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Bull. Ir. biogeog. Soc. No. 32 (2008)
97
IRISH SPIDERS (ARACHNIDA: ARANEAE) COLLECTED DURING A FIVE-
YEAR, ISLAND-WIDE STUDY INCLUDING 696 NEW COUNTY RECORDS
Anne Oxbrough
Department of Zoology, Ecology and Plant Science, University College Cork, Cork,
Ireland.
Summary
This paper presents new county records from one of the largest studies of Irish spiders
to date. Habitats surveyed included peatlands, grasslands and plantation forests of various
ages and tree species. Pitfall traps were used to sample spiders and specimens were also
identified from Malaise traps and a DVac Suction sampler in a subset of sites. Over half of
the known Irish spider species were sampled (219 species) and 696 new county records
are presented. Many of the species with new county records were associated with forested
habitats indicating the lack of previous research in this habitat type. It is recommended
that spider sampling should continue in a variety of habitat types across a wide
geographical range if a full Irish species distribution is to be realised.
Introduction
The study of Irish spiders has largely been limited to small-scale surveys that have
focused on the records of individual species (for instance see Mackie, 1963; Finlay, 1966;
Wanless, 1965). In more recent years, several authors have made significant additions to
the Irish species’ list and contributed much needed information on spider ecology and
species’ distributions. Particular consideration has been given to the spider fauna of Irish
bogs and fens (e.g. Higgins, 1985; Helsdingen, 1997, 1998; Nolan, 2002a) which probably
reflects the relative importance of this habitat in a national and international context. The
publication of van Helsdingen’s (1996a) County distribution of Irish spiders was the first
attempt to collate all known Irish records and to create systematic species lists for each
county. Such ‘baseline’ information is vital if we are to gain a true understanding of the
distribution and ecology of Irish spiders.
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Bull. Ir. biogeog. Soc. No. 32 (2008)
98
Large-scale biodiversity studies are necessary in order to help determine the broader
ecological associations of a species. Such studies will help establish the Irish spider fauna
in an international context and ensure that this important terrestrial group is not
overlooked in management and conservation schemes. Although ‘open’ habitats such as
bogs and fens, and grasslands and heathlands have received some attention (see above and
also Gibson, 1984 and McFerran et al., 1994), relatively little research has focused on
Irish forested habitats which represent approximately 10% of the country’s land cover
(Forest Service, 2004). Plantations constitute over 93% of this forest cover (Teagasc: Irish
Agriculture and Food Development Authority, 2005) compared to ~1% accounted for by
native woodlands (Department of Agriculture and Food, 2003). The species distributions
presented in this paper represent the largest single study of Irish spiders to have been
carried out from a range of habitats which included peatlands, grasslands and also
plantation forest of various ages and tree species. The data was collected as part of the
BIOFOREST project (http//www.ucc.ie/bioforest/) which aimed to examine the affect of
afforestation on Irish flora and fauna using various taxonomic groups (ground-dwelling
plants, canopy epiphytes, birds, spiders and hoverflies) as indicators of biodiversity (for
project reports see Smith et al., 2005, 2006; Iremonger et al., 2006a, b).
Survey design
In total, 102 sites in 21 counties were surveyed over five years (for site details see
Appendix 1) and were comprised of three distinct sub-projects:-
1. The survey of sites typically used during afforestation (sub-project 3.1.1) encompassed
three major Irish habitat types (peatlands, wet grasslands and improved grasslands). A
paired-site approach was used where 25 sites across the three habitat types were
sampled with adjacent ‘paired’ stands of five-year old Sitka spruce (Picea sitchensis)
matched for soil type, altitude, and pre-planting habitat type. Sample plots within these
sites were located in areas that may be of specific biodiversity value, such as wet
flushes, riparian areas and hedgerows as well as areas representative of the site as a
whole.
Bull. Ir. biogeog. Soc. No. 32 (2008)
99
2. The survey of plantation forests at various stages of the forest cycle (sub-project 3.1.2)
examined 32 sites in Sitka spruce plantations in the following age groups: 5 years (pre-
canopy closure), 8-15 years (canopy closure); 20-30 years (time of the first thinning)
and 35-50 years (commercial maturity). Ash (Fraxinus excelsior) was surveyed in
three age groups: 5 years, 8-15 years and 50+ years, and non-intimately mixed sites of
Sitka spruce and ash in the following age groups: 5 years, 8-15 years and 35-50 years.
Sampling areas within these sites were located in areas of homogenous vegetation
typical of the site as a whole.
3. The survey of open space within mature Sitka spruce plantations (sub-project 3.1.3)
sampled 12 sites that contained various types of open space (forestry road edges, rides
and glades). In addition to these sites, eight ‘experimental road width’ sites were
surveyed. These comprised recently felled stands which had been re-planted with Sitka
spruce. Within each site, a 100m section of forest road was established with a ‘wide
buffer zone’ of 30m width, adjacent to this a 100m section was also established with a
‘standard’ buffer of 15m. These buffer zones were defined as the distance from the
first tree row (base of tree) spanning the forest road, thus giving differing areas of
unplanted ground at the forest road edge. With the exception of the experimental road-
width sites all of the stands surveyed were first rotation.
Methods
Field surveys were carried out during the summers of 2001-2005. Spiders were
sampled primarily using pitfall traps, however in a subset of sites spiders were also
identified from Malaise traps and a DVac vortex suction sampler situated in areas adjacent
to pitfall traps. Pitfall traps consisted of plastic cups (7cm diameter by 9cm depth) filled to
1cm depth with ethylene glycol and with two drainage slits cut 1cm from the rim of the
cup. A bulb corer was used to make a hole in the ground for the plastic cup, which was
placed so that the rim of the cup was flush with the ground’s surface. The suction sampler
was used for five minutes in a 2x2m area. All sampling took place between 10th May and
18th August with pitfall and Malaise traps changed once every three weeks. Various
environmental and habitat variables were recorded at each sampling plot (i.e. cover
abundance of vegetation layers, litter cover and depth, deadwood cover and soil organic
Bull. Ir. biogeog. Soc. No. 32 (2008)
100
content).
Pitfall traps were located in sampling plots (between 5-6 plots per site) that were
separated by a minimum of 50m and were a minimum of 50m from the edge of the site. In
each plot, the traps were arranged in a 4x4m grid with one trap set at each corner and one
in the centre, though which linear features were sampled (i.e. riparian areas, hedgerows).
The traps were arranged in a line, each one separated by 2m. For surveys of open space
within mature plantations (sub-project 3.1.3), traps were set in three transects which were
arranged from the centre of the open space into the forest, and each transect being
separated by 2m.
Pitfall samples were stored in 70% alcohol and the spiders were sorted from the catch.
Identification of spiders to species level was carried out using a 50x magnification
microscope and nomenclature follows Roberts (1993). Juveniles were not identified due to
the difficulty involved in assigning them to species and published records were used to
establish new county records (Helsdingen, 1996a, b, 1998; McFerran, 1997; Smith and
Costello, 1998; Snazell and Jonsson, 1999; Merrett, 2000; Nolan, 2000a, b, 2002a, b,
2004; Cawley, 2001, 2004, 2007; Johnston and Cameron, 2002; Fahy and Gormally, 2003;
Nelson, 2005; McCormack, 2006; Oxbrough, 2007). Reference specimens are held in the
museum collection at University College Cork and can be obtained from the author. All
specimens were identified by the author and confirmations of difficult specimens were
made by Bob Johnston and Peter Merrett. The author is currently involved in analysing the
habitat associations of the species surveyed in this project and these data will be presented
in a separate paper.
Results
Over half of the known Irish spider fauna was found during the study with one new
Irish species and several rare species (reported in Oxbrough, 2007 and the present study).
In total 65,063 mature adults were identified representing 219 species from 17 families. Of
these species, 79% had new county records (NCRs) identified, a full list of which is given
in Appendix 2.
Bull. Ir. biogeog. Soc. No. 32 (2008)
101
Distribution of new county records
Overall, by far the most NCRs were identified in Counties Laois and Limerick (Table
1), whereas County Fermanagh had the lowest number of NCRs. Across the counties there
was a disproportionate number of sites surveyed and examination of the mean NCRs per
site within each county reveals that despite 10 or more sites being surveyed in Counties
Cork, Laois, Tipperary and Wicklow, there are relatively few NCRs per site. By contrast,
in Counties Dublin, Kildare and Leitrim where only 1 or 2 sites where surveyed, there
were over 20 NCRs identified per site.
TABLE 1. The frequency of new county records (NCRs) identified within each
county.
County (code) Total NCRs per county
Total sites surveyed per county
Mean NCRs per site
Limerick (LIM) 85 8 10.6 Laois (LAO) 57 11 5.2 Waterford (WAT) 49 4 12.3 Wicklow (WIC) 46 10 4.6 Kilkenny (KLK) 45 5 9.0 Tipperary (TIP) 45 11 4.1 Leitrim (LEI) 41 2 20.5 Kerry (KER) 41 9 4.6 Cork (COR) 37 13 2.8 Donegal (DON) 33 4 8.3 Mayo (MAY) 33 2 16.5 Galway (GAL) 32 5 6.4 Dublin (DUB) 24 1 24.0 Clare (CLA) 23 6 3.8 Kildare (KLD) 21 1 21.0 Westmeath (WES) 19 1 19.0 Offaly (OFF) 17 2 8.5 Sligo (SLI) 16 1 16.0 Carlow (CAR) 14 1 14.0 Wexford (WEX) 13 2 6.5 Fermanagh (FER) 8 2 4.0
The species with multiple new county records are shown in Table 2 and comprise 30%
of all NCRs. The most new county records were identified for the following species:
Lepthyphantes tenebricola, Micrargus herbigradus, Pocadicnemis juncea and Agyneta
Bull. Ir. biogeog. Soc. No. 32 (2008)
102
ramosa. The majority of these species’ NCRs were from plantation forest habitats,
however several species were found in a relatively high number of wet grassland sites
within each county such as Trochosa spinipalpis and P. juncea.
TABLE 2. Species with the most NCRs.
