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    Chapter Nine

    EXSC 326

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    +Metabolism

    Involves all chemical reactions ofbiomolecules within the body that encompasssynthesis anabolism! " brea#down

    catabolism!$hree factors a%ect total daily ener&y e'penditure

    $(EE!)

    *estin& metabolic rate+ consistin& of basal and

    sleepin& conditions plus the metabolic cost of arousal $hermo&enic e%ect of food consumed

    Ener&y e'pended durin& physical activity " recovery

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    +Component

    s of totaldaily ener&ye'penditure

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    +,asal " *estin& Metabolic *ates

    ,asal Metabolic *ate ,M*!

    Minimum level of ener&y to sustain vital

    functions in the wa#in& state

    *e-ects the body.s total heat production

    *estin& Metabolic *ate *M*!

    /lways sli&htly hi&her than ,M*

    In-uences) body si0e+ health1tness+muscle mass+ a&e+ hormones+ bodytemperature

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    +,asal " *estin& Metabolic *ates

    ,M* and *M* sum of metabolicprocesses re4uired to sustain normal

    regulatory balance and bodyfunctions durin& the restin& state

    5'y&en consumption values for ,M*

    ran&e between 67 and 287 m91min7:; to :

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    +Metabolic Si0e Concept

    Surface /rea 9aw fundamental relationshipbetween heat production and body si0e! ,M* and *M* vary in proportion to the s4uare of body

    surface area per hour #cal1m21h! =emales have > to 7? lower ,M* than males of the

    same a&e " si0e frommore fat " less fat@free mass

    $heconcept of metabolic si0erelates ,M* tobody mass raised to the 7:A> power ,M7:A>!

    ,M* e'pressed relative to ,M7:A>holds true forhumans " most mammals " birds that di%erconsiderably in si0e and shape

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    +9o&arithmicBlot of ,ody

    Mass "Metabolic*ate atts!

    for ,irds "Mammals of(i%erent

    Si0e andShape

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    +Metabolic *ates of Dumans)

    /&e and ender ComparisonsEstimates of ,M* or *M* fall within F7? of

    laboratory values

    Chan&es in body composition decrease in ==Mand1or increase in ?body fat! can e'plain the2 to 3? per decade ,M* reductionobservedfor adults

    =emales e'hibit an avera&e > to7? lower ,M*than males of same a&e (ue to less fat@free mass and more body fat fat has

    lower metabolic activity than muscle!

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    +,M* as a =unction

    of /&eand ender

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    +E%ects of *e&ular Bhysical

    /ctivity on ,M**e&ular physical activity endurance "

    resistance e'ercise! o%sets decreases in ,M*and *M* that occur with a&in&

    Estimatin& *estin& (aily Ener&y E'penditure*(EE!

    Gsually e'pressed in #cal1h or #cal1d Can be estimated from ,M* #cal1m21h! and surface

    area m2!

    Can be estimated from fat@free mass ==M!

    *(EE #cal1d! 3A7 H ==M #&!

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    +Estimates of absolute and

    relative 52upta#e of variousadult or&ans " tissues

    Organ

    OxygenConsumption (mL

    min1)

    % of RestingMetabolism

    9iver 6A 3A

    ,rain

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    +=ive =actors $hat /%ect $(EE

    : Bhysical /ctivity

    2: (iet@induced thermo&enesis

    3: Calori&enic e%ect of food on e'ercisemetabolism

    : Bre&nancy

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    +Bhysical /ctivity /%ects on

    $(EEBhysical activity e'erts by far the most profound

    e%ect on human ener&y e'penditure /ccounts for > to 37? of $(EE

    *e&ular physicalactivity stimulates restin&metabolism

    *e&ular endurance " resistance e'ercise o%sets

    the decrease in restin& metabolism that usuallyaccompanies a&in&

    Each @lb &ain in ==M increases *M* by A to 7#cal1d

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    +(iet@Induced $hermo&enesis

    (I$!or the $hermic E%ect of =ood$E=! =ood consumption increases ener&y

    metabolism 5bli&atory thermo&enesis) Ener&y re4uired to

    di&est+ absorb+ and assimilate food nutrients =acultative thermo&enesis) /ctivation of sympathetic

    nervous system and its stimulatin& in-uence onmetabolism: *eaches ma'imum within hour followin&a meal

    5verwei&ht individuals often have a bluntedthermic response lowered metabolism! thatcontributes to e'cess body fat accumulation

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    +Calori&enic E%ect of =ood onE'ercise Metabolism

    $he calori&enic e%ect of food on e'ercisemetabolism nearly doubles the food.s thermice%ect at rest

    (I$ of carbohydrate and protein e'ceedslipid(I$

    =or most individuals+ it seems reasonable

    to encoura&e moderate physical activity aftereatin& to possibly au&ment a diet@inducedincrease in caloric e'penditure for wei&htcontrol

