Fifth International Conference on Serpentine Ecology · Fifth International Conference on Serpentine Ecology ... Scott Angle, Lan Zhang, Rufus ... Fifth International Conference on

Fifth International Conference on Serpentine Ecology

University of Siena

Department of Environmental Science “G. Sarfatti”

Accademia dei Fisiocritici

Siena, Italy May 9-13, 2006 Third Circular


http://www.unisi.it/eventi/5-ICSE/index.htm

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Overview Ultramafic rocks (known as serpentine in the botanical and ecological literature), and the soils derived from them, occur throughout the globe. They form specialized habitats that host unique species and communities. In 1991, scientists studying ultramafic sites first gathered together on an international scale to share their research findings on the campus of the University of California, Davis, USA. The success of that first symposium, along with its attendant field trip to local ultramafic sites, inspired the several international conferences that have been held since then. These were situated at widely separated sites with nearby ultramafic locales: at ORSTOM headquarters in Nouméa, New Caledonia (1995), at Kruger National Park in South Africa (1999) and at the National Botanical Garden in Havana, Cuba in 2003. The 2006 conference will be held in Europe for the first time, at the University of Siena, Italy.

Organizing Committee Prof. Alessandro Chiarucci [email protected] Prof. Roberto Gabbrielli [email protected] Dr. Cristina Gonnelli [email protected] Prof. Mauro Mariotti [email protected] Dr. Andrea Saccani [email protected] Dr. Giulia Vercesi [email protected]

Scientific Committee Prof. Alan J.M. Baker [email protected] Prof. Rosalina Berazaín Iturralde [email protected] Prof. Robert Boyd [email protected] Prof. Alessandro Chiarucci [email protected] Prof. Roberto Gabbrielli [email protected] Prof. John Proctor [email protected] Prof. Nishanta Rajakaruna [email protected] Prof. Roger Reeves [email protected] Organizing secretariat

Scientific secretariat

Università di Siena Servizio Congressi Banchi di Sotto 46, 53100 Siena, Italy Tel +39-0577-232132 e-mail: [email protected]

Dr. Cristina Gonnelli Dipartimento di Biologia Vegetale Università di Firenze Via La Pira 4, 50121 Firenze, Italy Fax: +39 055 282358 e-mail: [email protected]



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Preliminary Program (definitive program will be available on the conference website by April 28th)

Lectures will be 20 minuts (15 min talk and 5 min discussion). Slide projector, overhead projector and computer facilities for powerpoint presentations will be available

9 May

18.00: Welcome Party

10 May

8.30 - 9.45: Registration

9.45 – 10.15: Opening Ceremony

10.15 - 10.45: Opening Lecture – Enrico Pandeli

10.45 – 11.15: Coffee Break

11.15 - 13.00: Session “Soils and microbiology” Soil and Vegetation Differences between Peridotite and Serpentinite Earl B. Alexander

Availability of Ni along an ultramafic toposequence in the Vosges Mountains (France) Guillaume Echevarria, Vanessa Chardot, Michel Gury, Stamatia Massoura, Hélène Molins, Jean Louis Morel

Chromium in Ultramafic soils from Niquelândia, Brazil Jeremie Garnier,Cecile Quantin , Eder de Souza Martins , Thierry Becquer

Water availability and soil chemistry as predictors of plant biomass and productivity in a subhumid tropical forest in southwestern Puerto Rico Jamie Horvath, Arthur Johnson

Degradation of high Ni biomass in soil Scott Angle, Lan Zhang, Rufus Chaney

13.00 – 14.30: Lunch

14.30 – 16.30: Session “Soils and microbiology” Dissolution of Chrysotile in the rhizosphere of the Ni hyperaccumulator Leptoplax emarginata Vanessa Chardot, Guillaume Echevarria, Cristophe Calvaruso, Marie-Pierre Turpault, Bruno Simon, Jean-Louis Morel

