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8/16/2019 FOSSIL PLANTS Updated
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FOSSIL PLANTS
RESEARCH AND PRESENTED BY:JOSHITA SAINI
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CONTENTSS.No Title Slide No.
1. What Are Fossils 1
. T!"es O# Fossils $
%. &eolo'i(al Ti)e S(ale *
$. What Are Pterido"h!tes+ ,
*. Fossil Pterido"h!tes -
,. What Are &!)os"er)s+ 1
-. Fossil &!)os"er)s
/. Re#ere(es
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WHAT ARE FOSSILS?
• Fossils derived from Latin word fossus, means literally“having been dug up”.
• Are the preserved remains or traces of animals, plants, andother organisms from the remote past.
• Totality of fossils, both discovered and undiscovered, andtheir placement in f ossiliferous (fossil-containing roc!formations and sedimentary layers (strata is !nown as thefossil record.
•
Palaeobotany is the study of plant fossils preserved inroc!s.
• The word “ Fossil ” has been defined as “any evidence of prehistoric life”.
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TYPES OF FOSSILS
• Petrifaction " #ormed because of the slow infiltration of tissues by someminerals li!e calcium, silica or magnesium.
•$rganic matter is replaced by mineral matter. Almost all the details ofeven cell walls are visible under the microscope.
•Compressions or mummification " %arts of the organism are flattened bythe vertical pressure of the overlying roc!s. $utlines of epidermal cells,hair, stomata, etc can be studied by these fossils.
• &o internal cellular details are preserved.
• Impressions " 'aterial of the plant disorganies but the impressionremains. #ormed because of the burial of the parts of the plant in the soil,later on harden into roc!. $nly e)ternal features of the plant are preserved.
•Casts or molds " formed when an organism, submerged in watercontaining lime, remains covered by a crust of mineral matter. *ecay ofsuch incrusted organisms leaves a cast or mold.
•+asts e)hibit nothing of the original tissues of the organisms.
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GEOLOGICAL TIME SCALE
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WHAT ARE PTERIDOPHYTES?
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FOSSIL PTERIDOPHYTES
• Division Psilophytopsida (e.g. hynia,
orneophyton, osterophyllum
• Lepidodendrales of Lycopsida
• Hyeniales (e.g. yenia, Sphenophyllales (e.g.
/phenophyllum of /phenopsida
• Cladoxylales of Pteropsida
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RHYNIA
• *iscovered from the middle devonian roc!s of Rhynie in Scotland (about 012 million
years ago• epresented by two species " R ma!or " R g#ynne$vaughani.
• %lant body sporophytic, sporophytes of . ma3or were larger than R g#ynne$
vaughani
• %lants had dichotomously branched rhiome. #rom the rhiome, developed many
dichotomously branched erect aerial shoots towards the upper side and many rhioidstowards the lower side.
• oots were absent.
• /hoots were photosynthetic in nature and had stomata all over.
• Reproductive Structures " sporangia present at the tips of some aerial shoots.
• $val to cylindrical in shape with distal pointed end and a broad basal end.
• Sporangia of R ma!or were larger than that of . gwynne-vaughani.
• Large no. of spores present inside, homosporous, arranged in tetrads.
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ANATOMY OF AERIAL SHOOT
• *ifferentiated into epidermis, corte) and stele.
• $utermost layer- epidermis covered by heavy cuticle. The continuity of epidermis was bro!en by many
stomata.
• 4nside the epidermis was a well-developed region of corte). 4t was differentiated into inner and outer corte).
• The outer corte) or hypodermis was 5-6 cells thic! and without any intercellular spaces. The inner corte) was
made up of many spherical cells having large intercellular spaces. 4nner corte) had a direct connection with
the stomata through these intercellular spaces.
• 7ndodermis and pericycle layer absent. /tele was simple protostele.
T.S aerial shoot Rh!ia
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HORNEOPHYTON
• Horneophyton lignieri, similar to hynia.
• /maller, more slender, more freely branched stems.
• Aerial parts were without any appendages.
• 8nderground portions had short-lobed tuberous corn-li!e structures.
•
/tele was protostelic.• %resence of large no. non-septate fungal hyphae in many tubers
• /porangia were terminal, sessile, present on the tips of stem 9 no
uniformity in shape.
• /porangia were surrounded by thic! wall and had a sterile columella.
• 4n bifid sporangium, colimella was also bifid.
• /pores were present in tetrads.
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YARRAVIA
• %arravia was a generic name given to the reproductive bodies.
• belongs to /ilurian period 9 discovered in Australia.
• &a!ed a)es terminated into large synangia.
• 7ach synangium consisted of many elongated sporangia
• &o spores were identified within sporangia.
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COOKSONIA
• :elongs to late /ilurian age 9 discovered in ;reat :ritain.
• /ome wor!ers give the generic name +oo!sonia to the
reproductive bodies of
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ZOSTEROPHYLLUM
• =ell-!nown vascular plants from Silurian age.
• epresented by only 0 species- myretonianum& ' australianum& " '
rhenanum
• /pecies described from (ld Red Sandstone of Scotland .
• %lant body consisted of a dichotomously branched vegetative system
bearing many fertile branches.
• #ertile branches terminated into sporangia-bearing spi!e li!e structures.
• . rhenanum was described from the Lower *evonian in ;ermany.
%artially submerged species of 'osterophyllum.
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LEPIDODENDRALES
• 7)tinct order of Lycopsida.
• %lants originated during the 8pper *evonian period and
e)tinct by the end of the %ermian period.
• %lants were big trees e.g. Lepidodendron
•
Lepidodendron- highly branched genus, branching wasdichotomous and ultimate branches bore small leaves.
