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CURRICULUM VITAE LORI K. SHEERAN, Ph.D.
Department of Anthropology and Museum Studies
Primate Behavior and Ecology Program
Central Washington University
400 East University Way
Ellensburg, WA 98926-7544
(509)963-1434
(509)963-3215
University Education
1993 Ph.D. Anthropology, Ohio State University
1987 M.A. Anthropology, Ohio State University
1984 B.S. Biology, Magna cum Laude, Wright State University
Research Specialization
Tibetan macaques, gibbons, tourism effects on primate behavior
Publications
Peer-Reviewed Academic Book Series
2001 Galdikas BMF; Briggs N; Sheeran LK; Shapiro GL; Goodall J. (co-editors) All Apes Great and
Small, Volume I: African Apes. Developments in Primatology: Progress and Prospects, Russell
H. Tuttle series editor. New York: Kluwer Publishers. 298 pages.
1995 Nadler RL; Galdikas BMF; Sheeran LK; Rosen N. (editors) The Neglected Ape. New York:
Plenum Publishers. 300 pages.
Peer-Reviewed Articles (student co-authors underlined)
Revising Dura E; Sheeran LK; Ruppert N; Arango C; Blue S. Mother-infant interactions in a wild
population of Southern pig-tailed macaques (Macaca nemestrina). Asian Primates. Accepted
pending revision.
Revising Fratellone G; Li JH; Sun LX; Sheeran LK; Wagner RS; Wang X. Social connectivity facilitates
collective decisions making in wild Tibetan macaques (Macaca thibetana). Primates. Accepted
pending revision.
Revising Rowe A; Sun LK; Sheeran LK; Li JH Wagner S; Xia DP; Uhey DA; Chen R. Collective decision
making in Tibetan macaques: How followers affect the rules and efficiency of group movement.
Animal Behavior. Accepted pending revision.
In Press Sun BH; Gu ZY; Wang X; Huffman MA; Garber PA; Sheeran LK; Zhang D; Zhu Y; Xia DP; Li
JH. Season, age, and sex affect the fecal mycobiota of free-ranging Tibetan macaques (Macaca
thibetana). American Journal of Primatology.
Updated May 2018
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2018 Wright K; Mayhew JA; Sheeran LK; Funkhouser J; Wagner RS; Sun LS; Li JH. Accepted
pending revision. Playing it cool: Social play, play signals, and bout termination by juvenile
Tibetan macaques (Macaca thibetana). Zoological Research 39(4):274:283.
2017 Xia DP; Wang X; Zhang QZ; Sun BH; Sun LX; Sheeran LK; Li JH. Progesterone levels and
sexual behavior in free-ranging Macaca thibetana. Mammal Research 10.1007/s13364-017-0342-
0 (8 pages).
2017 Sun LX; Xia DP; Sun S; Sheeran LK; Li JH. It pays to wait: Upward social mobility key to long-
term sustainability of a primate society. Scientific Reports 7(7082):1-8.
2016 Bernstein S; Sheeran LK; Wagner RS; Li JH; Koda H. The vocal repertoire of Tibetan macaques
(Macaca thibetana): A quantitative classification and a comparative perspective. American
Journal of Primatology 78:937-949. (2016 impact factor 2.435, ISI Journal Citation Reports
Ranking for 2014 17/153 [Zoology])
2016 Sun BH; Wang X; Bernstein S; Huffman MA; Xia DP; Gu ZY; Chen R; Sheeran LK; Wagner
RS; Li JH. Marked variation between winter and spring gut microbiota in free-ranging Tibetan
macaques (Macaca thibetana). Scientific Reports 6:26035. DOI: 10.1038/srep26035. (2014
Impact factor 5.578)
2016 Wang X; Sun LX; Sheeran LK; Sun BH; Zhang QX: Zhang D; Xia DP; Li JH. Social rank
versus affiliation: Which is more related to leadership during group movements in
Tibetan macaques (Macaca thibetana)? American Journal of Primatology 78:816-824. (2016
impact factor 2.435, ISI Journal Citation Reports Ranking for 2014 17/153 [Zoology])
2014 Bauer B; Sheeran LK; Matheson MD; Li JH; Wagner RS. Choice of infant bridging and female
affiliation partners by male Tibetan macaques (Macaca thibetana). Zoological Research.
35(3):222-230.
2014 Pritchard A; Sheeran LK; Gabriel K; Li JH; Wagner RS. Behaviors that predict personality
components in adult free-ranging Tibetan macaques (Macaca thibetana). Current Zoology
60(3):362-372. (2014 impact factor 1.797; top 20% (31/152) among 2013 JCR ZOOLOGY
journals)
2014 Usui L; Sheeran LK; Li JH; Sun LX; Wang X; Pritchard AP; DuVall-Lash A; Wagner RS. Park
rangers’ behaviors and their effects on tourists and Tibetan macaques (Macaca thibetana) at Mt.
Huangshan, China. Animals 4(3):546-561; doi:10.3390/ani4030546
2013 Jiang T; Li J; Sheeran LK; Zhu Y; Sun B; Xia DP; Wang X. Homosexual mounting in wild
Tibetan macaques (Macaca thibetana) at Huangshan, China. Life Science Journal 10(1):3192-
3197.
2013 Self S; Sheeran LK; Matheson MD; Li JH; Pelton O; Harding S; Wagner RS. Tourism and
infant-directed aggression in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
Anthrozoös 26(3):435-444. (2011 journal impact factor 0.86)
2012 Cheyne SM; Höing A; Rinear J; Sheeran LK. Sleeping site selection by agile gibbons: the
influence of tree stability, fruit availability, and predation risk. Folia Primatologica 83:299-311.
(2013 journal impact factor 1.000)
2012 Simons ND; Lorenz JG; Sheeran LK; Li JH; Xia DP; Wagner RS. Noninvasive saliva collection
for DNA analyses from free-ranging Tibetan macaques (Macaca thibetana). American Journal of
Primatology 74:1064-1070. (2013 journal impact factor 2.221)
Updated May 2018
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2010 Ruesto LA; Sheeran LK; Matheson MD; Li JH; Wagner RS. Tourist behavior and decibel levels
correlate with threat frequency in Tibetan macaques (Macaca thibetana) at Mt. Huangshan,
China. Primate Conservation 2010 (25): 99-104.
2010 Xia DP; Li JH; Zhu Y; Sun BH; Sheeran LK; Matheson MD. Seasonal variation and
synchronization of sexual behaviors in free-ranging male Tibetan macaques (Macaca thibetana)
at Huangshan, China. Zoological Research 31(5):509-515.
2009 McCarthy MS; Matheson MD; Lester JD; Sheeran LK; Li JH; Wagner RS. Sequences of Tibetan
Macaque (Macaca thibetana) Behaviors and Tourist Behaviors at Mt. Huangshan, China. Primate
Conservation 24:145-151.