Nickel tolerance and accumulation by bacteria from rhizosphere of nickel hyperaccumulators in serpentine ecosystem of Andaman, India Arundhati Pal, A. K. Paul

The Bacterial Flora of the Nickel-hyperaccumulator Plant Alyssum bertolonii Desv. Rita Barzanti, Alessio Mengoni, Francesca Ozino, Francesca Galardi, Chiara Mastretta, Cristina Gonnelli, Roberto Gabbrielli, Marco Bazzicalupo

Diversity of Cenococcum geophilum isolates from serpentine soils of NE Portugal assessed by AFLP and Ni in vitro tolerance Antonio Portugal, Susana Claudia Gonçalves, Maria Teresa Gonçalves, Rita Vieira, Maria Amélia Martins-Loução, Helena Freitas



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Nickel-induced protein pattern modifications in a serpentine tolerant isolate of the ectomycorrhizal fungus Cenococcum geophilum Susana C. Gonçalves, Elisa Bona, M. Amélia Martins-Loução, Helena Freitas, Graziella Berta

Relationships between the mycorrhizal status and Ni-hyperaccumulation by 8 endemic plant species from New Caledonian serpentine environments Amir Hamid, Perrier Nicolas, Collion Anne, Cigna Jerimy, Rigault Fréderic, Jaffré Tanguy


17.00 – 18.30: Poster Presentation - Session 1

11 May

9.00 – 10.30: Session “Conservation and management of serpentine areas” Bear Valley in the northern California Coast Ranges: Wildflowers, grazing, and conservation Jack Alderson, Ivana Noell

Lichens on asbestos-rich serpentinites: colonization, weathering and inactivation of chrysotile fibres Sergio Enrico Favero-Longo, Rosanna Piervittori

Conserving Soldier's Delight, the Largest Serpentine Area in Maryland, USA Marla McIntosh, Daniel Hochman

Local adaptation and gene flow across serpentine and non-serpentine habitats in two contrasting plant species Jessica Wright


11.15 – 13.00: Session “Physiological ecology of serpentine plants” Development of Molecular and Genetic Tools to Study the Evolution of Serpentine Adaptation, Endemism and Speciation in the Streptanthoid Complex, Brassicaceae Millie Burrell, Katherine Taylor, Robert Cantrell, Umesh Reddy, Alan Pepper

Differences in ion accumulation between Mimulus guttatus (Phrymaceae) populations differentially adapted to serpentine and non-serpentine soils Kristy U. Brady, H.D. Bradshaw, Jr., David E. Salt

Physiological adaptation of an Iranian serpentine endemic plant species, Cleome heratensis (Capparaceae) Tahmaseb Asemaneh, S. Majid Ghaderian, Alan J. M. Baker

Hyperaccumulation of nickel by two Alyssum species grown on Iranian ultramafics S. Majid Ghaderian, Ahmad Mohtadi, A.J.M. Baker

Test of a trade-off between elemental and organic defences against herbivores in natural populations of Thlaspi caerulescens Nausicaa Noret, Pierre Meerts, José Escarré

13.00 – 14.30: Lunch

14.30 – 16.30: Session “Physiological ecology of serpentine plants” Nickel storage, processing, and release in the hyperaccumulator Stackhousia tryonii: Preliminary observations from field, lab, and microscopy Ivana Noell



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Roots of Ni-hyperaccumulating and non-accumulating genotypes of Senecio coronatus differ in aspects of their cytology and elemental distribution Jolanta Mesjasz-Przybylowicz, Alban Barnabas, Wojciech Przybylowicz

Localisation & quantification of nickel in leaf and stem tissues of Hybanthus floribundus var. floribundus using micro-PIXE. Anthony Kachenko, Balwant Singh, Naveen Bhatia, Rainer Siegele

Nickel speciation in hyperaccumulating plants of New Caledonia Nicolas Perrier, Jean Paul Ambrosi, Jérome Rose, Tanguy Jaffré, Fabrice Colin