Species NCRs Lepthyphantes tenebricola 16 Pocadicnemis juncea 16 Micrargus herbigradus 15 Agyneta ramosa 14 Dicymbium tibiale 13 Walckenaeria vigilax 12 Lepthyphantes alacris 11 Asthenargus paganus 11 Ceratinella scabrosa 11 Maro minutus 11 Trochosa spinipalpis 11 Taranucnus setosus 10 Bathyphantes parvulus 10 Diplocephalus latifrons 10 Pirata ulignosus 10 Saaristoa abnormis 10 Saaristoa firma 10 Agyneta decora 9
Discussion
Overall, it should be expected that more new county records will be identified in
counties where a greater number of sites were surveyed. However as Table 1 shows, this
was not always the case. The high number of new county records presented is likely to be
a factor of both the large-scale nature of the study (102 sites across 21 counties) and the
range of habitats sampled, as well as the pattern in the recording of historical and recent
records.
Several rare species were collected during the study (see Oxbrough, 2007), however it
is likely that many of the new county records presented here are due to a lack of past
recording rather than actual species’ rarity. For example, A. ramosa has only recently been
added to the Irish list (Fahy and Gormally, 2003); whereas L. tenebricola has only been
Bull. Ir. biogeog. Soc. No. 32 (2008)
103
found in three Irish counties prior to this study, however both are known to be widespread
across Britain (Harvey et al., 2002). It is likely that these species’ preference for forested
habitats explains the few records to date. Indeed, several other species with a high number
of new county records are associated with forested habitats such as A. paganus and C.
scabrosa (Harvey et al., 2002). In Britain, both species have rather local and rare
distributions (Harvey et al., 2002) highlighting the need for further exploration in Irish
forested habitats. The geographical distribution of the new county records may reflect the
distribution of Irish plantation forests of which there are many in Counties Laois,
Limerick, Tipperary and Wicklow and towards which our sampling was directed.
However this can also be attributed to a ‘sampler bias’ effect related to where
arachnologists live or particular habitats which are interest to them. In County Dublin,
where one might expect the spider fauna to be relatively well known there were 24 new
county records identified from just one mature plantation site. However this probably
represents the lack of sampling carried out in forested habitats in this county.
Difficulty with species identification and changes in taxonomic classification may also
give a ‘false’ impression of species rarity. For example, Pocadicnemis juncea, with 16
new county records in the present study, was only relatively recently described as a
separate species from P. pumila (Millidge, 1975) and difficulty can be encountered when
distinguishing female specimens which may further distort records.
Although recent studies have greatly improved our knowledge of Irish spiders, it is still
vital to continue collecting baseline distribution information. In addition, sampling spiders
in a variety of habitat types using a wide range of sampling techniques will further
enhance our knowledge of their ecology.
Acknowledgements
I would like to thank my colleagues on the Bioforest project for help with survey
design, fieldwork and manuscript preparation. I would also like to thank Peter Merrett and
Robert Johnston for their help with identifying difficult specimens.
Bull. Ir. biogeog. Soc. No. 32 (2008)
104
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APPENDIX 1. Study site locations and corresponding habitat types; the sampling method
used is also given: P = pitfalls; S = Vortex suction sampler; M = Malaise trap. Further site
details including soil and vegetation information can be obtained from the aforementioned
project reports. ‘Planted’ refers to a 5 year old Sitka spruce plantation established on a
particular habitat type; OS = Open space; SS = Sitka spruce
Site code
Site name Grid Ref County Habitat Sampling method
Sub-project 3.1.1Agho Aghoney S547875 Laois Improved
grassland P
Balb Ballybeagh S349560 Kilkenny Planted improved grassland
P, S, M
Balp Ballynanoose R820686 Tipperary Planted improved grassland
P
Balu Ballynanoose R823684 Tipperary Improved grassland
P
Bght Ballybought N314325 Offaly Improved grassland
P
Bool Boolavaun R227820 Clare Wet grassland P Carp Carnamoyle C430294 Donegal Planted peatland P Caru Carnamoyle C434291 Donegal Peatland P Cast Castletown S279752 Kilkenny Improved
grassland P
Clar Clarbarracum S508862 Laois Wet grassland P Clop Cloonoughter R146464 Limerick Planted wet
grassland P
Clou Cloonoughter R148466 Limerick Wet grassland P Cooa Coolross R722649 Tipperary Planted wet
grassland P, S, M
Coog Coolsnaghtig W210559 Cork Wet grassland P Curr Curraghnaboul R723482 Limerick Wet grassland P Dong Donaghmore S266798 Laois Wet grassland P Doon Doon R978063 Tipperary Improved
grassland P
Flem Flemingstown R183064 Tipperary Improved grassland
P
Garv Garvoghil R230820 Clare Wet grassland P Gary Garyandrew S108354 Tipperary Improved
grassland P
Bull. Ir. biogeog. Soc. No. 32 (2008)
109
Geap Gearha V784729 Kerry Planted wet grassland
P
Geau Gearha V781725 Kerry Peatland P Glas Garryglas S508868 Laois Planted wet
grassland P
Glen Glenfield North R441147 Cork Planted wet grassland
P
Gore Gortnaree H187721 Fermanagh Planted wet grassland
P, S, M
Hanp Ballycahan N310325 Westmeath Planted improved grassland
P
Incp Inchamore R572993 Galway Planted wet grassland
P, S, M
Incu Inchamore R577997 Galway Peatland P, S, M Kilb Kilbraugh S295551 Tipperary Improved
grassland P, S, M
Kill Kilcullen S623410 Kilkenny Improved grassland
P
Knaw Knawhill R440148 Cork Wet grassland P Lead Knocklead S548861 Laois Planted improved
grassland P
Mntp Mountphillips R721648 Tipperary Wet grassland P Moaf Moanfune W987925 Waterford Improved
grassland P, S, M
Moig Moigh R720484 Limerick Wet grassland P Mull Mullanmeen
Under H162712 Fermanagh Wet grassland P, S, M
Muny Mungmacody S625415 Kilkenny Planted improved grassland
P
Ratr Rathreagh S281725 Kilkenny Planted improved grassland
P
Raup Curraun H102005 Leitrim Planted wet grassland
P
Rauu Curraun H102004 Leitrim Wet grassland P Slip Slievecorragh N969045 Wicklow Planted peatland P, S, M Sliu Slievecorragh N972046 Wicklow Peatland P, S, M Tiep Tieveclougher H124884 Donegal Planted wet
grassland P
Tieu Tieveclougher H124883 Donegal Peatland P Togp Toreenagowan R821146 Kerry Planted wet
grassland P, S, M
Togu Toreenagowan R840152 Kerry Peatland P, S, M Toop Toreenmore V523766 Kerry Planted wet
grassland P, S, M
Toou Toreenmore V519766 Kerry Peatland P
Bull. Ir. biogeog. Soc. No. 32 (2008)
110
Veep Ballyveeny F835050 Mayo Planted peatland P Veeu Ballyveeny F833050 Mayo Peatland P Sub-project 3.1.2 Bale Ballyea R197835 Clare Prethicket SS
plantation P
Baly Ballygiblin R461027 Cork Mature ash plantation
P
Barn Barnadown T139542 Wexford Mature ash plantation
P
Boky Buffanoky R811564 Limerick Mid-rotation SS plantation
P
Clyd Clydaghroe W182318 Kerry Thicket SS plantation
P
Comm Commeanaline W900538 Tipperary Thicket SS plantation
P
Cooa Coolross R720651 Tipperary Prethicket SS plantation
P
Cool Cooltmurraghy M774276 Galway Thicket ash plantation
P
Coon Cooneen Hill R916687 Tipperary Mature SS plantation
P
Corb Corbettstown N539394 Offaly Thicket ash plantation
P
Corr Corracloon R584913 Clare Mid-rotation SS plantation
P
Cumm Cummeenavrick W142816 Kerry Thicket SS plantation
P
Deme Demesne N831322 Kildare Mature ash plantation
P
Derr Derrybrien R625010 Galway Mature SS plantation
P
Doog Dooglaun R536916 Clare Thicket SS plantation
P
Fury Fuhiry W144734 Cork Mature SS plantation
P
Gfin Garrafin S291959 Laois Thicket SS plantation
P
Glyn Glynn’s Hill R551995 Clare Thicket SS plantation
P
Kduff Kilduffahoo R788555 Limerick Thicket SS plantation
P
Kila Kilalongford S967743 Carlow Mature SS plantation
P
Kilm Kilmacow W979926 Cork Prethicket SS plantation
P, S, M
Bull. Ir. biogeog. Soc. No. 32 (2008)
111
Lurg Lurgan Great M761227 Galway Prethicket SS plantation
P
Mary Marymount S260938 Laois Thicket SS plantation
P
Msop Monasop S278995 Laois Mature SS plantation
P
Mvan Moanvaun R891554 Tipperary Thicket ash plantation
P
Rath Rathcarrick G636349 Sligo Mature SS plantation
P
Reen Reenavanna R849544 Limerick Thicket ash plantation
P
Rinc Rincrew X081818 Waterford Mature ash plantation
P
Sagg Derrynasaggart W144800 Cork Mid-rotation SS plantation
P
Sinb Sinotts Bog T061663 Wexford Mature SS plantation
P
Suns Sunderlands T250809 Wicklow Mature SS plantation
P
Trum Trumra 292958 Laois Mid-rotation SS plantation
P
Sub-project 3.1.3 Athn Athdown O079158 Wicklow OS transect in
mature SS plantation
P
Bawn Bawnogue O016032 Wicklow Forest road edge in clearfelled plantation
P
Bmut Ballysmuttan O045145 Wicklow OS transect in mature SS plantation
P
Card Cardtown S277994 Laois Forest road edge in clearfelled plantation
P
Carr Carrigagulla W375836 Cork OS transect in mature SS plantation
P
Clea Cleanglass R234216 Limerick OS transect in mature SS plantation
P
Cloo Clootycarthy W208707 Cork Forest road edge in clearfelled plantation
P
Bull. Ir. biogeog. Soc. No. 32 (2008)
112
Cura Ballycurragh T052822 Wicklow OS transect in mature SS plantation
P
Foss Fossyhill S549891 Laois Forest road edge in clearfelled plantation
P
Gate Ballingate S976605 Wicklow Forest road edge in clearfelled plantation
P
Glan Glannaharee W454883 Cork OS transect in mature SS plantation
P
Gull Carrigagulla W371837 Cork Forest road edge in clearfelled plantation
P
Knoc Knocknagoum Q958215 Kerry OS transect in mature SS plantation
P
Lugg Lugg O032246 Dublin OS transect in mature SS plantation
P
Meen Meentinny R251135 Cork OS transect in mature SS plantation
P
More Lismore S027063 Waterford Forest road edge in clearfelled plantation
P
Muck Mucklagh One T085864 Wicklow OS transect in mature SS plantation
P
Rean Reanahoun R256200 Cork OS transect in mature SS plantation
P
Stoe Ballinastoe O180084 Wicklow OS transect in mature SS plantation
P
Toor Tooranahareen S128061 Waterford Forest road edge in clearfelled plantation
P
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
113
APP
EN
DIX
2. N
ew c
ount
y re
cord
s (N
CR
) of t
he sp
ecie
s sur
veye
d. P
aren
thes
es in
dica
te th
e si
te c
ode
and
num
ber o
f ind
ivid
uals
sam
pled
; Site
code
s fol
low
ed b
y ei
ther
‘S’ o
r ‘M
’ den
otes
a sp
ecim
en tr
appe
d w
ith ei
ther
a V
orte
x su
ctio
n sa
mpl
er (S
) or M
alai
se tr
ap
(M).
