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\ 1- v d It'lOan1'l11fll C HllJlJ'rI 7/2545 Technical Paper no. 7/ 2002 'W qGi f11'l3.J ad n1ad bGh EGG LAYING BEHAVIOR AND HATeHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray) Suvena Banyen Wonnat Hirunchulha 'II 91110 'II 'II 0- 7478 - 1299 Satun Inland Fisheries Station La- ngu Sarnn Province 9111 0 Tel. 0- 7478-129 9

I r1.LQ:::WLWl - Fisheries

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Page 1: I r1.LQ:::WLWl - Fisheries

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W qGi f11l3J f11l11~1ad bb~~bWl~r1n1ad bGh n1~f)1t1

EGG LAYING BEHAVIOR AND HATeHING METHODS OF

SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen

Wonnat Hirunchulha

~mi1u)~~ll1~

1~~ijnrr~~lIGi II

tl Gi~j ~~lIGi 91110 II II

1~) 0 - 7478 - 1299

Satun Inland Fisheries Station

La- ngu Sarnn Province 9111 0

Tel 0 - 7478-129 9

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LAVING IlEHA VIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagltr baska (Gray)

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

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fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

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II II I I

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24

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~

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

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q U q q

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Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

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3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

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Page 2: I r1.LQ:::WLWl - Fisheries

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mmn11l11fllHlUUrl 712545 Technical Paper no 72002

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

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23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

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fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

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~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

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IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

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1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

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bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

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~

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tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

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QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

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QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

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I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

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11

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q

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nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

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Cb

It)mYl Hll ~ 6 ~

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li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

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lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

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Page 3: I r1.LQ:::WLWl - Fisheries

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

~1nJ~ II

ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

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1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

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q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

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Page 4: I r1.LQ:::WLWl - Fisheries

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

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1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

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l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

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lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

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QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

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~ OQl Q q OQl

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~~P11il~HWltlHWltlYi 5 bull I )I

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QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

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~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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nj jlJf11j V-lYlJ~W 1J 111bull

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II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

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U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

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~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

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6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

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24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

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6 21600 510 54 5

7 21600 411 0 510

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10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

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20 23800 530 55 0

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22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

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14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

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10 5008 64 64 5371 72 68 4672 62 60

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16 49 20 64 6 1 4523 65 64

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Page 5: I r1.LQ:::WLWl - Fisheries

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

YrWi~U~nHrlW~ 1 A(jmA~vnW lV1rJ~mJ~WinWhnJ1Vh~~UtJ1~nilj~lJltj 50 1liWamplllJ~l 1~V~1~viW

tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

tJlI1imWf)~fflJnU lUllelnWl~vlhlJ1tl 1 1~f)W IIllJUWI11~lWrJff1llllll1rl-Jl1-JU l~mllJl~lll~el

~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

1lJJ mJt~1il~ )ll1~ooflJJllllW l[~ 1 l~Cl~1Jj~JJl(1I 3-5 yjOJ Uru~J1Jl~l~Ul1~~~~~VYljltlnuj~11l~

1 ~ ll~ Cl ~yjo~vt 0 11Jl 111 ~ flj ~ YllJflW 1~ tI 1 ~ 11 J~ tIJ nW 0 V 1 W1 ClJJ U UI~ ~ mJt~ 111 ~ ov 1J 0 0 fl JJlmh~1111J

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

J1lJ il1flflljffmnmJt~lm~iJll-lliilUjmll1~ lrl0111~ 9 -Bl-1l1fllJ 2542 ~jln1J~I-J 2 ~ll$lOW 1 l~tI1~JJ JI SI

IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

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Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

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o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

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1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

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q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

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m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

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cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

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15250623

50

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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v v d r 12 tlf)Jrn~UlJYlW fl IYllm ~ ~1W

~flfJtll~Ull-WJ TIl~lm~mWVlf1llJVlJ iiBe)Wflh llJflll rHllJ-WelVWh 1l111llJ1J ffU11 ~hjJ~ 6 o I 10 QI d

~

SI

~

dQI QI QI

llflfl1jYlgtHItJJ~llll11Vlll~ I 1J JU 29 1l1 Wll11IfllHW 23040plusmn396 m11 fl11lJfl11Jm~1l0JmM

Q d Q QI d d

5J97plusmn279 llSlJIlI11Ilj IlCl~ml11V11m~lltJJlHW 5403plusmn247 1lSWIlI11Ilj llJlllilJr-lWlflYl 2 1l~~11fl1111 QI cv rI I jI QI I QI rI 2 11lJVlW1ij~I111Jfll111flJlJm~lltJJl~~mmlflUtJJl11Vl11~ W = 7333CW-15081 R = 04362 l~~fll111

~

ff11-WJTIj~WJ1Jfl~liJV1Jm~lltJJll~~111l1irflUOJ1l1l-WWi W = 62948CL-I0986 R2 = 02549 f1J~jVi 7

jtJ~ 6 ~fllltll~lll1-wWjI~lm~mll~

~

~1vlJn(nf~)

30000 x W = 7333CWmiddot 15081

25000 R =04362

20000

x 15000

10000 I R = 0254)

5000

o tl111JVllmllJfl11~(9lJ )

40 42 44 46 48 50 52 54 56 58 60

~ ~ I do QI QI rI I jI 0 QI 0 QI

j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

YrWi~U~nHrlW~ 1 A(jmA~vnW lV1rJ~mJ~WinWhnJ1Vh~~UtJ1~nilj~lJltj 50 1liWamplllJ~l 1~V~1~viW

tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

tJlI1imWf)~fflJnU lUllelnWl~vlhlJ1tl 1 1~f)W IIllJUWI11~lWrJff1llllll1rl-Jl1-JU l~mllJl~lll~el

~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

1lJJ mJt~1il~ )ll1~ooflJJllllW l[~ 1 l~Cl~1Jj~JJl(1I 3-5 yjOJ Uru~J1Jl~l~Ul1~~~~~VYljltlnuj~11l~

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1 jJ 1 1 jJ d 1 In 1 jJ ~ 1 d 1 jJl1inOOflIJ1~lt1 lllJl~lllJ lU~OllJOJ I1l1jiI ~v IJ1l-1 iI1mWU1~ClOlJ 1Cl~il1flUIJP1111~ lU1jiI lIU1

~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

1ClJ~mn1VflCl1J1lIJ11~ U1CllrHJV 1 lI11J10flflj~UYlfl1lJJ11~ U lWO 1lUU ~tJ1lJ1~1 liU11Wlfll

VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

J1lJ il1flflljffmnmJt~lm~iJll-lliilUjmll1~ lrl0111~ 9 -Bl-1l1fllJ 2542 ~jln1J~I-J 2 ~ll$lOW 1 l~tI1~JJ JI SI

IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

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U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

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q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

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m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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j ~Wh~fl~lJ 173753 2 86876 071 NS

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

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Page 7: I r1.LQ:::WLWl - Fisheries

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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v v d r 12 tlf)Jrn~UlJYlW fl IYllm ~ ~1W

~flfJtll~Ull-WJ TIl~lm~mWVlf1llJVlJ iiBe)Wflh llJflll rHllJ-WelVWh 1l111llJ1J ffU11 ~hjJ~ 6 o I 10 QI d

~

SI

~

dQI QI QI

llflfl1jYlgtHItJJ~llll11Vlll~ I 1J JU 29 1l1 Wll11IfllHW 23040plusmn396 m11 fl11lJfl11Jm~1l0JmM

Q d Q QI d d

5J97plusmn279 llSlJIlI11Ilj IlCl~ml11V11m~lltJJlHW 5403plusmn247 1lSWIlI11Ilj llJlllilJr-lWlflYl 2 1l~~11fl1111 QI cv rI I jI QI I QI rI 2 11lJVlW1ij~I111Jfll111flJlJm~lltJJl~~mmlflUtJJl11Vl11~ W = 7333CW-15081 R = 04362 l~~fll111

~

ff11-WJTIj~WJ1Jfl~liJV1Jm~lltJJll~~111l1irflUOJ1l1l-WWi W = 62948CL-I0986 R2 = 02549 f1J~jVi 7

jtJ~ 6 ~fllltll~lll1-wWjI~lm~mll~

~

~1vlJn(nf~)

30000 x W = 7333CWmiddot 15081

25000 R =04362

20000

x 15000

10000 I R = 0254)

5000

o tl111JVllmllJfl11~(9lJ )

40 42 44 46 48 50 52 54 56 58 60

~ ~ I do QI QI rI I jI 0 QI 0 QI

j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

YrWi~U~nHrlW~ 1 A(jmA~vnW lV1rJ~mJ~WinWhnJ1Vh~~UtJ1~nilj~lJltj 50 1liWamplllJ~l 1~V~1~viW

tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

tJlI1imWf)~fflJnU lUllelnWl~vlhlJ1tl 1 1~f)W IIllJUWI11~lWrJff1llllll1rl-Jl1-JU l~mllJl~lll~el

~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

1lJJ mJt~1il~ )ll1~ooflJJllllW l[~ 1 l~Cl~1Jj~JJl(1I 3-5 yjOJ Uru~J1Jl~l~Ul1~~~~~VYljltlnuj~11l~

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1 jJ 1 1 jJ d 1 In 1 jJ ~ 1 d 1 jJl1inOOflIJ1~lt1 lllJl~lllJ lU~OllJOJ I1l1jiI ~v IJ1l-1 iI1mWU1~ClOlJ 1Cl~il1flUIJP1111~ lU1jiI lIU1

~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

1ClJ~mn1VflCl1J1lIJ11~ U1CllrHJV 1 lI11J10flflj~UYlfl1lJJ11~ U lWO 1lUU ~tJ1lJ1~1 liU11Wlfll

VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

J1lJ il1flflljffmnmJt~lm~iJll-lliilUjmll1~ lrl0111~ 9 -Bl-1l1fllJ 2542 ~jln1J~I-J 2 ~ll$lOW 1 l~tI1~JJ JI SI

IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

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U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

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q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

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m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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j ~Wh~fl~lJ 173753 2 86876 071 NS

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

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Page 8: I r1.LQ:::WLWl - Fisheries

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2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

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tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

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~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

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Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

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1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

f1l j Ifl rl tliW 1tlVl BlIlJllUli IlJ fIJj 11 I~ nJfll1lJ m ~Yl 1-1 m 1Yl tlU IU tl~ il1tl BlilLY ~ fl rll tlf1l j

q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

Q[t11 bL~I~1 U ~lLU ~~UQpoundI~1t~ ~Q~tQ~ClU11 Uqlr1l0L JIALArtflWl n~lmpoundCllfLULlwg ~LMIL~ wL n~1

m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

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Page 9: I r1.LQ:::WLWl - Fisheries

