K Fellow/Grantee Award Start Date Krista Kaasik, PhD 2007 David B. Kaback, MD, PhD 1976 Natasha M. Kablaoui, PhD 1997 Balint Kacsoh, PhD 2020 David Kadosh, PhD 1998 Susan M. Kaech, PhD 1999 Björn F.C. Kafsack, PhD 2010 Jon M. Kaguni, PhD 1980 Fred Marvin Kahan, PhD 1961 Jason A. Kahana, PhD 1999 Kenneth Kahn, MD 1956 Patricia Kahn, PhD 1981 Brian Paul Kaine, PhD 1981 Akira Kaji, PhD [G] 1972 George F. Kalf, PhD [G] 1972 Klaus Kallman, PhD 1950 Naoyuki Kamatani, MD 1981 C. Alexander Kamb, PhD 1988 Barton A. Kamen, MD, PhD 1979 Robert I. Kamen, PhD 1971 Mark S. Kaminski, MD 1983 Mark P. Kamps, PhD 1988 Ryuichi Kanayama, MD 1970 Chul-Hee Kang, PhD 1989 Yunsik Kang, PhD 2018 Dr. Seppo J. Kangas [G] 1961 Camilla Mei-fang Kao, PhD 1997 Alan P. Kaplan, PhD 1992 Daniel Lee Kaplan, PhD 2001 Henry S. Kaplan, MD [G] 1956 Kenneth B. Kaplan, PhD 1994 Leonard J. Kaplan, MD, PhD 1990 Paul L. Kaplan, PhD 1983 Paul M. Kaplan, PhD 1971 Geoffrey M. Kapler, PhD 1989 Tarun M. Kapoor, PhD 1998 Dietmar J. Kappes, PhD 1988 Alan Mitchell Kapuler, PhD [G] 1971 John Karanicolas, PhD 2004 Katrin Karbstein, PhD 2004 Lucien Karhausen, MD 1959 John J. Karijolich, PhD 2012 David A. Karnofsky, MD 1952 Susan Karplus, PhD 1965

Howard T. Karsner, MD [G] 1949 Nobuhiko Katunuma, MD [G] 1967 Flora Naomi Katz, PhD 1981 Joseph J. Kaufman, MD [G] 1971 John F. Kaumeyer, PhD 1978 John Edward Kay, PhD 1967 Michael S. Kay, MD, PhD 1999 Samuel C. Kayman, PhD 1978 Paul Sanford Kayne, PhD 1989 Thomas A. Keating, PhD 1998 Corrine Keaty, PhD [G] 1950 Jung-Min Kee, PhD 2009 Jo-Ann N. Keene, PhD 1984 Scott Neal Keeney, PhD 1994 Marilyn Rose Kehry, PhD 1980 Philip C. Kelleher, MD [G] 1973 Paul Malcolm Keller, PhD 1977 Rebecca Kellum, PhD 1991 Keith H. Kelly, MD 1949 Teresa Keng, PhD 1983 Byrl James Kennedy, MD, MSc 1949 Grace E. Kenney, PhD 2019 Kinneret Keren, PhD 2005 Leeat Yankielowicz- Keren, PhD 2017 Ian Macpherson Kerr, PhD 1963 Walter Kerr, MD [G] 1952 Ellen N. Kersh, PhD 2000 Richard H. Kessin, PhD 1971 Vineet N. Kewalramani, PhD 1996 Avinash Khanna, PhD 2014 Kamal Mohan Khanna, PhD 2005 Raja W. Khuri, MD 1962 Diane R. Kiino, PhD 1983 Joanne Killinger, PhD 1970 Bill A. Kilpatrick, PhD 1978 Lary J. Kilton, MD 1978 Annette S. Kim, MD, PhD 1998 Daniel H. Kim, PhD 2009 Joon Kim, PhD 1990 Sally A. Kim, PhD 2006 Tae Kook Kim, PhD 1995 Yumi Kim, PhD 2011 Bryan C. King, PhD 2015 Charles Miller King, PhD [G] 1972 Gerald W. King, MD [G] 1973 Mary E. King, MD 1972 Mary Lou King, PhD 1976

Thomas D. Kinney, MD [G] 1949 Elaine Kinney-Thomas, PhD 1979 Riojun Kinosita, MD, PhD [G] 1958 Thomas J. Kipps, MD, PhD 1982 Serkan Kir, PhD 2013 Karla Ann Kirkegaard, PhD 1983 Paul T. Kirschmeier, PhD 1979 Werner H. Kirsten, MD [G] 1968 Takashi Kei Kishimoto, PhD 1988 Dr. Donald E. Kizer [G] 1971 Daniel Joseph Klein, PhD 2005 George Klein, MD, PhD 1974 Hannah L. Klein, PhD 1976 Peter S. Klein, MD, PhD 1990 William H. Klein, PhD 1974 William Lee Klein, PhD 1973 Ralph E. Kleiner, PhD 2016 Paul Klemperer, MD [G] 1955 Leonard Klevan, PhD 1978 Morton Kligerman, MD [G] 1952 Dr. Harlan Klug [G] 1951 Anne K. Knecht, PhD 1997 Laura Jean Knoll, PhD 1995 Marcel Knossow, DSc 1982 Peter James Knudsen, PhD 1980 Akira Kobata, PhD [G] 1972 Naoko Kobayashi, PhD 1991 Henry J. Koch, MD 1950 Ronald L. Koch, PhD 1976 Carla M. Koehler, PhD 1995 Kathryn M. Koeller, PhD 2001 Heinz K. Kohler, MD, PhD [G] 1972 Kazunori Koide, PhD 1998 Michael A. Koldobskiy, MD, PhD 2016 Alex Leo Kolodkin, PhD 1987 Andrew Kolodziej, PhD 1993 Zacharias Komninos, MD 1952 Hisato Kondoh, PhD 1976 James B. Konopka, PhD 1985 Michael P. Koonce, PhD 1988 Milan J. Kopac, PhD [G] 1969 Edward David Korn, PhD 1951 Stephen Kornfeld, MD, PhD 1991 Dr. Nora Kory 2017 Peter L. Kosa, PhD, MBA 1998 Steven A. Kosciuk, PhD 1983 Raymond A. Koski, PhD 1977