Spec
ies
No.
N
CR
New
Cou
nty
Rec
ords
Fam
ily S
eges
triid
ae
Sege
stri
a se
nocu
lata
(Lin
naeu
s)
2 LI
M (C
urr=
3, M
oig=
1); O
FF (B
ght=
3)
Fam
ily G
naph
osid
ae
Dra
ssod
es c
upre
us (B
lack
wal
l) 1
WA
T (M
ore=
1)
Dra
ssod
es la
pido
sus (
Wal
cken
aer)
1
KLK
(Cas
t=1)
H
aplo
dras
sus s
igni
fer (
C.L
. Koc
h)
2 C
OR
(Clo
o=1,
Gul
l=2,
Mee
n=1,
Rea
n=2)
M
icar
ia p
ulic
aria
(Sun
deva
ll)
1 LA
O (C
ard=
1)
Fam
ily C
lubi
onid
ae
Agro
eca
prox
ima
(O.P
.-Cam
brid
ge)
3 C
OR
(Mee
n=5)
; KER
(Cly
d=1)
; MA
Y (V
eep=
1)
Clu
bion
a br
evip
es (B
lack
wal
l) 1
TIP
(Kilb
M)
Clu
bion
a co
mta
(C.L
. Koc
h)
4 C
OR
(Kilm
M=1
); LE
I (R
aup=
1); L
IM (C
lop=
1, C
lou=
2, C
urr=
1); O
FF (B
ght=
1)
Clu
bion
a di
vers
a (O
.P.-C
ambr
idge
) 2
TIP
(Mnt
pM=2
, Mva
n=1)
; WIC
(Slip
M=1
, Sliu
=2, S
liuM
=1)
Clu
bion
a lu
tesc
ens (
Wes
tring
) 2
KLK
(Rat
r=1)
; TIP
(Coo
aM=1
2, M
ntpM
=19)
C
lubi
ona
triv
ialis
(C.L
. Koc
h)
5 C
LA (B
ale=
1); K
ER (G
eap=
1, T
oguM
=3, T
oopM
=2, T
oou=
11);
LIM
(Cle
a=1)
; MA
Y (V
eeu=
1);
TIP
(Coo
aM=1
) Fa
mily
Zor
idae
Zo
ra sp
inim
ana
(Sun
deva
ll)
2 LA
O (F
oss=
1); W
AT
(Mor
e=24
)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
114
Fam
ily T
hom
isid
ae
Ozy
ptila
trux
(Bla
ckw
all)
2 K
LK (C
ast=
11, K
ill=4
, Mun
y=6)
; LIM
(Cle
a=6,
Clo
p=1,
Clo
u=3,
Cur
r=4,
Moi
g=7,
Ree
n=4)
Ph
ilodr
omus
ces
pitu
m (W
alck
enae
r)
1 TI
P (C
ooaM
=1, K
ilbM
=1, M
ntpM
=33)
Ti
bellu
s mar
itim
us (M
enge
) 3
CO
R (C
loo=
1); L
AO
(Fos
s=1)
; WA
T (M
ore=
1)
Tibe
llus o
blon
gus (
Wal
cken
aer)
1
MA
Y (V
eeu=
1)
Xyst
icus
cri
stat
us (C
lerc
k)
1 K
LK (C
ast=
3, K
ill=1
) Xy
stic
us e
rrat
icus
(Bla
ckw
all)
1 D
ON
(Tie
u=1)
Xy
stic
us u
lmi (
Hah
n)
1 K
ER (
Gea
p=1)
Fa
mily
Sal
ticid
ae
Neo
n re
ticul
atus
(Bla
ckw
all)
1 C
OR
(Car
r=1)
Fa
mily
Lyc
osid
ae
Alop
ecos
a pu
lver
ulen
ta (C
lerc
k)
4 K
LK (C
ast=
2, K
ill=1
, Mun
y=1,
Rat
r=1)
; LA
O (A
gho=
1, C
ard=
11, F
oss=
7); L
EI (R
aup=
2,
Rau
u=2)
; WES
(Han
p=1)
Ar
ctos
a pe
rita
(Lat
rielle
) 1
LAO
(Fos
s=1)
Pa
rdos
a ag
rest
is (W
estri
ng)
1 TI
P (K
ilb=1
, Mnt
p=1)
Pa
rdos
a ag
rico
la (T
hore
ll)
1 TI
P (M
ntp=
1)
Pard
osa
amen
tata
(Cle
rck)
1
KLK
(Cas
t=54
, Kill
=45,
Mun
y=31
, Rat
r=72
) Pa
rdos
a ni
gric
eps (
Thor
ell)
3 LA
O (C
lar=
3, G
las=
1, L
ead=
5); L
IM (C
lea=
5, C
lop=
10, C
lou=
2, R
een=
3); W
ES (H
anp=
1)
Pard
osa
palu
stri
s (Li
nnae
us)
4 K
LK (C
ast=
14, K
ill=2
0); L
AO
(Car
d=2,
Cla
r=17
, Fos
s=11
, Mso
p=1)
; LIM
(Clo
p=1,
Clo
u=83
, C
urr=
17, M
oig=
35);
WA
T (M
oaf=
9, M
ore=
2, T
oor=
3)
Pira
ta h
ygro
philu
s (Th
orel
l) 2
MA
Y (V
eeu=
1); W
AT
(Too
r=1)
Pi
rata
latit
ans (
Bla
ckw
all)
4 C
LA (D
oog=
6); G
AL
(Inc
p=1)
; LA
O (C
ard=
9, M
ary=
1); L
IM (M
oig=
2)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
115
Pira
ta u
ligin
osus
(Tho
rell)
9
CO
R (C
arr=
24, C
loo=
2, F
ury=
1, G
lan=
1, K
ilm=3
, Mee
n=1,
Rea
n=15
); D
ON
(Tie
p=6)
; GA
L (I
ncp=
7) K
ER (G
eap=
79, G
eau=
44, K
noc=
4, T
ogp=
8, T
ogu=
5, T
oou=
5); L
AO
(Car
d=18
); LI
M
(Cle
a=7,
Clo
u=1)
; MA
Y (V
eep=
31, V
eeu=
20);
WA
T (M
ore=
7); W
ES (H
anp=
1); W
IC (B
mut
=1,
Slip
=11)
Tr
ocho
sa sp
inip
alpi
s (O
.P.-
Cam
brid
ge)
11
CLA
(Boo
l=32
, Gar
v=3)
; CO
R (G
len=
8, K
naw
=7);
DU
B (L
ugg=
1); G
AL
(Inc
u=3)
; KER
(G
eap=
1, G
eau=
3); K
LK (M
uny=
2); L
AO
(Cla
r=1,
Gla
s=1)
; LEI
(Rau
p=17
, Rau
u=34
); TI
P (B
alp=
1, C
ooa=
1, D
oon=
1, M
ntp=
1); W
AT
(Mor
e=1)
; WIC
(Gat
e=2)
Tr
ocho
sa te
rric
ola
(Tho
rell)
2
LAO
(Agh
o=1,
Cla
r=5,
Don
g=1,
Fos
s=6,
Gfin
=1, G
las=
1, L
ead=
1); L
EI (R
aup=
1,R
auu=
3)
Fam
ily P
isau
ridae
D
olom
edes
fim
bria
tus (
Cle
rck)
1
LIM
(Cur
r=8)
Pi
saur
a m
irab
ilis (
Cle
crk)
2
LAO
(Car
d=2)
; LEI
(Rau
p=1)
Fa
mily
Dic
tyni
dae
Cry
phoe
ca si
lvic
ola
(C.L
. Koc
h)
5 C
OR
(Kna
w=1
); D
ON
(Tie
p=1)
; LA
O (F
oss=
1); L
IM (C
urr=
2); O
FF (B
ght=
1)
Fam
ily H
ahni
idae
An
tiste
a el
egan
s (B
lack
wal
l) 2
DO
N (C
arp=
1, C
aru=
12, T
iep=
2, T
ieu=
12);
LIM
(Cle
a=2,
Cur
r=3)
H
ahni
a m
onta
na (
Bla
ckw
all)
1 W
IC (S
liu=2
) H
ahni
a na
va (B
lack
wal
l) 2
CO
R (C
lar=
1); L
IM (C
lea=
1)
Fam
ily M
imet
idae
Er
o ca
mbr
idge
i (K
ulcz
ynsk
i) 3
LIM
(Clo
p=1,
Cur
r=1)
; MA
Y (V
eep=
5); W
IC (G
ate=
2, S
lip=1
) Er
o fu
rcat
a (V
iller
s)
1 LI
M (C
lea=
1)
Fam
ily T
herid
iidae
Eu
ryop
sis f
lavo
mac
ulat
a (L
. Koc
h)
2 M
AY
(Vee
u=30
); W
AT
(Mor
e=8)
Ph
olco
mm
a gi
bbum
(Wes
tring
) 4
DO
N (C
arp=
1); L
IM (C
lop=
1); T
IP (B
alp=
2); W
IC (B
awn=
2, B
mut
=1, S
lip=2
) Ro
bert
us a
rund
inet
i (O
.P.-
Cam
brid
ge)
2 C
OR
(Gla
n=1)
; KER
(Gea
p=2,
Tog
p=1,
Tog
u=1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
116
Ro
bert
us li
vidu
s (B
lack
wal
l) 2
LAO
(Agh
o=1,
Car
d=31
, Fos
s=21
, Gfin
=16,
Gla
s=4,
Lea
d=35
, Mar
y=44
, Mso
p=10
, Tru
m=4
0);
LEI (
Rau
p=2)
Ro
bert
us n
egle
ctus
(O.P
.-C
ambr
idge
) 2
CLA
(Doo
g=5)
; KLK
(Kill
=1)
Theo
noe
min
utis
sim
a (O
.P.-
Cam
brid
ge)
7 C
AR
(Kila
=1);
FER
(Gor
e=1)
; KER
(Cum
m=7