2

EGG LAYING BEHAVIOR AND HATCHING METHODS OF SOUTHERN SALTWATER TERRAPIN Batagur baska (Gray)

Suvena Banyen Wonnat Hirunchulha

Satun Fisheries Station Kampang La-ngu Satun Province 91110 Thailand

ABSTRACT

Eggs laying behavior and hatching methods of Southern Saltwater Terrapin Bwagllr baska (Gray) was conducted between September 1999 to September 2000 at Satun Inland Fisheries Station Relationship of 29 male and female broodstocks between weight (W)

rapace wid th (CW) carapace length (CL) were linear regression In the same weight female roodslOcks had always carapace width and carapace length bigger than male broodstock The _gs obtained from 14 female broodstocks varied between 9-27 and total eggs were 305

Eggs vere in cubated by 3 methods of earth pond concrete pond and styrofoam The obtained hatching rate was 4082 5816 5510 respectively Incubation period of hatching i5 100-110 74-84 90- 100 days average temperature 288degC 305degC 297degC In cubation - eriod were not showed a significant different The growth rate specific growth rate relative

w rate vere not showed a signi ficant different The survival rate of all young turtle were

ords Batoglll baska (Gray) Egg Laying Behavior Hatching

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~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

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16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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1 1 d ltjJd~d Is) d~dt I IIJI

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

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1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

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lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

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_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

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QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

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QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

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tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

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6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

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24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

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5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

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10 4815 54 64 4571 52 60 4098 46 55

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14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

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8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

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12 5126 58 60 5109 61 58 4502 60 58

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14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

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Page 10: I r1.LQ:::WLWl - Fisheries

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m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

~1nJ~ II

ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

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23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

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f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

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ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

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Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

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q

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Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

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2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

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(flflJ) (lflJ ) (lflJ )

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2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

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Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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3 4563 52 56 5213 57 67 4029 49 53

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5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

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2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

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6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

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249 038 031 202 049 041 250 0 30 027

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 11: I r1.LQ:::WLWl - Fisheries

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~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

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tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

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23

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

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m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

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IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

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bull bull bull

22

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4 ~ t QI J I

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

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U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

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bull bull

24

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~

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

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ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

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li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

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7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

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23 15000 440 490

24 4S5 490

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27 15800 470 520

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6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

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19 23600 550 540

20 23800 530 55 0

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22 24500 505 505

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24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

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17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

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Page 12: I r1.LQ:::WLWl - Fisheries

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

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1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

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lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

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QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

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q q

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QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

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~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

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tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

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12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

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2 18400 470 520

3 IX400 470 520

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5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

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3 4563 52 56 5213 57 67 4029 49 53

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5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

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6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

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Page 13: I r1.LQ:::WLWl - Fisheries

6

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llflfl1jYlgtHItJJ~llll11Vlll~ I 1J JU 29 1l1 Wll11IfllHW 23040plusmn396 m11 fl11lJfl11Jm~1l0JmM

Q d Q QI d d

5J97plusmn279 llSlJIlI11Ilj IlCl~ml11V11m~lltJJlHW 5403plusmn247 1lSWIlI11Ilj llJlllilJr-lWlflYl 2 1l~~11fl1111 QI cv rI I jI QI I QI rI 2 11lJVlW1ij~I111Jfll111flJlJm~lltJJl~~mmlflUtJJl11Vl11~ W = 7333CW-15081 R = 04362 l~~fll111

~

ff11-WJTIj~WJ1Jfl~liJV1Jm~lltJJll~~111l1irflUOJ1l1l-WWi W = 62948CL-I0986 R2 = 02549 f1J~jVi 7

jtJ~ 6 ~fllltll~lll1-wWjI~lm~mll~

~

~1vlJn(nf~)

30000 x W = 7333CWmiddot 15081

25000 R =04362

20000

x 15000

10000 I R = 0254)

5000

o tl111JVllmllJfl11~(9lJ )

40 42 44 46 48 50 52 54 56 58 60

~ ~ I do QI QI rI I jI 0 QI 0 QI

j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

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tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

tJlI1imWf)~fflJnU lUllelnWl~vlhlJ1tl 1 1~f)W IIllJUWI11~lWrJff1llllll1rl-Jl1-JU l~mllJl~lll~el

~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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J1lJ il1flflljffmnmJt~lm~iJll-lliilUjmll1~ lrl0111~ 9 -Bl-1l1fllJ 2542 ~jln1J~I-J 2 ~ll$lOW 1 l~tI1~JJ JI SI

IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

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I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

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~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

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u q U q

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30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

f1l j Ifl rl tliW 1tlVl BlIlJllUli IlJ fIJj 11 I~ nJfll1lJ m ~Yl 1-1 m 1Yl tlU IU tl~ il1tl BlilLY ~ fl rll tlf1l j

q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

Q[t11 bL~I~1 U ~lLU ~~UQpoundI~1t~ ~Q~tQ~ClU11 Uqlr1l0L JIALArtflWl n~lmpoundCllfLULlwg ~LMIL~ wL n~1

m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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v 0 CV ltV V

VlrnU-Itl1l1fl4 tJllPlfl (fli]J) mllJfl1HmtpHl~ (lflJ) flJlllVlJflitltjn)~ (lfll) lJEl~t1mYllmmtJ

fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

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Page 14: I r1.LQ:::WLWl - Fisheries

7

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40 42 44 46 48 50 52 54 56 58 60

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2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

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tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

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~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

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nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

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11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

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It)mYl Hll ~ 6 ~

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li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

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UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

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1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

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rI 4Q1 lel Q rI I IJrI QI

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91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

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liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

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207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

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q dav d v 0 dCloCV rI ds Q ~ a

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Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

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(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

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ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

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illtJlt1fl~lJ l1~Y1lJlI

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j~rtl1~fl~lJ 8004 2 4002 263 NS

illtJltlfl~lJ l1~rtlJII

76102

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Page 15: I r1.LQ:::WLWl - Fisheries

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19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

~1nJ~ II

ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

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q

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m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

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nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

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12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

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24 4S5 490

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7 21600 411 0 510

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10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

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20 23800 530 55 0

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Page 16: I r1.LQ:::WLWl - Fisheries

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40 42 44 46 48 50 52 54 56 58 60

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j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

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tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

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~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

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~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

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VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

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Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

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~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

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1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

f1l j Ifl rl tliW 1tlVl BlIlJllUli IlJ fIJj 11 I~ nJfll1lJ m ~Yl 1-1 m 1Yl tlU IU tl~ il1tl BlilLY ~ fl rll tlf1l j

q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

Q[t11 bL~I~1 U ~lLU ~~UQpoundI~1t~ ~Q~tQ~ClU11 Uqlr1l0L JIALArtflWl n~lmpoundCllfLULlwg ~LMIL~ wL n~1

m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

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Page 17: I r1.LQ:::WLWl - Fisheries

10

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~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

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ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

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n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

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u q U q

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d ~ ~ ~ ~~ ~ d d

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(lI Q ~ t)I

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IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

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bull bull bull

22

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4 ~ t QI J I

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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ltu

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nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

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24

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

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Page 18: I r1.LQ:::WLWl - Fisheries

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CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

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IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

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lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

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C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

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QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

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~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

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ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

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QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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23

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fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

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~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

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12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

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20 430 480

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10 22000 520 550

11 22000 490 520

12 22000 540 570

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Page 19: I r1.LQ:::WLWl - Fisheries

12

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5J97plusmn279 llSlJIlI11Ilj IlCl~ml11V11m~lltJJlHW 5403plusmn247 1lSWIlI11Ilj llJlllilJr-lWlflYl 2 1l~~11fl1111 QI cv rI I jI QI I QI rI 2 11lJVlW1ij~I111Jfll111flJlJm~lltJJl~~mmlflUtJJl11Vl11~ W = 7333CW-15081 R = 04362 l~~fll111

~

ff11-WJTIj~WJ1Jfl~liJV1Jm~lltJJll~~111l1irflUOJ1l1l-WWi W = 62948CL-I0986 R2 = 02549 f1J~jVi 7

jtJ~ 6 ~fllltll~lll1-wWjI~lm~mll~

~

~1vlJn(nf~)

30000 x W = 7333CWmiddot 15081

25000 R =04362

20000

x 15000

10000 I R = 0254)

5000

o tl111JVllmllJfl11~(9lJ )

40 42 44 46 48 50 52 54 56 58 60

~ ~ I do QI QI rI I jI 0 QI 0 QI

j1JI1 7 m 111 [(lJl~lI1i j ~11 1 1J m 11Jfl11J m ~II tJJ 1lCl ~W 11111 fl fl 1111V 11m ~1l OJLlCl~11111~1 fl II

I QI tI I

U t) lU11YfU ~mm~mW

l1n A11lJ btlJ~tJ1jl ~lrhrYi rJYrtJfiUC1~ mJ~W TII~hm ~tJ1W WUl11 WUW1~tJ111Jf1l YllnWll1 q

QJod )) 6d J ))IIQlo QI

W W 1i q il~ lJA 11lJ n 11-J m ~~ rJ-J Uel ~m1lJV1 1m~~ rJ-J]J 1n f111W rJWW 1i

bull I fflJrJ llel~tJ-JlJW Ill W nlJl nUW A 11lJ f11 N

UrJ-Jm~~rJ-Jil~i-Jll~n~1-Jll~m1lJV11m~~rJ-JnllWl 1wlJ~I~lmll~tJ-JnW ~-Jlil~ 8 Uel~lil~ 9

t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

t111-1oJn(nfl-J)

30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

2 Yl q~ f)11llf) n ~HnJyrUfn ~1JU()~vh f)1 () HI

11 n~el f11l ffnY1l~ 11~lJrr~V~Wlrl11 J1-JI~rJmJmAlJf1-JI~ rJWWqf1~f11VW 2542 1~tJvirJlll1 I I II

YrWi~U~nHrlW~ 1 A(jmA~vnW lV1rJ~mJ~WinWhnJ1Vh~~UtJ1~nilj~lJltj 50 1liWamplllJ~l 1~V~1~viW

tJ11 ri~lliJ VII ~11l1A f) n ~ ril W 11lU f) -J~lliJ v111mn W il111~ rJ~fflJYrmJ 11 tl-J l1n~fflJ~mJn Wll~1lll1l~ 11nU q q