• $n the lower side, the trun! was anchored by four main
horiontal radiating arm li!e a)es. /everal spirally
arranged lateral appendages were present on thesubdivisions of these branches. These are called
stigmarian appendages or stigmarian roots.
• Lepidodendrales were possibly heterosporous plants.
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HYENIA ELEGANS
• *iscovered from the )iddle Devonian roc*s.• ad stout horiontal rhiome bearing sterile and fertile erect branches
• /terile branches bore for!ed appendages on their nodes, which functioned
as leaves.
• #ertile branches bore whorls of sporangiophores
• Two segments of each fertile whorl were refle)ed and each of them bore
two sporangia.
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SPHENOPHYLLUM
•
Appeared in 8pper *evonian and became e)tinct in the Triassic period.• +entral a)is of the aerial shoots was slender, irregularly branched.
• /tems were delicate and divisible into nodes and internodes.
• Leaves present on the nodes, arranged in whorls, in multiples of 0.
• >-51 leaves were present at each node.
• 7ntire as well as dichotomously divided leaves were also present on thesame shoot in some species, showing heterophyllous nature.
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INTERNAL ORGANIZATION OF STEM
•/tem had triangular mass of primary )ylem in the centre.
•/tele was actinostelic and the exarch protoxylem was present at
the angles of the actinostele.
•+orte) consisted of peripheral region of thic! walled cells and
an inner region of thin walled cells.
•/econdary growth by means of cambium was seen in the stems.
•$utside the primary )ylem developed a ring of cambium.
•%itted trachieds and )ylem parenchyma were present in the
secondary wood.
•%eriderm developed in the inner portion of the corte) after the
start of cambial activity.
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CALAMITALES
•'embers of this fossil order appeared in +pper Devonian and
became e)tinct in early ,riassic period .
• %lant body was sporophytic& sporophytes were very large and
tree-li!e.
•/tems and branches showed considerable secondary growth.
• =horls of sporangiophore, usually alternating with the sterile
bracts, were present in strobili
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CLADOXYLON
• %lants from 'iddle *evonian
• Thic! stem, branched irregularly.
• /ome of the branches had fan shaped, deeply divided leaves showing a
series of dichotomies.
•
/porangium present at the tip of each dichotomously diving branch of sonfan shaped leaves.
• ighly comp
• le) type of polystelic vascular system.
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COENOPTERIDALES
•$ccurred in the roc!s of 8pper *evonian, carboniferous and Permian era
• #ossil group of fascinating fern-li!e plants.
• /howed wide range of growth habits, some had creeping stems
(:otryopteris cylindrica, had erct trun!s (ygopteris primaria.• Lac! of clear cut differentiation between the leaves and stem in most
of the members.
• %lants had large frond li!e spirally arranged appendages. #ronds
were 0* in their branching pattern.
• ?ascular system of stem was usually protostelic.
• /porangia were generally large, oval, pedicellate and attached
terminally on the fertile branches.
• All members were homosporous, e)cept Stauropteris burntislandica
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WHAT ARE GYMNOSPERMS?
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Fossil '!)os"er)s
• Ar(haeo"teris
• 0!'io"teris oldha)ia
•
&losso"teris• Willia)soia
• Peto!lo
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ARCHAEOPTERIS
• eported from the roc!s of the upper *evonian period of &orth America, ussia.
• ad large fern li!e fronds.
• #erns were bipinnately branched had all the pinnae developed in one plane
• 7ach pinnule had a single vascular bundle, which soon became dichotomously
branched .
• The fertile fronds had sporangia
• The sporangia replaced the pinnules of some of the lower pinnae in the fertile fronds.• 'ost of the species appeared to be homosporous.
• - latifolia was heterosporous and possessed both microspores and mega spores on
the same plant.
• 4t possessed two !inds of sporangia .
• )egasporangia possessed about 5> spores and microsporangia about 522 or morespores.
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Lyginopteris Oldhamia
• %lants are vine-li!e with large fronds.• The stem was erect, branched, @mmto6mm in diameter.
• The leaves were bi-or tripinnately compound and arranged
spirally on the stem.
• The pinnae were borne at right angles to rachis and were
arranged opposite to each other.
• The pinnae bore pinnules.
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T.S STEM
Lyginopteris Oldhamia
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T.S OF STEM
• &e)t to 7pidermis is the outer corte) which consists of
radially broadened fibrous strands that form a vertical
networ!.
• The inner corte) consists of ordinary parenchymatous
cells. &e)t to the corte) is the pericycle which consistsof sclerotic cells called sclerotic nests.
• &e)t to the pericycle are five strands of primary
vascular bundles. These are separated by
parenchymatous areas. 7ach vascular bundle is mesarchand consists of primary phloem towards the outer side.
• 4n the centre there is a large pith made up of
parenchymatous cells.
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MALE FRUCTIFICATION:
CROSSOTHECA
• is 'eerall! re'arded as the )ale #r2(ti#i(atio
• I Crossotheca, a 2)3er o# 3oot sha"ed 3ilo(2lar )i(ros"ora'ia
4ere arra'ed radiall! o the 2derside o# dis(s.
• 5i(ros"ora'ia 4ere 3ore as elo'ate6 "edat a""eda'es o
the 2lti)ate "i2le o# the #rod. The #ertile #rods rese)3le )i2tehair3r2shes.
• The s"ora'ia la(7 a2l2s ad rese)3les those o# C!(as. O
dehis(e(e6 the )i(ros"ores 4ere (arried 3! 4id. The )i(ros"ores
are s"heri(al 4ith the tri8radiate )ar7i's 9trilete.
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L.S OVULE OF THE
LYGINOPTERIS OLDHAMIA
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FEMALE FRUCTIFICATION
( LAGENOSTOMA OLDHAMIA)
• dis(o;ered #or)