraveldaytoCockoftheRockLodgeSwainson’sThrush–CatharusustulatusOneseenatPantiacollaLodgeWhite-earedSolitaire-EntomodestesleucotisOneseenattheMiradoraboveCockoftheRockLodgeChiguancoThrush-TurduschiguancochiguancoVerycommonbirdofAndesGreatThrush-TurdusfuscaterockenderiCommoninthecloudforestontheManuroadBlack-billedThrushTurdusignobilisThecommonlowlandThrush–seenatPantiacollaLodgeandManuWildlifeCenterWhite-neckedThrush–TurdusalbicollisSeenatPantiacollaLodge

TANAGERSANDALLIESRed-cappedCardinal-ParoariagularisCommoninthelowland–seenatCamungoandBlancoox-bowLakeMagpieTanager-CissopisleverianaQuietcommoninthelowlands–seenbellowCockoftheRockLodgeandPantiacollaLodgeBlack-earedHemispingus-HemispingusmelanotisberlepschiSeenatCockoftheRockLodge

Page 44: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Rust-and-YellowTanager-ThlypopsisruficepsSeenatRocotalandalsoonourtraveldayfromCuscotoCockoftheRockLodgeYellow-crestedTanager-Tachyphonuscristatus5intotalintwoconsecutivedaysatPantiacollaLodgeWhite-shoulderedTanager-TachyphonusluctuosusAtPantiacollaLodgeinamixedflockWhite-wingedShrike-Tanager-LanioversicolorThecanopyflockleaderseenatPantiacollaLodge.ThecaniopyflockleaderMasked-crimsonTanager-RamphocelusnigrogulariSeenatManuWildlifeCenterSilver-beakedTanager(Ramphoceluscarbo)CommonintheAmazonialowlandsbutnotinprimaryforestHoodedMountain-Tanager-Buthraupismontana3seenaboveWayqechaonourtraveldaytoCockoftheRockLodgeScarlet-belliedMountain-Tanager-AnisognathusigniventrisAnotherattractivebird–seenalsoaboveWayqechaonourtraveldaytoCORLFawn-breastedTanager-Pipraeideamelanonota1seenatRocotalBlueandYellowTanager-PipraeideabonariensisSeenatHuacarpayLakesandalsoontheManuroadonourtraveldaytoCORLOrange-earedTanager-ChlorochrysacalliparaeaCommonatCockoftheRockLodgeinthegardenandalongtheroadBlue-GrayTanager-ThraupisepiscopusAcommonbirdinopenareasintheAmazon.Episcopus–areferencetotheepiscopalblueplumageofthisspecies.

PalmTanager-ThraupispalmarumCommonintheLowlandsMaskedTanager-TangaranigrocinctaSeeninthegardenofPantiacollaLodgeBlue-neckedTanager-TangaracyanicollisAnothercommonbirdincloudforest–seenatCockoftheRockLodge

Page 45: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Beryl-spangledTanager-TangaranigroviridisAnothercommonbirdincloudforestwithmixedflocksTurquoiseTanager-TangaramexicanaSeenatPantiacollaLodgeandManuWildlifeCenterParadiseTanager-TanagarachilensisStunningTanagerfoundinthefoothillsandlowlandsofManu.NotfoundinChile!

ParadiseTanager–CarolineSimpson

Opal-crownedTanager-Tangaracallophrys1seenatPantiacollaLodgeSaffron-crownedTanager-TangaraxanthocephalalamprotisAnothernicecolorfulbirdseeninthecloudforest–herethesubspecieswithanorange….notsaffroncrownGreenandGoldTanager-TangaraarthusSeenatPantiacollaLodgeGoldenTanager-TangaraarthusVerycommoninthecloudforest