- ~ ~ QI III I )) ~6 ~ ~ I II) 1lt) I0 I I

tJlI1imWf)~fflJnU lUllelnWl~vlhlJ1tl 1 1~f)W IIllJUWI11~lWrJff1llllll1rl-Jl1-JU l~mllJl~lll~el

~ll~mY1l 1 ill11~ 2-3 Ai-J lll1l~lm tJ1l1ff~-JWqamplm llJffnlllll1~-Jl1-J1j~-Jll~ln~~l IW f1-JI~v-JflW

amp Q lrl rJ tJ1 f11f1~ lJ IV tJ el-J lll1l~ 1U~I1 ellJ 11-J 1j UW1~ i1C1lJ nl1-J Ill~v 90 1liW amplllJ~ l elf1lQ eltJ 67 1liW~llJ~l qq

16

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1 ~ r)(1 tlfl l1mV 1 iI tY tI ~Il~ W ~OJJVfl1J ~ llUI dTU nl-HlcrlU ~ llCl ~1 VOI-J1J~nrudlU nuClllvtmJ ~ Otl 1 ~Cl~ 1J d

1lJJ mJt~1il~ )ll1~ooflJJllllW l[~ 1 l~Cl~1Jj~JJl(1I 3-5 yjOJ Uru~J1Jl~l~Ul1~~~~~VYljltlnuj~11l~

1 ~ ll~ Cl ~yjo~vt 0 11Jl 111 ~ flj ~ YllJflW 1~ tI 1 ~ 11 J~ tIJ nW 0 V 1 W1 ClJJ U UI~ ~ mJt~ 111 ~ ov 1J 0 0 fl JJlmh~1111J

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1 jJ 1 1 jJ d 1 In 1 jJ ~ 1 d 1 jJl1inOOflIJ1~lt1 lllJl~lllJ lU~OllJOJ I1l1jiI ~v IJ1l-1 iI1mWU1~ClOlJ 1Cl~il1flUIJP1111~ lU1jiI lIU1

~ jJ l II l jJ jJ ~l I lll I 1 I 1 II II 0

1ClJ~mn1VflCl1J1lIJ11~ U1CllrHJV 1 lI11J10flflj~UYlfl1lJJ11~ U lWO 1lUU ~tJ1lJ1~1 liU11Wlfll

VW~Wilt1fl1~~WJJ~lJ1fl 3middot5 19fl-I~IIJm flj~11YlfldJltYtI~Ill1r)(Vu1flflJj~lJ1UI 10-20 1IJ~j mr~il1fltl~

1 1 d ltjJd~d Is) d~dt I IIJI

1~ln1~ UllCl~m~UYlfl11middotHlllClli1JfltJUtIYljltl llhmJ 1 OflflHlW0Jfltl~jO~jOtl1l1Jl1~U UCl~1Cl1J

CJ1oW1jo~ntlrr~l 1~JillmrW~~o~iJJ 1 JJ~lJltll 30 mVi l-WOJi1J1lJ~nUO~~l~flltJ~l~1~OV 1 f1CllWClJ

J1lJ il1flflljffmnmJt~lm~iJll-lliilUjmll1~ lrl0111~ 9 -Bl-1l1fllJ 2542 ~jln1J~I-J 2 ~ll$lOW 1 l~tI1~JJ JI SI

IJ~1lJJI11U~nCll 1900 lUlrl~l~mlJl~tiJtJnCll 2130 1-1 1Jl1l1flI11rlOI-J11J1~ 23800 fliJJ UCl~1~111J Ll~ 22000 flilJ l1n~~11r)1 19 yloJ 11m1rfl1~ 72 fliJJ f1111Jfll1lUCl~f111JJ(Jl11~ 42 llCl~ 6 7

19fl-l~I1Jm ~lJJiilPi1J

il1flflljYl~ClO~1l1l1~lm~01I-J 29111 mlJljm1~1~lwV~ 14 Ill 111l1~lUt111l1flLQ~VrlOI-J11l

1~ 2240714plusmn1973 85 flilJ UCl~1~11N1Jut1l1tifl1Q~tI 2084286plusmn186824 flilJ t1l1l1m~fl~ll

J~111Jrlmrm1~lm~Ir1Jl1J1~ 7902J4G98 flilJ hltJ1~1~11hjl11-J~HfI~V 7593i684 flilJ f111JJfllll

m~v 431plusmn010 19fU~1l1~j m~m1JJVnm~V 6 85plusmn037 19fU~llJ~j J11l1fldl1J~mvlJUo~1l1l1~lrlOI-J 111 1 ~IICl~1 ~111J 1 ~l~ 11 dl11111 1U ClJ 1 ~ llllltrl ~lrlf11rflUtl~Utlll1 Cll~I1U 00fl1J1 ll1l1~ 111~Cl~11111 ~ 1 ~

JI iI I

1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

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23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

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~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

U~ltY~tllll-l~

_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

f1l j Ifl rl tliW 1tlVl BlIlJllUli IlJ fIJj 11 I~ nJfll1lJ m ~Yl 1-1 m 1Yl tlU IU tl~ il1tl BlilLY ~ fl rll tlf1l j

q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

Q[t11 bL~I~1 U ~lLU ~~UQpoundI~1t~ ~Q~tQ~ClU11 Uqlr1l0L JIALArtflWl n~lmpoundCllfLULlwg ~LMIL~ wL n~1

m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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v 0 CV ltV V

VlrnU-Itl1l1fl4 tJllPlfl (fli]J) mllJfl1HmtpHl~ (lflJ) flJlllVlJflitltjn)~ (lfll) lJEl~t1mYllmmtJ

fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

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10 5008 64 64 5371 72 68 4672 62 60

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12 5126 58 60 5109 61 58 4502 60 58

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16 49 20 64 6 1 4523 65 64

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19 5332 70 68 45 80 67 66

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 20: I r1.LQ:::WLWl - Fisheries

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14

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40 42 44 46 48 50 52 54 56 58 60

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R =0436210000 i

X limu~w 560 llCWmiddot 10490

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18

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2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

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q dav d v 0 dCloCV rI ds Q ~ a

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IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

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(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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j ~Wh~fl~lJ 173753 2 86876 071 NS

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fllV 1 ltJbull

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(lill) (1111) (lIlll

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

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WllIW jJClflClWmlPl miClj 1~lJ -vshy

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

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4 5281 65 65 5168 66 64 4551 64 62

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7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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i2

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Page 21: I r1.LQ:::WLWl - Fisheries

ill

bull bull W= 62948CLmiddot 10986

14

v v d r 12 tlf)Jrn~UlJYlW fl IYllm ~ ~1W

~flfJtll~Ull-WJ TIl~lm~mWVlf1llJVlJ iiBe)Wflh llJflll rHllJ-WelVWh 1l111llJ1J ffU11 ~hjJ~ 6 o I 10 QI d

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SI

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dQI QI QI

llflfl1jYlgtHItJJ~llll11Vlll~ I 1J JU 29 1l1 Wll11IfllHW 23040plusmn396 m11 fl11lJfl11Jm~1l0JmM

Q d Q QI d d

5J97plusmn279 llSlJIlI11Ilj IlCl~ml11V11m~lltJJlHW 5403plusmn247 1lSWIlI11Ilj llJlllilJr-lWlflYl 2 1l~~11fl1111 QI cv rI I jI QI I QI rI 2 11lJVlW1ij~I111Jfll111flJlJm~lltJJl~~mmlflUtJJl11Vl11~ W = 7333CW-15081 R = 04362 l~~fll111

~

ff11-WJTIj~WJ1Jfl~liJV1Jm~lltJJll~~111l1irflUOJ1l1l-WWi W = 62948CL-I0986 R2 = 02549 f1J~jVi 7

jtJ~ 6 ~fllltll~lll1-wWjI~lm~mll~

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30000 x W = 7333CWmiddot 15081

25000 R =04362

20000

x 15000

10000 I R = 0254)

5000

o tl111JVllmllJfl11~(9lJ )

40 42 44 46 48 50 52 54 56 58 60

~ ~ I do QI QI rI I jI 0 QI 0 QI

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t111-nJrI(rlfl-J)

30000

25000

20000

15000 1

~X X

bull bull

bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

11Jl1 8 A11lJ fflJllW fj l ~ I111-J A 11lJ f111-J m ~ ~ rJ -J llel~ tJ Ill W nU rJ-JYWWW 1i tiel ~ lllJW W fj l~1 m ~ tJlW q q

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30000 bull I bull25000

20000

15000

10000

R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

1ilfi 9 Al1lJbtlJllW 1h ~I111-Jm11JV11m ~~rJ-Jllel~tJ111JnUrJ-Jvi m~W TIlm~lll1~tJ TII~lm ~tJ1W 1J q q

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16

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1I-JlI11 9-27 yjol m~v 2179plusmn5 69 YJOl ~Olllll~l 1 Ill jlJJLl~l~ll1~iYI-J 305 yjOlIll~111~Yi 1 ll~ 2

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J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

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Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

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(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

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lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

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2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

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C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

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529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

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20

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q q

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t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

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I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

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q

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m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

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nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

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11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

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Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

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bull u bull

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q U q q

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liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

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207 ml

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99-305

2

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q dav d v 0 dCloCV rI ds Q ~ a

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IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

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Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

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fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

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5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

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249 038 031 202 049 041 250 0 30 027

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Page 22: I r1.LQ:::WLWl - Fisheries

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25000

20000

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bull ~ bull u~iu~ W = 7333CWmiddot 15081

R =0436210000 i

X limu~w 560 llCWmiddot 10490

25000 R = 06171

ml~rIlt)Il(l~) 40 42 44 46 48 50 52 54 56 58 60

~ 1=1 QI QI rI I 3J 0 QI I QI rI I QI 0 I

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20000

15000

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R 07middot196 5000

ml~m)(l~)

40 42 44 46 48 50 52 54 56 58 60

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16

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19fl-l~I1Jm ~lJJiilPi1J

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Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

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25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

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(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

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lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

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lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

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C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

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q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

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23

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mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

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~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

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u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

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Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

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4 ~ t QI J I

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

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24

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

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ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

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7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

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24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

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76

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5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

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10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

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20 23800 530 55 0

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22 24500 505 505

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25 27200 550 560

26 27800 58 0 590

27 23000 550 560

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13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

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9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

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12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

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16 49 20 64 6 1 4523 65 64

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Page 23: I r1.LQ:::WLWl - Fisheries

16

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18

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2 19 6 0 0 0 0 4 6667

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4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