Chitra V. Kotwaliwale, PhD 2008 Gerald B. Koudelka, PhD 1984 Timothy F. Kowalik, PhD 1990 Marilyn Sue Kozak, PhD 1973 David Kramish, MD [G] 1960 Stephen Krauss, MD [G] 1972 Thane Kreiner, PhD 1988 Willi Kreis, MD, PhD [G] 1972 Norman Kretchmer, MD, PhD [G] 1957 Michael Paul Kriegler, PhD 1980 David Kristofferson, PhD 1982 Dr. Henry Kritzler [G] 1959 Dr. Heinrich Kroeger [G] 1967 Kristen L. Kroll, PhD 1995 Theodore G. Krontiris, MD, PhD 1980 Kristin A. Krukenberg, PhD 2010 Raju S. Kucherlapati, PhD 1973 Diane Jane S. Kuehn, PhD 1979 Margarethe J. Kuehn, PhD 1994 Brian Kuhlman, PhD 1999 Warakorn Kulalert, PhD 2018 Jerzy K. Kulski, PhD 1980 Soma Kumar, PhD [G] 1971 Mondira Kundu, MD, PhD 2000 Samuel Kunes, PhD 1988 Ching-Hung Kuo, PhD 1972 Janet Ann Kurjan, PhD 1980 László Kürti, PhD 2007 Marvin Kuschner, MD [G] 1955 Young Kwon, PhD 2009 Jack E. Kyte, PhD 1972

L Fellow/Grantee Award Start Date Thomas R. LaBar, PhD 2019 Alain Laederach, PhD 2004 Lindsay M. LaFave, PhD 2017 Jean-Gabriel Lafontaine, PhD 1954 Johanna Lahdenranta, PhD 2006 Robert S. Lahue, PhD 1985 Jamie Lahvic, PhD 2018 Eric C. Lai, PhD 2000 Ming-Zong Lai, PhD 1985 Brian J. Laidlaw, PhD 2016 Laimonis A. Laimins, PhD 1981 Patricia L. Lakin-Thomas, PhD 1985 Paul A. Lalor, PhD 1984

Ngan Lam, PhD 2005 Eric J. Lambie, PhD 1988 John P. Lambooy, PhD [G] 1966 David G. Lambright, PhD 1992 Michael E. Lamm, MD [G] 1973 Arthur Lampert, PhD 1976 Gayle K. Lamppa, PhD 1981 Joseph V. Landau, PhD 1952 Gabriel C. Lander, PhD 2010 Charles Daniel Lane, PhD 1973 Stanley Earl Lane, PhD 1971 Dennis Robert Lang, PhD 1971 Lewis L. LanierPhD 1979 Anne E. Lanjuin, PhD 2005 Lawrence H. Lanzl, PhD [G] 1964 Jessica P. Lao, PhD 2013 Christopher P. Lapointe, PhD 2018 Liana F. Lareau, PhD 2009 Frederick Joseph LaRiviere, PhD 2002 Dr. Michael Laskowski [G] 1966 Dr. Daniel Laszlo [G] 1953 Raymond Latarjet, MD, PhD [G] 1967 Raffael Lattes, MD [G] 1953 John K. Lattimer, MD [G] 1958 Allison N. Lau, PhD 2015 Ken Siu-Kwong Lau, PhD 2009 Andrew P. Laudano, PhD 1982 John S. Laughlin, PhD [G] 1956 Charles T. Lauhon, PhD 1992 Dr. Pekka Lauren [G] 1962 Suzanne Laurin, MD 1960 Sean M. Lavelle, MD [G] 1965 David Halcro Lavrin, PhD 1963 Edwin A. Lawrence, MD [G] 1951 Walter Lawrence, Jr., MD [G] 1963 Benjamin Lax, PhD [G] 1969 Jerome Lazar, PhD 1974 Paul B. Lazarow, PhD 1973 Beth A. Lazazzera, PhD 1996 Franklin R. Leach, PhD [G] 1968 Jess Herter Leber, PhD 2004 David E. Lebwohl, MD 1986 Robert J. Lechleider, MD 1992 Mary Lee S. Ledbetter, PhD 1973 Nada Ledinko, PhD [G] 1970 Brian Lee, PhD 2005 Dr. Chae Koo Lee [G] 1958