2006 Matheson MD; Sheeran LK; Wagner RS; Li JH. Tourist impact on Tibetan macaques.
Anthrozoös 19(2):158-168. (2008 journal impact factor 0.612)
2005 Mootnick AR; Baker E; Sheeran LK. Familiarity during immaturity: implications for the captive
propagation of gibbons. International Journal of Primatology 26(6):1417-1433. (2011 journal
impact factor 1.538)
1998 Sheeran LK; Zhang YZ; Poirier FE; Yang DH. Preliminary report on black gibbon (Hylobates
concolor) behavior. Tropical Biodiversity 5(2):113-125.
1994 Sheeran LK; Poirier FE. The black-crested gibbon of China. Primate Conservation 11:20-22.
1994a Jiang X; Ma S; Wang Y; Sheeran LK; Poirier FE. Human encounter and predator avoidance
behavior in black-crested gibbon, Hylobates concolor. Acta Anthropologica Sinica 13(2):182-
188.
1994b Jiang X; Ma S; Wang Y; Sheeran LK; Poirier FE. The mating system of black-crested gibbons
(Hylobates concolor) and its relationships with ecology, behavior, and phylogeny. Acta
Anthropologica Sinica 13(2):344-352.
1994c Jiang X; Ma S; Wang Y; Sheeran LK; Poirier FE. Group size and composition of black-crested
gibbon (Hylobates concolor). Zoological Research 15(2):15-22.
Manuscripts in Preparation for Submission to Peer-Reviewed Journals (student co-authors underlined)
Under Review Funkhouser E; Mayhew JA; Sheeran LK; Mulcahy JB; Li JH. Investigations of context-
dependent dominance in captive chimpanzees (Pan troglodytes) and wild Tibetan macaques
(Macaca thibetana). Nature: Scientific Reports.
Under Review Funkhouser E; Mayhew JA; Mulcahy JB; Sheeran LK. Human caregivers occupy prominent
positions in captive chimpanzee (Pan troglodytes) social networks. Journal of Applied Animal
Welfare Science.
In Prep Bernstein S; Sheeran LK; Wagner RS; Li JH; Koda H. A sex-specific audience effect on
female Tibetan macaque copulation calls. Animal Behavior.
In Prep Wang X; Sun LX; Garber PA; Sheeran LK; Sun BH; Kyes RC; Xia DP; Li JH. Triadic
associations predict female leadership of group movement in Tibetan macaques (Macaca
thibetana).
Updated May 2018
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Peer-Reviewed Abstracts (student co-authors underlined)
2015 Bell MC; Sheeran LK; Skollár G. Javan gibbons (Hylobates moloch) vary gesture use by
recipient’s attentional state at the Gibbon Conservation Center, CA. American Journal of
Primatology 77(S1):61.
2015 Clifton CJ; Sheeran LK; Wagner RS; Li JH. Female-female bridging behavior in Tibetan
macaques (Macaca thibetana) at Mt. Huangshan, China. American Journal of Primatology
77(S1):63.
2015 Fratellone GP; Sun L; Sheeran LK; Wagner RS; Li JH. Leadership in the collective movements
of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. American Journal of
Primatology 77(S1):116.
2014 Pritchard A; Sheeran LK; Garbiel K; Li JH; Wagner RS. Discriminating behaviors to measure
personality traits in free-ranging Tibetan macaques (Macaca thibetana). American Journal of
Physical Anthropology 153(S58):211-212.
2013 Dunayer ES; Berman CM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Short term steepness
changes in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. American Journal of
Primatology 75(S1):89.
2012 Dunayer ES; Matheson MD; Sheeran LK; Beck D; Li JH; Wagner RS. Does tourism have a
market effect on the grooming for tolerance interchange in Tibetan macaques (Macaca thibetana)
at Mt. Huangshan, China? American Journal of Primatology 74(S1):27.
2012 Hendershott R; Matheson M; Sheeran L; Wagner R; Li J. Sociosexual behaviors of Tibetan
macaques (Macaca thibetana). American Journal of Primatology 74(S1):28.
2012 Kibler-Campbell A; Thompson C; Sheeran LK. Causal inference examined in Cebus apella
through a hammer-and-anvil task. American Journal of Primatology 74(S1):29.
2011 Simons ND; Lorenz JG, Sheeran LK; Matheson MD; Li JH; Wagner RS. Noninvasive saliva
collection from free-ranging Tibetan macaques (Macaca thibetana) in Huangshan, China.
American Journal of Primatology 73(S1):96.
2010 Sheeran LK; Matheson M; Li JH; Wagner, S. Age and sociability in free-living Tibetan
macaques. Symposium on University Research and Creative Expression, Central Washington
University, Ellensburg, WA.
2010 Rinear J; Sheeran LK; Cheyne SM. Effects of tree characteristics on sleeping tree choice in
Bornean agile gibbons (Hylobates albibarbis). American Journal of Primatology 72(S1):37.
2010 Self S; Sheeran LK; Matheson M; Li JH; Harding S; Pelton O; Wagner RS. Tourist density and
infant-directed aggression in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
American Journal of Primatology 72(S1):59-60.
2009 Sheeran LK; Matheson M; Li JH; Wagner RS. Age and potential social partners in Tibetan
macaques (Macaca thibetana). Symposium on University Research and Creative Expression,
Central Washington University, Ellensburg, WA.
2008 Jones AM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Aggression and habituation toward
humans in two troops of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
American Journal of Primatology 70(S1):61.
2008 Mack HM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Grooming behavior of Tibetan
macaques (Macaca thibetana) in the presence of tourists at Mt. Huangshan, China. American
Journal of Primatology 70(S1):59.
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2008 Matheson MD; Mack HM; Sheeran LK; Yenter TA; Schulz P. Captive ex-pet Macaca
fascicularis use hair and dental floss to floss their teeth. American Journal of Primatology
70(S1):56.
2008 Yenter TA; Matheson MD; Sheeran LK; Li J; Wagner RS. Self directed behaviors, tourist
density and proximity in a free living population of Tibetan macaques (Macaca thibetana) at an
ecotourism destination in Anhui Province, China. American Journal of Primatology 70
(S1):52.
2007 Matheson MD; Hartel J; Whitaker C; Sheeran LK; Li JH; Wagner RS. Self-directed behavior
correlates with tourist density in free-living Tibetan macaques (Macaca thibetana) at the Valley
of the Wild Monkeys, Mt. Huangshan, China. American Journal of Primatology 69 (S1):41-42.
2007 McCarthy M.S; Matheson MD; Lester JD; Sheeran LK; Li JH; Wagner RS. Sequences of
Tibetan macaque (Macaca thibetana) behaviors and tourist behaviors at Mt. Huangshan, China.
American Journal of Primatology 69 (S1):42.