Phytoextraction of Ni by native populations of A. murale on an ultramafic site in Albania Aida Bani, Guillaume Echevarria, Sulejman Sulçe, Jean Louis Morel

Testing the anti-herbivory hypothesis of Ni hyperaccumulation in Alyssum pintodasilvae on Tribolium castaneum and Porcellio dilatatus Sandra Silva, António Portugal, Susana C. Gonçalves, José P. Sousa, M. Teresa Gonçalves


17.00 – 18.30: Poster session – Session 2

12 May

Mid-Conference field trip

08.00: Departure from Siena to the Cecina Valley area. 10.30 - 16.30: Guided visit to the “Monterufoli Nature Reserve,” including a lunch-buffet (possibly in the field). 18.30: Return to Siena.

13 May 9.00 – 10.30: Session “Biodiversity: biogeographical and ecological issues”

Biology of ultramafic rocks and soils: current status and research goals for the future Arthur Kruckeberg, Robert Boyd

Geomorphology and floristic biodiversity of two ophiolitic outcrops (Pietra Parcellara and Monte Mangiapane, Northern Appenines). G.V. Vercesi, G. Santamaria, M. Perracino

Soils and Vegetation in Mont Avic Natural Park (Valle d'Aosta, Italy) Michele D'Amico

Floristic Diversity on Ultramafic Rocks in New Caledonia Tanguy Jaffré, Nicolas Perrier


11.00 – 12.00: Session “Biodiversity: biogeographical and ecological issues” The Flora and Biogeochemistry of the Ultramafics of Goiás State, Brazil: An Update Roger Reeves, Alan Baker, Thierry Becquer, Guillaume Echevarria



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Taxonomic and geographic patterns in the serpentine endemic flora of California Hugh Safford, Susan Harrison

Plant-Soil Relations of a Serpentine Site in the Southern Coast of Sri Lanka Mohomed Cassim Iqbal, Lakmali Kulasekara, Nishanta Rajakaruna, Sithy Sabeera Iqbal

12.00 – 12.40: Session “Serpentine faunas: interactions with soils and plants” The Defense Hypothesis of Metal Hyperaccumulation: Challenges and New Directions Robert Boyd

Micro-PIXE studies of elemental distribution in sap-feeding insects associated with Ni hyperaccumulator, Berkheya coddii Jolanta Mesjasz-Przybylowicz, Wojciech Przybylowicz, Pawel Migula, Miroslaw Nakonieczny, Maria Augustyniak, Monika Tarnawska, Elzbieta Glowacka

13.00 – 14.30: Lunch

14.30 – 16.30: Session “Population and community ecology” Community and Ecosystem Consequences of Nickel Hyperaccumulation: New Questions, Few Answers Robert Boyd

Understanding Community Assembly on Serpentine: A Study of Functional Traits Relating to Serpentine Tolerance Nishanta Rajakaruna, David D. Ackerly

California ultramafic pine forests and woodlands: diversity and geobotanical approach Daniel Sánchez-Mata, Michael G. Barbour

Dynamics of serpentine vegetation under a Mediterranean climate: a 12-year experiment Alessandro Chiarucci

Effect of increasing Zn concentrations on life history traits in metalliferous and non-metalliferous populations of Thlaspi caerulescens Dechamps Caroline, Lefèbvre Claude, Meerts Pierre

A populational perspective of the Mn-hyperaccumulative trait in the eastern Australian tree, Gossia bidwillii (Myrtaceae) Denise R. Fernando, George N. Batianoff, Ian E. Woodrow, Estelle J Bakkaus, Richard N. Collins, Alan J Baker

Genetic variation in nickel hyperaccumulation in the Alyssum serpyllifolium complex from the Iberian Peninsula Joe Pollard, Christine Donhardt


17.00 – 18.00: Closing Ceremony

20.00: Social dinner

14-17 May

Post conference field trip



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Posters Poster should be mounted during the morning of May 10 and removed during the afternoon of May 13. The maximum poster size allowed is 70 (w) x 100 (h) cm.