Cheng-Yu Lee, PhD 2003 Chih-Yung Lee, PhD 2013 Cynthia K. Lee, PhD 1982 Gloria Lee, PhD 1979 Janet Susan Lee, PhD 1979 June-Yong Lee, PhD 2015 Kenneth K. Lee, PhD 2003 Sang He Lee, PhD 1977 Siu Sylvia Lee, PhD 2001 Teh H. Lee, PhD [G] 1965 Tzoong-Chyh Lee, PhD 1969 Wei-Lih Lee, PhD 2001 Robert S. Lees, MD 1970 Stanley S. Lefkowitz, PhD [G] 1963 David W. Lehman, PhD 1976 Herndon Lehr, MD 1957 Gerard M. Lehrer, MD [G] 1963 John A. Leigh, PhD 1983 Joseph Leighton, MD [G] 1969 Leslie Anne Leinwand, PhD 1978 Jonathan P. Leis, PhD 1971 Ihor R. Lemischka, PhD 1984 Mark A. Lemmon, PhD 1993 Dr. Petra Lenta [G] 1971 John D. Leonard, PhD 2016 Stanley P. L. Leong, MD, MS, FACS 1981 Kathrin Leppek, PhD 2015 George V. LeRoy, MD [G] 1955 Jayne F. Lesley, PhD 1972 Jun-Yi Leu, PhD 2000 Kerstin K. Leuther, PhD 1994 Hilde Levi, PhD [G] 1967 Michael M. Levi, MD [G] 1969 Daniel H. Levin, PhD [G] 1966 David E. Levin, PhD 1984 Petra Anne Levin, PhD 1996 Tera C. Levin, PhD 2015 Jedd F. Levine, MD 1983 Dr. Laurence Levine 1963 Reuben Levisohn, PhD 1965 Daniel J. Lew, PhD 1990 Alfred S. Lewin, PhD 1979 James Bryan Lewis, PhD 1971 Jay Todd Lewis, PhD 1980 Robert C. Lewis, MD, PhD [G] 1954 Chuan Li, PhD 2018 Haoxin Li, PhD 2020

Hung-Wen Li, PhD 2001 Julia Su Zhou Li, PhD 2020 Kaijun Kevin Li, PhD 1997 Lewyn Li, PhD 2003 Min Chiu Li, MD 1953 Renhao Li, PhD 1999 Rong Li, PhD 1993 Shang Li, PhD 2001 Ting-kai Li, MD 1963 Yen Li, PhD 1983 Chun Liang, PhD 1994 Zhuobin Liang, PhD 2016 Heather Hong Liao, PhD 1999 Sha-Mei Liao, PhD 1988 Xiaoling Charlene Liao, PhD 1992 Louis James Libertini, PhD 1971 Stuart S. Licht, PhD 1998 Michael J. Lichten, PhD 1982 Dr. Louise Ann Lichtenberg 1970 Yung S. Lie, PhD 2001 Judy Lieberman, MD, PhD 1984 Michael W. Lieberman, MD, PhD 1968 Miriam Lieberman, PhD 1956 Seymour Lieberman, PhD [G] 1956 Eric C. Liebl, PhD 1991 Evan C. Lien, PhD 2017 Jeffrey D. Lifson, MD 1983 William Lijinsky, PhD 1954 Brendan N. Lilley, PhD 2006 Daniel H. Lin, PhD 2018 Jim Jung-Ching Lin, PhD 1979 Jing-Jer Lin, PhD 1992 Qing Lin, PhD 2000 Shuibin Lin, PhD 2013 Steven Lin, PhD 2013 Weei-Chin Lin, MD 1999 Dr. Carl C. Lindegren [G] 1962 Erik Lindh, MD, PhD 1976 Anders Mikael Lindroth, PhD 2001 Tullia Lindsten, MD, PhD 1983 Geoffrey Shiu Fei Ling, MD, PhD 1982 Elwood Albert Linney, PhD 1973 Peter N. Lipke, PhD 1976 Fritz A. Lipmann, MD, PhD [G] 1952 Stuart W. Lippincott, MD [G] 1947 Muriel M. Lippman, PhD [G] 1964 Allan Lipton, MD [G] 1972

R. Michael Liskay, PhD 1975 Mortimer Litt, MD [G] 1961 Charles D. Little, PhD 1978 Clarence Cook Little, DSc [G] 1947 Sheila Parks Little, PhD 1978 Alvin Y. Liu, PhD 1981 Chih-Ping Liu, PhD 1970 Edison Tak-Bun Liu, MD 1983 Jun O. Liu, PhD 1991 Kaixian Liu, PhD 2020 Lucy Liu, PhD 2018 Paul Z. Liu, PhD 2005 Qinghua Liu, PhD 2001 Rihe Liu, PhD 1997 Xiaoxiao Shawn Liu, PhD 2013 Xuan Liu, MD, PhD 1991 Yanshun Liu, PhD 2003 Yin Liu, PhD 2014 Zhongchi Liu, PhD 1991 Kenneth J. Livak, PhD 1979 George Pietro Livi, PhD 1983 Cecilia W. Lo, PhD 1979 Wilson Hwei-yuen Lo, MD 1952 Albert F. LoBuglio, MD [G] 1970 Leslie F. Lock, PhD 1986 David J. Lockhart, PhD 1990 Dr. William R. Lockhart [G] 1956 Virgil Loeb, Jr., MD 1950 Jennifer Lynn Loebach, PhD 1995 John B. Loefer, PhD [G] 1951 Robert B. Loftfield, PhD 1952 Catriona Y. Logan, PhD 1998 Stewart N. Loh, PhD 1993 Allan Richard Lohe, PhD 1979 John D. Loike, PhD 1976 Patrick J. Loll, PhD 1990 A.J. Lomant, MD 1972 Dr. Paul S. Lombardi 1969 Leon Lombrozo, PhD 1961 Irving M. London, MD [G] 1956 David G. Longfellow, PhD 1972 Jane Ellen R. Lopilato, PhD 1984 Muriel R. Loran, PhD [G] 1953 Albert B. Lorincz, MD [G] 1969 Allan L. Lorincz, MD 1951 Jon R. Lorsch, PhD 1995 David J. Loskutoff, PhD 1972