2002 Klein C; Sheeran LK. Survival rates in captive lar gibbons and siamangs. Caring for Primates.
Abstracts of the XIXth Congress of the International Primatological Society. Beijing:
Mammalogical Society of China, Pp. 125.
2001 Sheeran LK. Ethical issues in primatological research: Captive settings. American Journal of
Primatology 54 (S1):27-28.
2001 Shur M; Sheeran LK. Affiliation behaviors among captive female chimpanzees in a naturalistic
captive environment. American Journal of Primatology 54 (S1):49.
1994 Mootnick AR; Baker E; Sheeran L. Inbreeding in captive gibbons (Hylobates). American
Journal of Primatology 31(4):22.
1993 Sheeran LK. Behavior of wild black gibbons in Yunnan Province, PRC. American Journal of
Primatology 30(4):348.
1992 Sheeran LK. Intergroup dueting behavior of black crested gibbons during two seasons. American
Journal of Primatology 25(2):125.
Conference Proceedings
1995 Ogden JO; Perkins LA; Sheeran LK. (co-editors) Partial Proceedings from the International
Orang utan Conference. San Diego, CA: Zoological Society of San Diego. 270 pages.
Encyclopedia Entries, Book Chapters, and Reviews (student co-authors underlined)
2018 Sheeran LK. Review of The New Chimpanzee. Choice connect 56(1):
http://choiceconnect.org/review/magprint/209766
2018 Sheeran LK. Review of Bonobos: Unique in Mind, Brain, and Behavior. Choice connect 55(11):
http://www.choiceconnect.org/webclipping/208468/jp-d80-f54g2oru1wvny6l--
9dd885ffididbhi7-pwn7ecrzp
2018 Sheeran LK. Review of The Great Apes. Choice connect 54(11):
http://www.choiceconnect.org/review/magprint/208469
2017 Sheeran LK. Review of Monkeytalk. Choice connect 54(11):
http://www.choiceconnect.org/review/magprint/203095
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2017 Sheeran LK. Review of Ethnoprimatology: Primate Conservation in the 21st Century. Choice
connect 54(7): http://www.choiceconnect.org/review/magprint/201010
2016 Sheeran LK. Review of Primate Behavior and Human Origins. Choice connect 54(3):
http://www.choiceconnect.org/review/magprint/198872
2016 Sheeran LK. Review of Mahale Chimpanzees: 50 Years of Research. Choice connect 53(8):
http://www.choiceconnect.org/review/magprint/196527
2016 Sheeran LK. Review of Primates in the Real World: Escaping Primate Folklore and Creating
Primate Science. Choice connect 53(8): http://www.choiceconnect.org/review/magprint/195310
2016 Sheeran LK. Review of Dispersing Primate Females: Life History and Social Strategies in Male-
Philopatric Species. Choice connect 53(5):
http://www.choiceconnect.org/review/magprint/194066
2015 Sheeran LK. Review of Storytelling Apes. Choice connect 53(2):
http://www.choiceconnect.org/review/magprint/192576
2015 Sheeran LK. Review of Howler Monkeys: Adaptive Radiation, Systematics, and Morphology.
Choice connect 52(11): http://www.choiceconnect.org/review/magprint/190182
2015 Sheeran LK. Review of Extractive Industries and Ape Conservation. Choice connect 52(9):
http://www.choiceconnect.org/review/magprint/188129
2013 Sheeran LK. Tibetan macaques. In: Handbook of the Mammals of the World, Volume 3 Primates,
R.A. Mittermeier and D.E. Wilson (eds.). Lnyx Edicions, Rockville Center, NY, Pp. 642-643.
2013 Sheeran LK. Review of The Primate Mind: Built to Connect with Other Minds. Journal of
Anthropological Research 69: 120-121.
2010 Sheeran LK. Lack of seasonal influence on duet duration in black gibbons (Nomascus concolor
jingdongensis). In: Collected Readings in Biological Anthropology in Honor of Professor LS
Penrose and Dr. Sahrah B. Holt, Sudip Datta Banak (ed.). Unas Letras Industria Editorial,
Merida, Yucata, Pp. 359-367.
2010 Sheeran LK; Matheson MD; Li JH; Wagner RS. A preliminary analysis of aging and potential
social partners in Tibetan macaques (Macaca thibetana). In: Collected Readings in Biological
Anthropology in Honor of Professor LS Penrose and Dr. Sahrah B. Holt, Sudip Datta Banak
(ed.). Unas Letras Industria Editorial, Merida, Yucata, Pp. 349-358.
2006 Jensvold ML; Sheeran LK. Ape and human cognition. In: Encyclopedia of Anthropology, H.J.Birx (ed.). Sage Publications, Pp. 207-212.
2005 Sheeran LK. Apes no less. Zoo View Winter, 2005:16-17.
2002 Sheeran LK; Shur MA. Review of Tree of Origin: What Primate Behavior Can Tell Us about
Human Social Evolution. American Journal of Human Biology 14(1):82-84.
2001 Sheeran LK. Preparing students for primatological fieldwork: Reducing risks. The Apes:
Challenges for the 21st Century Brookfield, Illinois, Pp. 250-258.
2001 Sheeran LK. Report on the Gibbon Conservation Workshop. The Apes: Challenges for the 21st
Century Brookfield, Illinois, Pp. 349-350.
2001 Perkins L; Chivers D; Eudey A; Galdikas BMF; Sheeran L; Supriatna J; van Schaik C. Asian ape
roundtable discussion. The Apes: Challenges for the 21st Century Brookfield, Illinois, Pp. 352.
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1999 Mootnick AR; Sheeran LK. A description of the pileated gibbon (Hylobates pileatus).
Australasian Primatology 13(3):14-16.
1997 Sheeran LK. Asian apes: Gibbons. In History of Physical Anthropology: An Encyclopedia,
Spencer F. (editor). New York: Garland Press, Pp. 112-118.
1997 Sheeran LK. A review of gibbon behavior, ecology, and evolution. In: Beacham’s International
Threatened, Endangered, and Extinct Species, Walton Beacham (ed.). Washington, D.C.:
Beacham Publishers, Pp. 46-53.
1997 Sheeran, LK; Mootnick AR. The crested gibbon. In: Beacham’s International Threatened,
Endangered, and Extinct Species, Walton Beacham (ed.). Washington, D.C.: Beacham
Publishers, Pp. 57-60
1996 Lan DY; Sheeran LK. The status of black bibbons at Xiaobahe, Wuliang Mountains, Yunnan
Province, China. Asian Primates 5(1-2):2-4.
1995 Sheeran LK. Behavior of wild black gibbons (Hylobates concolor jingdongensis). In Primate
Research and Conservation, Xia W; Zhang Y (editors). Beijing: China Forestry Publishing
House, Pp. 221-225.
1995 Mootnick AR; Sheeran LK. The future of the gibbon. The Keeper 5:26-27.
1995 Rosen N; Sheeran LK; Nevadomsky J. Report on the International Conference on the Orang
Utan: The Neglected Ape. Current Anthropology 36(2):353-354.
