FOSSIL PLANTS Updated

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    FOSSIL PLANTS

    RESEARCH AND PRESENTED BY:JOSHITA SAINI

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    CONTENTSS.No Title Slide No.

    1. What Are Fossils 1

    . T!"es O# Fossils $

    %. &eolo'i(al Ti)e S(ale *

    $. What Are Pterido"h!tes+ ,

    *. Fossil Pterido"h!tes -

    ,. What Are &!)os"er)s+ 1

    -. Fossil &!)os"er)s

    /. Re#ere(es

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    WHAT ARE FOSSILS?

    •  Fossils  derived from Latin word fossus, means literally“having been dug up”.

    • Are the preserved remains or traces of animals, plants, andother organisms from the remote past.

    • Totality of fossils, both discovered and undiscovered, andtheir placement in f ossiliferous (fossil-containing roc!formations and sedimentary layers (strata is !nown as thefossil record.

     Palaeobotany is the study of plant fossils preserved inroc!s.

    • The word “ Fossil ” has been defined as “any evidence of prehistoric life”.

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     TYPES OF FOSSILS

    • Petrifaction " #ormed because of the slow infiltration of tissues by someminerals li!e calcium, silica or magnesium.

    •$rganic matter is replaced by mineral matter. Almost all the details ofeven cell walls are visible under the microscope.

    •Compressions or mummification " %arts of the organism are flattened bythe vertical pressure of the overlying roc!s. $utlines of epidermal cells,hair, stomata, etc can be studied by these fossils.

    • &o internal cellular details are preserved.

    • Impressions " 'aterial of the plant disorganies but the impressionremains. #ormed because of the burial of the parts of the plant in the soil,later on harden into roc!. $nly e)ternal features of the plant are preserved.

    •Casts or molds " formed when an organism, submerged in watercontaining lime, remains covered by a crust of mineral matter. *ecay ofsuch incrusted organisms leaves a cast or mold.

    •+asts e)hibit nothing of the original tissues of the organisms.

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    GEOLOGICAL TIME SCALE

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    WHAT ARE PTERIDOPHYTES?

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    FOSSIL PTERIDOPHYTES

    •  Division Psilophytopsida (e.g. hynia,

    orneophyton, osterophyllum

    •  Lepidodendrales of Lycopsida

    •  Hyeniales (e.g. yenia, Sphenophyllales (e.g.

    /phenophyllum of /phenopsida

    • Cladoxylales of Pteropsida

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    RHYNIA

    • *iscovered from the middle devonian roc!s of Rhynie in Scotland (about 012 million

    years ago• epresented by two species " R ma!or " R g#ynne$vaughani.

    • %lant body sporophytic, sporophytes of . ma3or were larger than R g#ynne$

    vaughani

    • %lants had dichotomously branched rhiome. #rom the rhiome, developed many

    dichotomously branched erect aerial shoots towards the upper side and many rhioidstowards the lower side.

    • oots were absent.

    • /hoots were photosynthetic in nature and had stomata all over.

    •  Reproductive Structures " sporangia present at the tips of some aerial shoots.

    • $val to cylindrical in shape with distal pointed end and a broad basal end.

    • Sporangia of R ma!or were larger than that of . gwynne-vaughani.

    • Large no. of spores present inside, homosporous, arranged in tetrads.

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    ANATOMY OF AERIAL SHOOT

    • *ifferentiated into epidermis, corte) and stele.

    • $utermost layer- epidermis covered by heavy cuticle. The continuity of epidermis was bro!en by many

    stomata.

    • 4nside the epidermis was a well-developed region of corte). 4t was differentiated into inner and outer corte).

    • The outer corte) or hypodermis was 5-6 cells thic! and without any intercellular spaces. The inner corte) was

    made up of many spherical cells having large intercellular spaces. 4nner corte) had a direct connection with

    the stomata through these intercellular spaces.

    • 7ndodermis and pericycle layer absent. /tele was simple protostele.

      T.S aerial shoot Rh!ia

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    HORNEOPHYTON

    •  Horneophyton lignieri, similar to hynia.