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13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

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19

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529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

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20

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IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

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q I QI 0 QI 0 QI QI cS

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11

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

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n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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d ~ ~ ~ ~~ ~ d d

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(lI Q ~ t)I

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IV - shy

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640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

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4 ~ t QI J I

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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ltu

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nj jlJf11j V-lYlJ~W 1J 111bull

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II II I I

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24

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

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1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

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1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

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liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

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207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

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~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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j ~Wh~fl~lJ 173753 2 86876 071 NS

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(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

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YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

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5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

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Page 24: I r1.LQ:::WLWl - Fisheries

11m mlll(Jll

iJ

1900 2130 23-800 9474 19 72 42 67

Iif) 2542 IJU 6 fll i1HlW 1 1845 2030 8400 7368 19 68 42 66

J ljf) 2542 IJU 11 1930 2100 21600 20600 3703 27 70 42 67

11 fl 2542 IJU 3 til lltf1ClW I 2000 2145 22000 21000 4347 ) J 70 44 63

IifI 2542 iU 15 til iiiClW 1 1830 2230 21800 20AOO 5j85 27 74 42 70

Iif]2542iJU 15fll tflt)U 1 L8-30 2300 21600 19000 12381 ) 4 70 43 63 -

1ifI 2542 iJU 15l1 t~t)U I 830 2330 23600 22000 6400 25 79 44 70

Iifl 2542 IJU 15 til tltf1E)U I 1830 2400 24800 23000 8182 22 78 43 66

25 vfI 2542 Uill 3 til lllt)U I 2100 0200 24500 22800 6800 25 68 43 69

iif 2542lmJ 8 1845 2125 24000 22200 7200 25 85 44 74

2 1IfI 2543 UilJ II 2400 0300 19000 18AOO 5154 II 86 43 74

41LfI 2543 lmJ 13 til iltflE)U I 2230 0130 24600 23AOO 4800 25 88 44 74

(I) 1111 2543 11 I f11 l~BU 2 2130 2400 7407 27 75 43 68

61111 2543 2 2200 0100 20000 22222 9 80 47 69

7902 2179 7593 431 685

plusmn1 ~86824 plusmn569 plusmnOIO plusmn037

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

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Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

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100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

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ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

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in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

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fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

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II II I I

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24

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~

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QI QI If I QI

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IQlr d Qlvrl I j) QI

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1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

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q U q q

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Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

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3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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v 0 CV ltV V

VlrnU-Itl1l1fl4 tJllPlfl (fli]J) mllJfl1HmtpHl~ (lflJ) flJlllVlJflitltjn)~ (lfll) lJEl~t1mYllmmtJ

fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

8318658

15250623

50

52

1663732

5129714

Y1liljfll-t1f)~7 ilml~MWJl1Jmh UJ 1W DYlJ lfllJll~tlJl~tJ 1~~lrn~

m1~~mllJmhjl U

j~11l1~fl~lJ

SS

12595

df

2

MS

6297

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244NS

fllUlt1fl~lJ 11n1lJlI

129143

14173S

50

52

25 83

mjHfIl-t1f)~ 811ml~ljfl111JUU)Un1Dmlfllj l)~rulfitJ lYl-cYmiYlTI u

SS df MS Furtri ~fll11JUlh j lU

j~rtl1~fl~lJ 8004 2 4002 263 NS

illtJltlfl~lJ l1~rtlJII

76102

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50

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 25: I r1.LQ:::WLWl - Fisheries

18

0=1 f J) cl 3 l1ifll1hfl UWllfl1~el1WYJmlJl~tflJ

lrl tHlli lvl lUvl~ Ii111j i ~ u~l1lJlvJ mnu i ~ lvl 111U vitYi j 1~ i ~lvl1 iivl Bj tl1 fllft lriB i ~B BfllJ 1

l di cI 0 1 l II ~ 1 ~ I 1 1 1 ~ bl1lJ 1 UlllljlijllhIlWfl1lJi1UBjmBmH)~ m hJlJ1 1lI1jVlj b1 12 Jil1lJj lUlli1jlfl1~llllmj Ill IlBj

Inu i ~ i lil r1fl j ~ VlU fl j ~ IVi BU II ~ 1 1lfl~Bjl1lJ1V 11l Vlfl ~BU V1V i ~1tJrIj1WflEl ru~l~ lJlvJ B fll1lJ tJ BIl DV

UBjlil~ml llflflljf1fltJlYiui11lJjB~lVlflllh11i1 40 ill iielmlflljVlfl 4082 111Bldlfl111 1Wj~v~11mVlfl

100-110 ll1 ~etlll1fllJm~tJ 288 Bjftlllim9fV(Y 1WjBflBlfl1IlYlfll~W1i1 57 1i1 iiellljlflljYlfl 5816 bull ltJ

11JBIO~W11 lln~v~nmVlfl 74-84 111 ~Drut1fllJm~v 305 tljft11liI9fV(Y 1Wflritlj lvllJYlflL~Wlil 54 1i1ii bull ltu

elllj1fl1jVlfl 5510 ItJBlcUWiltWj~v~nmYlfl90-IOO 1W ~Bflll1fllJm~V 297 OjftWlJI9fV(Y ~jmjN~ 2 bull ltu

lri m11 ~ 11 tJ BldlfU 11fln Yl fH ~ U Ii1U Bj 11i Lvl 1m ~ tllU ~ Yl fll Ul~ ~i~Vl ~ Ojilm1~ 11fll1lJ II tJ j tJj lU Vllj I I I

(y~~YilJhtJBldlfUl1flljYlm~j 3 11) lliiifll1lJUllflvl1jnW mlljiitr mhltyVl1j(Y~~vh ~i11Jm1lJlllmjW 95

I ad a ~QI

luBWlfWIlIljIl1j1jmnflVl 3

Vln1j~ 2 ~1Ulli~~lj1jVlfllWllvl~11i (yiBj) ll~hUlUlli~YlfllthlIi11Wllvl~iTI j~v~nm

1Wfl WNfl (1W) ll~ BtlllWlJ m ~VIl tlm~V~nlYlfl (tljft11liI9ftm)bull ltu

lliiyj ~lWnj1~()-j()~) ~ljl~jY1 fll~~j~~~l1i -- shylimlJ~ n~Ylnj i~~~1 lirlfl~jnl~ nri()~ 1)~lJ

lI~nlli ~ ~ ill

19 6 2 3333 4 66 67 4 6667

2 19 6 0 0 0 0 4 6667

3 27 I 4 4444 9 100 9 100

4 23 7 5 7143 5 7142 4 5714

5 27 9 6 6667 8 8889 5 5556

6 21 7 7143 0 0 2 2857

7 25 8 4 50 00 7 8750 6 7500

8 22 7 0 0 0 0 0 0

9 25 8 2 2500 3 3750 3750

10 25 8 6 7500 8 100 7 8750

II II 3 0 0 0 0 0 0

12 25 M 2 2500 6 7500 3 3750

13 27 9 4 4444 7 7778 7 7778

14 9 3 0 0 0 0 0 0

5700 54 00nlJ 305 98 4000

Oll llflu n fl llhhll(ll]oIWt1) 4082 5816 5510

74-g4 90-100

305 297

19

lrimlfH9l1i~plJlnG ~n111U ~C1fH~ 11~ l~lUilaf) hil1~tlBm1illf) 1 ~ C1fH~ 1tlltJ 1U ildj~mHH~cu co U q