1995 Zhang Y; Quan G; Yang D; Liu Z; Sheeran LK. Population parameters of the black gibbon in
China. IN: Primate Research and Conservation, Wuping X: Zhang Y (eds.). Beijing: China
Forestry Publishing House, Pp. 203-221.
1994 Zhang YZ; Sheeran LK. Current status of the Hainan black gibbon (Hylobates concolor
hainanus). Asian Primates 3(3-4):3.
1993 Sheeran LK. Field study on the black gibbons in Yunnan and Hainan. Chinese Primate Research
and Conservation News 2(1):2-3.
1988 Poirier FE; Sheeran LK. Review of P.C. Else and J. Lee Primate Ecology and Conservation.
American Journal of Physical Anthropology 78(4):626-8.
Reports
2012-16 Annual reports submitted to National Science Foundation, International Research Experiences
for Students (NSF-IRES) grant.
2007-10 Annual and final reports submitted to National Institutes of Health, Bridges to the
Baccalaureate grant.
Grants Funded or Under Review
2018 LSB Leakey Foundation Grant, Anthropogenic Change on Aacaca thibetana (co-PIs Sofia
Bernstein, Jessica Mayhew, Lori Sheeran), under review, $12,465
2017 CWU Proposal Preparation Grant, $3,445 (co-PIs Lori Sheeran, Jessica Mayhew)
Updated May 2018
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2010-16 National Science Foundation International Research Experiences for Students (NSF-IRES),
$149,930 (PI Lori Sheeran; Co-PIs Audrey Huerta, Megan Matheson, Steve Wagner); 3 year
grant, one year no-cost extension, and one year NSF-approved extension
2015 CWU Faculty Summer Graduate Program Development Award, $4,726
2009 CWU Faculty Summer Writing Award, $2,000
2006 National Institutes of Health, Bridges to the Baccalaureate: Linking YVCC and CWU, $230,000
(PI Lori Sheeran; Co-PI Mary Lee Jensvold); 3 year grant
2005 CWU Faculty Seed Grant, $1,900 (Co-PIs Lori Sheeran and Mary Lee Jensvold)
2004 OISP and ISPAC, International Small Grant, $500 (Co-PIs Lori Sheeran, Megan Matheson,
Steve Wagner)
2004 CWU Foundation, Instruction/Research Equipment Proposal, $7,041 (PI Lori Sheeran)
2004 CWU Foundation, Len Thayer Small Grant, $500 (PI Lori Sheeran)
2004 CWU Foundation, Len Thayer Small Grant, $750 (Co-PIs Lori Sheeran, Megan Matheson,
Steve Wagner, Tracy Andrews)
2001 Center for Spatially Integrated Social Science/The National Science Foundation, to attend
workshop Accessibility in Space and Time: A GIS Approach, Ohio State University, $500, 16-20
July (PI Lori Sheeran)
1998 The National Science Foundation ILI-IP Grant, $49,200 ($98,400 with institutional match) (PI
Susan Parman, Co-PI Lori Sheeran)
1996 The California State University, Fullerton Foundation Faculty Enhancement and Instructional
Improvement Award, $150 and 3 workload units (PI Lori Sheeran)
1996 California Campus Compact Grant, $2,500 (Co-PIs Lori Sheeran, Jacob Pandian, Susan Parman)
1995 American Society of Primatologists Conservation Grant, $500 (Co-PIs Lori Sheeran, Zhang
Yongzu)
1995 Primate Conservation Incorporated, $3,500 (Co-PIs Lori Sheeran, Zhang Yongzu)
1995 The California State University, Fullerton Foundation Faculty Enhancement and Instructional
Improvement Award, $500 (PI Lori Sheeran)
1994 The California State University, Fullerton Foundation, Junior Research Award, $3,500 (PI Lori
Sheeran)
1994 The California State University, Fullerton Foundation, Affirmative Action Award, 3 workload
units (PI Lori Sheeran)
1993 The National Geographic Society, $20,288 (PI Zhang Yongzu, Co-PI Lori Sheeran)
1990 The National Geographic Society, $18,000 (PI Frank Poirier, Co-PI Lori Sheeran)
1990 The Charles A. Lindbergh Fund, $10,000 (PI Frank Poirier, Co-PI Lori Sheeran)
1990 The Wenner Gren Foundation for Anthropological Research, $5,000 (PI Lori Sheeran)
1990 The Ohio State University Graduate Research Award, $1,500 (PI Lori Sheeran)
Updated May 2018
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Professional Presentations
International Conferences (student co-authors underlined)
2017 Sheeran LK. Female-female bridging at the Valley of the Wild Monkeys, Mt. Huangshan, China,
Invited lecture, Huangshan International Symposium, Nonhuman Primates: Insights in to Human
Behavior and Society, Hefei, Anhui Province, China, 21-25 July.
2017 Sheeran LK; Mayhew JA; Sun LX; Wagner RS; Xia DP; Li JH. Duan Wei Hou Research
Project: Fieldwork with Tibetan macaques, Invited lecture, University of Victoria, BC, Canada,
16 January.
2016 Bernstein, SK; Sheeran LK; Wagner RS; Li JH; Koda H. A quantitative analysis of the vocal
repertoire of a free-ranging group of Tibetan macaques (Macaca thibetana) in the Valley of the
Wild Monkeys, Mt. Huangshan, China, Invited lecture, Joint Meeting of the International
Primatological Society and the American Society of Primatologists, Chicago, 21-27 August.
2016 Sheeran LK. Human and Tibetan macaque (Macaca thibetana) interactions in the Valley of the
Wild Monkeys, Huangshan, China. 5th International Symposium on Primatology and Wildlife
Science, Inuyama, Japan, 3-6 March.
2013 Dunayer ES; Berman CM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Short term steepness
changes in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. 36th Annual Meeting
of the American Society of Primatologists, San Juan, Puerto Rico, 19-22 June.
2013 Usui R; Sheeran LK; Sun L; Li JH; Wang X; Pritchard A; Duvall-Lash A; Wagner RS.
Behaviors of park rangers and their effects on Tibetan macaques (Macaca thibetana) at Mt.
Huangshan, China. MPJ2013: Joint Meeting of the 29th Japan Primate Society and the 2013
Mammal Society of Japan, Okayama, Japan, 6-9 September.
2012 Jones AM; Matheson MD; Sheeran LK; Li JH; Wagner RS. The use of infants in triadic
interactions among adult male and female Tibetan macaques (Macaca thibetana): testing the
agonistic buffering hypothesis. 24th Congress of the International Primatological Society, Cancun,
Mexico, 12-17 August.
2011 Winters SA; Sheeran LK; Matheson MD; Li JH; Wagner R. Maternal response to infant
disability: a case study of traumatic injury to an infant Tibetan macaque (Macaca thibetana).