Page 46: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

SwallowTanager-TersinaviridisSeenatPantiacollaLodgeandManuWildlifeCenterBlack-facedDacnis-DacnislineataAlsoseenatPantiacollaLodgeandManuWildlifeCenterYellow-belliedDacnis-DacnisflaviventerAtPantiacollaLodgeandManuWildlifeCenterPurpleHoneycreeper-CyanerpescaeruleusAfemaleoneseenatPantiacollaLodgeGreenHoneycreeper-ChlorophanesspizaSeveralrecordedinthelowlandsGuiraTanager-HemithraupisguiraSeenatPantiacollaLodgeYellow-backedTanager-Hemithraupisflavicollis2seenatPantiacollaLodgeCinereousConebill-ConirostrumcinereumcinereumSeenonthehighercloudforestontheManuroadCappedConebill-ConirostrumalbifronsAmaleoneatRocotalMoustachedFlowerpiercer-Diglosamystacalisalbilinear2seenonourtraveldayonthehighercloudforestoftheManuroadBlack-throatedFlowerpiercer-DiglossopisbrunneiventrisCommononthehighercloudforestoftheManuroadMaskedFlowerpiercer-DiglossopiscyaneaCommoninthecloudforestPeruvianSierra-Finch-PhrygiluspunensisSeenonourtraveldaytoCockoftheRockLodgefromCusco-namedforthesouthernPeruviandepartmentofPunoMourningSierra-Finch-PhrygilusfruticetiAtHuacarpayLakesandalsoontheManuroadAsh-breastedSierra-Finch-PhrygilusplebejusSeenonthehighermountainsoftheManuroad

Page 47: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

EChestnut-breastedMountain-Finch–PoospizaCaesarGoodviewsofthisPeruvianendemiconthehighermountainoftheManuroadonourtraveldaytoCockoftheRockLodgefromCuscoBlue-blackGrassquit–Volatiniajacarina2seenontheagriculturalfieldnearafterPatriaonourtraveldaytoPantiacollaLodgeSlate-coloredSeedeater–SporophilaschistaceaAmaleoneseenatBlanquillomacawclaylickBlackandWhiteSeedeater–SporophilaluctuosaAfemaleoneseenatCockoftheRockLodgeDouble-collaredSeedeater-SporophilacaerulescensOnemaleseenatBlanquillomacawclaylickBlack-billedSeedFinch–Sporophilaatrirostris2atCamungoox-bowLakeChestnut-belliedSeed-Finch–SporophilaangolensisAmaleoneseenattheBlanquillomacawclaylickBand-tailedSeedeater-CatameniaanalisSeenatHuacarpayLakesandalsoontheManuroad–acommonbirdoftheAndesBananaquit-CoerebaflaveolaSeenatCockoftheRockLodge–notcommoninManuBuff-throatedSaltator-SaltatormaximusCommon in the lowlands – seen at Cock of the Rock Lodge and Pantiacolla Lodge.Klicka et al. (2007) found strong genetic support for a sister relationship betweenSaltator and core Thraupidae. Sushkin (1924) proposed that Saltatorwas thraupine,not emberizine/cardinaline. SACC proposal passed to transfer Saltator fromCardinalidae to Incertae Sedis. SACC proposal to transfer to Thraupidae did notpass.Barkeretal. (2013)foundthatSaltatorandSaltatriculawereembeddedintheThraupidae. SACC proposal passed to transfer to Thraupidae. Chavez et al. (2013)found that relationshipswithin the genus are not consistentwith the current linearsequenceofspecies.SACCproposalpassedtoreviselinearsequence.GrayishSaltator-SaltatorcoerulescensSeenatMacawclaylickandCamungoox-bowLakeGolden-billedSaltator–SaltatoraurantiirostrisWesawthisatHuacarpayLakes-acommonbirdoftheAndesSlate-coloredGrosbeak–SaltatorgrossusAmaleoneseenatPantiacollaLodge