, Gea
p=1,
Kno
c=5,
Tog
p=3,
Too
p=1,
Too
pS=5
, To
ouS=
2); L
AO
(Mso
p=1)
; LIM
(Cle
a=25
); W
AT
(Too
r=1)
; WEX
(Sin
b=3)
Th
erid
ion
bim
acul
atum
(Lin
naeu
s)2
TIP
(Mnt
pM=1
); W
AT
(Mor
e=2,
Too
r=2)
Th
erid
ion
impr
essu
m (L
. Koc
h)
3 FE
R (M
ullM
=1);
KLK
(Bal
bM=1
); TI
P (M
ntpM
=2)
Ther
idio
n in
stab
ile (O
.P.-
Cam
brid
ge)
6 K
LK (R
atr=
1); L
AO
(Gfin
=1, L
ead=
1); L
EI (R
aup=
1); O
FF (C
orb=
1); T
IP (C
ooa=
3); W
IC
(Slip
M=1
) Th
erid
ion
vari
ans (
Hah
n)
1 TI
P (M
ntpM
=1)
Fam
ily N
estic
idae
N
estic
us c
ellu
lanu
s (Th
orel
l) 1
WIC
(Ath
n=1)
Fa
mily
Tet
ragn
athi
dae
Met
a m
enge
i (B
lack
wal
l) 4
CA
R (K
ila=1
); D
UB
(Lug
g=1)
; LIM
(Clo
p=2,
Kdu
f=1)
; WA
T (T
oor=
1)
Met
a m
eria
nae
(Sco
pli)
1 LI
M (B
oky=
1, K
duf=
1)
Met
a se
gmen
tata
(Cle
rck)
4
KLK
(Bal
bM=3
, Kill
=1, M
uny=
1); L
AO
(Gla
s=1)
; LIM
(Moi
g=1)
; TIP
(Mnt
pM=6
) Pa
chyg
nath
a de
geer
i (Su
ndev
all)
2 LE
I (R
auu=
2); L
IM (C
lop=
8, C
lou=
17, C
urr=
2, M
oig=
20)
Tetr
agna
tha
mon
tana
(Sim
on)
2 G
AL
(Inc
pM=3
, Inc
uM=1
2); L
IM (M
oig=
1)
Fam
ily A
rane
idae
Ar
anie
lla o
pist
ogra
pha
(Kul
czyn
ski)
2 G
AL
(Inc
uM=1
); TI
P (K
ilbM
=1)
Epis
inus
trun
catu
s (La
triel
le)
1 C
OR
(Kilm
=1)
Hyp
osin
ga p
ygm
aea
(Sun
deva
ll)
1 D
ON
(Car
p=1)
Fa
mily
The
ridio
som
atid
ae
Ther
idio
som
a ge
mm
osum
(Koc
h)
2 LI
M (C
lop=
1); T
IP (C
ooaM
=1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
117
Fam
ily L
inyp
hiid
ae
Agyn
eta
caut
a (O
.P.-C
ambr
idge
) 4
GA
L (I
ncu=
1), K
ER (G
eap=
5), M
AY
(Vee
p=12
) Ag
ynet
a co
nige
ra (O
.P.-C
ambr
idge
) 7
KER
(Tog
p=1)
, KLK
(Mun
y=1)
, LIM
(Bok
y=1,
Clo
u=1,
Kdu
f=1)
, MA
Y (V
eep=
1), O
FF
(Bgh
t=2)
, TIP
(Com
m=4
, Fle
m=1
, Kilb
=1),
WIC
(Slip
=1, S
liu=1
) Ag
ynet
a de
cora
(O.P
.-Cam
brid
ge)
9 C
LA (B
ool=
1), C
OR
(Coo
g=3)
, KLK
(Kill
=1),L
AO
(Agh
o=5,
Cla
r=3,
Fos
s=3)
, LIM
(Kdu
f=1,
M
oig=
1, R
een=
2), M
AY
(Vee
p=40
, Vee
u=25
), TI
P (B
alu=
9, C
omm
=1, D
oon=
1, M
van=
1),
WA
T (M
ore=
68, T
oor=
1), W
ES (H
anp=
1)
Agyn
eta
oliv
acea
(Em
erto
n)
9 C
LA (G
arv=
1), C
OR
(Clo
o=1,
Gla
n=19
, Kilm
=1, M
een=
5, R
ean=
7), G
AL
(Inc
p=39
), K
ER
(Cly
d=2,
Gea
p=29
, Gea
u=12
, Kno
c=30
, Tog
p=2,
Tog
u=24
, Too
u=5)
, LA
O (C
lar=
1), L
EI
(Rau
u=1)
, LIM
(Cle
a=3,
Kdu
f=1,
Moi
g=1)
, MA
Y (V
eep=
123,
Vee
u=10
), W
AT
(Rin
c=1,
To
or=1
) Ag
ynet
a ra
mos
a (J
acks
on)
14
CA
R (K
ila=1
1), C
LA (B
ool=
24, C
orr=
1, G
arv=
13 ),
CO
R (C
lar=
6, C
loo=
6, C
oog=
1, G
lan=
25,
Gul
l=6,
Kna
w=7
, Mee
n=5,
Rea
n=4)
, DO
N (C
aru=
9, T
ieu=
3), D
UB
(Lug
g=4)
, KER
(Gea
p=1,
K
noc=
102,
Too
u=1)
, KLK
(Mun
y=1,
Rat
r=1)
, LA
O (A
gho=
1, C
ard=
2, C
lar=
4, F
oss=
8, G
las=
8,
Lead
=1),
LEI (
Rau
p=1,
Rau
u=1)
, LIM
(Bok
y=4,
Cle
a=17
), M
AY
(Vee
p=8)
, TIP
(Bal
p=6,
B
alu=
3, M
ntp=
1), W
AT
(Mor
e=16
, Too
r=21
), W
EX (S
inb=
1), W
IC (A
thn=
5, B
awn=
61,
Bm
ut=9
, Cur
a=5,
Gat
e=5,
Muc
k=12
) Ag
ynet
a su
btili
s (O
.P.-C
ambr
idge
) 8
KLD
(Dem
e=10
), K
LK (K
ill=3
), LA
O (C
ard=
7, F
oss=
11, G
fin=5
, Gla
s=15
, Mar
y=1,
Mso
p=13
, Tr
um=4
), LE
I (R
aup=
17, R
auu=
6), L
IM (B
oky=
2, C
lea=
111,
Cur
r=6,
Kdu
f=2,
Moi
g=5,
R
een=
1), S
LI (R
ath=
11),
WA
T (M
ore=
60, T
oor=
37),
WES
(Han
p=1)
Al
lom
enge
a sc
opig
era
(Gru
be)
1 K
ER (C
umm
=1)
Allo
men
gea
vidu
a (L
. Koc
h)
1 LI
M (C
lop=
1)
Aphi
leta
mis
era
(O.P
.-Cam
brid
ge)
2 C
OR
(Gla
n=1)
; KER
(Cly
d=2,
Cum
m=3
) Ar
aeon
cus c
rass
icep
s (W
estri
ng
3 G
AL
(Inc
uM=1
); K
ER (T
oouM
=1);
MA
Y (V
eeu=
16)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
118
As
then
argu
s pag
anus
(Sim
on)
11
CLA
(Coo
r=10
, Doo
g=2,
Gly
n=10
); D
UB
(Lug
g=18
); FE
R (G
ore=
1); G
AL
(Der
r=22
); K
LD
(Dem
e=1)
; KLK
(Cas
t=2)
; LA
O (C
ard=
5, F
oss=
4, G
fin=3
, Lea
d=1,
Mar
y=15
, Mso
p=8,
Tr
um=1
3); L
IM (B
oky=
16, C
lea=
2); T
IP (C
omm
=6, C
oon=
12);
WEX
(Sin
b=15
); W
IC
(Ath
n=15
, Baw
n=14
, Bm
ut=1
2, C
ura=
11, G
ate=
3, M
uck=
31, S
toe=
10, S
uns=
11)
Bary
phm
a tr
ifons
(O.P
.-Cam
brid
ge)
5 K
LK (B
alb=
1, K
ill=1
, Mun
y=3)
; LA
O (C
lar=
1, G
las=
2, L
ead=
2); L
EI (R
aup=
2, R
auu=
3); L
IM
(Clo
p=2,
Cur
r=2)
; MA
Y (V
eeu=
1)
Bath
ypha
ntes
app
roxi
mat
us (O
.P.-
Cam
brid
ge)
3 K
LK (R
atr=
2); L
AO
(Lea
d=1)
; LIM
(Clo
u=8,
Cur
r=1)
Bath
ypha
ntes
gra
cilis
(Bla
ckw
all)
1 LI
M (B
oky=
3, C
lea=
8, C
lop=
22, C
lou=
55, C
urr=
18, K
duf=
13, M
oig=
17, R
een=
35)
Ba
thyp
hant
es n
igri
nus (
Wes
tring
) 5
LAO
(Car
d=1,
Cla
r=3,
Fos
s=5,
Gla
s=12
, Lea
d=26
); LE
I (R
aup=
4, R
auu=
1); L
IM (C
lea=
7,
Clo
p=3,
Clo
u=1,
Kdu
f=1,
Ree
n=3)
; OFF
(Bgh
t=2,
Cor
b=1)
, SLI
(Rat
h=1)
Ba
thyp
hant
es p
arvu
lus (
Wes
tring
) 10
C
LA (B
ale=
22, B
ool=
4, D
oog=
4, G
arv=
16, G
lyn=
4); C
OR
(Cla
r=2,
Clo
o=2,
Coo
g=2,
Gle
n=19
, K
ilm=2
1); D
UB
(Lug
g=4)
; GA
L (C
ool=
16, I
ncp=
3, In
cu=2
, Lur
g=43
); K
ER (C
umm
=1, G
eap=
3,
Gea
u=3,
Tog
p=9,
Tog
pS=2
, Tog
u=6,
Too
p=2,
Too
u=1,
Too
uM=1
); K
LK (B
alb=
7,
Bal
bM=4
,Cas
t=4,
Kill
=8, M
uny=