List of Posters (arranged by thematic issues)

Conservation and management of serpentine areas Phytoextraction of nickel in Albanian ultramafic soils Aida Bani, Sulejman Sulçe, Guillaume Echevarria, Jean Louis Morel

Threatened plants of Cuban ultramafic areas Rosalina Berazain Iturralde, Luis R. Gonzalez-Torres, Fabiola Areces Berazain, Julio C. Lazcano Lara

Conservation of Melocactus actinacanthus (Cactaceae) Alejandro Palmarola-Bejerano, Luis R. Gonzalez-Torres, Jesus Matos, Alicia Rodriguez, Alberto Torres

Technical Procedure for the Ecological Restoration of the lateritic nickel deposit Koniambo Massif (New Caledonia) Fabien Roumagnac, Nicolas Perrier, Sebastien Sarramegna

Present syntaxonomical knowledge about the lichen and vascular vegetation of ultramafic soils in Tuscany and future conservation perspectives Daniele Viciani, Lorenzo Lastrucci, R. Benesperi, B. Foggi, V. Gonnelli

Soils and microbiology Tolerance of fungi from ultramafic sites in Ireland to increasing nickel concentrations Francis Brearley, Michelle Conboy

Chromium-bearing phases and availability in natural ultramafic soils Garnier Jérémie, Becquer Thierry, Ettler Vojtech, Gasser Ubald, Quantin Cécile

Podzolization over ophiolitic materials in Mont Avic Natural Park (Aosta, Italy) Michele D'Amico, Francesca Julitta, Davide Cantelli

Understanding N and P plant nutrition in lateritic soils of New Caledonia: methodological approach Alexandre Lagrange, Hamid Amir, Jean-Paul Ambrosi, Laurent L'Huillier, Marc-Henry Delrieu

Nuclear microprobe studies of mycothallic gametophytes of a fern from ultramafic area Jolanta Mesjasz-Przybyłowicz, Elżbieta Orłowska, Wojciech Przybyłowicz, Przemysław Ryszka, Katarzyna Turnau (note that the special characters for Polish names don’t show up here or in next abstract)

Influence of arbuscular mycorrhizal colonization on growth and elemental uptake by Ni-hyperaccumulator Berkheya coddii Roessler Elżbieta Orłowska, Jolanta Mesjasz-Przybyłowicz, Wojciech Przybyłowicz, Katarzyna Turnau

Metallophiles from Serpentine Ecosystem of Andaman and their Interaction with Heavy Metals Arundhati Pal, A. K. Paul

Decomposition of Senecio coronatus leaves is affected by leaf Ni concentration Robert Boyd, Micheal Davis, Kevin Balkwill



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Physiological ecology of serpentine plants Trace element composition of ferns and their allies growing on serpentine outcrops in the Ligurian region Laura Cornara, Enrica Roccotiello, Remo Bernardello, Giuliana Drava, Vincenzo Minganti, Mauro Mariotti

Plant biodiversity in the Botanic Alpine Garden of Pietra Corva (Northern Apennines, Italy) Anna Betto

Mineralogic and petrographic characterization of Pietra Corva ophiolitic soils (Northern Apennines, Italy) Anna Betto

Morphometric variability in Thlaspi rotudifolium styles is influenced by Mg-availability in serpentine or non-serpentine substrates Sergio Enrico Favero Longo, Annalisa Ingaramo, Consolata Siniscalco

Intra-specific differences in mineral element composition in populations of the Ni-hyperaccumulator Alyssum bertolonii Desv. Francesca Galardi, Alessio Mengoni, Rita Barzanti, Miluscia Arnetoli, Cristina Gonnelli, Roberto Gabbrielli