Lemelson/Society for Psychological Anthropology Conference, Spokane, WA, 19-21 May.
2010 Matheson MD; Jones AM; Sheeran LK; Li JH; Wagner RS. Interactions between park wardens
and Tibetan macaques (Macaca thibetana) at a tourist site in Huangshan, China. 23rd Congress of
the International Primatological Society, Kyoto, Japan, 12-18 September.
2007 Sheeran LK. The anthropology of aging. CWU-Anhui University Interdisciplinary Research
Symposium, Huangshan, China, 3 August.
2004 Sheeran LK. The anthropological perspective on primatology: Modeling human behavior and
evolution. CWU-Anhui University Interdisciplinary Research Symposium, Hefei, China, 13
August.
2002 Klein C; Sheeran LK. Differential mortality in captive lar gibbons and siamangs. XIXth Meeting
of the International Primatological Society, Beijing, China, 4-9 August.
2000 Sheeran LK. Preparing students for primatological fieldwork: Reducing risks. The Apes:
Challenges for the 21st Century, Brookfield Zoo, Chicago, IL, 10-13 May.
Updated May 2018
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1998 Sheeran LK. Black gibbon (Hylobates concolor) group size: Implications for gibbon mating
systems. Great Apes of the World Conference II, Kuching, Malaysia, 29-31 July
1998 Mootnick AR; Sheeran LK. An overview of gibbon (Hylobates) taxonomy for use by breeding
facilities. Great Apes of the World Conference II, Kuching, Malaysia, 29-31 July.
1994 Zhang Y; Quan G; Yang D; Liu Z; Sheeran LK. Population parameters of the black gibbon.
VIIIth Meeting of the International Primatological Society, Bali, Indonesia, 12 August.
National Conferences (student co-authors underlined)
2017 Sun L; Xia DP; Sun S; Sheeran LK; Li JH. It pays to wait: Intragenerational social mobility
key to long-term stability in Tibetan macaque society. Evolution 2017 (Joint meeting of
the American Society of Naturalists, the Society for the Study of Evolution, and the
Society of Systematic Biologists, Portland, OR, 23-27 June.
2015 Bell MC; Sheeran LK; Skollár G. Javan gibbons (Hylobates moloch) vary gesture use by
recipient’s attentional state at the Gibbon Conservation Center, CA. 38th Annual Meeting of the
American Society of Primatologists, Bend, OR, 17-20 June.
2015 Clifton CJ; Sheeran LK; Wagner RS; Li JH. Female-female bridging behavior in Tibetan
macaques (Macaca thibetana) at Mt. Huangshan, China. 38th Annual Meeting of the American
Society of Primatologists, Bend, OR, 17-20 June.
2015 Fratellone GP; Sun L; Sheeran LK; Wagner RS; Li JH. Leadership in the collective movements
of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. 38th Annual Meeting of the
American Society of Primatologists, Bend, OR, 17-20 June.
2014 Pritchard A; Sheeran LK; Garbiel K; Li JH; Wagner RS. Discriminating behaviors to measure
personality traits in free-ranging Tibetan macaques (Macaca thibetana). 83rd Annual Meeting of
the American Association of Physical Anthropologists, Calgary, Alberta, 8-12 April.
2012 Dunayer ES; Matheson MD; Sheeran LK; Beck D; Li JH; Wagner RS. Does tourism have a
market effect on the grooming for tolerance interchange in Tibetan macaques (Macaca thibetana)
at Mt. Huangshan, China? 35th Annual Meeting of the American Society of Primatologists,
Sacramento, CA, 20-23 June.
2012 Hendershott R; Matheson M; Sheeran L; Wagner R; Li J. Sociosexual behaviors of Tibetan
macaques (Macaca thibetana). 35th Annual Meeting of the American Society of Primatologists,
Sacramento, CA, 20-23 June.
2012 Kibler-Campbell A; Thompson C; Sheeran LK. Causal inference examined in Cebus apella
through a hammer-and-anvil task. 35th Annual Meeting of the American Society of
Primatologists, Sacramento, CA, 20-23 June.
2011 Simons NA; Lorenz JL; Sheeran LK; Matheson MD; Li JH; Wagner RS. Noninvasive saliva
collection from free-ranging Tibetan macaques (Macaca thibetana) in Huangshan, China. 34th
Annual Meeting of the American Society of Primatologists, Austin, TX, 16-19 September (poster
presentation).
2010 Rinear J; Sheeran LK; Cheyne SM. Effects of tree characteristics on sleeping tree choice in
Bornean agile gibbons (Hylobates albibarbis). 33rd Annual Meeting of the American Society of
Primatologists, Louisville, KY, 15-19 June (poster presentation).
Updated May 2018
11
2010 Self S; Sheeran LK; Matheson M; Li J; Harding S; Pelton O; Wagner RS. Tourist density and
infant-directed aggression in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
33rd Annual Meeting of the American Society of Primatologists, Louisville, KY, 15-19 June.
2008 Jones AM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Aggression and habituation toward
humans in two troops of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. 31st
Annual Meeting of the American Society of Primatologists, West Palm Beach, FL, 18-21 June.
2008 Mack HM; Matheson MD; Sheeran LK; Li JH; Wagner RS. Grooming behavior of Tibetan
macaques (Macaca thibetana) in the presence of tourists at Mt. Huangshan, China. 31st Annual
Meeting of the American Society of Primatologists, West Palm Beach, FL, 18-21 June .
2008 Matheson MD; Mack HM; Sheeran LK; Yenter TA; Schulz P. Captive ex-pet Macaca
fascicularis use hair and dental floss to floss their teeth. 31st Annual Meeting of the American
Society of Primatologists, West Palm Beach, FL, 18-21 June.
2008 Yenter TA; Matheson MD; Sheeran LK; Li JH; Wagner RS. Self-directed behaviors, tourist
density and proximity in a free living population of Tibetan macaques (Macaca thibetana) at an
ecotourism destination in Anhui Province, China. 31st Annual Meeting of the American Society
of Primatologists, West Palm Beach, FL, 18-21 June.
2007 Matheson MD; Hartel J; Whitaker C; Sheeran LK; Li JH; Wagner RS. Self-directed behavior
correlates with tourist density in free-living Tibetan macaques (Macaca thibetana) at the Valley
of the Wild Monkeys, Mt. Huangshan, China. 30th Annual Meeting of the American Society of
Primatologists, Winston-Salem, NC, 20-23 June.
2007 McCarthy M; Matheson MD; Sheeran LK; Lester JD; Li JH; Wagner RS. Sequences of Tibetan
macaque behaviors and tourist behaviors at Mt. Huangshan, China. 30th Annual Meeting of the
American Society of Primatologists, Winston-Salem, NC, 20-23 June.