107,
Rat
r=68
); LE
I (R
aup=
18,
Rau
u=1)
; LIM
(Cle
a=1,
C
lop=
21, C
lou=
7, C
urr=
29, K
duf=
1, M
oig=
9, R
een=
5); M
AY
(Vee
p=1,
Vee
u=32
); TI
P (B
alp=
10, B
alu=
23,
Coo
n=7,
Coo
a=23
, Coo
aS=1
, Coo
aM=2
, Doo
n=1,
Fle
m=6
1, K
ilb=6
, M
ntp=
3, M
ntpM
=1, M
van=
4)
Bath
ypha
ntes
setig
er (O
.P.-
Cam
brid
ge)
1 G
AL
(Inc
u=1)
Cen
trom
erita
con
cinn
a (T
hore
ll)
1 LA
O (C
ard=
3, C
lar=
2)
Cen
trom
erus
arc
anus
(O.P
.-C
ambr
idge
) 1
WA
T (M
ore=
2, T
oor=
1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
119
Cen
trom
erus
dilu
tus (
O.P
.-C
ambr
idge
) 7
FER
(Gor
e=2)
; KER
(Cly
d=4,
Cum
m=7
, Kno
c=11
, Too
p=1)
; KLD
(Dem
e=2)
; LIM
( Bok
y=4,
C
lea=
24);
MA
Y (V
eep=
1); S
LI (R
ath=
2); W
AT
(Moa
f=1,
Mor
e=3)
C
entr
omer
us p
rude
ns (O
.P.-
Cam
brid
ge)
1 LI
M (C
lea=
1)
Cen
trom
erus
sylv
atic
us (B
lack
wal
l) 3
LIM
(Clo
u=1)
; TIP
(Bal
p=1)
; WIC
(Baw
n=1,
Gat
e=1,
Sto
e=1)
C
erat
inel
la b
revi
pes (
Wes
tring
) 3
DO
N (C
arp=
58, C
aru=
77, T
iep=
60, T
ieu=
38);
LIM
(Cle
a=8,
Clo
p=1,
Cur
r=6,
Moi
g=1)
; WA
T (M
oaf=
1, M
ore=
30, T
oor=
8)
Cer
atin
ella
bre
vis (
Wid
er)
6 C
AR
(Kila
=1);
GA
L (I
ncp=
3); L
EI (R
aup=
1, R
auu=
3); L
IM (C
lea=
1); M
AY
(Vee
p=5)
; WA
T (M
oaf=
6, T
oor=
6)
Cer
atin
ella
scab
rosa
(O. P
.-C
ambr
idge
) 11
C
OR
(Kilm
=2, K
naw
=1);
KLD
(Dem
e=7)
; KLK
(Cas
t=9,
Kill
=2, M
uny=
8, R
atr=
5); L
AO
(C
lar=
4, F
oss=
1, G
las=
1, T
rum
=2);
LEI (
Rau
p=14
, Rau
u=8)
; LIM
(Clo
p=4,
Clo
u=36
, Cur
r=26
, M
oig=
27);
OFF
(Bgh
t=9)
; WA
T (M
oaf=
10);
WES
(Han
p=17
); W
EX (B
arn=
1, S
inb=
1); W
IC
(Slip
=1, S
uns=
4)
Cne
phal
ocot
es o
bscu
rus (
Bla
ckw
all)
4 D
ON
(Car
u=3,
Tie
u=22
); D
UB
(Lug
g=2)
; LEI
(Rau
u=2)
; OFF
(Cor
b=1)
D
icym
bium
nig
rum
(Bla
ckw
all)
4 C
OR
(Coo
g=1,
Gle
n=2,
Gul
l=1,
Kilm
=2, K
naw
=4, M
een=
1); L
EI (R
aup=
4, R
auu=
15);
LIM
(C
lop=
2, C
lou=
3, C
urr=
2, M
oig=
6); W
AT
(Moa
f=1,
Mor
e=7)
D
icym
bium
tibi
ale
(Bla
ckw
all)
13
CLA
(Bal
e=3,
Cor
r=1,
Doo
g=21
); C
OR
(Bal
y=34
, Cla
r=4,
Clo
o=1,
Gla
n=6,
Gul
l=3,
Kilm
=25,
M
een=
4, R
ean=
7); D
UB
(Lug
g=7)
; FER
(Gor
e=1)
; GA
L (C
ool=
4, In
cp=1
, Lur
g=3)
; KER
(C
lyd=
2, C
umm
=90,
Kno
c=1)
; KLD
(Dem
e=2)
; LA
O (C
ard=
10, C
lar=
2, F
oss=
20, G
fin=1
, M
sop=
1); L
IM (C
lea=
2, K
duf=
1, M
oig=
1, R
ean=
33);
OFF
(Cor
b=15
); SL
I (R
ath=
48);
TIP
(Com
m=2
, Coo
a=5,
Kilb
=2, M
ntp=
1, M
van=
12);
WEX
(Bar
n=19
, Sin
b=34
) D
iplo
cent
ria
bide
ntat
a (E
mer
ton)
1
WA
T (M
oafS
=1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
120
D
iplo
ceph
alus
latif
rons
(O.P
.-C
ambr
idge
) 10
C
OR
(Bal
y=96
, Cla
r=58
, Clo
o=2,
Fur
y=49
, Gla
n=81
, Gul
l=19
, Kilm
=1, M
een=
127,
Rea
n=97
, Sa
gg=6
5); D
ON
(Tie
u=1)
; GA
L (D
err=
18);
LAO
(Car
d=3,
Fos
s=18
, Gfin
=17,
Gla
s=9,
Mar
y=31
, M
sop=
68, T
rum
=70)
; LIM
(Bok
y=46
, Cle
a=86
, Kdu
f=23
); SL
I (R
ath=
37);
TIP
(Bal
u=2,
C
omm
=13,
Coo
n=36
, Gar
y=3,
Kilb
=2, M
ntp=
1, M
van=
1); W
AT
(Mor
e=10
, Too
r=2)
; WEX
(B
arn=
20, S
inb=
106)
; WIC
(Ath
n=7,
Baw
n=8,
Bm
ut=5
7, C
ura=
119,
Gat
e=11
, Muc
k=12
1,
Stoe
=43,
Sun
s=14
5)
Dip
loce
phal
us p
erm
ixtu
s (O
.P.-
Cam
brid
ge)
3 D
UB
(Lug
g=2)
; LIM
(Cle
a=3,
Clo
p=1,
Moi
g=15
); M
AY
(Vee
u=2)
Dip
loce
phal
us p
icin
us (B
lack
wal
l) 4
KLD
(Dem
e=1)
; LEI
(Rau
u=2)
; OFF
(Bgh
t=2)
; WIC
(Sun
s=3)
D
iplo
styl
a c
onco
lor (
Wid
er)
3 LA
O (F
oss=
22, L
ead=
2); W
AT
(Too
r=5)
; WES
(Han
p=1)
D
ism
odic
us b
ifron
s (B
lack
wal
l) 8
DO
N (C
arp=
4, C
aru=
1, T
iep=
7, T
ieu=
6); K
LK (B
alb=
1, B
albM
=27,
Cas
t=5,
Kill
=9, M
uny=
11,
Rat
r=19
); LE
I (R
aup=
6, R
auu=
4); L
IM (C
lea=
2, C
lop=
10, C
lou=
6, C
urr=
3, K
duf=
10, M
oig=
2,
Rea
n=1)
; OFF
(Bgh
t=10
); SL
I (R
ath=
2); W
AT
(Moa
f=3,
Mor
e=19
, Too
r=29
); W
ES (H
anp=
3)
Dre
pano
tylu
s unc
atus
(O.P
.-C
ambr
idge
) 1
DO
N (T
ieu=
1)
Erig
one
dent
ipal
pis (
Wid
er)
3 K
LK (C
ast=
17, K
ill=5
7); L
AO
(Agh
o=6,
Car
d=3,
Cla
r=4,
Don
g=1,
Fos
s=16
);WIC
(Ath
n=1,
G
ate=
2, S
liuM
=1, S
toe=
9)
Erig
one
long
ipal
pis (
Sund
eval
l) 3
CA
R (K
ila=1
); TI
P (C
omm
=3, C
oon=
1, K
ilb=2
); W
AT
(Moa
f=1)
Er
igon
ella
hie
mal
is (B
lack
wal
l) 5
CO
R (G
ull=
1); K
ER (K
noc=
1, T
oou=
1); K
LD (D
eme=
9); L
AO
(Car
d=2,
Don
g=1,
Fos
s=7,
Le
ad=1
); LI
M (M
oig=
2, R
ean=
1)
Erig
onel
la ig
nobi
lis (O
.P.-
Cam
brid
ge)
2 K
ER (G
eap=
1, G
eau=
1); W
AT
(Mor
e=1)
Gon
gylid
iellu
m la
tebr
icol
a (O
.P.-
Cam
brid
ge)
3 D
ON
(Car
p=1)
; GA
L (I
ncp=
7, In
cu=2
); K
ER (G
eap=
4, K
noc=
2, T
ogp=
1, T
ogu=
4, T
oop=
1,
Toou
=1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
121
Gon
gylid
iellu
m v
ivum
(O.P
.-C
ambr
idge
) 5
CO
R (C
arr=
8, C
oog=
3, C
loo=
1, F
ury=
12, G
lan=
29, G
len=
1, G
ull=
6, K
ilm=7
6, K
naw
=4,
Mee
n=9,
Rea
n=9,
Sag
g=5)
; DU
B (L
ugg=
16);
KLD
(Dem
e=3)
; LIM
(Bok
y=6,
Cle
a=19
, Clo
p=5,
C
lou=
9, C
urr=
6, K
duf=
25, M
oig=
1, R
ean=
16);
WIC
(Ath
n=5,
Baw
n=31
, Bm
ut=2
, Cur
a=7,
G
ate=
9, M
uck=
6, S
lip=8
, Sliu
=3, S
liuM
=1, S