    • /maller, more slender, more freely branched stems.

    • Aerial parts were without any appendages.

    • 8nderground portions had short-lobed tuberous corn-li!e structures.

    /tele was protostelic.• %resence of large no. non-septate fungal hyphae in many tubers

    • /porangia were terminal, sessile, present on the tips of stem 9 no

    uniformity in shape.

    • /porangia were surrounded by thic! wall and had a sterile columella.

    • 4n bifid sporangium, colimella was also bifid.

    • /pores were present in tetrads.

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    YARRAVIA

    • %arravia was a generic name given to the reproductive bodies.

    •  belongs to /ilurian period 9 discovered in Australia.

    •  &a!ed a)es terminated into large synangia.

    • 7ach synangium consisted of many elongated sporangia

    •  &o spores were identified within sporangia.

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    COOKSONIA

    • :elongs to late /ilurian age 9 discovered in ;reat :ritain.

    • /ome wor!ers give the generic name +oo!sonia to the

    reproductive bodies of

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    ZOSTEROPHYLLUM

    • =ell-!nown vascular plants from Silurian age.

    • epresented by only 0 species- myretonianum& ' australianum& " '

    rhenanum

    • /pecies described from (ld Red Sandstone of Scotland .

    • %lant body consisted of a dichotomously branched vegetative system

     bearing many fertile branches.

    • #ertile branches terminated into sporangia-bearing spi!e li!e structures.

    • . rhenanum was described from the Lower *evonian in ;ermany.

    %artially submerged species of  'osterophyllum.

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    LEPIDODENDRALES

    • 7)tinct order of Lycopsida.

    • %lants originated during the 8pper *evonian period and

    e)tinct by the end of the %ermian period.

    • %lants were big trees e.g. Lepidodendron

    Lepidodendron- highly branched genus, branching wasdichotomous and ultimate branches bore small leaves.

    • $n the lower side, the trun! was anchored by four main

    horiontal radiating arm li!e a)es. /everal spirally

    arranged lateral appendages were present on thesubdivisions of these branches. These are called

    stigmarian appendages or stigmarian roots.

    •  Lepidodendrales were possibly heterosporous plants.

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    HYENIA ELEGANS

    • *iscovered from the )iddle Devonian roc*s.• ad stout horiontal rhiome bearing sterile and fertile erect branches

    • /terile branches bore for!ed appendages on their nodes, which functioned

    as leaves.

    • #ertile branches bore whorls of sporangiophores

    • Two segments of each fertile whorl were refle)ed and each of them bore

    two sporangia.

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    SPHENOPHYLLUM

    Appeared in 8pper *evonian and became e)tinct in the Triassic period.• +entral a)is of the aerial shoots was slender, irregularly branched.

    • /tems were delicate and divisible into nodes and internodes.

    • Leaves present on the nodes, arranged in whorls, in multiples of 0.

    • >-51 leaves were present at each node.

    • 7ntire as well as dichotomously divided leaves were also present on thesame shoot in some species, showing heterophyllous nature.

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    INTERNAL ORGANIZATION OF STEM

    •/tem had triangular mass of primary )ylem in the centre.

    •/tele was actinostelic and the exarch protoxylem was present at

    the angles of the actinostele.

    •+orte) consisted of peripheral region of thic! walled cells and

    an inner region of thin walled cells.

    •/econdary growth by means of cambium was seen in the stems.

    •$utside the primary )ylem developed a ring of cambium.

    •%itted trachieds and )ylem parenchyma were present in the

    secondary wood.

    •%eriderm developed in the inner portion of the corte) after the

    start of cambial activity.

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    CALAMITALES

    •'embers of this fossil order appeared in +pper Devonian and

     became e)tinct in early ,riassic period .

    • %lant body was sporophytic& sporophytes were very large and

    tree-li!e.

    •/tems and branches showed considerable secondary growth.

    • =horls of sporangiophore, usually alternating with the sterile

     bracts, were present in strobili

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    CLADOXYLON

    • %lants from 'iddle *evonian

    • Thic! stem, branched irregularly.