rlm--u1~f)fllJllC1~llm~lio~1~~~~t1a~ 11~~~U1JHljlumjlw 5-71U Ygtl~ju~ 10

_(i4c7~ ~ - ~ ~ r~~~jjf STl(fij~i~~~~

JlJ~ 1 On f)l~ 1~n1~~Ylf) a af) ~ 1f)1~Il C1~ fl fH~ 1tllV 1 111 ~ a af) il1n1~ ii~f)1] tll~rl allU 1 m C1lJH C1dJO ~ 1~ cu q q

~~~11tlJll9lOlJlnvldm(Jl11 5-7111bull

~ I ~ bull

C1 f) 1191111~ 11 1J YI Vi 1~~ 1 f) f111 Vl n11 i ti a ~ 11 Li a fl aUf)~ Illl fl ~ f) C1 a ~1vllJ iiU111 lf) LQ ~V

QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

~1nJ~ II

ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

1IlJfl11Jl~mtl 95 lUtll~ljy) iijPllHHnnfl~ 5 6 ll middotCl~ 7

23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

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II II I I

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24

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QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

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1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

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2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

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5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

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2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 26: I r1.LQ:::WLWl - Fisheries

19

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QJ 0 Q) v I 5028plusmn447 4756plusmn427 4l51 plusmn4 79 tlJ1J 1l11JC11mJ m1lJm1jm~~ajmC1j 553plusmn065 569plusmn052

529plusmn082 19JlIALlJIlj IlllJ~1Ygtl1J m11J1Jllm~j 616=046 625plusmn038 581plusmnOAO 19JlJ~llJm IlllJcl1f11J

~j9ll11jNlnf1Yl 4

20

~t IV I ~QI JJ J 11HltlW1JlnilIbYW f(~ flliVl~1H)~fIiU 15 1W nfllP11lJntlltlj~fll1lJ 1f1-~tltl~m ~~tl ~lJltltlW

q q

~1nJ~ II

ltJn 11 nfll~lljjtlffWf(~fllill~ntl~tllV 15-1jU U q q

t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

l~l~Y1mJltltitl~j litlfltlWtl~P1ltn~tl~tl~l~lJ 6 26 969 tln~ 914 t1J8t~W~ P11lJ~1~U nJtltcliW~fll1lJ ~ J od 0 0 QI J od 0 ~

tll1~tVllJ 1898 1493 Un~ 2098 It tlitCJfjP1 P111J[1WIU It OitCJfUP1f111lJVllIVllJ 487 128 Un~ 740

tlJtll~mf P11lJih~u 5P1jltlliil~UJl~ lJ lP1~lYm (specific growth rate SGR) llJtlcliU~~tl-1W

ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

ICJfU9lllJP1j9ltlljfll1lJVllm~~tl~ 00010 00053 Un~ 00020 ICJfU9lUJP1j9ltl1U 1ll1JmmJ ~~9lnl~Vl 3

11

21

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a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

bull gt bull

BPI) 1fll1 IJ~rul~1J lP1UtlJOfH~lVi ~111 J 1flflliVlmlJ 3 ill ~hJ~m1lJllPIfl~1 HlWmhJ~tmi1firu111JLlG~Vi U GJ tJ

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23

l f - 3 OnYlOtll1IinllnO 1IJlnmel1h1t11t1lJ1iUJ

lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

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fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

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~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

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j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

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bull bull

24

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~

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ICVd1 q QlQld I 11 - QI

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QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

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QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

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ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

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I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

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11

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m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

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liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

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cI

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

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1663732

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 27: I r1.LQ:::WLWl - Fisheries

20

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t]flI~ltllq 15 lj ~Vltlilltllitl~W litlfltljtl~P1 lW~tl~tl-l1~lJ nJll1rfllQ~V 5343plusmn249

QI 0 QI V ~ 5217plusmn202 Un~ 4749plusmn250 flilJ P11lJm~U mllJml~m~~tl~mM 658plusmn038 654plusmn049 Un~ 640plusmn030

~ OQl Q q OQl

[CJfWP1tlJP1i P11lJnl~U mllJVllm~~tl~mnV 646plusmn031 633plusmn041 Un~ 624plusmn027 lCJfWP1tlJl1i P11lJm~U

~~P11il~HWltlHWltlYi 5 bull I )I

I-11~1t1~vYil~mfhBP1iltlljtil~qjl~hJ 1P1 (growth rate GR) t1Jtlt~W~ I-1mumVjlJtltllt]tl

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ttJtlt95U~~tl-1jtltl~Jmrfll~lilltllitl~U litlfltlUfl~P1 tW~ tl~tl~ l~lJ 020 027 Iln~ 027 ttJOl~U~~tl 0 00 01 QI jJ od I cv 0 QItl tl 1W P11lJm~U t tJjICJflJP1P1tll1jUtl~mllJml~m~~tl~ 053 040 Un~ 060 t tliICJfUP1P1tllU IlllJm~u

1 od 0 QI I od til QI 0 cv cv amp) q 1 lutJjtCJfjP1P1tl1UmllWllm~~tl~ 013 007 Un~ 027 lutliICJfj9lP1tllU 9l1lJm~U O9liltllHilH1Jt9liJ 9l

IV QI 0 bull IV I QI 0 IV I t QI filJVIVlti (relative growth rate RGR) mlJ9lfllUUtlnnl1JtltllJllll 00034 00045 ln~ 00020 mlJ9ltl

QI 0 cJ ~ I QI jJ I QI 0 QI

1W Illmn~u CJfWP1llJ9liP1tllUmllJml~m~~tl~ 00039 00030 tln~ 00043 ICJfjP1llJP1j9ltl1U P11lJm~U

q I QI 0 QI 0 QI QI cS

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11

21

V1nlj~ 3 BPI i 1fllHIl~rul~hJ1PI BPIi 101i 111 ~rut~J [PI ~ llvn~ llrl~BPIi1fll) tll~rul~U1PI u u u

a a I d a

LllJVIVlli Utl HlfHPI1m~tl1~ YHJ1V 15 TW ~

OIIllfll j[ J~ lJl~ll~ O~1i lfllllJ~tlJ~U 1~ 5m lfllm)~tl~U i~nllJ~YlTj

in fmwn (GR) ~lYil~ (SGR) (RGR) 1JII]l hflin flil~ IJl1( ) lhlnin nil~ U11( lill1lin nil~ un

() () (111-1) (1111l 111) (flllJl1ll) (lUJl111) (lIlJl111) WI

tlB~j 526 1898 487 020 053 013 00034 00039 00010

tlBf)ellJn~~ 969 1493 128 027 040 007 00045 00030 00053

n~B~ h1lJ 914 2098 740 027 060 027 00020 00043 00020

PIrl tl~ fllJ VI ~rl tlJrl fH~ 11JBPIi 1fllii tl~PI1V I00 IUtl19hJ~ ~11~ le)PIi1flli IIl~rul~U1PI BPIi 1 ~ u

fl1 i I J ~ rul~111 PI ~ 1lVi 1~ II rl ~)PIi 1fl1)t 1l1rul~ J [PI ftjJI~ VI fi VI ~LltlJ ilfl i 1~rfl 1 llJttU) Ui 1WVI1J ffG~VlJ 11 u u

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23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

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~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

V

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d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

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JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

y

ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

ihll1fllJ 2542 fi~ lJm1fllJ 2543 1~tllIllrl~191111~1~1Ulh~ 9-27 ~B~

~ QQ ~~ I I ~ 3 f)llffflil111ifl1h1f) Ulmm~~l~YIlmJl~tYlJ

f1l j wtll ~hJ1iBihI UBflmJtl191llrl~tlcitl~ 1 ~lJ hhl9ltlll 1~tlUYl1~bIi1~~j ~v1Um1lJl~tl~hI 95

11JtllcJjtj~ (pgt0 05) djmj1~tlllllYil~iJ1tlf1ljVltlr1~ 3 111 B4UlrlmU 15 lUnti9lj1f1ljjtl~9I1tl 100

I U tl Ictlhl ~ ti91 j 1 f1lj I il ~tul~1J 191 ti91 j 1 f1lj I iJ ~tu l~U 191 ~ 11Vl1~ llrl~ til j 1f1l j IiJ ~tul~U 191 ~lJWYlli1lJtI9I tlll 1~ U U U

tlhlV1l~LYO~ ~j~v1UfIlllJl~mrll 95 IUtllcJjhl~ (pgt005)

11

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_IQ ~ I I v d I v ~ t

1 01 l1JlUViVI tll~VI~ ~ 1I1~lIlJnl ~1V1l m ~ tll~

~ I I QI ftr1 I q JJV QI ~ d I I

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q

11~~ Im~hltj~11~1~WJjtlUYiljYil11r1mUmtliJ1tlflU ~9Illll~ 1l1b1~l~ 1i1t1~ I~U~U

IJ ))J~1-Ut~ ~~q1trt~ lpLLl)JtQfbl~l fl1 U~lLU ~~l1rJl~ ~U11Q ~

Q[t11 bL~I~1 U ~lLU ~~UQpoundI~1t~ ~Q~tQ~ClU11 Uqlr1l0L JIALArtflWl n~lmpoundCllfLULlwg ~LMIL~ wL n~1

m iCl1fLUL lwg wl n~lmiCl t fLUL lwgIJLnMQfLU 111 ~Q~tQl~tlQlU~fLULlwg~~tt~tJ~r1Qk1~U~fLUU~U

~r1~rrt~IJt1 LWtU~kMt~MULU lLU 111 ~l0t r1Ub~CI ClLQt JG ~~1 rtLrlW~ClWw11QIltQnULCI U~fLUrtLmmQr1 ~1 s=r I 0 1=76 l I r JI(I ltt JI I 17 ~

nr1 ~fbl~1~CI w~u11Q~Q[1~t1 ~2mLlAtL~UWllrtLl~~rt1tQIJ ~1A~t t t~W~uUUL CI wuttraquo Qk1

r1l~~1U~rtQUMb MQ~l~M~~llIt1 tIJttw~rt~CllfWrt~~UQ~~lJl1 r1l~tm~r1LW~QffLUL lWg~L ~1

U~Ul LlACLQLrtUQQ~Ullumnbrtl tr1tW1t11 dJULU~LfMltQlALf1UtW1t11 dJlUWtQW~IJt1 ~1 11111nQr1LW (f) b I lrt 11) lit (fj ($ fA (f) lit I bl 0 I I

CLQL~IU~~LfMl UQQL~lU~~U1WlibtQ~~ClrQW~tQ ~1~1 ULI1UQ(W UrtL)Jrtl~l1l~t1Utu(WUWll~1UQpoundfb1