Marie Lossky, PhD 1988 Robert Alan Love, PhD 1985 Scott E. Loveless, PhD 1980 Ky Tsing Lowenhaupt, PhD 1979 Jerold M. Lowenstein, MD [G] 1967 David F. Lowry, PhD 1992 Bertram A. Lowy, PhD [G] 1956 Chao Lu, PhD 2014 Hua Lu, MD, PhD 1994 Hua Lu, PhD 2011 Wan-Jin Lu, PhD 2011 Xiaowei Lu, PhD 2000 Ying Lu, PhD 2011 Zhipeng Lu, PhD 2015 Joseph J. Lucas, PhD 1972 Dr. Paul P. Luger [G] 1964 Christine M. Lukacs, PhD 1998 Thomas J. Lukas, PhD 1978 Roger H. Lumb, PhD 1970 Ellen A. Lumpkin, PhD 1999 Erik Arthur Lundquist, PhD 1996 Jingchuan Luo, PhD 2020 Shijing Luo, PhD 2012 Wenyu Luo, PhD 2013 Patrick James Lupardus, PhD 2006 Salvador E. Luria, MD [G] 1961 Andrew D. Luster, MD, PhD 1991 Terry Paul Lybrand, PhD 1985 John R. Lydeard, PhD 2010 Susan K. Lyman, PhD 1996 Henry T. Lynch, MD [G] 1969 Dr. Kenneth M. Lynch, Jr. [G] 1950 Dr. Carmine Lyons 1952 Costas A. Lyssiotis, PhD 2010

M Fellow/Grantee Award Start Date Le Ma, PhD 2000 Lifu Ma, PhD 1997 David Maag, PhD 2006 Ernst-Gunter Maasz, MD 1966 David M. MacAlpine, PhD 2001 Paul M. Macdonald, PhD 1984 Frederick M. MacDonnell, PhD 1992 Lorne A. MacHattie, PhD 1962 Laura M. Machesky, PhD 1993 David P. Mack, PhD 1990

Paul S. Maddox, PhD 2004 Sergio I. Magalini, MD 1955 Jesse Mager, PhD 2004 Nicole M. Mahoney, PhD 2000 Ivan P. Maillard, MD, PhD 2005 Paul E. Mains, PhD 1983 Henri Emile Maisin, MD, PhD 1952 Richard M. Maizels, PhD 1977 Nizarali R. Makan, MD 1977 Dr. Sajiro Makino [G] 1966 Jocelyn E. Malamy, PhD 1993 Vivek Malhotra, PhD 1985 Jarema Malicki, PhD 1993 Leonard I. Malkin, PhD [G] 1971 M. Frank Mallette, PhD [G] 1956 Debora Mancini-DiNardo, PhD 2001 H. George Mandel, PhD [G] 1960 Ofer Mandelboim, PhD 1997 Jeffrey G. Mandell, PhD 2000 Ines Mandl, PhD [G] 1968 James J. Manfredi, PhD 1983 Leon H. Manheimer, MD [G] 1964 Z. Thaddeus Mankowski, MD [G] 1964 Dr. David E. Mann, Jr. [G] 1963 David A. Mann, PhD 2000 Thomas H. Mann, PhD 2019 Dr. Gilbert B. Manning [G] 1960 Timothy Manser, PhD 1982 Junhao Mao, PhD 2003 Kai Mao, PhD 2015 Ayelet Maor-Shoshani, PhD 2004 York Marahrens, PhD 1994 Eugene E. Marcantonio, MD, PhD 1986 John J. Marchalonis, PhD 1973 Philip I. Marcus, PhD [G] 1974 Kurt W. Marek, PhD 2005 Jeanne P. Margolskee, PhD 1979 Elizalde Adolfo Mariano, MD 1962 Elizabeth Christine Marin, PhD 2005 George Edward MarkIII, PhD 1971 Vernon Mark, MD 1951 Gabor Markus, MD, PhD [G] 1965 Mary Ann K. Markwell, PhD 1976 Florence L. Marlow, PhD 2004 Elchanan Maron, PhD 1972 Keith R. Marotti, PhD 1980 Till Marquardt, PhD 2002

Michael Thomas Marr, PhD 2001 Philippa C. Marrack, PhD 1971 Lorraine Marsh, PhD 1986 Dr. Lawrence M. Marshall 1953 Phillip A. Martens, PhD 1978 Dr. Alfredo Marti [G] 1965 Klaus D. Martignoni, PhD 1971 Dr. Daniel S. Martin [G] 1960 Hayes Martin, MD [G] 1958 Stuart S. Martin, PhD 1998 Timothy D. Martin, PhD 2014 James Martinell, PhD 1977 Eric Martz, PhD 1969 Maria Teresa Mas, PhD 1985 Richard S. Maser, PhD 2002 Francis Masin, MD [G] 1959 Marianna Masin, MD 1966 Earl James Mason, MD, PhD 1954 Howard S. Mason, PhD [G] 1957 William S. Mason, PhD 1971 John J. Masters, PhD 1994 Andrea Marie Mastro, PhD 1971 A. Gregory Matera, PhD 1991 Maria K. Mateyak, PhD 2001 Gladys M. Mateyko, PhD [G] 1968 Rebecca S. Mathew, PhD 2010 Dr. Robert Mathewson [G] 1964 Diane Mathis, PhD 1977 Pedro M. Matias, PhD 1987 Ryota Matsuoka, PhD 2012 Adam G.W. Matthews, PhD 2008 Richard F. Mattingly, MD [G] 1970 Lida H. Mattman, PhD [G] 1968 David Q. Matus, PhD 2007 Robert J. Matusik, PhD 1976 Gerd G. Maul, PhD [G] 1971 Russell A. Maurer, PhD 1979 Lisa Maves, PhD 1998 Penelope Nazos Mavromara, PhD 1984 E. Stuart Maxwell, PhD 1977 John T. May, PhD 1971 Brian H. Mayall, MD 1963 Klaus Mayer, MD 1955 Jacob Allen Mayfield, PhD 2002 Kelly Edward Mayo, PhD 1983 Heather D. Mayor, PhD [G] 1974 Esteban O. Mazzoni, PhD 2007