toe=
13, S
uns=
11)
Gon
gylid
um ru
fipes
(Lin
naeu
s)
4 G
AL
(Inc
uM=1
); K
ER (C
lyd=
1, C
umm
=2, G
eau=
1); K
LK (M
uny=
1); W
IC (A
thn=
1)
Hila
ira
exci
sa (O
. P.-C
ambr
idge
) 8
CA
R (K
ila=1
); D
ON
(Car
p=1)
; DU
B (L
ugg=
1); G
AL
(Der
r=1)
; KER
(Cly
d=1)
; LA
O (G
fin=1
, M
ary=
1); L
IM (C
lea=
1, R
ean=
2); T
IP (C
omm
=24,
Coo
n=3,
Mva
n=3)
H
ypse
liste
s jac
kson
i (O
.P.-
Cam
brid
ge)
3 D
ON
(Car
p=2,
Car
u=1,
Tie
p=1,
Tie
u=7)
; KER
(Gea
u=2)
; WIC
(Baw
n=5)
Kae
stne
ria
pulla
ta (O
.P.-C
ambr
idge
) 7
DO
N (T
iep=
3, T
ieu=
7); G
AL
(Lur
g=2)
; KER
(Too
p=2,
Too
pM=8
, Too
u=7)
; KLK
(Bal
b=1,
M
uny=
4); M
AY
(Vee
u=1)
; OFF
(Cor
b=4)
; TIP
(Mva
n=6)
Le
pthy
phan
tes a
lacr
is (B
lack
wal
l) 11
D
UB
(Lug
g=35
); K
LD (D
eme=
1); K
LK (C
ast=
1); L
AO
(Car
d=1,
Gfin
=2, L
ead=
9, M
sop=
10,
Trum
=10)
; LEI
(Rau
p=3)
; LIM
(Bok
y=25
, Cle
a=12
1, K
duf=
2); M
AY
(Vee
u=1)
; SLI
(R
ath=
56);T
IP (B
alp=
3, B
alu=
1, C
omm
=2, C
oon=
6, D
oon=
3, F
lem
=1);
WEX
(Bar
n=1,
Sin
b=9)
; W
IC (A
thn=
34, B
awn=
1, B
mut
=48,
Cur
a=24
, Gat
e=3,
Muc
k=49
, Slip
=2, S
toe=
10, S
uns=
14)
Lept
hyph
ante
s cri
stat
us (M
enge
) 4
LAO
(Agh
o=1,
Car
d=1,
Cla
r=1,
Gla
s=5,
Lea
d=3)
; LEI
(Rau
p=3)
; TIP
(Bal
p=6,
Bal
u=1,
Coo
a=1,
M
van=
3); W
AT
(Too
r=2)
Le
pthy
phan
tes e
rica
eus (
Bla
ckw
all)
3 D
ON
(Car
p=14
, Car
u=22
, Tie
p=8,
Tie
u=15
); K
LK (B
alb=
5, B
albM
=1, B
albS
=3, C
ast=
2,
Mun
y=9,
Rat
r=28
); LE
I (R
aup=
8, R
auu=
5)
Lept
hyph
ante
s fla
vipe
s (B
lack
wal
l) 7
CLA
(Bal
e=1,
Doo
g=2,
Gly
n=3)
; KER
(Cum
m=1
, Too
p=1)
; KLD
(Dem
e=2)
; KLK
(Kill
=2);
LEI
(Rau
p=6)
; LIM
(Cle
a=4,
Kdu
f=3)
; SLI
(Rat
h=97
) Le
pthy
phan
tes m
enge
i (K
ulcz
ynsk
i) 3
LEI (
Rau
p=4,
Rau
u=7)
; LIM
(Cle
a=8,
Clo
p=5,
Clo
u=6)
; WA
T (M
ore=
11, T
oor=
11)
Lept
hyph
ante
s neb
ulos
us
(Sun
derv
all)
1 O
FF (C
orb=
1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
122
Lept
hyph
ante
s obs
curu
s (B
lack
wal
l) 5
GA
L (D
err=
1), K
LD (D
eme=
1); L
IM (C
lea=
8, K
duf=
1); T
IP (B
alu=
1, C
omm
=3, C
oon=
1,
Kilb
M=2
, Mnt
p=2,
Mnt
pM=1
); W
IC (A
thn=
9, B
awn=
2, B
mut
=1, C
ura=
3, M
uck=
2, S
lipM
=1,
Sliu
M=1
, Sto
e=6)
Le
pthy
phan
tes p
allid
us (O
.P.-
Cam
brid
ge)
7 K
LK (B
alb=
2); L
AO
(Car
d=1,
Gfin
=1, L
ead=
2, M
sop=
2, T
rum
=1);
LEI (
Rau
p=1)
; LIM
(B
oky=
1, C
lea=
1); S
LI (R
ath=
2); W
AT
(Too
r=2)
; WIC
(Ath
n=2,
Baw
n=3,
Sun
s=1)
Le
pthy
phan
tes t
eneb
rico
la (W
ider
) 16
C
AR
(Kila
=100
); C
LA (C
orr=
12, D
oog=
7, G
lyn=
10);
DU
B (L
ugg=
72);
GA
L (D
err=
1, In
cpM
=1,
Lurg
=1);
KER
(Cly
d=1,
Kno
c=1,
Tog
uS=1
); K
LD (D
eme=
13);
KLK
(Bal
b=1)
; LA
O (C
ard=
12,
Cla
r=1,
Fos
s=1,
Gfin
=5, G
las=
6, M
ary=
2, M
sop=
51, T
rum
=25)
; LEI
(Rau
p=2,
Rau
u=1)
; LIM
(B
oky=
33, K
duf=
2, M
oig=
1); S
LI (R
ath=
88);
TIP
(Bal
p=1,
Bal
u=2,
Com
m=5
, Coo
n=31
, D
oon=
2, F
lem
=2);
WA
T (M
oaf=
3, R
inc=
23, T
oor=
11);
WES
(Han
p=1)
; WEX
(Bar
n=4,
Si
nb=2
8), W
IC (A
thn=
20, B
awn=
2, B
mut
=15,
Cur
a=12
, Gat
e=20
, Muc
k=30
, Sto
e=21
, Sun
s=23
) Le
pthy
phan
tes t
enui
s (B
lack
wal
l )
1 D
ON
(Car
p=8,
Car
u=3,
Tie
p=3,
Tie
u=2)
Le
pthy
phan
tes z
imm
erm
anni
(B
ertk
au )
1 LI
M (B
oky=
193,
Cle
a=14
7, C
lop=
14, C
lou=
4, C
urr=
3, K
duf=
31, M
oig=
1)
Lept
porh
optr
um ro
bust
um
(Wes
tring
) 8
CO
R (G
len=
1, K
naw
=2);
KLK
(Bal
b=2,
Kill
=6, C
ast=
7, R
atr=
1); L
EI (R
aup=
3, R
auu=
3); L
IM
(Cle
a=7,
Clo
p=2,
Clo
u=6,
Cur
r=1,
Kdu
f=4,
Moi
g=2,
Rea
n=3)
; OFF
(Bgh
t=22
, Cor
b=4)
; SLI
(R
ath=
2); W
AT
(Moa
f=2,
Mor
e=1)
; WES
(Han
p=6)
Li
nyph
ia h
orte
nsis
(Sun
deva
ll)
1 K
LD (D
eme=
1)
Liny
phia
tria
ngul
aris
(Cle
rck)
1
KLD
(Dem
e=1)
Lo
phom
ma
punc
tatu
m (B
lack
wal
l) 5
DO
N (C
arp=
10, C
aru=
4, T
iep=
3, T
ieu=
4); K
LK (B
alb=
1, R
atr=
1); L
EI (R
aup=
4, R
auu=
3); L
IM
(Cle
a=2,
Clo
p=1,
Clo
u=1,
Kdu
f=1,
Moi
g=3
Rea
n=2)
; MA
Y (V
eeu=
3)
Mac
rarg
us ru
fus (
Wid
er)
2 LA
O (M
sop=
1); W
IC (B
mut
=2)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
123
M
aro
min
utus
(O.P
.-Cam
brid
ge)
11
CA
R (K
ila=1
3); C
LA (B
ool=
6, C
orr=
2, G
arv=
2); C
OR
(Car
r=18
, Gla
n=4,
Gul
l=5,
Mee
n=1,
R
ean=
4); D
UB
(Lug
g=63
); K
ER (C
umm
=12;
Gea
u=7;
Tog
p=2)
; KLD
(Dem
e=2)
; LA
O
(Car
d=34
, Fos
s=4,
Mso
p=9,
Tru
m=1
3); L
IM (B
oky=
1, C
lea=
23, C
lop=
1, C
lou=
1); M
AY
(V
eep=
1); T
IP (B
alb=
3, C
omm
=1, C
oon=
6, F
lem
=1);
WIC
(Ath
n=8,
Baw
n=5,
Bm
ut=1
, C
ura=
15, G
ate=
9, M
uck=
6, S
lip=1
, Sliu
=1, S
toe=
15, S
uns=
10)
Mas
o su
ndev
alli
(Wes
tring
) 2
WA
T (M
oaf=
1); W
IC (B
mut
=1, C
ura=
1, M
uck=
1, S
lip=1
, Slip
S=1,
Sliu
=4)
Mei
onet
a be
ata
(O.P
.-Cam
brid
ge)
1 D
ON
(Tie
u=15
) M
eion
eta
rure
stri
s (C
. L. K
och)
1
CO
R (C
loo=
1)
Mei
onet
a sa
xatil
is (B
lack
wal
l) 9
CLA
(Cor
r=1,
Doo
g=3,
Gla
n=1)
; DU
B (L
ugg=
6); G
AL
(Inc
u=1)
; KLK
(Cas
t=2,
Mun
y=2)
; LA
O
(Agh
o=3;
Don
g=1,
Fos
s=1,
Gla
s=2,
Lea
d=1)
; OFF
(Bgh
t=7,
Cor
b=1)
; TIP
(Bal
u=7,
Gar
y=1)
; W
AT
(Too
r=2)
; WES
(Han
p=3)
M
etop