~LClll~HnL~IU~~LfMt L ~r1~fM~I~Whflllt flgt1 U~rtQ1-WLClum~rulJlltrt~l~lJl1 l~rtg1U~L~IU~W11ULY

nWQ~l Mu~mt L tL~WrtUlMtW~111lN~kt ~1ULClUQQUt-tlu~M1I-qUIJMLIJ~L~IU~klbQ~tl U~Q[1mllirl ~1 tLlrtIJ U~ll~mttPbtLt~IJ~ULtttQ~l~It~IUt-fLU W11 ULyn w~u1 L~IUIJQ[JI fIgt 1 Ut-~1t1

11Ulr1WmJ1IWl r1)J1f1 l-ll m1tLnQlll1J1rtLd~nUl~lnl I1U1ULrt 11) s=r I (f) ltt J7 I lrtft b I ~f 1= I In 0

Wr1~I~IJ~nL f IA~IJt1 ~1rt~rtLlllt~IJm)Jl1rtLnl~r1libt~rtQIlt)JbLIltt-lH~ uuf1JU p ULrtr1~lf1gt1tt1 IJt1tptQ

11grtLllltnpound~1 rt~Qfb11b L ~1~1tt b L U~ rt~1tQ~U~IJl1W~UttQ~QfbI1QJum~111lLIltt-l H ~rt1~IJl1 tl1-WQ IJW~

rt ~lrt1~1WWllt)J lln11 ~Qfbl1 br1L f lA WL Utt b Lit L t-t rt ~ nQl~t1 rt~~tQ I1~Qfbl1b~IJ llUL rtU W n~ t 175Z rt ~Ln

t-~)~ 11ttL IJrtU 11Q ~lt-L ~ U~f ~1 U t-L I1l1~r1~ ft-lfb It1t-QIJ ~IA~[InbIt~Qfb 111 r1L fIA t-QI 11~ rt Ll~

Cb

It)mYl Hll ~ 6 ~

m j illnl nlt1J iltl liWI~ UC1~tllfllJ 1-Ji rn 2542 Uil JV~iJ r--H1vltH)(PIj 1fllj liln 1li(1~fllj tl4 Ul~lVtlll11j

li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

I d dOl ~cr~ Q QI I Q QI 0 I

iJilJli1V 1li111JVll f1lfiJ l(i)-Jf1~~tl ljjtljcy) iIjilJ9Wi1~ Uf1~ViftlJV 1Vi-JIfI 2529 fllJlll~Itl~(i)l

m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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VlrnU-Itl1l1fl4 tJllPlfl (fli]J) mllJfl1HmtpHl~ (lflJ) flJlllVlJflitltjn)~ (lfll) lJEl~t1mYllmmtJ

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UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

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cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

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Page 28: I r1.LQ:::WLWl - Fisheries

11

21

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~V1 ntJ nllIVi 1~ ~jfilvl1flj tlllJ 11J mJ nWlh ~m 39(6) 629-637

1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

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liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

Q113NNt10n 1 ll1llfwn nnUmHm2~tl~ tll2mllJVldm~IN YH)YII-~t~lmtJlt

fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

Hi1 ~ ilftl1lJUlhtll lW df MS F

j ~Wh~fl~lJ 173753 2 86876 071 NS

4765512 39 122193

493lt)265 41 --~

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v 0 CV ltV V

VlrnU-Itl1l1fl4 tJllPlfl (fli]J) mllJfl1HmtpHl~ (lflJ) flJlllVlJflitltjn)~ (lfll) lJEl~t1mYllmmtJ

fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

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15250623

50

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5129714

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m1~~mllJmhjl U

j~11l1~fl~lJ

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12595

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2

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129143

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SS df MS Furtri ~fll11JUlh j lU

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 29: I r1.LQ:::WLWl - Fisheries

23

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lu l~lmtllU~W fl1JUtl~hJ UClfl flllfl1 9l fl ritlJ 1vhJ iJU9li 1fll jwn 4082 t1Jtlt~U~ 5816

tlltltcUU~ 5510 nJtllcUtli 9l11JiiWlU 1~tl1ctlrHn 100-110 1U 74-84 111 90-100 1U ~1l1Jii1~J ~ ltCl do do do OQlamp I f

fJfllI1fJ1J 288 tlJfflI9HH9W~ 305 ClJfflI9HWlitl~ 297 tlJfflt9HH9W~ mmn~u 91J1IIlnIl1Hl1n tJtlJbltl

QI Qamp dQl rd r~ d ltflflUl (2529) 91JWmnmlllVli1t1VlClIlnWn 50-60 aJClH9itJIl tUmn 90-120 1U VWUll1f11J 60-70 tlJfll 19lfllclftJ(1 ViJ-wVi1JhiJum 1miWmlf1jtlllf11Wn 1 ncil~tlJnUll~iJfJUll1fJiJ~ lJnU1J1n l~tlYi~JJrJ Un

mu (2529) j1~JlUl11~1lf11Wn 80-100 lU Vitlflll1f1iJl1n1Jwnlu 60-70 tlJfflI9lnclftl~ ViJ-Wtl1l~UJ l~11fllj1~CJflI11fJiJtl1lvhmj1~~thlj1tl l1~ClU~lllll~iU~J ll1m11JrtlU~J ~JJU1Unwvln iu~J~ClJ

914 amp doJ Q d 4 ~ 9 ~ t1J 111 n flYHlmJil1j 91~lJm1JJ151mflflllll1 fi1JVltltll11J1 1tJflljVl~ M)Jt1J nn 20 t9lUllllJm 111 Hn ~tl~

~ dI1 II bull bull 1 In1 d 1flnCl~nu bl1lJn IInfllll (2533 n) 91JYl~flCl~Yin lJl9l1mtl1U UnCltl~ YilJ1Jtlllj1mjYin 4508

cJ 9 d d 111 ~ ltJml9llJIl lUllf11 92-150 11J Vlflflll1f1lJ 305 tl~ffll9iCll9ltl~ mjYin 1Jj1111~YiflIlCl~flCltll11JW9ljll~ v

ffll ~ ~ 1J 1 iJl11Ii11J1 Vi 1f11 tllu llflm 1 l ~Vl j 1tlVi liw n lu 111 nlj 1tllll1 ~ l W In W i ll~ OJ Vi jlJtJ1 n~ hhl~

Iitl~lliIl11llHnUl1J m1JmjVl~fltl~ lJt1Hilfl (2535 1) fljjtlim lin tllfl1J (2542) mil1l1fl11lJ91WiJHCI bull u

~ tln wvl ntltlm~tJ~lJtl ~~ll1nIlYmnj 1 U9lj 1fl1jwnIJtl J~1tJ1~~lJm111nl1l~ 1Vl1f1 1l1lJ j1tlJ lWlJtl~

~Vi l1 (2528) j 1tlJ 1W 11i 1m11J 1 lA lVlln llfl Ii j jlJ151~Wn1Jf1ri tl~ hhJ iJum 1wm~W~1 697-83 2 bull 111 f)Ic]jtJ~ miiJ 5m 1i Cl~ n~ ~VllCI ~1 mm 1tl ~ 11~ lltlJ1ilfl(253 5 fl lIfI J) n ri rd W10 lllh~tJ l~1Vl1f111 ~

1~flj11[ltlJ1n 1 ii5o)j1mi-Wflfl~tl 8323 llltlIc]jU~ lW1f11 47 1U ~6rnl1f1iJ 3161 tl~flll9lnl95(m ll~iJ QI I lIt~d cld cIf IQI ~ 9J d Clm 1H)~C1~~Yl1f1 1~IYW~ 000 i-0002 lthm9l11l Jm)tJmmfl1Vi~~lll~nmJlJClJVllnVll11J1~1J lln

~9l11~tltlf)f11Wri1Wll1DJWflYiCllll1f1iJlTlciI~tljtllJ mlJmjUIf)1 ICI mfllJ (2542) lC1 tlYlWll (2537) jltl

u q U q

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J11J fl1 j Vl Tl i u1lVi111111oWl rfil~11 11J llCl ururu1 (2536) n ri 111101~tl)f)1 j lj lu11 flliJ tl~tl ~wnYi flUII1 fliJ q u u q qJ

d ~ ~ ~ ~~ ~ d d

30 tl~flll9in91tJ~ llnmlle~f)1jljlJIViflrl IltljYinVltlfll1f11J 31 ClJflll9lm9ltl~

Q

JWl~JCl~(f1lA1~WnCltlnlln lunmA1mrJ I 1W llmJtl~W UtlfltlUn11l 1lnnrie~ lvllJ iJ~l

)I I I

o (lI (lI d ~ do d d (lI

Wll1UnlllflC1tl m11Jm1~f)J~Cl~m(tl lC1m11Jtl11f)J~tl~mntl mlWl~ 5028plusmn4A7m1J 553plusmn065

19lW~llJlli 616plusmnOA6 19l11~IlJ9lj 4756plusmn427 n~1J 569plusmn052 19l1~llJm 625plusmn038 19lW~lJllj lln

(lI Q ~ t)I

4351plusmn479 mlJ 529plusmn082l9iUllllJ9lj 581plusmnOAO 91W(9111Jllj ~tl~ilntl~mJf)1jVl~fltl~lJtlJ (bll1Jn 2533 1)

nri1111C1f1l~1~Wnlln liJJmlf1ll1~tJ 508 n-rlJ m11Jnl1~m~o~m~tl 664 19lW~1J9lJ m11Jtl11

m~ Cl~m ~tl 632 ml1J1n~f)l~1J1tlml1 1jtlf11mr~~tl1J~H~lJHmj~51lj1ri1U I I 11 ~jllnm~U1mjn v

~lCl1tl 15 1U nf1l~liJ59ljntl~llltl 100 ltJtlc]jW~ IJWl~mfttJhl1ln fl111Jnl1jm~tl~1lflm1lJtl11 m~Cl~lJtl~nf1l~lllmitl~lJ JtlflCllln11l lnnritl~ lvl1J fie 5343+249 n-rlJ 658+038 19lW~llJm

IV - shy

646plusmn03Il9lU~llJm 5217plusmn202 n1J 654plusmn049 19lU~llJ9lj 633plusmnOAl 19lU~llJ9lj 1ln 47A9plusmn250 n1J

640plusmn030 19lW~lJ~j 624plusmn027 19l1j~lJm Irlmh~lm~tlIJtl~)f1l~1~l~lJ~U111UIlj1nUll~tlJl~U 19l (GR)

bull bull bull

bull bull bull

22

Q d ltOf

1 illm~tHln rt mn

4 ~ t QI J I

1 fll a rt Oll HIf)1I W YUH 1iJ Vi U ~ IVIl m tll U

~f1EJ ~ vi D ul1 yi1J jb~ 1 m D11J ~fi lfl11lJ ~lJ-W1J1r1l1lJ ylJ fl1 j 1~1J mJ hHJ~ fl11lJ ~lJ-W1Jbull I ~ ~ CV I CV d v cv d I

j I111J fl11lJ O11J f)j f1D ~u l1111J nU fJ WHlVi1J 1i W = 56011 C W -10490 U fl11lJ YlJVi1J 1i j 1111 ~ bull QI I QI rI QI QI d I ~

fl11lJVl1f)j f1 D~ bb 1J1111J f1 U tl~Vi tlViW ~ W = 71275 CL -20416 u fl11lJ lJViW li j 1111 ~fl11lJ 011 ~jI o V I QI rI QI QI tI I

f)jf1 D~ UW Ill W nU J ~ bblJIIW ~ W = 7333 CW -15081 bb fl11lJ lJViW 1i j I111Jfl11lJ rJ11f)jf1 tl ~U

- QI levd I QlQI~1 ~ W111Wf1UfJ~LlJViW~ W = 62948 CL -10986 Lyf1n1Uill1l111nmbVilJUWfl11lJ011~Lfl11lJrJl1

f1 jf1 fJ ~ Ob ~lJ~W1l1lJ ~f1 fill1 IlJ tlL 1 W UW 1f1 t11 11 tmvi 1tlW O~fl11lJ rJ11U tlJf)j f1 fJJlJl f1 f1 11fl11lJ

I QId c J d cv ~ ld QIf111 ~ blJfJ UIl W 1I1W f1 rJJIVi1UlJlJ1f1fl11lJrJl1f)jf1 tl ~ 0 lJ fJll j 1f11 j bVilJfH1 ~f111fl1j bVilJVl1 ~fl11lJ 011 ~ f1 ~ tWb~wi1b~rJYiB111~ ~ fi DW Ii1~ o~ f)jf1 tl ~ f1 lJ u bij mYimJj 111 H vim~W ~uul1wW (VilJh 1W U 1J1f1

tl1 11 tm l11 tlW ~1b~ rJ 0 ~ fl111J rJ11 u fl11lJ f1 11 ~ m f1 fJ~lJl f1f1 11~H41 lJD fl11lJ ~lJ-WWVi 1~ 0 n

ltu

lh lrJlful W fl1 j flf1 btl Df1fl11lJ lJ p tlrUD ~ mJ~W ~ u 1 ~11lW IifJ~~bf1Pl lViff~ lViffl~ rJ 1 W fl1 j 1~1 j lJVit)~

nj jlJf11j V-lYlJ~W 1J 111bull

i Q QI J ~ r 2 fll Irt f)1IlW~V1 f)j1iJf)11mnJVi~1~11~ lUIVIlf)Ieml

II II I I

f11 j 1 H hiUtl ~l~1f)j tlllHll f) f11 HI 11 cHJ ~ fl i~ ii ulh~11lil~lVtlYi 111 9 yhn u lJ1f1l1 111 27

lID~ yDlIflD~tluYimjrJ IWfltl 2529 ~~f1~11hl9hf)jmW1Ml~ 12-25 yjD~ ~tlul1 J111tf1UD~Ul1l~1 I )I

Yiuf1 ~riDW11~ I ~lbm1~1l~ llim~rJ 7902plusmn4698 f1ilJ 11llJtl111tf1 1~m~mlliyjD~ 7593plusmn684 f1ilJ

lf1I~rJ~tluuwli1rJ Ufl( (2529) ~Jf1~11111~uliyjfJJI1tf1 6971 f1ilJ J111tmhWf)W~~11lWri1Wbull u

U D-l lij fJ mb U fJ ~bI1 1~ ~ ntl ~tl tiD Df1lJ1 ill f1u1111i 1 1f1mli wi1m O11-l I~ 9 TI1J 11fllJ 2542 Un ~1lf1 r11rJ

1 ~d 0 ltk I lIJ nl od 1l~ i U 6 lJf)j1fllJ 2543 bf1~fW~mJ f1lfjj (2529) Cf~f1m111 bIl1f)jmWO11~ bUujlJ1illbf1DWliW11fllJ