Grant Alan McArthur, MB BS (Hons) PhD, FRACP 1995 Monica E. McCallum, PhD 2017 Kay L. McCarthy, PhD 1976 Duncan L. McCollester, MD, PhD [G] 1969 Paul R. McCurdy, MD [G] 1961 Timothy K. McDaniel, PhD 1997 Dr. John McDonald 1950 Joseph J. McDonald, MD, PhD 1951 Dr. Margaret Ann McDowell [G] 1957 Michael J. McEachern, PhD 1988 Joan E. McEwen, PhD 1980 Jon McGinn, PhD 2020 Robert K. McGinty, MD, PhD 2012 Dr. Thomas R. McGowan 1959 Patrick T. McGrath, PhD 2006 Joseph S. McGuire, MD [G] 1974 Dr. Lorne McHattie 1950 Alesia N. McKeown, PhD 2016 Kara L. McKinley, PhD 2017 Robert G. McKinnell, PhD [G] 1968 Colleen N. McLaughlin, PhD 2020 Cory Y. McLean, PhD 2012 Seamus McMillan, PhD 1980 Ronald McNamara, PhD 1992 Alexander McPherson, PhD 1970 James J. McSharry, PhD [G] 1973 Brian M. McStay, PhD 1984 R. McWhirter, CBE, DMRE, FRCSE, FRCR [G] 1951 Susan O. Meakin, PhD 1988 Joan C. Mecsas, PhD 1994 Roy B. Mefferd, Jr., PhD 1952 Paul C. Megee, PhD 1995 Joseph V. Meigs, MD [G] 1952 Pier Melanotte, MD 1955 Thomas E. Melendy, PhD 1989 Cornelis J.M. Melief, MD, PhD 1973 Susan J. Mellette, MD 1957 Robert C. Mellors, MD, PhD 1950 Lynn V. Mendelman, PhD 1989 John Mendelsohn, MD [G] 1972 Mortimer L. Mendelsohn, MD, PhD 1951 Enrique Méndez Cormán, PhD 1971 Marc D. Meneghini, PhD 2001 Dr. Gordon Menn 1970 Eric H. Mercer, PhD 1993 Christian G. Merkel, PhD 1976 D. Scott Merrell, PhD 2002

Karl A. Merrick, PhD 2012 Melanie M. Merriman, PhD 1979 Alexey J. Merz, PhD 2000 I. Meschan, MD [G] 1951 Herbert Mescon, MD 1949 Joseph L. Messina, PhD 1983 Luigi Messineo, PhD 1958 Donald J. Messner, PhD 1986 Lauri Meurman, MD [G] 1965 Richard J. Meyer, PhD 1972 Frederick J. Meyers, MD 1982 Marian B. Meyers, PhD 1978 Matthew L. Meyerson, MD, PhD 1995 W. Matthew Michael, PhD 1997 Tamar Michaeli, PhD 1987 George Albert Michaels, PhD 1967 Julia M. Michelotti, PhD 1991 Rhett J. Michelson, PhD 2000 Maria M. Mihaylova, PhD 2013 Ross B. Mikkelsen, PhD 1973 Charles P. Miles, MD [G] 1966 Alexander Miller, PhD 1956 Allan Malcolm Miller, PhD 1986 Brian Gene Miller, PhD 2002 I. George Miller, MD 1968 Jacques F.A.P. Miller, PhD [G] 1973 Jeffery F. Miller, PhD 1986 Kenneth B. Miller, MD 1977 Mathew G. Miller, PhD 2005 Matthew P. Miller, PhD 2014 Rand M. Miller, PhD 2015 Richard A. Miller, MD, PhD 1977 Tamara P. Miller, MD 2016 Virginia L. Miller, PhD 1985 Frank Millich, PhD 1960 Josselin Milloz, PhD 2009 Joan T. Mills, PhD 1981 G. Todd Milne, PhD 1996 Jeremy David Milton, PhD 1965 Terry C. Mincey, PhD 1979 Jeffrey H. Miner, PhD 1992 Aaron Peter Mitchell, PhD 1984 Alva D. Mitchell, PhD 1976 James R. Mitchell, PhD 2001 Pamela J. Mitchell, PhD 1986 Jyotirmay Mitra, PhD [G] 1967 Kaneatsu Miyamoto, MD 1967

Merle Mizell, PhD [G] 1970 Stephen C. Mockrin, PhD 1973 Gregory R. Moe, PhD 1985 Raymond E. Moellering, PhD 2011 Kenny C. Mok, PhD 2005 Antoine Molaro, PhD 2014 Kivie Moldave, PhD [G] 1956 Goran Moller, MD, PhD [G] 1969 Peter L. Molloy, PhD 1976 Lisa Molz, PhD 1992 Norie Momiyama, PhD 2005 Alfonso Mondragon, PhD 1985 Gabriela C. Monsalve, PhD 2014 Dr. F. Gomez Mont [G] 1957 Gaetano T. Montelione, PhD 1987 Craig Montell, PhD 1983 Philip O'Bryan Montgomery, Jr., MD [G] 1967 Taiowa A. Montgomery, PhD 2009 Sylvan E. Moolten, MD [G] 1954 Alice E. Moore, MD [G] 1955 Dan H. Moore, PhD [G] 1954 George E. Moore, MD, PhD [G] 1959 Hsiao-Ping Hsu Moore, PhD 1982 Kevin W. Moore, PhD 1979 Dr. Robert A. Moore [G] 1951 Sean D. Moore, PhD 2002 Susan Ann Moore, PhD 1977 Sheri L. Moores, PhD 1998 Paul S. Moorhead, PhD 1955 John Richard Moran, PhD 1983 John V. Moran, PhD 1995 Katarina Moravcevic, PhD 2012 Robert Byrl Moreland, PhD 1983 Edward L. Morgan, PhD 1978 Jeffrey R. Morgan, PhD 1983 William D. Morgan, PhD 1983 Kazuo Mori, MD [G] 1968 Vitor Mori, PhD 2020 Dr. Pablo Mori-Chaves [G] 1964 Alex O. Morla, PhD 1990 Harold Moroson, PhD [G] 1970 Helen G. Morris, PhD 1961 John McLean Morris, MD 1951 Kevin N. Morris, PhD 1991 Vincent Lynn Morris, PhD 1973 Martin Morrison, PhD [G] 1971 Trudy G. Morrison, PhD 1972