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    +Climate

    Environmental factors in-uence metabolic rate

    *M* of people in a tropical climate avera&es > to 27?hi&her than those livin& in temperate areas

    E'ercisin& in hot weather causes about a >? hi&her 52

    consumption

    *esults from an elevated core temperature+ ener&y forsweat &land activity and altered circulatory dynamics

    Cold environments increase ener&y metabolism durin&rest and e'ercise

    (epends on body fat content and e%ectiveness ofclothin&

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    +Bre&nancy

    /dded ener&y cost of wei&ht@bearin&locomotion durin& pre&nancy results

    primarily from)/dditional wei&ht transported

    *educed economy of e%ort fromencumbrance

    of fetal tissue

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    +Classication of Bhysical/ctivities by Ener&yE'penditure:Classifyin& strenuousness of physical tas#s) Intensity how KhardL you perform can be

    based on ?52ma'!

    (uration how lon& you perform activity!

    2:Bhysical /ctivity *atio B/*! Classies wor# as the ratio of ener&y re4uired for a

    tas# to the restin& ener&y re4uirement

    9i&ht wor#) Gp to 3 times the restin&re4uirementDeavy wor#) 6 to ; times restin&

    metabolismMa'imal wor#) 8 times or more above rest

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    +$he ME$

    ME$+ metabolic e4uivalent)Multiples of *M*

    5ne ME$ e4uals restin& o'y&en consumptionor 2>7 m91min for men and 277 m91min forwomen

    E'ercise at 2 ME$s re4uires twice the restin&

    metabolism+ 3 ME$s e4uals three times rest+etc:

    $o consider variations in body si0e+ e'pressME$s in 52per unit body mass

    ME$ e4uals 3:> m91#&1min

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    +=ive@9evel Classication of B/,ased on Ener&y E'penditure

    Leel!nergy !xpen"iture

    #cal min1 L min1 mL #g min1 M!$

    Men

    9i&ht 2:7:7A:< :77::322:8 :8

    Deavy A:>8:8 :>7:88 23:737:6 6:7A:8

    ery heavy 7:72:< 2:772: O2:>7 O3;:< O7:7

    &omen

    9i&ht :>3:< 7:377:68 >: :22:A

    Moderate 3:>>:< 7:A7:78 2:68:; 2:;:>A:< :7:7:;8 2A:23

    Gnduly heavy O8:> O:87 O3 OA:6

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    +(aily *ates of /vera&e Ener&yE'penditure

    $he avera&e man8@>7y! e'pends2877 #cal1d the

    avera&e womene'pends 2277 #cal1d Most people spend nearly

    A>? of their day in li&htener&y e'penditureactivities P;h sleep1lyin&down 6h standin& 2hwal#in&1recreationalactivitiesQ

    'en"er ge an" !nergy !xpen"iture

    ge (y)

    !nergy!xpen"iture

    (#cal)

    Males >@; 3777

    8@2< 2877

    2>@>7 2877

    >H 2377

    *emales >@; 2277

    8@2< 2277

    2>@>7 2277

    >H 877

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    +Ener&y Cost of Dousehold+Industrial+ " *ecreational/ctivitiesSee /ppendi' =+ online at

    http)11thepoint:lww:com1m##;e

    for e'amples of ener&y e'penditures e'pressed

    as #cal1#& body mass for common householdactivities+ selected industrial tas#s+ and popular

    recreational and sports activities

    http://thepoint.lww.com/mkk8ehttp://thepoint.lww.com/mkk8e
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    +In-uence of ,ody Mass

    Increases in body mass raise ener&y e'pendedin many activities+ especially in wei&ht@bearin&e'ercise

    ith wei&ht@supported e'ercise e:&: stationary cyclin&!+the in-uence of body mass on ener&y cost decreasesconsiderably

    =or overwei&ht persons+ wei&ht@bearin& e'ercise&enerates a considerablecaloric e'penditure

    E'pressin& ener&y cost per #& of body massreducesener&y cost di%erences betweenindividuals re&ardless of a&e+ race+ &ender+ andbody mass

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    +*elationship,etween,ody Mass "5'y&en

    Consumption(urin&al#in&

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    +Deart *ate to Estimate Ener&yE'penditure

    =or each individual+ heart rate D*! and 52

    relate linearly over theran&e of e'erciseintensities to about ;7? of ma'imum

    E'ercise D* can provide an estimate of 52and thus

    ener&y e'penditure durin&aerobic e'ercise

    Many factors independent of and in addition toe'ercise limit the use of e'ercise D* to estimate

    ener&y e'penditure environmental temperature+ emotions+ food inta#e+

    body position+ musculature+ continuous vs:discontinuous mode+ static or dynamic movements

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    +E'ample

    9inear relationshipbetween heart rate"52for two female

    bas#etball players

    of di%erent aerobictness levels

    Note di%erences inthe slopes of thetwo lines+ indicatin&

    di%erent tnesslevels yellow linemore t person!