2001 Shur MA; Sheeran LK. Affiliative behaviors of captive female chimpanzees. 24th Annual
Meeting of the American Society of Primatologists, Savannah, GA, 8-11 August.
1997 Collier MK; Sheeran LK. Participants in Workshop Human to Nonhuman Primate Disease
Transmission. 20th Annual Meeting of the American Society of Primatologists, San Diego, CA,
27 June-1 July.
1994 Mootnick AR; Baker E; Sheeran LK. Inbreeding in captive gibbons (Hylobates). 31st Annual
Meeting of the Animal Behavior Society and 17th Annual Meeting of the American Society of
Primatologists, Seattle, WA, 23-30 July.
1993 Sheeran LK. Behavior of wild black gibbons in Yunnan Province, PRC. 16th Annual Meeting of
the American Society of Primatologists, Sturbridge, MA 18-22 August.
Regional Conferences (student co-authors underlined)
2016 Sheeran LK; Wagner RS; Sun LX; Li JH; Xia DP. Human and Tibetan macaque (Macaca
thibetana) interactions in the Valley of the Wild Monkeys, Huangshan, China. 69th Annual
Northwest Anthropological Conference, Tacoma, WA, 23-26 March.
2016 Voight, B; Sheeran LK; Wang X; Wagner RS; Li JH; Sun LX. A comparison of proximity of
Tibetan macaques (Macaca thibetana) near provisioned food sources at Mt. Huangshan, China.
69th Annual Northwest Anthropological Conference, Tacoma, WA, 23-26 March.
Updated May 2018
12
2016 Wright K; Sheeran LK; Mayhew J; Wagner RS; Sun LX; Li JH. Tibetan macaque play locations
preference and causes of play termination at Mt. Huangshan, China. 69th Annual Northwest
Anthropological Conference, Tacoma, WA, 23-26 March. (poster presentation).
2015 Bell MC; Sheeran LK; Skollár G. Juvenile Javan gibbons (Hylobates moloch) vary gesture use
by recipient’s attentional state at the Gibbon Conservation Center (Santa Clarita, CA). 68th
Annual Northwest Anthropological Conference, Eugene, OR, 26-28 March (poster presentation).
2015 Clifton, CJ; Sheeran LK; Wagner RS; Li JH. Female-female bridging behavior in Tibetan
macaques (Macaca thibetana) at Mt. Huangshan, China. 68th Annual Northwest Anthropological
Conference, Eugene, OR, 26-28 March.
2015 Fratellone GP; Sun L; Sheeran LK; Wagner RS; Li JH. Movement progression in the collective
movements of Tibetan Macaques (Macaca thibetana) at Mount Huangshan, China. 68th Annual
Northwest Anthropological Conference, Eugene, OR, 26-28 March (poster presentation).
2014 Clifton, CJ; Sheeran LK; Wagner RS; Li JH. Female-female bridging behavior in Tibetan
Macaques (Macaca thibetana) at Mt. Huangshan, China. 25th Regional Conference of
Undergraduate Research, M.J. Murdock Charitable Trust, Vancouver, WA 14-15 November
(poster presentation).
2014 Price E; Sheeran LK; Li JH. Vocalizations of Tibetan macaques (Macaca thibetana) at Mt.
Huangshan, China. 67th Annual Northwest Anthropological Conference, Bellingham, WA, 26-29
March.
2014 Staven A; Sheeran LK; Li JH; Wagner RS. Interactions Between tourists and juvenile Tibetan
macaques at Mt. Huangshan, China. Annual Meetings of the Society for Northwestern Vertebrate
Biology, Pasco, WA, 3-7 February.
2013 Link J; Murphy A; Li JH; Sheeran, LK. Maternal investment in free-ranging Tibetan macaques
(Macaca thibetana). 66th Annual Northwest Anthropological Conference, Portland, OR, 27-30
March (poster presentation).
2013 Pritchard AJ; Sheeran, LK; Jensvold ML; Gabriel K; Li JH; Wagner RS. Measuring personality
traits in provisioned Tibetan macaques (Macaca thibetana), Mt. Huangshan, China. 66th Annual
Northwest Anthropological Conference, Portland, OR, 27-30 March (poster presentation).
2012 Bauer B; Sheeran LK; Matheson MD; Wagner R; Li JH. Male Tibetan macaque choice of infant
bridging and female affiliation partners. 65th Annual Northwest Anthropological Conference,
Pendleton, OR, 15-17 March (poster presentation).
2012 Burwell L; Sheeran LK; Matheson MD; Wagner RS; Li JH. Competition for provisioned corn in
Macaca thibetana at Mt. Huangshan, China. 65th Annual Northwest Anthropological Conference,
Pendleton, OR, 15-17 March (poster presentation).
2011 Batts CB; Sheeran LK; Matheson MD; Li JH; Wagner RS. Impact of eco-tourism on infant and
juvenile play behaviors in Tibetan macaques (Macaca thibetana). 20th Regional Conference of
Undergraduate Research, M.J. Murdock Charitable Trust, Seattle, WA 11-12 November (poster
presentation).
2011 Blue M; Sheeran LK; Wagner RS. The effects of matrilineal dominance hierarchies on birth-sex
ratios within a group of free-living Tibetan macaques (Macaca thibetana). 20th Regional
Conference of Undergraduate Research, M.J. Murdock Charitable Trust, Seattle, WA 11-12
November (poster presentation).
Updated May 2018
13
2011 Hendershott RL; Matheson MD; Sheeran LK; Wagner RS; Li JH. Sociosexual Behaviors of
Tibetan Macaques (Macaca thibetana). 64th Annual Northwest Anthropological Conference,
Moscow, ID, 21-23 April.
2011 Simons ND; Lorenz JG; Sheeran LK; Matheson MD; Wagner RS; Li JH. Methods and
implications of the noninvasive collection of saliva from nonhuman primates. 64th Annual
Northwest Anthropological Conference, Moscow, ID, 21-23 April.
2011 Winters S; Matheson MD; Sheeran LK; Wagner RS; Li JH. Social Recruitment in Tibetan
Macaques (Macaca thibetana) at Mt. Huangshan, China. 64th Annual Northwest Anthropological
Conference, Moscow, ID, 21-23 April.
2011 Winters S; Sheeran LK; Matheson MD; Li JH; Wagner RS. The Influence of Infant Physical
Disability on Mother-Infant Attachment: A case study of limb loss in an infant Tibetan macaque
(Macaca thibetana). 64th Annual Northwest Anthropological Conference, Moscow, ID, 21-23
April.
2011 Wescliff JB; Matheson MD; Sheeran LK; Wagner RS; Li JH. How close is too close? Social
proximity dynamics of Macaca thibetana. 64th Annual Northwest Anthropological Conference,
Moscow, ID, 21-23 April.
2010 Crnich C; Matheson MD; Gabriel K; Sheeran LK. Visitor effect and environmental enrichment
use in former pet monkeys. 63rd Northwest Anthropology Conference, Ellensburg, WA 24-27
March.