Ultrastructural studies on the nickel hyperaccumulator Streptanthus polygaloides Gray (Brassicaceae) Daniel Sánchez-Mata, Vicenta de la Fuente, Ricardo Amils, Lourdes Rufo, Nuria Rodríguez, María del Pilar Rodríguez-Rojo

Micromorphological and ultrastructural comparative characters of three Mediterranean Quercus (Fagaceae) shrubs growing on ultramafic soils Vicenta de la Fuente, Ricardo Amils, Lourdes Rufo, Nuria Rodríguez, Daniel Sánchez-Mata, Blanca Díez-Garretas, Alfredo Asensi

Serpentine faunas: interactions with soils and plants Influence of leaf age on Ni concentrations of some hyperaccumulator and accumulator species from New Caledonia Robert Boyd, Tanguy Jaffré

Plant-herbivore studies of the high-Ni insect Stenoscepa sp. (Orthoptera: Pyrgomorphidae), an herbivore of the South African Ni hyperaccumulator Berkheya coddii (Asteraceae) Robert Boyd, Micheal Davis, Michael Wall, Kevin Balkwill

Host-herbivore studies of Chrysolina clathrata (Coleoptera: Chrysomelidae) from Mpumalanga, South Africa Robert Boyd, Micheal Davis, Michael Wall, Kevin Balkwill

Pesticides as additional stressing factors for Chrysolina pardalina, a feeder of the Ni-hyperaccumulating plant Berkheya coddii Maria Augustyniak, Paweł Migula, Jolanta Mesjasz-Przybyłowicz, Monika Tarnawska, Miroslaw Nakonieczny, Agnieszka Babczynska, Wojciech Przybylowicz, Michal Augustyniak (note that the special characters for Polish names don’t show up here)

Responses of Eisenia fetida (Oligochaeta) on different levels of organization to ultramafic soils R.A. Maleri, A.J. Reinecke, J. Mesjasz-Przybylowicz, S.A. Reinecke, W.J. Przybylowicz

Population and community ecology Elemental patterns among ultramafic populations and plant organs of the South African Ni hyperaccumulator Senecio coronatus



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Robert Boyd, Micheal Davis, Kevin Balkwill

Ectomycorrhizal Fungal Communities from Serpentine and Non- Serpentine Mediterranean Oak Forests Sara Branco

The potential effects of herbivory and metal hyperaccumulation on the fitness of Thlaspi montanum Martha Palomino Tovar

Storage and movement of nickel in ultramafic ecosystems of North Carolina and Puerto Rico Kelly Wilson, Kartikeya Singh, Solimar Marrero, Eva Davila, Joe Pollard

Biodiversity: biogeographical and ecological issues Nickel localization in tissues of hyperaccumulator species of Rubiaceae from ultramafic areas of Cuba Rosalina Berazaín, Vicenta de la Fuente, Lourdes Rufo, Nuria Rodríguez, Daniel Sánchez-Mata, Ricardo Amils

Nickel localization in tissues of hyperaccumulator species of Euphorbiaceae from ultramafic areas of Cuba Rosalina Berazaín, Vicenta de la Fuente, Lourdes Rufo, Nuria Rodríguez, Ricardo Amils, Blanca Díez-Garretas, Daniel Sánchez-Mata, Alfredo Asensi

Regional patterns of Cuban ultramafic plants Luis Roberto González Torres, Rosalina Berazaín Iturralde

The serpentine flora in Bulgaria – distribution, composition and diversity Dolja Pavlova

Plant-Soil Relations on Serpentine Outcrops of Deer Isle, Maine in Northeastern United States Andrew J. Thrall, Nathaniel S. Pope, Eva C. Dannenberg, Kathleen M. Tompkins, Peter G. Pavicevic, Tanner B. Harris, Nishanta Rajakaruna

