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Begins at puberty
Continues throughout
life
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Triggered
by FSH
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2) Enzymes released by exocytosis
) !crosomal process gro"s and binds to a receptor on #itelline layer$) Fusion of membranes causes opening of %a& channels
resulting in depolarization of membrane 'fast bloc( to
polyspermy)
) Entry of sperm causes cortical reaction* Signal transduction
causes Ca& release from E+ "hich causes cortical granules released
by exocytosis* Causes a s"elling and hardening of #itilline layer'becomes sperm proof fertiliztion membrane)
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Clea#age
,rapid cell di#isions creating
many small cells'basically s(ip -. and -2
phases)
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/n humans fertilization and clea#age occurs in the
o#iduct
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Some animals 'frogs) ha#e
une0ual clea#age
1ol( concentrates in #egetal
pole,creates axes of body
,dorsal #s #entral
,anterior #s posterior
,left #s* right
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-astrulation
Formation of layered embryo
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Gastrulation
"It is not birth, marriage, or death, but gastrulation, which is truly
the most important time in your life."
e"is 3olpert '.456)7uring gastrulation8 cell mo#ements result in a massi#e
reorganization of the embryo from a simple spherical ball of cells8 the
blastula8 into a multi,layered organism* 7uring gastrulation8 many of
the cells at or near the surface of the embryo mo#e to a ne"8 moreinterior location*
The primary germ layers'endoderm8 mesoderm8 and ectoderm) are
formed and organized in their proper locations during gastrulation*
Endoderm8 the most internal germ layer8 forms the lining of the gut
and other internal organs* Ectoderm8 the most exterior germ layer8
forms s(in8 brain8 the ner#ous system8 and other external tissues*
Mesoderm8 the middle germ layer8 forms muscle8 the s(eletal
system8 and the circulatory system*
N t h d f
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9granogenesis
,first to form are:,notochord
,neural tube
,somites
Notochordfrom
dorsal mesoderm ;ust
abo#e archenteron
,forms
#ertebral column
%eural tube from
neural plate that
rolls o#er on
itself,forms C%S
Somites : bloc(s of
mesoderm that run
along the notochord ,gi#e rise to
#ertebrae and
muscles of axial
'trun() s(eleton
#ideo
http://biology.kenyon.edu/courses/biol114/Chap14/Chapter_14.htmlhttp://biology.kenyon.edu/courses/biol114/Chap14/Chapter_14.html7/26/2019 Development 05
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Classic example of organogenesis and
induction in the de#elopment of the
#ertebrate eye
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3ill form embryo and embryonic
membranes
Secretes enzymes that brea( do"n endometrium8 bathing
blastocyst in blood
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Chorion surrounds all
other membranes
!mnion forms
amniotic sac
1ol( sac contains
no yol( 'early site
of blood cell
production)
!llantosis forms
umbilical chord
$ extaembryotic membranes
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$ "ee(s
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"ee(s
6 "ee(s
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6 "ee(s
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.6 "ee(s