2010 Ginn L; Sheeran LK; Matheson MD; Li JH; Wagner RS. The influence of affiliative relations on
copulation strategies of Tibetan macaques (Macaca thibetana). 63rd Northwest Anthropology
Conference, Ellensburg, WA 24-27 March.
2010 Hall B; Sheeran LK; Matheson MD; Li JH; Wagner RS. Maternal stress and infant attachment in
free-living Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. 63rd Northwest
Anthropology Conference, Ellensburg, WA 24-27 March.
2010 Rinear J; Sheeran LK; Cheyne S. Effects of tree characteristics in sleeping tree choice in
Bornean agile gibbons (Hylobates albibarbis). 63rd Northwest Anthropology Conference,
Ellensburg, WA 24-27 March.
2010 Self S; Sheeran LK; Matheson MD; Li JH; Wagner RS; Harding S; Pelton O. Tourist density and
infant-directed aggression in Macaca thibetana at Mt. Huangshan, China. 63rd Northwest
Anthropology Conference, Ellensburg, WA 24-27 March.
2010 Sheeran LK; Matheson MD; Li JH; Wagner RS. Sociability and aging in free living Tibetan
macaques (Macaca thibetana). 63rd Northwest Anthropology Conference, Ellensburg, WA 24-27
March.
2009 Ginn L. The influence of affiliative relations on copulation strategies of Tibetan macaques
(Macaca thibetana); LK Sheeran; MD Matheson faculty mentors. 18th Regional Conference of
Undergraduate Research, M.J. Murdock Charitable Trust, Spokane, WA 30-31 October (poster
presentation).
2009 Sheeran LK; Matheson M; Li JH; Wagner RS. Age and potential social partners in Tibetan
macaques (Macaca thibetana). 62nd Annual Meeting of the Northwest Anthropological
Association, Newport, OR, March (poster presentation).
2008 Jones A; Matheson M; Sheeran LK; Li JH; Wagner RS. Interactions between humans and
Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. 61st Annual Meeting of the
Northwest Anthropological Association, Victoria, B.C., Canada, April.
Updated May 2018
14
2007 Wallin J; Jensvold ML; Sheeran LK. Play, Laughter, and Humor in Captive Chimpanzees (Pan
troglodytes). 60th Annual Meeting of the Northwest Anthropological Association, Pullman, WA,
March (poster presentation).
2006 Jensvold ML; Sheeran LK; Halberg R; Keyser J. Laughter, Number of Play Partners, & Play
Bout Duration in Captive Chimpanzees (Pan troglodytes). 59th Annual Meeting of the Northwest
Anthropological Association, Seattle, WA, March.
2006 Ruesto L; Sheeran L; Matheson M; Li JH; Wagner S. Investigation of possible impacts of tourist
density, behavior, and decibel level on Tibetan macaque aggression. 59th Annual Meeting of the
Northwest Anthropological Association, Seattle, WA, March.
2005 Wagner RS; Sheeran LK; Matheson MD. Biodiversity and habitat characteristics of Monkey
Valley amphibians, Huangshan, China. Annual Meeting of the Society of Northwestern
Vertebrate Biology, Corvallis, OR, February (poster presentation).
2005 Matheson MD; Sheeran LK; Wagner RS. Preliminary report on tourist-Tibetan macaque
(Macaca thibetana) interactions at Huangshan, China. 58th Annual Meeting of the Northwest
Anthropological Association, Spokane, WA, March.
2001 Klein C; Sheeran LK. Differential mortality in captive lar gibbons and siamangs, 15th Semi-
Annual Meeting of the Southern California Primate Research Forum, Fullerton, CA, 10
November.
1998 Sheeran LK. Black gibbon (Hylobates concolor) group size and population density. 69th Meeting
of the Southwestern Anthropological Association, Sacramento, CA, 16-18 April.
1996 Sheeran LK. Development of sanctuaries for Thai gibbons. 5th Semi-annual Meeting of the
Southern California Primate Research Forum, Long Beach, CA, 16 November.
1995 Sheeran LK. Gibbon rehabilitation and reintroduction projects. Annual Meeting of the Southern
California Academy of Sciences, Fullerton, CA, 5-6 May.
1994 Sheeran LK. Endangerment of gibbons in China. 1st Semi-annual Meeting of the Southern
California Primate Research Forum, Claremont, CA, 12 November.
Updated May 2018
15
Organized Paper Sessions, Workshops, and Conferences
International Conferences
2000 Organizer of paper session, The Small Apes. The Apes: Challenges for the 21st Century,
Brookfield Zoo, Chicago, IL, 10-13 May
2000 Co-organizer of workshop, Gibbon Conservation Issues. The Apes: Challenges for the 21st
Century, Brookfield Zoo, Chicago, IL, 10-13 May (with M.E. Pruett-Jones and A. Varsik)
2000 Invited Member of Asian Ape Roundtable Discussion, The Apes: Challenges for the 21st Century,
Brookfield Zoo, Chicago, IL, 10-13 May
1998 Co-organized session, The Lesser Apes. Great Apes of the World Conference II, Kuching,
Malaysia, 29-31 July (with Dr. Jatna Supriatna)
1994 Host and organizer of The International Conference on Orang Utans: The Neglected Ape,
Fullerton, CA, 05-08 March
National Conferences
2001 Organizer of workshop, Ethical Issues in Primatology. 24th Annual Meeting of the American
Society of Primatologists, Savannah, GA, 8-11 August
Regional Conferences 2010 Organizer of paper session, Primate Behavior. 63rd Northwest Anthropology Conference,
Ellensburg, WA, 24-27 March
2008 Moderator for two sessions (Visual Anthropology and Resource Management) at the Symposium
on University Research and Creative Expression, Central Washington University, Ellensburg,
WA, 21 May
2001 Host and co-organizer of the 15th Semi-Annual Meeting of the Southern California Primatology
Research Forum, Contemporary Views of Gibbon Behavior and Ecology, Fullerton, CA, 10
November
1999 Host and organizer of the 70th Annual Meeting of the Southwestern Anthropological Association,
Fullerton, CA, 16-18 April
1999 Host and organizer of the 10th semi-annual meeting of the Southern California Primatology
Research Forum, Primate Culture, Fullerton, CA, 18 April
1998 Organized session, Primate Behavior, Southwestern Anthropological Association 69th Annual
Meeting, Sacramento, CA, 16-18 April
1996 Host and organizer of the 4th semi-annual meeting of the Southern California Primatology
Research Forum, Primate Conservation in the 21st Century, Fullerton, CA, 27 April
Updated May 2018
16
Teaching & Professional Experience
2006- present Director, Primate Behavior and Ecology MS & BS programs, Central Washington