Saxicolous and Terricolous Lichens of a Peridotite Outcrop in the Northeastern United States: A baseline study exploring the lichen-substrate relationship Tanner B. Harris, Fred C. Olday, Nishanta Rajakaruna

The Ultramafics of the Santa Elena Peninsula, Costa Rica: A Preliminary Biogeochemical Study Roger D. Reeves, Alan J.M. Baker, Rosaura Romero



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Post-Conference Excursion (14-17 May 2006)

In 2001, a network of 11 Italian protected areas was created to protect serpentine habitat (Mont Avic Regional Park, Beigua Regional Park, Capanne di Marcarolo Regional Park, Aveto Regional Park, Monte Prinzera Regional Reserve, Rupe di Campotrera Regional Reserve, Monteferrato Protected Area, Pieve S. Stefano Serpentine Protected Area, Monti Rognosi Nature Reserve, Montenero Provincial Reserve, Monterufoli-Caselli Provincial Reserve). These Protected Areas are located within 5 different Regions (Aosta Valley, Piedmont, Liguria, Emilia-Romagna and Tuscany). A coordination system was created by means of an Agreement Protocol (namely the Coordination of Ophiolitic Protected Areas or “C.A.P.O.”), with the purpose of comparing and exchanging scientific information and management experience, and promoting common actions in the fields of promotion, communication and educational. Because of this relationship between scientific research and environmental conservation in serpentine habitats, the mid-Conference field trip and a post-Conference field trip are organised to visit some of the Protected Areas participating to the C.A.P.O. network. Further information about the C.A.P.O. can be found at www.parks.it/aree.protette.ofiolitiche. 14 May 2006 09.00: Departure from Siena to the Upper Tiber Valley (Province of Arezzo, Tuscany). 10.30-15.00: Guided visit to the “Monti Rognosi Regional Nature Reserve,” including a lunch-

buffet (possibly in the field). 15.00-18.00: Visit to local villages and sites of cultural interest. 18.00-20.00: Accommodation in farm house or hotel - free time. 20.00: Dinner (typical local food). After dinner it will be possible to visit other local sites of cultural interest. 15 May 2006 08.00: Departure from the Upper Tiber Valley (a brief stay for a landscape overview of the “Protected Serpentine Nature Area of Pieve S. Stefano” may be possible) to the Emilian Apennine area (Taro Valley - Province of Parma, Emilia-Romagna). 11.30-17.00: Guided visit to the “Monte Prinzera Regional Nature Reserve,” including a lunch-buffet (possibly in the field). 17.00-19.00: Visit to local villages and sites of cultural interest. 19.00-20.00: Accommodation in farm house or hotel - free time. 20.00: Dinner (typical local food). After dinner it will be possible to visit other local sites of cultural interest or participate in an evening astronomic observation. 16 May 2006 08.00: Departure from the Emilian Apennine area to Ligurian-Piedmontese Apennine area (Scrivia Valley - Provinces of Genova and Alessandria, Regions Liguria and Piemonte).



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11.00-15.00: Guided visit to the “Beigua Regional Nature Park,” including a lunch-buffet (possibly in the field). 15.00: Short drive (about 0.45 h) 16.00-19.00: Guided visit to the “Capanne di Marcarolo Regional Nature Park.” 19.00-20.00: Accommodation in farm house or hotel - free time. 20.00: Dinner (typical local food). After dinner it will be possible to visit other local sites of cultural interest. 17 May 2006 08.00: Departure from the Ligurian-Piedmontese Apennine area to the Northern Tuscan area (Province of Prato, Region Toscana) 11.30-17.00: Guided visit to the “Monteferrato Nature Protected Area,” including both naturalistic and cultural-artistic sites, and a lunch-buffet in an historic estate (typical local food). Afterwards travel may be arranged to Florence or Pisa for overnight stays (not included or organised by the Conference) or departure from airports (half an hour to Florence Airport and 1.30 to Pisa Airport). It will be possible to arrange special shuttle service for earlier departures (please contact organisation or communicate your needs at the conference desk) Useful information, warnings and advice for the Post-Conference field excursion • Please see the map on the next page to see the route of travel and locations of stops. • The visited areas are in a Mediterranean climatic region. However, since in mid-May the

weather can be still unsettled (fresh or rainy) and the elevation can reach 500-1000 m above sea level, adequate mountain clothes and boots are recommended.