Anthony T.E. Morrissey, PhD 1970 K. John Morrow, Jr., PhD [G] 1971 Cynthia Casson Morton, PhD 1983 Diane Gail Morton, PhD 1983 Aron A. Moscona, PhD [G] 1963 Donald E. Mosier, MD, PhD 1971 Gerald Moss, MD, PhD [G] 1967 Joseph D. Mougous, PhD 2005 Mohammad Movassaghi, PhD 2001 Kerri A. Mowen, PhD 2001 Ryan H. Moy, MD, PhD 2020 Aaron L. Moye, PhD 2019 Jan Ailene Moynihan, PhD 1984 David Mu, PhD 1994 Michael M. Mueckler, PhD 1983 Jo Anne Mueller, PhD [G] 1972 Kanai Lal Mukherjee, PhD 1960 Carel Mulder, PhD [G] 1971 Mandy M. Muller, PhD 2014 John T. Mulligan, PhD 1988 Equinn Munnell, MD [G] 1963 Hamish N. Munro, MD [G] 1971 Tariq M. Murad, MD, PhD [G] 1971 Dr. Joseph Murdock 1970 Andrew C. Murley, PhD 2017 John E. Murphy, PhD 1990 Martin J. Murphy, Jr., PhD 1969 Mary Lois Murphy, MD 1952 Robert F. Murphy, PhD 1979 Margaret R. Murray, PhD [G] 1952 Mark J. Murray, PhD 1980 Lyndsay M. Murrow, PhD 2015 Piero Mustacchi, MD 1949 Dale Muzzey, PhD 2010 Benjamin R. Myers, PhD 2009 Lawrence C. Myers, PhD 1995 Richard M. Myers, PhD 1982 David Myerson, MD, PhD 1981 Kyungjae Myung, PhD 1999

N Fellow/Grantee Award Start Date Anders M. Näär, PhD 1995 George W. Nace, PhD [G] 1956 Sigrid Nachtergaele, PhD 2015 Maxence V. Nachury, PhD 2003 Marion R. Nadel, PhD 1977

Ulrich Naf, PhD 1953 Takayuki Nagasaki, PhD 1990 Andrew R. Nager, PhD 2013 Barbara T. Nagle, PhD 1977 Shruti Naik, PhD 2014 Roderick Nairn, PhD 1976 Hiroshi Nakai, PhD 1984 Noburu Nakajima, PhD 1983 Toru M. Nakamura, PhD 1999 Yunsun Nam, PhD 2007 Chaya Nanavati, PhD 1993 Nagarajan Nandagopal, PhD 2018 Vikram Narasimhan, PhD 1990 Geeta J. Narlikar, PhD 1998 Larry Nathanson, MD [G] 1970 Inke S. Nathke, PhD 1992 Gioacchino Natoli, MD 1998 Georges Natsoulis, MD, PhD 1986 Gian-Carlo Nava, MD 1959 William W. Navarre, PhD 2000 Marc Navre, PhD 1984 Leon William Nawrocki, PhD 1985 B. Nedelkoff, MD [G] 1962 Jelena Nedjic, PhD 2010 Eva J. Neer, MD 1973 Norma F. Neff, PhD 1976 Jasper Neggers, PhD 2019 Robert S. Negrin, MD 1988 Aaron Michael Neiman, PhD 1994 Eric Mercer Nelson, PhD 1984 Norton Nelson, PhD [G] 1955 Muriel N. Nesbitt, PhD [G] 1972 Gaylen J. Neufeld, PhD 1966 Dr. Earl H. Newcomer [G] 1961 Dianne K. Newman, PhD 1998 Phillip Allan Newmark, PhD 1995 Donald D. Newmeyer, PhD 1983 Elizabeth M. Newton, PhD 1997 Thomas Hans Newton, MD 1958 Huck Hui Ng, PhD 2000 Vu Ngo, PhD 2001 Alexandra Nguyen, PhD 2019 Don X. Nguyen, PhD 2005 Duy P. Nguyen, PhD 2014 Jack Nguyen, PhD 2000 Thuy-Linh Nguyen, PhD 2002 Trang Nguyen, PhD 2018

