Development 05

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    Begins at puberty

    Continues throughout

    life

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    Triggered

    by FSH

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    2) Enzymes released by exocytosis

    ) !crosomal process gro"s and binds to a receptor on #itelline layer$) Fusion of membranes causes opening of %a& channels

    resulting in depolarization of membrane 'fast bloc( to

    polyspermy)

    ) Entry of sperm causes cortical reaction* Signal transduction

    causes Ca& release from E+ "hich causes cortical granules released

    by exocytosis* Causes a s"elling and hardening of #itilline layer'becomes sperm proof fertiliztion membrane)

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    Clea#age

    ,rapid cell di#isions creating

    many small cells'basically s(ip -. and -2

    phases)

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    /n humans fertilization and clea#age occurs in the

    o#iduct

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    Some animals 'frogs) ha#e

    une0ual clea#age

    1ol( concentrates in #egetal

    pole,creates axes of body

    ,dorsal #s #entral

    ,anterior #s posterior

    ,left #s* right

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    -astrulation

    Formation of layered embryo

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    Gastrulation

    "It is not birth, marriage, or death, but gastrulation, which is truly

    the most important time in your life."

    e"is 3olpert '.456)7uring gastrulation8 cell mo#ements result in a massi#e

    reorganization of the embryo from a simple spherical ball of cells8 the

    blastula8 into a multi,layered organism* 7uring gastrulation8 many of

    the cells at or near the surface of the embryo mo#e to a ne"8 moreinterior location*

    The primary germ layers'endoderm8 mesoderm8 and ectoderm) are

    formed and organized in their proper locations during gastrulation*

    Endoderm8 the most internal germ layer8 forms the lining of the gut

    and other internal organs* Ectoderm8 the most exterior germ layer8

    forms s(in8 brain8 the ner#ous system8 and other external tissues*

    Mesoderm8 the middle germ layer8 forms muscle8 the s(eletal

    system8 and the circulatory system*

    N t h d f

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    9granogenesis

    ,first to form are:,notochord

    ,neural tube

    ,somites

    Notochordfrom

    dorsal mesoderm ;ust

    abo#e archenteron

    ,forms

    #ertebral column

    %eural tube from

    neural plate that

    rolls o#er on

    itself,forms C%S

    Somites : bloc(s of

    mesoderm that run

    along the notochord ,gi#e rise to

    #ertebrae and

    muscles of axial

    'trun() s(eleton

    #ideo

    http://biology.kenyon.edu/courses/biol114/Chap14/Chapter_14.htmlhttp://biology.kenyon.edu/courses/biol114/Chap14/Chapter_14.html
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    Classic example of organogenesis and

    induction in the de#elopment of the

    #ertebrate eye

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    3ill form embryo and embryonic

    membranes

    Secretes enzymes that brea( do"n endometrium8 bathing

    blastocyst in blood

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    Chorion surrounds all

    other membranes

    !mnion forms

    amniotic sac

    1ol( sac contains

    no yol( 'early site

    of blood cell

    production)

    !llantosis forms

    umbilical chord

    $ extaembryotic membranes

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    $ "ee(s

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    "ee(s

    6 "ee(s

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    6 "ee(s

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    .6 "ee(s