obac
trus
pro
min
ulus
(O.P
.-C
ambr
idge
) 9
CA
R (K
ila=1
); C
LA (B
ale=
5, G
arv=
10, G
lyn=
1); C
OR
(Bal
y=1,
Cla
r=3,
Gla
n=1,
Gul
l=1)
; DO
N
(Car
p=3,
Car
u=13
, Tie
p=7,
Tie
u=1)
; KER
(Cum
m=1
8, G
eap=
3, G
eau=
1, T
ogp=
4, T
ogu=
1,
Toop
=8);
KLK
(Kill
=1, M
uny=
19);
LAO
(Agh
o=1,
Lea
d=1)
; TIP
(Bal
b=1,
Doo
n=47
, Fle
m=1
, K
ilb=1
, Mva
n=1)
; WEX
(Sin
b=1)
M
icra
rgus
her
bigr
adus
(Bla
ckw
all)
15
CA
R (K
ila=2
); C
OR
(Car
r=2,
Clo
o=2,
Fur
y=1,
Gla
n=3,
Gul
l=2,
Kilm
=2, M
een=
2, R
ean=
2,
Sagg
=2);
DO
N (C
arp=
1, C
aru=
5, T
iep=
6, T
ieu=
6); D
UB
(Lug
g=3)
; GA
L (I
ncp=
4, L
urg=
4);
KER
(Cly
d=14
, Cum
m=1
2, G
eap=
1, G
eau=
1, K
noc=
3, T
ogp=
1, T
ogpS
=1);
KLD
(Dem
e=1)
; LA
O (C
ard=
2, F
oss=
1, G
fin=1
, Gla
s=3,
Lea
d=1,
Tru
m=1
); LE
I (R
aup=
3, R
auu=
2); L
IM
(Bok
y=2,
Cle
a=5,
Clo
p=3,
Clo
u=1,
Kdu
f=12
); M
AY
(Vee
p=5,
Vee
u=1)
; SLI
(Rat
h=4)
; TIP
(C
oom
=3, C
ooa=
6, C
oon=
3, M
ntpS
=1, M
van=
2); W
AT
(Mor
e=2,
Too
r=2)
; WEX
(Sin
b=2)
M
icra
rgus
suba
equa
lis (W
estri
ng)
8 C
OR
(Clo
o=3,
Kilm
=23,
Kilm
S=1,
Kna
w=1
); G
AL
(Lur
g=4)
; KLK
(Cas
t=1,
Kill
=8, R
atr=
2);
LAO
(Agh
o=4,
Cla
r=1,
Don
g=3,
Fos
s=2,
Gla
s=1,
Lea
d=1)
; LIM
(Kdu
f=2,
Ree
n=1)
; WA
T (M
ore=
2, T
oor=
1); W
ES (H
anp=
1); W
IC (G
ate=
2)
Mic
rolin
yphi
a pu
silla
(Sun
deva
ll)
1 LI
M (C
lea=
1, K
duf=
1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
124
Mic
rone
ta v
iari
a (B
lack
wal
l) 1
KLD
(Dem
e=6)
M
iller
iana
iner
rans
(O.P
.-C
ambr
idge
) 1
KER
(Tog
pM=1
)
Min
yrio
lus p
usill
us (W
ider
) 2
WA
T (M
ore=
2); W
IC (B
awn=
1, G
ate=
3)
Mon
ocep
halu
s cas
tane
ipes
(Sim
on)
3 C
OR
(Car
r=1,
Mee
n=1)
; SLI
(Rat
h=1)
; WIC
(Ath
n=1,
Bm
ut=1
, Muc
k=1)
M
onoc
epha
lus f
usci
pes (
Bla
ckw
all)
5 LA
O (C
ard=
3,C
lar=
5, F
oss=
19, G
fin=2
5, G
las=
18, L
ead=
16, M
ary=
62, M
sop=
26, T
rum
=33)
; LE
I (R
aup=
42, R
auu=
21);
LIM
(Bok
y=29
, Cle
a=90
, Clo
p=9,
Clo
u=17
, Cur
r=70
, Kdu
f=64
, M
oig=
20, R
ean=
5); W
AT
(Moa
f=4,
Mor
e=13
, Rin
c=1)
; WES
(Han
p=6)
N
erei
ne c
lath
rata
(Sun
deva
ll)
1 LE
I (R
aup=
6, R
auu=
7)
Ner
iene
mon
tana
(Cle
rck)
2
LEI (
Rau
u=1)
; LIM
(Clo
u=1)
N
erie
ne p
elta
ta (W
ider
) 2
CO
R (C
arr=
4, G
lan=
3, G
len=
2); L
IM (B
oky=
2, C
lou=
2, C
lea=
2, K
duf=
1)
Oed
otho
rax
fusc
us (B
lack
wal
l) 2
LAO
(Agh
o=23
, Car
d=7,
Cla
r=51
, Don
g=25
, Fos
s=4,
Gfin
=7, M
sop=
1); W
IC (B
awn=
1, G
ate=
2,
Slip
=2, S
lipM
=1, S
liuM
=1, S
toe=
1, S
uns=
5)
Oed
otho
rax
gibb
osus
(Bla
ckw
all)
6
DU
B (L
ugg=
4); K
ER (C
lyd=
1, C
umm
=5, G
eap=
35, G
eau=
2, K
noc=
14, T
ogp=
12, T
ogu=
7,
Toop
=6, T
oopM
=1, T
oou=
11, T
oouM
= 2)
; KLK
(Cas
t=3,
Kill
=1, M
uny=
9, R
atr=
6); L
IM
(Cle
a=35
, Clo
p=1,
Clo
u=6,
Cur
r=6,
Kdu
f=2,
Moi
g=2,
Rea
n=16
); M
AY
(Vee
p=11
, Vee
u=29
); TI
P (C
omm
.=4,
Coo
n=2,
Doo
n=2,
Fle
m=3
, Kilb
=12)
O
edot
hora
x re
tusu
s (B
lack
wal
l) 4
DO
N (T
iep=
2); K
LD (D
eme=
1); L
IM (C
lop=
7, C
lou=
13, K
duf=
3, M
oig=
2, R
een=
4); W
AT
(Moa
f=2,
Mor
e=3,
Rin
c=1)
Pe
leco
psis
men
gei (
Sim
on)
2 LE
I (R
aup=
1, R
auu=
1); W
IC (S
liu=3
) Pe
leco
psis
nem
oral
is (B
lack
wal
l) 4
CLA
(Cor
r=1)
; CO
R (G
lan=
1, G
ull=
1, M
ean=
1, R
een=
1); L
IM (B
oky=
1, C
lea=
3, C
urr=
1,
Kdu
f=1)
; WIC
(Ath
n=1,
Bm
ut=3
, Cur
a=1,
Muc
k=1,
Sto
e=1)
Pe
leco
psis
par
alle
la (W
ider
) 4
DO
N (T
iep=
1, T
ieu=
4); K
ER (G
eap=
1); L
IM (B
oky=
1); W
IC (B
mut
=3, M
uck=
1, S
uns=
1)
Pepo
nocr
aniu
m lu
dicr
um (O
. P.-
Cam
brid
ge)
3 G
AL
(Inc
p=8,
Incp
S=1,
Incp
M=3
, Inc
u=1;
Incu
S=1)
; LIM
(Cle
a=1)
; WA
T (M
ore=
4)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
125
Poca
dicn
emis
junc
ea (
Lock
et &
M
illid
ge)
16
CLA
(Bal
e=12
, Boo
l=7,
Doo
g=3
, Gar
v=3,
Gly
n=1)
; CO
R (C
arr=
1, G
len=
44, G
ull=
1, K
ilm=1
0,
Kna
w=7
4, R
ean=
1); D
ON
(Car
p=5,
Tie
p=5,
Tie
u=1)
; DU
B (L
ugg=
1); F
ER (G
ore=
3, M
ull=
1,
Mul
lM=1
); G
AL
(Coo
l=6,
Incp
=15,
Incu
=1, L
urg=
11);
KER
(Gea
p=8,
Gea
u=1,
Tog
p=6,
To
gpM
=2, T
ogpS
=1, T
ogu=
2, T
oop=
3, T
oou=
1, T
oouM
=1);
KLK
(Bal
b=4,
Bal
bM=2
1;
Bal
bS=2
2, C
ast=
22, K
ill=5
3, M
uny=
296,
Rat
r=18
); LA
O (A
gho=
9, C
ard=
15, C
lar=
5, D
ong=
3,
Foss
=16,
Gla
s=32
, Lea
d=29
); LE
I (R
aup=
27, R
auu=
15);
LIM
(Clo
p=43
, Clo
u=2,
Cur
r=21
, M
oig=
6, R
een=
2); M
AY
(Vee
p=6)
; TIP
(Bal
p=15
, Bal
u=8,
Coo
a=5,
Doo
n=4,
Fle
m=4
, Kilb
=3,
Mnt
p=4,
Mva
n=15
); W
AT
(Mor
e=2,
Too
r=1)
; WES
(Han
p=34
); W
IC (A
thn=
4, B
awn=
37,
Bm
ut=2
, Gat
e= 4
, Muc
k=2,
Slip
=29,
Slip
M=6
, Sliu
=26)
Po
cadi
cnem
is p
umila
(Bla
ckw
all)
3 K
ER (C
lyd=
20, C
umm
=40,
Gea
p=83
, Gea
u=55
, Kno
c=50
, Tog
p=26
, Tog
pM=1
9, T
ogpS
=3,
Togu
=20,
Too
p=25
, Too
pM=2
, To
opS=
2, T
oou=
17, T
oouM
=3, T
oouS
=1);
LEI (
Rau
p=5,
R
auu=
1); L
IM (B
oky=
1, C
lea=
24, C
lop=
23, C
lou=
8, C
urr=
8, K
duf=
3, M
oig=
2, R
een=
2)
Poec
ilone
ta g
lobo
sa (B
lack
wal
l) 1
LIM
(Clo
p=1,
Clo
u=1)
Po
rrho
mm
a ca
mpb
elli
(O. P
.-C
ambr
idge
) 3
CO
R (C
arr=
1); G
AL
(Ker
r=1)