Vu Quang Nguyen, PhD 2015 Marguerite Sykes Nichols, MD 1951 Warren W. Nichols, MD, PhD [G] 1969 Eric M. Nicholson, PhD 1999 Martin J. H. Nicklin, PhD 1985 James J. Nickson, MD [G] 1951 John E. Niederhuber, MD [G] 1975 Robert L. Nielsen, MD 1955 Dr. Robert Nielson 1960 Mikko Niemi, MD [G] 1968 Erich A. Nigg, PhD 1981 Evgenia N Nikolova, PhD 2013 Olivier Nolan-Stevaux, PhD 2004 Kikuo Nomoto, MD 1966 Kenneth D. Noonan, PhD [G] 1973 Stig Nordling, MD, PhD [G] 1971 Jared T. Nordman, PhD 2009 Thomas M. Norman, PhD 2015 Ingrid C. Northwood, PhD 1993 Rolf Nothiger, PhD 1967 Ajit Nott, PhD 2005 Jamison Leigh Nourse, PhD 1997 Bennett G. Novitch, PhD 1999 Abraham Novogrodsky, MD, PhD 1964 Maja Nowakowski, PhD 1975 Robert W. Noyes, MD [G] 1955 Vasilis Ntziachristos, PhD 2001 Walter J. Nungester, MD [G] 1952 Ysbrand Nusse, PhD 2020

O Fellow/Grantee Award Start Date Natasha M. O'Brown, PhD 2016 Michael B. O'Connor, PhD 1984 Shigeyoshi Odashima, MD [G] 1968 Dr. Lester D. Odell [G] 1951 Eileen R. O'Donnell, PhD 1975 Kathryn A. O'Donnell, PhD 2006 Elsa H. O'Donnell-Alvelda, DSc [G] 1966 Max Oeschger, PhD [G] 1971 Herbert F. Oettgen, MD [G] 1968 Richard Ogden, PhD 1970 Peter O'Hare, PhD 1983 Mochihiko Ohashi, MD, PhD 1967 Shinpei Ohki, PhD [G] 1969 Douglas H. Ohlendorf, PhD 1979 Atsushi Oikawa, PhD 1964

Akinyemi I. Ojesina, MBBS, PhD 2008 Curtis Toshio Okamoto, PhD 1990 Dr. Chiyeko Okawa 1953 Nils Oker-Blom, MDhc [G] 1966 Jonathan D. Oliner, MD, PhD 1994 Baldomero M. Olivera, PhD 1966 Salvatore Oliviero, PhD 1989 Sarah Naomi Olsen, PhD 2018 Arthur J. Olson, PhD 1976 Kenneth Barrie Olson, MD [G] 1965 Tracy S. Olson, PhD 1988 Wilma K. Olson, PhD 1971 Katy Ong, PhD 2020 Tong-man Ong, PhD 1970 Gary Mark Onik, MD 1984 Siew Loon Ooi, PhD 2004 Joseph T. Opferman, PhD 2001 Enid T. Oppenheimer, BSc [G] 1960 Dr. Jeannette C. Opsahl 1952 Paulo Kristian Orberg, PhD 1984 Jose Ordovas-Montanes, PhD 2016 Esteban A. Orellana Vinueza, PhD 2019 Virginie Orgogozo, PhD 2003 Sean M. O'Rourke, PhD 2002 Jeffrey W. Orr, PhD 1995 Frank R. Orsini, PhD 1970 Louis G. Ortega, MD 1953 Mary Osborn, PhD 1968 Barbara A. Osborne, PhD 1979 Dr. Richard H. Osborne [G] 1960 Timothy F. Osborne, PhD 1983 Erin A. Osborne Nishimura, PhD 2011 E. Michael Ostap, PhD 1994 Hanne L. Ostergaard, PhD 1989 Elaine A. Ostrander, PhD 1987 Jeffrey M. Ostrove, PhD 1981 Irene M. Ota, PhD 1989 George A. O'Toole, PhD 1995 Renee Otten, PhD 2012 Guangshuo Ou, PhD 2007 Abigail E. Overacre-Delgoffe, PhD 2019 Tristan Wold Owens, PhD 2020 Hironobu Ozaki, PhD 1965 Harvey Leon Ozer, MD 1964

P Fellow/Grantee Award Start Date

Dr. Donald M. Pace [G] 1963 Michael E. Pacold, MD, PhD 2012 Jacques Padawer, PhD [G] 1956 Michael R. Paddy, PhD 1984 Richard A. Padgett, PhD 1981 Juhee Pae, PhD 2019 Scott L. Page, PhD 1998 Cara Theresia Pager, PhD 2007 Ji-Hye Paik, PhD 2006 Debkumar Pain, PhD 1983 Philip Lowell Paine, PhD 1971 Deborah J. Palestrant, PhD, MBA 2001 Lydia Codetta Pan, PhD 1987 Tao Pan, PhD 1991 Sandra Panem, PhD 1970 Antonito T. Panganiban, PhD 1982 Jaswant S. Pannu, MD [G] 1961 George N. Papanicolaou, MD, PhD [G] 1961 B.W. Papermaster, PhD [G] 1967 Michael F. Para, MD 1979 Luis F. Parada, PhD 1985 Suzanne Paradis, PhD 2000 Joel D. Pardee, PhD 1978 Roberta A. Parente, PhD 1986 Horace V. Parker, MD 1957 Keith L. Parker, MD, PhD 1984 Joel Grant Parkes, PhD 1971 Francisco Parra, MD 1960 Devin H. Parry, PhD 2004 Dr. Mary S. Parshley [G] 1968 Bryce Mark Paschal, PhD 1992 Steven E. Passmore, PhD 1990 Ardem Patapoutian, PhD 1996 Anand G. Patel, MD, PhD 2020 Nikit B. Patel, PhD 2020 Nipam H. Patel, PhD 1991 Alan Robb P. Paterson, PhD 1955 Dushyant Pathak, PhD 1989 Gentry N. Patrick, PhD 2000 Phillip A. Patten, PhD 1990 David Patterson, PhD 1971 R. Bruce Patterson, Jr., PhD 1989 Thomas A. Patterson, PhD 1984 Roland A. Pattillo, MD [G] 1970 Dr. Dietrich Paul 1971 Dr. Izhak Jacob Paul 1972 Dr. J.S. Paul 1972