~~ ~ W 1fllJ 1-1 li1 ~ l1 b1 rJU ~y rf1U n~ m f11ffl1 til 1 fl1 j 11 ~ I ~ bli 1tl1~1I il ~ f1 11 ~ lJ 1liW b~ rJ1tlUj 1rJ ~ ll U fJ ~

llllljOu u 1JSrJ (2525) bl4qJlI1rJ bl flf1 (2529) bblibbllf11i1~01f1b1i1Vlt1l ~O~f1Vlj1rJnWI1~lJf1l1~ r II I II

f111lhf111~lJl1b~rJf111 UWI1Jtl bb11~1~tWli1~tl1b~lJrl~ IlllJlJlf1tJ bmy1fJrul1 (2531) -n~iimOO

bfJ U1n lbli 1Vl mijtl~ lJ lUll IilJ 1f1~~Vi11~UI1 lJ f11~t~ tl j V ~ iJtl~ lJ1 ill I ~ uli IIi 1Sl~ f1 OUf1 11 lJ f1 ll~ bb 1 bull II II

finWH lJil Uf111 lJb~f1ef1iWbllWbVij1~ili ltl1~f1 t 111 ~limbl1~1W1WltW 1 Wfli ~iiUl1bIi1f)j fl1W 29 ~1

ll~~l~bY1rJ~ 14 ~1 lf1rJ 10 ~1 11~1~hlliN~W Uyf1~1111~~1JVlll1ijrmiUf11Wf11jl1~1~UD~blilJl~f1 lJ5lJb~ rJ1tllJ ylJlf1rJ Un ylfJO 11 (253 1) ~~ f1 ~1111bb ~ ~WVltW 11lf1 l~ftW111ijri1W 1 W f11j jU O1W f11 j

J VI 1 U u ~ lvl 1Y1~ fI

bull bull

24

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ti9lj11iJ~tul~U 191~-llll~ (S(1O tim1f1ljLiJ~tul~U 191-fflJWYlpound (RGR) IijBYl~bIBUYl1 middotr(1O~r1~ 3 b~ Yl~HW~ U U q

~

VlUi1r1~ 3 i11 hju9ltlIl1~ntHJJ1~lhjfHhqjYi 95 1lltlctlU~

tYltl

~ v r I v G 1 f1l j ff f) In tl f) il W~YHlIIlJ VI U 1itVll m ~ ell ~

ICVd1 q QlQld I 11 - QI

Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

IIl 1m ~BlU flbllJWU1rj~d1~I~BU~mfllJ fi~ I~BUVltjf~f1ltlU 2542 11~1~j~I1i1~I$iBU

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4 12000 40S 460

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6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

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5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

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10 4815 54 64 4571 52 60 4098 46 55

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12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

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10 5008 64 64 5371 72 68 4672 62 60

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16 49 20 64 6 1 4523 65 64

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III 4960 62 58 4713 67 64

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Page 30: I r1.LQ:::WLWl - Fisheries

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Vl BVliJ lil9l1m~mUlJflJ1lj(1lJVlhlli j~I111~flJ1lJm1~m~~ B~llrl~U 1l1U tl W = 56011 CW-q

QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

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q U q q

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q dav d v 0 dCloCV rI ds Q ~ a

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fl1fUml0

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1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

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2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

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d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

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5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

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4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

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249 038 031 202 049 041 250 0 30 027

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 31: I r1.LQ:::WLWl - Fisheries

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24

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QI QI If I QI

10490 fll1lJbIlJVlijlij~I111~m1lJtJllm~~B~llrl~iJll1Utl W = 71275 CL -20416

IQlr d Qlvrl I j) QI

UlJVlhllil9l1m~mU1JflJ11J fflJVlhlli j ~n1~ m1lJm1~m~~B~IW~tJll1h1tl W = 7333CWshyq

QlQld1 0 QI 15081 WlllJbIlJVlUli j~I111~fll11Jtl1lm ~~B~lrl~iJll1U tl W = 62948CL -10986

~Q QJG 4 2 f)l jff f)illVIt VI f) j j 1J f)l j fl tY1JV1 ~~ 1I1~f)l j 11-3 IIll tl-3 IVll m ~ ~1~

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9 12400 420 460

10 13000 440 470

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13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

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24 4S5 490

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10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

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20 23800 530 55 0

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27 23000 550 560

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18 476 62 62 6013 71 69

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  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 32: I r1.LQ:::WLWl - Fisheries

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207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

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TllJrl tiiHrul Hl tltTW~ ti111rulJrf ff1~Cl 1ri~l~Vl~ llCl~fhYm tlVllJ rltwlWlVl 2533 fl fllJVlVlCltlJYlfl ltI u V II~ ~

~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

~llJrl T1JUJ1Jfl tlWll(l t1l1HfNfl ~~Cl 11JWW1Vl IICl~ fllYiCl QVllJfltwlWlVl 2533 U fllJVlVlrltlJ~fl

IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

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fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

Q 0 ~v ~ I(V(I

1iI1ll-lf-nrlflYl2 Wll1Wmn rnllJmlJm~ltlJ lLomllJVllm~lt)J tllIY1W1illillmt)lWbull

ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

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fllV 1 ltJbull

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fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

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11 11( ~ m llJl1lh ntl SS df MS F

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j~rtl1~fl~lJ 8004 2 4002 263 NS

illtJltlfl~lJ l1~rtlJII

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Page 33: I r1.LQ:::WLWl - Fisheries

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It)mYl Hll ~ 6 ~

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li14(i)vl1-J 1 Jtl-J(PI~Inulhvrtl~1I9li11tl 11Hnjnliih~lJ~52(5)457-463 shy

rhli j U(i)~(1~ ~1J 2529 nlj 0mJ11utl-Jl9ltl nlj 11-J 1liJtl -Jlvl1m~tll1 11JCY1j nlJth~lJ-J 39(5)535-539 Q Q d Qc ~ jJ 0 QJ Q 0 Q 0

il(PI (PI 1J1 tllV(PI(PI ~ n~ 2536 lil 0 (PI Utl ~(PI tl CY1i1 J lJ1Vl V1 fll (i)j nliih ~ lJ -J l f1 ~ lVltJ1fll MlJ Vl1 wl~m m lJbull

lh~lJ~ ~2 11th hV1IPJJl1 f(PItl(i)l(i)l1J lJiflltJcy) i1fl1l1wi1~ ihlij iltOJftn~ W~ llDVlVi6 ilttjf1n~ 2541 nlJtll-lJ1f1

(1 f1lvl1m ~ V1tll tl utlflVtl n~ (PI 1(i) V1ltUtl1111 J vl H lil1(i)tlll ltl n 1J ili1flliinJU~ 2312541 f1 tl-J

1 1 $I0 uJ~lJ-Jmil(i) miJuJ~lR 18 i1m

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m~ tl1tll tlU ml1(i)YiVlJllf1~I11(i) li j J1Jli1~ 11j li nllUl ~lJ -J 39(4 )391 -421

1~ OJ ftfl r--mrn 2535 n 9i1iVltJ1 J V-J vl1Vl~ 1f1Lf1 ~ ~hiVltJ1Jtl -J nlJ II IiU V1VYrtl iU tl -J vl1(P1 ~ l- 1tluJ ~ Vl fI

1Vw l1HY1jnljUj~JR 45(1 )603-646

1ltUlftfl mn 2535 J nlJnl1~I~V-Jvl1Vl~lf1hnh ~1Vlfll ilV llilJmJUJ ~lJ -J 45(2)717-741

bull u bull

UtlJUJl eJ -J U J ~ fll flj 253 G 91(PJ[ 1~ tl HUU V -J tl11V eJ-JUJ ~ fllflj 44 11 vhlJ il J ~ 1j1l(1~ltl ~9I1 u u bull

lJVliih1nlJ 84 mh WiTv I-JIfI tJtlJlllV llilllrVll ihll(1 V(i)lJflUlltl1Vl lm~ ii1liJ l(i)-Jf1~~tl 2529nlJftn1J11uV-Jl9ltJ

q U q q

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1Vi lJiltl nltjiltj1J11tJvl UJ~llJ l~ih1f lJli1V iTtJtltJtJ(PIVlfVi6 U(1~ ~1Vl6 U95ftlJ 2536 n fllJ

Vl (i) f1 V-Jl U~VUYi Vl1 tltl1Jlf1 J fllvl1(i) ~t~ ~lVtl1i1li li11 (i)vl1 ~ 1 11J m J fll J 11 J ~ lJ -J 46(1 ) 33 -38

tVi hiltl nltjiltJ1J11tJvl ih~liJ lWiiYtli 1l(1~ lJli1V lTtJV tJwlVlfVi6 2536 U nlJtl4uT~r~f1lvl191l-

1tJ U tl fI Vtl f1~ (i)Vli -J li1~ rJlJ~ J ~1iJm1 iJ i1m m tJ~h-J tltl11J 1J fllJ UJ ~ lJ -J 46(2) I 17-186

rI 4Q1 lel Q rI I IJrI QI

hlhiltl ViJi1lJltJtJVl 1lf1 lliV tlVWllliljtlJ 2525 fllJtlUltJVitJlimtJ1il(i)ljfltl-J(1~~ tlf1-J l eu u q qJ cu

91(1 lliCYlinlJlhlJ-J 35(5)513-526 OJ

VWtJvllllJltJumflJ1Jpound 2537 fl1JVi1~I~V-J9I~Vi11JihYilimll1li1JnlJUJ~lJ~ 47(3)253-261s bull

liJ (1 Jj Vl1 J 11V 2536 fllJ 1H llNllJV~1tJ tlli1lium ~ 11J 1J n li UJ~lJ ~ 46(4) 323-330

~ QI 0 QI QI rI S 0 d Q Vi I ~ 1 lJli1V IfjtJ1Jl9IVlHH

a lC1 ~ 1(JDll1 H)-JlJtl~fllm 2531 lil1VlV1fllJ11~ iUlC1~1Vi1~Yn Jl(i)lVl~f1

tJ~l1rufl1~m1lJ illim~~ J1(J~1W1)~lJmI11li1f11JmlJlh~lJ~ 15 11th btln ~ltJm~ lJ fl~ltl n ~n1J flrJfl~lil~ 2543 Ivl11 IUl ~lVlfll Vl V ff1Um1tJl(1 J 1TI fllj j 1Vf1 f llJ tl 91l

207 ml

ilinllJ ~lIln lmufffa] 15~8 fl1 ) n~Clv j11~ VlnLl~I~1i1~lnlUn~tl~hjlJ11jn1jflljUj~lJ~ 38(4)

99-305

2

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~lvhflJ~tll~llWflritJJ 1vJlJ JltJJlWfllJiYlJlJWlilflfllJ1h~~lU 2533 flJlJIh~lJJ 1~1 745-752

q dav d v 0 dCloCV rI ds Q ~ a