    • /ome of the branches had fan shaped, deeply divided leaves showing a

    series of dichotomies.

    /porangium present at the tip of each dichotomously diving branch of sonfan shaped leaves.

    • ighly comp

    • le) type of polystelic vascular system.

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    COENOPTERIDALES

    •$ccurred in the roc!s of 8pper *evonian, carboniferous and Permian era

    • #ossil group of fascinating fern-li!e plants.

    • /howed wide range of growth habits, some had creeping stems

    (:otryopteris cylindrica, had erct trun!s (ygopteris primaria.• Lac! of clear cut differentiation between the leaves and stem in most

    of the members.

    • %lants had large frond li!e spirally arranged appendages. #ronds

    were 0* in their branching pattern.

    • ?ascular system of stem was usually protostelic.

    • /porangia were generally large, oval, pedicellate and attached

    terminally on the fertile branches.

    • All members were homosporous, e)cept Stauropteris burntislandica

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    WHAT ARE GYMNOSPERMS?

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    Fossil '!)os"er)s

    •  Ar(haeo"teris

    • 0!'io"teris oldha)ia

    &losso"teris• Willia)soia

    • Peto!lo

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    ARCHAEOPTERIS

    • eported from the roc!s of the upper *evonian period of &orth America, ussia.

    • ad large fern li!e fronds.

    • #erns were bipinnately branched had all the pinnae developed in one plane

    • 7ach pinnule had a single vascular bundle, which soon became dichotomously

     branched .

    • The fertile fronds had sporangia

    • The sporangia replaced the pinnules of some of the lower pinnae in the fertile fronds.• 'ost of the species appeared to be homosporous.

    •  - latifolia was heterosporous and possessed both microspores and mega spores on

    the same plant.

    • 4t possessed two !inds of sporangia .

    • )egasporangia possessed about 5> spores and microsporangia about 522 or morespores.

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     Lyginopteris Oldhamia

    • %lants are vine-li!e with large fronds.• The stem was erect, branched, @mmto6mm in diameter.

    • The leaves were bi-or tripinnately compound and arranged

    spirally on the stem.

    • The pinnae were borne at right angles to rachis and were

    arranged opposite to each other.

    • The pinnae bore pinnules.

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    T.S STEM

     Lyginopteris Oldhamia

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     T.S OF STEM

    •  &e)t to 7pidermis is the outer corte) which consists of

    radially broadened fibrous strands that form a vertical

    networ!.

    • The inner corte) consists of ordinary parenchymatous

    cells. &e)t to the corte) is the pericycle which consistsof sclerotic cells called sclerotic nests.

    •  &e)t to the pericycle are five strands of primary

    vascular bundles. These are separated by

     parenchymatous areas. 7ach vascular bundle is mesarchand consists of primary phloem towards the outer side.

    • 4n the centre there is a large pith made up of

     parenchymatous cells.

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     MALE FRUCTIFICATION:

    CROSSOTHECA 

    • is 'eerall! re'arded as the )ale #r2(ti#i(atio

    • I Crossotheca, a 2)3er o# 3oot sha"ed 3ilo(2lar )i(ros"ora'ia

    4ere arra'ed radiall! o the 2derside o# dis(s.

    • 5i(ros"ora'ia 4ere 3ore as elo'ate6 "edat a""eda'es o

    the 2lti)ate "i2le o# the #rod. The #ertile #rods rese)3le )i2tehair3r2shes.

    • The s"ora'ia la(7 a2l2s ad rese)3les those o# C!(as. O

    dehis(e(e6 the )i(ros"ores 4ere (arried 3! 4id. The )i(ros"ores

    are s"heri(al 4ith the tri8radiate )ar7i's 9trilete.

     

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    L.S OVULE OF THE

    LYGINOPTERIS OLDHAMIA

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    FEMALE FRUCTIFICATION

     ( LAGENOSTOMA OLDHAMIA)

    • dis(o;ered #or)