Page 48: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

NEWWORLDSPARROWSANDALLIESRufous-collaredSparrow-ZonotrichiacapensisAverycommonbirdoftheAndesthroughoutPeruandSouthAmericaYellow-browedSparrow-AmmodramusaurifronsSeenbellowCockoftheRockLodgePectoralSparrow-ArremontaciturnusSeenatPantiacollaLodgeRRBlack-facedBrushfinch-AtalaptesmelanolaemusCommon in the cloud forest – seen at Rocotal. The Atlapetes genus has beencompletelyrevampedbasedonbio-chemicaldataandworkdoneatCopenhagenFieldMuseum. Formerly considered part of Rufous-napped Brush-Finch complex A.rufinucha. Atlapetes melanolaemus was formerly (Hellmayr 1938, Paynter 1970a,MeyerdeSchauensee1970,Ridgely&Tudor1989,Sibley&Monroe1990)consideredasubspeciesofA.rufinucha,butseeGarcía-Moreno&Fjeldså(1999).CommonChlorospingus–ChlorospingusflavopectusCommonintheManucloudforestShort-billedChlorospingus-ChlorospingusparvirostrisSeen above Cock of the Rock Lodge. Genetic data (REFS, Burns et al. 2002, 2003)indicatethegenusChlorospingusisnotamemberoftheThraupidae,but(Klickaetal.2007)amemberoftheEmberizidae.SACCproposalpassedtotransfertoEmberizidae.Barkeretal.(2013)furtherconfirmedtheplacementofthegenuswiththeNewWorldsparrows.FrankPitelka(inTordoff1954a)longagonotedtheemberizine-likebehaviorofChlorospingus.SACCproposalpassedtochangeEnglishnamesofthespeciesinthegenusfrom“Bush-Tanager”to“Chlorospingus”.Yellow-throatedChlorospingus-ChlorospingusflavigularisSeenatCockoftheRocklodgeandQuebradaQuitacalzon-commoninmixedflocks

CARDINALGROSBEAKSRed-crownedAnt-Tanager-HabiarubicaSeenatPantiacollaLodgeandManuWildlifeCenterBlue-blackGrosbeak-Cyanocompsacyanoides*

NEWWORLDWARBLERSBlackburnianWarbler–SetophagafuscaAfemaleseenatRocotal

Page 49: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Two-bandedWarbler-MyiothlypisbivittatusCommonaroundCockoftheRockLodgeEGolden-bellied(Cuzco)Warbler-MyiothlypischrysogasterSeenaroundQuebradaQuitacalzononourtraveldayfromCockoftheRockLodgetoPantiacolla Lodge. Ridgely & Greenfield (2001) treated the northern subspecieschlorophrys as a separate species – ChocoWarbler - fromMyiothlypis chrysogasterbased on differences in descriptions of songs; see Zimmer (1949) for rationale forconsidering them sister taxa. This Golden–bellied Warbler becomes a PeruvianendemicRusset-crownedWarbler-MyiothlypiscoronatusSeenatRocotalinamixedflockThree-striped(Yungas)Warbler-BasileuterustristriatuspunctipectusSeenbellowCockoftheRockLodge.Gutiérrez-Pintoetal. (2012)foundthatbroadlydefined Basileuterus tristriatus consists of 10 distinct clades, one of which is B.trifasciatus, which is the sister group to B. tristriatus tacarcunae of theDarién. Donegan (2014) provided evidence that the Bolivian punctipectus group(includingcanens)shouldbetreatedasaseparatespecies.Slate-throatedRedstard(Whitestart)-MyioborusminiatusSeenaroundCockoftheRockLodgeSpectacledRedstart(Whitestart)-MyioborusmelanocephalusSeenaroundRocotal2individualsinamixedflock

BLACKBIRDSCrestedOropendola-Psarocoliusdecumanusmaculosus5seenbellowCORLonourtraveldayfromCockoftheRockLodgeDusky-greenOropendola-PsarocoliusatrovirensSeenatCockoftheRockLodgeRusset-backedOropendola-PsarocoliusangustifronsalfrediCommonintheAmazonlowlandsOliveOropendola-PsarocoliusbifasciatusSeenatPantiacollaLodgeandManuWildlifeCenterCasquedCacique-Cacicusoseryi10+seenatManuWildlifeCenterYellow-rumpedCacique-CacicuscelaVerycommonbirdintheAmazonlowlandforestalwaysnestingnearthelodges