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IvllflJ ~ tllW 1VlfJl ~tlll1 n ~UVlflli lJihL J ltlJlWfllJiYlJlJWlilflfllJh~hlJ 2533 flJlJIJnm

jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

d QlQ v Iltvlt

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fl11~J11flJ~tH

(fl~hJ) (tl))

1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

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ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

Ju

d u nmHlIWti1flfl31lmlti1fl111111jJj lf1lj rlfH-tltJ911JtJ~1~t~lfli flltJ

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fllV 1 ltJbull

UEl)ltJ UElfl flrlfl~ hJll

fll1JJ fllUJ[Jll 11111110 f1ll1J fll111l1) lj linin f11111 f11111l11

(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

5611 65 66 5168 62 68 4098 49 55

2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

amp 0 QI QI rv I

YlH3NtA1f)Y1 5 lHI1Wf) (mlJ) mllJf111jm~~Hlj (9flJ) mllJfJl1m~lItJ3 (9flJ) UtJllf)lPl1m~tIlW

tilt 15 illbull

cI

WllIW jJClflClWmlPl miClj 1~lJ -vshy

unnjo fll11Jtl11~ fll1lJUll Ii 11110 f1111J011~ fll1lJVl1 ullnTo fl11lJ 011~ fll1UVll

(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

bull bull

i2

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Page 34: I r1.LQ:::WLWl - Fisheries

2

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q dav d v 0 dCloCV rI ds Q ~ a

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jI -I bull 1Wl 161-767

Nikolsky GV 1963 The Ecology of Fishes Academic Press New York 351 p

fl1fUml0

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1200 400 460

2 1J 41S 44S

3

12000 420 470

4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

76

29

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ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

26 27800 58 0 590

27 23000 550 560

28 2S800 540 560

29 29200 510 510

IQftV 23040plusmn396 5197plusmn279 5403plusmn247

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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Page 35: I r1.LQ:::WLWl - Fisheries

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2 1J 41S 44S

3

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4 12000 40S 460

S 410 46S

6 12200 410 460

7 12400 410 460

8 12400 400 445

9 12400 420 460

10 13000 440 470

II 13000 46S 485

12 13200 420 480

13 13400 42S 475

14 13600 430 490

15 13800 430 475

16 14000 440 490

17 14200 no 47S

18 14400 44S 47S

19 14600 425 490

20 430 480

21 14600 4S0 49S

22 440 SOO

23 15000 440 490

24 4S5 490

2S ISOOO 450 500

26 15400 435 490

27 15800 470 520

28 15800 440 490

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29

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ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

17000 460 410

2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

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27 23000 550 560

28 2S800 540 560

29 29200 510 510

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(ohll tl111 (liIL) (tlill) il111 (lill) (tlill) o1H (1111)

(lill) (1111) (lIlll

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3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

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5583 69 66 5168 68 66 4682 67 64

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3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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Page 36: I r1.LQ:::WLWl - Fisheries

29

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ftlvlUl1 1-1 linin fll)lJll )Hm~~t)~ mllJVl1m~~(H

(flflJ) (lflJ ) (lflJ )

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2 18400 470 520

3 IX400 470 520

4middot bull 19000 510 510

5 19000 54 0 54 0

6 21600 510 54 5

7 21600 411 0 510

8 21800 500 54 0

9 22000 50 5 560

10 22000 520 550

11 22000 490 520

12 22000 540 570

13 22400 520 550

14 22600 53 0 560

15 22800 510 540

16 nXO() 53 0 54 0

17 22 800 520 54 0

18 21600 550 54 0

19 23600 550 540

20 23800 530 55 0

21 21000 540 565

22 24500 505 505

23 24600 53 5 570

24 2middot1800 530 540

25 27200 550 560

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27 23000 550 560

28 2S800 540 560

29 29200 510 510

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(lill) (1111) (lIlll

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2 4595 52 58 5660 68 62 4175 47 61

3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

7 51lt) 61 64 4270 52 61 3912 41 54

8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

II 51 12 56 62 4420 51 60 4439 49 59

12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

20 4458 58 64

5028 553 616 4756 569 625 4351 529 581

plusmn447 plusmnO65 plusmnO46 plusmn427 plusmnO52 plusmnO38 plusmn479 plusmn082 plusmnO40

bull bull

JI

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tilt 15 illbull

cI

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(flilJ) (lU) ( ~lJ) (flfU) (lIJ) (~lJ) (ofu) (llJ ) (JilJ )

5583 69 66 5168 68 66 4682 67 64

2 54 67 66 66 566 73 68 4857 65 64

3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

7 5354 68 65 5661 68 66 5137 68 65

8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

II 5242 66 64 5085 51 53 5039 64 62

12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

III 4960 62 58 4713 67 64

19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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Page 37: I r1.LQ:::WLWl - Fisheries

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3 4563 52 56 5213 57 67 4029 49 53

4 5298 57 63 4826 54 66 3992 47 54

5 4308 45 57 51 12 62 68 4488 49 56

6 4948 48 60 4776 48 64 4158 45 55

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8 5624 58 69 4510 56 59 4445 48 56

9 5606 65 68 5009 63 63 4046 54 55

10 4815 54 64 4571 52 60 4098 46 55

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12 4970 50 58 5109 61 63 4402 48 58

13 5343 63 63 4916 55 63 4311 50 59

14 4402 48 54 4332 52 58 4900 61 62

15 5181 59 57 4397 62 60

16 3820 51 55 4726 64 63

17 4375 55 61 4274 63 60

18 476 62 62 6013 71 69

19 4632 60 69 3955 62 59

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tilt 15 illbull

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3 5157 66 62 521) 67 67 5069 63 64

4 5281 65 65 5168 66 64 4551 64 62

5 5678 70 68 5441 67 65 5 56 67 64

6 5172 65 64 5111 65 64 4735 64 64

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8 5350 66 64 5282 64 62 4982 64 62

9 5808 72 72 5009 64 63 4568 56 55

10 5008 64 64 5371 72 68 4672 62 60

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12 5126 58 60 5109 61 58 4502 60 58

13 5573 67 65 5216 70 67 45 11 62 61

14 5008 59 60 5010 59 58 4520 61 60

15 5181 65 62 4797 67 64

16 49 20 64 6 1 4523 65 64

17 5175 65 63 4641 63 64

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19 5332 70 68 45 80 67 66

20 525ll 66 64

1l~O 5343plusmn 65llplusmn 646plusmn 5217plusmn 654plusmn 633plusmn 4749plusmn 640plusmn 6 24plusmn

249 038 031 202 049 041 250 0 30 027

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Page 38: I r1.LQ:::WLWl - Fisheries

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8 5350 66 64 5282 64 62 4982 64 62

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20 525ll 66 64

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i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

8318658

15250623

50

52

1663732

5129714

Y1liljfll-t1f)~7 ilml~MWJl1Jmh UJ 1W DYlJ lfllJll~tlJl~tJ 1~~lrn~

m1~~mllJmhjl U

j~11l1~fl~lJ

SS

12595

df

2

MS

6297

F

244NS

fllUlt1fl~lJ 11n1lJlI

129143

14173S

50

52

25 83

mjHfIl-t1f)~ 811ml~ljfl111JUU)Un1Dmlfllj l)~rulfitJ lYl-cYmiYlTI u

SS df MS Furtri ~fll11JUlh j lU

j~rtl1~fl~lJ 8004 2 4002 263 NS

illtJltlfl~lJ l1~rtlJII

76102

84106

50

52

15 22

~ j

  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน
Page 39: I r1.LQ:::WLWl - Fisheries

bull bull

i2

VIljl Unnf)~ 6 ilfl)l~Mm1JUU)U)lW e1Yl) lfllH~5tlJl~tJ lYl

11 11( ~ m llJl1lh ntl SS df MS F

j~1dm)~lJ 6931965 2 3465982 20S NS

illtJlt1fl~lJ l1~Y1lJlI

8318658

15250623

50

52

1663732

5129714

Y1liljfll-t1f)~7 ilml~MWJl1Jmh UJ 1W DYlJ lfllJll~tlJl~tJ 1~~lrn~

m1~~mllJmhjl U

j~11l1~fl~lJ

SS

12595

df

2

MS

6297

F

244NS

fllUlt1fl~lJ 11n1lJlI

129143

14173S

50

52

25 83

mjHfIl-t1f)~ 811ml~ljfl111JUU)Un1Dmlfllj l)~rulfitJ lYl-cYmiYlTI u

SS df MS Furtri ~fll11JUlh j lU

j~rtl1~fl~lJ 8004 2 4002 263 NS

illtJltlfl~lJ l1~rtlJII

76102

84106

50

52

15 22

~ j

  • พฤตกรรมการวางไขและเพาะฟกไขเตากระอาน