• The Management Corporations of the visited Protected Areas will provide to the participants all possible logistic, technical and scientific support, including an indication of alternative opportunities in the case of rainy weather.

• Because we will be visiting protected conservation areas, collecting a limited amount of specimens of minerals, rocks, soil, flora or fauna will be possible only upon request to the local guide or managers.

• Documentation about visited areas and their essential scientific aspects will be provided to the participants, thanks to the Organizing Committee in association with the Coordination of Ophiolitic Protected Areas (C.A.P.O.).

• If necessary because of particular reasons, the Organizing Committee may make changes to the program outlined above. In case, the participants will be informed as soon as possible.



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14.05.06 Tiberina Valley Protected Areas

17.05.06 Monteferrato

Protected Area

16.05.06 Ligurian-Piedmontese

Apennine Protected Areas

15.05.06 Emilian Apennine Protected Areas

12.05.06 Cecina Valley

Protected Areas

Mont Avic

Marcarolo Aveto

Prinzera Campotrera

Montenero

Monterufoli

Beigua

Monteferrato

Serpentine Monti Rognosi

Post Conference Field Trip

Mid Conference Field Trip

Protected Areas adhering to the Coordination of Ophiolitic Protected Areas (C.A.P.O.)

Visited territories (overnight stay: Tiberina Valley Protected Areas, Emilian Apennine Protected Areas and Ligurian-Piedmontese Apennine Protected Areas)

Legend

Mid Conference and Post Conference Field Trip map



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Conference Documentary A unique feature of this Fifth International Conference will be the production of a documentary that will record highlights of the event.? The documentary will be filmed and produced by Ms. Megan Erskine of Grand Ledge, Michigan, USA.? Ms. Erskine holds a B.A. degree from Kalamazoo College.? She has created documentaries that have appeared in the East Lansing Film Festival (USA), has served as Assistant Director on a documentary that was awarded the 2nd place prize for Best Documentary at the 2005 East Lansing Film Festival, and served as Production Assistant in a tennis documentary that will appear at the US Open in September 2006.? As Documentarian for the Fifth International Conference on Serpentine Ecology, Ms. Erskine?s documentary will allow delegates to recall the excitement of the conference and its field trips long after they have returned to their home countries.

Program Updates

Please check the conference website for updates and last-minute changes in the conference program and list of contributions.



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This conference was supported by:

Provincia di Arezzo

Provincia di Parma

Provincia di Prato

C.M. Alta Val di Cecina

Parco delle Capanne di Marcarolo

Parco Regionale del Beigua

ANPIL del Monte Ferrato

www.monteferrato.it

www.parcobeigua.it

www.parcocapanne.it

http://www.cm-valtiberina.toscana.it/

http://www.cmtaroceno.pr.it/

http://www.comune.siena.it/

http://www.comune.terenzo.pr.it/

http://www.provincia.arezzo.it/

http://www.regione.toscana.it/

http://www.mps.it/

http://www.unisi.it/

http://www.provincia.parma.it/

http://www.comune.fornovo-di-taro.pr.it/comune/

http://www.comune.terenzo.pr.it/

http://www.cm-valdicecina.pisa.it/index.asp





http://www.parks.it/aree.protette.ofiolitiche/index.html

http://www.parks.it/aree.protette.ofiolitiche/index.html

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Fifth International Conference on Serpentine Ecology · Fifth International Conference on Serpentine Ecology ... Scott Angle, Lan Zhang, Rufus ... Fifth International Conference on