Page 50: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Red-rumpedCacique-CacicushaemorrhousSeenatManuWildlifeCenter(Southern)MountainCacique-Cacicuschrysonotus2 seen in theManu cloud forest on our travel day from Cusco to Cock of the RockLodge.Thenorthern(leucoramphus)andsouthern(chrysonotus)groupsofsubspecieswere treated as separate species by Blake (1968b), but most classifications havetreated them as a single species (e.g., Hellmayr 1937, Meyer de Schauensee 1966,1970, Ridgely & Tudor 1989, Fjeldså & Krabbe 1990; and usually as Cacicusleucoramphus,anerror,becausechrysonotushaspriority)becausespecimensnearthecontactzoneshowsomesignsofgeneflow(Hellmayr1937,Bond1953).However,seeJaramillo & Burke (1999) for possible reasons for ranking them as species; this wasfollowedbyRidgely&Greenfield (2001),Hilty (2003),andFraga(2011).Powelletal.(2014) found that the two were deeply divergent genetically, more so than someCacicustreatedasspecies,butdidnotsamplepopulationsanywherenearthecontactzone.SACCproposaltotreatleucoramphusasaseparatespeciesdidnotpass.Hosneretal.(2015b)foundevidenceforintergradationbetweenthetaxainAyacucho,Peru.EpauletOriole-IcteruscayanensisSeenatCamungoCanopytower-agoodviewOrange-backedTroupial-IcteruscroconotusSeenatCamungooxbowlakeRRPale-eyedBlackbird-Agelaiusxanthophthalmus3intotalseenatCamungoandBlancoox-bowLakes-verygoodviewsYellow-wingedBlackbird-AgelaiusThiliusCommonatHuacarpayLakesGiantCowbird-MolothrusoryzivoraCommononMadredeDiosRiverislands–abroodparasite

FINCHESHoodedSiskin-SpinusmagellanicaurubambensisVerycommonbirdoftheAndesseenatHuacarpayLakesandontheManuroadOlivaceousSiskin-SpinusolivaceaSeenatCockoftheRockLodgeThick-billedEuphonia-EuphonialaniirostrisSeenatManuWildlifeCenterWhite-loredEuphonia-EuphoniachryssopastaSeenatPantiacollaLodgeandManuWildlifeCenter

Page 51: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

White-ventedEuphonia–EuphoniaminutaOnemaleseenatPantiacollaLodgeOrange-belliedEuphonia-EuphoniaxanthogasterSeenatCockoftheRockLodgeBlue-nappedChlorophonia-Chlrophoniacyanea4seenatCockoftheRockLodge

MAMMALSMONKEYSUrubambaBrownTiti-Monkey–CallicebusurubambensisSeenatPantiacollaLodge.ENDANGEREDBlack-cappedSquirrelMonkey–SaimiriboliviensisSeentwicearoundManuWildlifeCenterLarge-headedCapuchin–Sapajusmacrocephalus.SeenatPantiacollaLodgeandManuWildlifeCenter

Shock-headedCapuchin–CarolineSimpson

Page 52: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

Shock-headedCapuchin–CebuscuscinusSeenatManuWildlifeCenterGray’sBald-facedSaki–PitrheciairrorataGeatviewofoneoncollpatrailatManuWildlifeCenterColombianRedHowlerMonkey–AlouattasemiculusAfamilygroupseenatPantiacollaLodge

ColombianRedHowlerMonkeyhavingabadmorning!-CarolineSimpson

BlackSpiderMonkey–AteleschamekSeenatPantiacollaLodgeandManuWildlifeCenter.ENDANGERED

DOGFAMILYCulpeo(AndeanFox)–LycalopexculpaeusOneseenatHuacarpayLakes

RACCOONFAMILYGiantOtter–Pteronurabrasilensis3individualsseenatCamungoOx-bowLake

Page 53: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

PECCARIESCollaredPeccary–Tayassutajacu2seenonthetrailtoBlancoox-bowLake

SQUIRRELSSanborn’sSquirrel–SciurussanborniSeenatCockoftheRockLodge

LARGERODENTSBrownAgouti–DasyproctavariegateCommonatCockoftheRockLodgeinthegarden

BATSLong-nosedBat–RhynchonycterisnasoSeenatBlancoOxbowLakeverycommonroostingintheroofoftheboathouse

REPTILES&AMPHIBIANSSide-neckedTurtle–PodocnemisunifilisSeenontheMadredeDiosRiverTreeBoa–Corallushortulanus1seenjustoutsidethedinnerroomatManuWildlifeCenter

TreeBoa–CarolineSimpson

Page 54: MANU EXPEDITIONS BIRDING TOURS · DAY-BY-DAY ACTIVITIES Oct 08th: A full day at Huacarpay Lakes and night in the Inca Capital Cusco Oct 09/10th: A two day trip to Machu-Picchu ( birds

StickInsect–CarolineSimpson