Amanda Paulovich, MD, PhD 2002 James Rolf Paulson, PhD 1976 Michael Ross Payne, PhD 1978 Shelley M. Payne, PhD 1978 Farhang Payvar, PhD 1978 Bret J. Pearson, PhD 2005 Chad G. Pearson, PhD 2005 Olaf Hjalmer Pearson, MD 1949 Joseph Y. Peary, ScD 1954 Larry R. Pease, PhD 1977 Ariana Peck, PhD 2019 Lawrence Jess Peck, PhD 1984 Dehua Pei, PhD 1992 Bernard M. Pejsachowicz, MD 1970 Russell E. Pemberton, MD 1968 Richard Pencharz, PhD [G] 1951 Hanjing Peng, PhD 2014 Jeffrey W. Peng, PhD 1993 Jay Stuart Pepose, MD, PhD 1983 Michael A. Pereira, PhD 1971 Karen K. Perkins, PhD 1987 Margaret E. Perkins, PhD 1976 Hans Peter Perlmann, PhD [G] 1967 Benjamin Perman, PhD 1999 Louis Pernokas, MD 1956 Fernand G. Peron, PhD [G] 1961 Lester Persky, MD 1950 Anders Persson, MD, PhD 1977 Ruth Pertel, PhD 1968 Bernard M. Pessac, MD 1970 Blake Robert Peterson, PhD 1995 Barbara Petrack, PhD 1958 Lilli M. Petruzzelli, MD, PhD 1992 Russell C. Petter, PhD 1983 Howard R. Petty, PhD 1980 Sarah J. Pfau, PhD 2017 Karl E. Pfeifer, PhD 1989 Julie K Pfeiffer, PhD 2002 Douglas H. Phanstiel, PhD 2012 Lennart Philipson, MD [G] 1967 Carolyn M. Phillips, PhD 2008 Peter C. Phillips, MD 1984 Dr. Philip G. Phillips 1971 Scott Thomas Phillips, PhD 2004 Eric M. Phizicky, PhD 1983 Fabio Piano, PhD 1996 Albert I. Pick, MD [G] 1971

Leslie Pick, PhD 1986 Jacqueline W. Pierce, PhD 1984 John Sturtevant Pierce, PhD 1969 Sarah B. Pierce, PhD 1998 Beverly J. Piggott, PhD 2013 Elizabeth Mary K. Pillers, MD 1952 Maxim Pimkin, MD, PhD 2018 Gregory S. Pincus, ScD [G] 1954 Henry Pinkerton, MD [G] 1951 David J. Pintel, PhD 1979 Ann L. Pinto, PhD 1987 Douglas W. Piper, MD [G] 1965 Paula M. Pitha-Rowe, PhD [G] 1973 Helen M. Piwnica-Worms, PhD 1984 Jeffrey A. Pleiss, PhD 2001 Otto J. Plescia, PhD [G] 1958 Diane J. Plotkin, PhD 1978 Marian L. Plunkett, PhD 1983 Dr. Meir Podberezec 1952 Alberto D. Podjarny, PhD 1978 Kit J. Pogliano, PhD 1993 Claudia L. Poglitsch, PhD 1992 Beatriz Teresa Garcia Pogo, MD, DMSc 1964 Frank M. Polack, MD 1960 Jon R. Polansky, MD 1976 Deborah A. Polayes, PhD 1984 Herman Polet, MD 1963 Alex Pollen, PhD 2013 Arthur W. Pollister, PhD [G] 1966 Roberta R. Pollock, PhD 1981 Brian A. Pollok, PhD 1983 Charles M. Pomerat, PhD [G] 1951 Laura Pontano Vaites, PhD 2012 Jan Ponten, PhD [G] 1967 Maximilian W. Popp, PhD 2012 James W. Poppell, MD 1952 Mark A. Poritz, PhD 1992 Daniel A. Portnoy, PhD 1983 Joseph Post, MD [G] 1968 Magdalena E. Potok, PhD 2015 Christopher S. Potten, PhD 1968 Christopher J. Potter, PhD 2003 Kedar N. Prasad, PhD 1970 Elizabeth D. Prescott, PhD 2004 John C. Prescott, PhD 1994 Carolyn M. Price, PhD 1985 Gerald B. Price, Jr., PhD 1971

James M. Price, MD, PhD [G] 1956 Maureen Gail Price, PhD 1980 Richard Price, MD 1973 Avi Priel, PhD 2008 Aaron Prigot, MD [G] 1961 Jeffrey J. Prior, PhD 1981 Cynthia Pritchard, PhD 1980 Martin L. Privalsky, PhD 1979 Carol L. Prives, PhD 1968 Max Rowland Proffitt, PhD 1971 Albert T. Profy, PhD 1984 Samuel Proger, MD [G] 1958 Basil S. Proimos, PhD [G] 1970 Haydn Mark Prosser, PhD 1991 George Russell Prout, Jr., MD [G] 1960 Peter M. Pryciak, PhD 1992 Cristina Puchades, PhD 2020 Holde Puchtler, MD 1955 Theodore T. Puck, PhD [G] 1947 Larry Deloss Puckett, PhD 1973 Jennifer A Punt, VMD, PhD 1993 Ellen Puré, PhD 1982 Dorothy Jane M. Purifoy, PhD 1969 John A. Putkey, PhD 1982 Christopher D. Putnam, PhD 2002 Frank W. Putnam, PhD [G] 1956 Hugh M. Pyle, MD [G] 1971