; WIC
(Muc
k=1)
Porr
hom
ma
conv
exum
(Wes
tring
) 1
CO
R (M
een=
1)
Porr
hom
ma
palli
dum
(Jac
kson
) 9
CA
R (K
ila=1
); C
OR
(Car
r=11
, Clo
o=2,
Fur
y=1,
Gla
n=3,
Gul
l=1,
Mee
n=6,
Rea
n=4,
Sag
g=4)
; D
UB
(Lug
g=8)
; GA
L (D
err=
5); L
AO
(Fos
s=3,
Gfin
=2, M
ary=
3, T
rum
=2);
LIM
(Bok
y=4,
C
lea=
14);
SLI (
Rat
h=1)
; TIP
(Com
m=4
, Coo
n=7)
; WEX
(Sin
b=3)
Po
rrho
mm
a py
gmae
um (B
lack
wal
l) 1
WIC
(Bm
ut=3
, Sto
e=1)
Sa
aris
toa
abno
rmis
(Bla
ckw
all)
10
DO
N (C
arp=
4, C
aru=
1, T
iep=
2); D
UB
(Lug
g=13
); K
LD (D
eme=
3); K
LK (B
alb=
2, R
atr=
1);
LAO
(Car
d=33
, Fos
s=8,
Gfin
=40,
Lea
d=3,
Mar
y=64
, Mso
p=52
, Tru
m=8
); LE
I (R
aup=
1); L
IM
(Bok
y=26
, Cle
a=21
, Clo
p=5,
Clo
u=1,
Kdu
f=40
); TI
P (B
alp=
3, C
omm
=71,
Coo
n=81
, Fle
m=1
, M
van=
1, M
ntp=
1); W
AT
(Mor
e=8,
Too
r=9)
; WEX
(Bar
n=1,
Sin
b=66
)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
126
Saar
isto
a fir
ma
(O.P
.-Cam
brid
ge)
10
CLA
(Cor
r=1,
Gly
n=1)
; DU
B (L
ugg=
4); K
ER (C
lyd=
7, C
umm
=3, K
noc=
5); K
LD (D
eme=
1);
KLK
(Bal
b=1)
; LA
O (M
ary=
2, T
rum
=1);
LIM
(Cle
a=4,
Kdu
f=2)
; TIP
(Bal
p=1;
Coo
n=1,
Fl
em=1
); W
AT
(Too
r=3)
; WIC
(Muc
k=1,
Sliu
=1, S
toe=
1)
Salo
ca d
icer
os (O
.P.-C
ambr
idge
) 1
CLA
(Gly
n=1)
Si
lom
etop
us e
lega
ns (O
.P.-
Cam
brid
ge)
7 G
AL
(Inc
p=1,
Incu
=6);
KER
(Gea
p=90
, Gea
u=16
, Tog
p=8,
Tog
pM=2
, Tog
pS=1
, Tog
u=23
, To
guS=
1, T
oop=
1, T
oop=
9); K
LK (B
alb=
1); L
AO
(Agh
o=3,
Car
d=1,
Cla
r=5,
Don
g=1,
Fos
s=4,
G
las=
6, L
ead=
3); L
IM (C
lea=
9); M
AY
(Vee
p=21
, Vee
u=33
); W
AT
(Mor
e=88
, Too
r=3)
Ta
llusi
a ex
pert
a (O
.P.-C
ambr
idge
) 2
LIM
(Clo
u=6)
; MA
Y (V
eeu=
1)
Tapi
nocy
ba in
sect
a (L
. Koc
h)
2 K
LD (D
eme=
1); L
EI (R
auu=
1)
Tapi
nocy
ba p
alle
ns (O
.P.-
Cam
brid
ge)
6 D
ON
(Tie
p=2,
Tie
u=1)
; DU
B (L
ugg=
7); K
ER (T
ogu=
1); L
AO
(Car
d=2,
Fos
s=10
, Mso
p=1,
Tr
um=5
); TI
P (C
oon=
1); W
IC (A
thn=
9, B
awn=
3, B
mut
=2, G
ate=
22, S
toe=
3)
Tapi
nocy
ba p
raec
ox (O
. P.-
Cam
brid
ge)
1 LI
M (C
lea=
1)
Tara
nunc
us se
tosu
s (Si
mon
) 10
D
ON
(Car
p=1,
Tie
p=1)
; DU
B (L
ugg=
1); F
ER (G
ore=
1); G
AL
(Inc
p=2,
Lur
g=6)
; KLK
(Mun
y=1,
R
atr=
1); L
AO
(Fos
s=1,
Gla
s=3,
Lea
d=15
); LI
M (C
lou=
2); M
AY
(Vee
p=10
); TI
P (B
alp=
1,
Bal
u=1,
Kilb
=1);
WIC
(Slip
=2)
Tiso
veg
ans (
Bla
ckw
all)
5 LA
O (A
gho=
1, D
ong=
1, F
oss=
2); L
EI (R
auu=
1); L
IM (C
lop=
1); W
IC (A
thn=
1, G
ate=
1,
Slip
M=1
, Sliu
=12,
Sto
e=47
) Tr
icho
pter
na th
orel
li (W
estri
ng)
2 G
AL
(Inc
u=2)
; MA
Y (V
eeu=
96)
Trox
ochr
us sc
abri
culu
s (W
estri
ng)
3 D
ON
(Tie
u=1)
; KLK
(Cas
t=12
, Kill
=2, R
atr=
4); L
AO
(Don
g=8)
W
alck
enae
ria
acum
inat
a (B
lack
wal
l) 4
LIM
(Bok
y=7,
Cle
a=8,
Clo
p=4,
Clo
u=1,
Cur
r=10
, Kdu
f=13
, Moi
g=1)
; SLI
(Rat
h=5)
; WA
T (M
oaf=
3, M
ore=
16, T
oor=
2); W
ES (H
anp=
11)
Wal
cken
aeri
a at
robt
ibia
lis (O
. P.-
Cam
brid
ge)
9 C
LA (C
orr=
1, D
oog=
2, G
lyn=
5); D
ON
(Car
u=6)
; GA
L (D
err=
1, In
cp=3
6, In
cu=1
); K
ER
(Gea
p=28
, Gea
u=11
, Kno
c=13
, Tog
u=1,
Too
p=3,
Too
u=16
); K
LK (R
atr=
2); L
AO
(Car
d=1,
D
ong=
2); M
AY
(Vee
p=2,
Vee
u=2)
; WA
T (M
ore=
31, T
oor=
7); W
IC (S
liu=4
) W
alck
enae
ria
clav
icor
nis (
Emer
ton)
1
DO
N (T
ieu=
1)
Bull.
Ir. b
ioge
og. S
oc. N
o. 3
2 (2
008)
127
Wal
cken
aeri
a cu
spid
ata
(Bla
ckw
all)
4 C
OR
(Car
r=1,
Gla
n=8,
Rea
n=5)
; DO
N (C
arp=
4, C
aru=
10, T
iep=
5); L
EI (R
aup=
1, R
auu=
6);
WA
T (M
ore=
4, T
oor=
7)
Wal
cken
aeri
a dy
sder
oide
s (W
ider
) 9
CA
R (K
ila=8
); C
LA (G
lyn=
1); C
OR
(Car
r=1,
Clo
o=28
, Gla
n=1,
Gul
l=3)
; DU
B (L
ugg=
5); K
ER
(Kno
c=1)
; TIP
(Bal
u=1)
; WA
T (M
ore=
1); W
EX (S
inb=
); W
IC (A
thn=
32, B
mut
=40,
Gat
e=5,
Sl
iu=1
) W
alck
enae
ria
koch
i (O
. P.-
Cam
brid
ge)
2 C
LA (G
arv=
1); D
ON
(Car
p=1)
Wal
cken
aeri
a no
dosa
(O.P
.-C
ambr
idge
) 4
CA
R (K
ila=1
); K
ER (C
lyd=
1); L
IM (C
lea=
1); M
AY
(Vee
u=1)
Wal
cken
aeri
a nu
dipa
lpis
(Wes
tring
) 8
CLA
(Boo
l=1,
Cor
r=9,
Doo
g=3)
; GA
L (D
err=
4, In
cp=2
, Inc
u=1)
; LA
O (C
ard=
11, F
oss=
3,
Gfin
=8, M
ary=
1, M
sop=
8, T
rum
=5);
LIM
(Bok
y=5,
Cle
a=2,
Moi
g=5,
Rea
n=1)
; SLI
(Rat
h=3)
; TI
P (C
omm
=1, C
oon=
3); W
AT
(Mor
e=3)
; WES
(Han
p=1)
W
alck
enae
ria
unic
orni
s (O
.P.-
Cam
brid
ge)
6 FE
R (G
oreM
=1);
LEI (
Rau
p=1)
; LIM
(Clo
p=2,
Clo
u=2,
Moi
g=1)
; MA
Y (V
eeu=
2); O
FF
(Bgh
t=1)
; WIC
(Slip
M=1
) W
alck
enae
ria
vigi
lax
(Bla
ckw
all,
1851
) 12
C
AR
(Kila
=23)
; CO
R (C
arr=
5, C
loo=
5, G
lan=
4, G
len=
2, G
ull=
11, M
een=
2, R
ean=
3); K
ER
(Cly
d=14
, Cum
m=1
8, G
eap=
3, G
eau=
14, K
noc=
7, T
ogp=
6, T
ogu=
22, T
oguM
=1, T
oou=
7); K
LK
(Mun
y=4)
; LA
O (C
ard=
32, C
lar=
11, F
oss=
5, G
las=
2, M
sop=
2); L
IM (C
lea=
7, C
lop=
31,
Clo
u=12
, Cur
r=7,
Kdu
f=4,
Moi
g=2,
Rea
n=3)
; MA
Y (V
eeu=
16);
OFF
(Cor
b=1)
; TIP
(Bal
p=1,
B
alu=
3, C
omm
=5, F
lem
=1, M
ntp=
2, M
van=
26);
WA
T (M
ore=
21, T
oor=
5); W
ES (H
anp=
3);
WIC
(Ath
n=13
, Baw
n=27
, Cur
a=3,
Bm
ut=1
2, G
ate=
2, M
uck=
12, S
lip=2
, Sliu
=39,
Sto
e=35
)