University (CWU)
2011-present Professor of Anthropology, CWU
2005-2011 Associate Professor of Anthropology, CWU (tenured)
2003-2005 Assistant Professor of Anthropology, CWU
1999-2003 Associate Professor of Anthropology, California State University-Fullerton (tenured)
1999 Visiting Assistant Professor of Anthropology, University of Southern California
1993-1999 Assistant Professor of Anthropology, California State University-Fullerton
1993 Lecturer in Anthropology, Ohio State University
Courses Taught at CWU
General Anthropology (ANTH 107) [General Education course]
Introduction to Biological Anthropology (ANTH 110) [General Education course]
Introduction to Biological Anthropology Lab (ANTH 110L) [General Education course]
Anthropological Principles and Assessments (ANTH 301)
Primate Social Behavior (ANTH 313)
Modern Human Variation (ANTH 314) [General Education course]
Long Term Primate Studies (ANTH 412)
Primate Evolution (ANTH 418)
Primate Conservation (ANTH 411/PRIM 511)
Research Methods in Primatology (ANTH 413/PRIM 513)
China Field School (ANTH 493)
Introduction to Primatology (PRIM 501)
Program and Curriculum Development at CWU
2017 Development of graduate certificate program in Captive Primate Care (intended to serve graduate
and post-baccalaureate students)
2015 Development of certificate program in Captive Primate Care (intended to serve matriculated
undergraduate and non-matriculated students)
2009 Approval of Primate Behavior MS proposal by Western Interstate Commission on Higher
Education to include the program in the Western Regional Graduate Research Program
2008 Approval of Primate Behavior MS proposal by WA State Higher Education Coordinating Board
Undergraduate Advisees 20 undergraduate students spring quarter 2017
M.S. Student Advisement
Thesis Chair, Primate Behavior Program: Kalli Kohnen, Amanda Osborne, Josefine Holm, Samantha
Jones, Kirili Stauch, Hilary Hemmes-Kavanaugh, Emily Dura, Brenda Gilpin, Courtney Martinez,
Alexandra Sheldon, Kaitlin Wright, Allison Farley, Emily Veitia, Briana Schnepel, Melanie Bell, Erika
Price, Natasha Zientek, Alexander DuVall-Lash, Alexander Pritchard, Rie Usui, Sofi Bernstein, Allison
Kibler-Campbell, Laura Burwell, Briana Bauer, Grace Enlow, John Rinear, Brian Hall, Sydney Self
Thesis Chair, Resource Management Program: Mary Jo Austin, Lucy Ruesto
Updated May 2017
17
Thesis Committee Member: Kristin Ocasio-Rodriguez, Kaylen Kilfeather, Erin Connelly, Carly Batist,
Krista Banda, Katherine MacDonald, Lilian Stolar, Holly Soubiea, Whitney Emge, Kayley Sullins,
RyAnn Stafford, Gregory Fratellone, Anne Salow, Asa Staven, Anthony Carr, Ricardo Fernandes,
Benjamin Gombash, Jennifer Humphreys, Debra Lewis, Erica Dunayer, Austin Leeds, Hilaree Sorenson,
Sandra Winters, Julie Wescliff, Kelli Poe, Rebecca Hendershott, William Ramirez, Elizabeth Gibbons,
Tracy (Herigstad) Campion, Lindsay Zager, Jackie Buckner, Cassandra Crnich, Meredith Sherrill, Laurel
Marburg, Jennifer Keyser, Julia Galluci, Heather Mack, Tennyson Egan, Amber Jones, Jacquelyn Smith,
Shannon (Reider) Wallin, Cleve Hicks
Outside Thesis Committee Member: Amanda Layton, Sue Litchfield, Kevin Russell, Susan Braun, Lisa
Marie Lohse-Miranda, Catherine Tucker, Caitlyn Cornell, Elizabeth Dickson, Kimberly Bonsen, Amber
Green
Undergraduate Science Honors Committee Chair or Co-Chair: Jason Wallin, Laura Ginn, Terrence
Yenter, Michael Blue, Grant Clifton
Undergraduate Science Honors Committee Member: Courtney Batts, Glee Larsen
Ph.D. Committee Member: Samantha Bricknell, Victoria Buresch, Benjamin Rawson (all at Australian
National University); Sofia Bernstein (Kyoto University)
Students’ Awards
2018 College of the Sciences Undergraduate Research Award, Catherine McAdams
2018 CWU Master’s Research or Creative Activity Fellowship, Samantha Jones
2017 National Science Foundation-East Asia and Pacific Summer Institutes Award, Kirili Stauch
CWU Graduate Student Summer Research or Creative Activity Fellowship, Alexandra Sheldon
2016 SOURCE Outstanding Graduate Poster Presentation, Allison Farley
CWU Graduate Student Summer Research or Creative Activities Fellowship, Courtney Martinez
CWU Master’s Research or Creative Activity Fellowship, Courtney Martinez
CWU Graduate Student Summer Research Fellowship, Brenda Gilpin
CWU Master’s Research or Creative Activity Fellowship, Brenda Gilpin
2015 SOURCE Undergraduate Scholar of the Year, Grant Clifton
CWU Graduate Student Summer Research or Creative Activity Fellowship, Allison Farley
CWU Graduate Student Summer Research or Creative Activity Fellowship, Brianna Schnepel
CWU Graduate Student Summer Research or Creative Activity Fellowship, Kaitlin Wright
2014 CWU Graduate Student Summer Research or Creative Activity Fellowship, Melanie Bell
College of the Sciences Undergraduate Research Award, Grant Clifton
Science Honors Research Program, Grant Clifton
2013 CWU Graduate Student Summer Research or Creative Activity Fellowship, Natasha Zientek
SOURCE Award for Outstanding Student Oral Presentation, Alexander Pritchard
2012 CWU Graduate Student Summer Research or Creative Activity Fellowship, Alexander Duvall-
Lash
2011 Science Honors Research Program, Michael Blue
McNair Scholar, Alicia Faulkner
CWU Graduate Student Summer Research or Creative Activity Fellowship, Briana Bauer
CWU Graduate Student Summer Research or Creative Activity Fellowship, Allison Kibler-
Campbell
2009 Science Honors Research Program, Laura Ginn (co-mentor with Megan Matheson)
2007 Departmental Alumni Scholarship, Laura Ginn
2007 Science Honors Research Program, Terrence Yenter (co-mentor with Megan Matheson)
Updated May 2017
18
2007 SOURCE Award for Outstanding Undergraduate Student Oral Presentation, Jason Wallin (co-
mentor with Mary Lee Jensvold)
2007 Departmental Alumni Scholarship, Tim Held
2006 Science Honors Research Program, Jason Wallin (co-mentor with Mary Lee Jensvold)
2005 CWU Graduate Student Summer Research Fellowship, Lucy Ruesto
2002 California Predoctoral Fellow, Brenda Royce
1999 McNair Scholar, Nicole Kanbara
1998 California Predoctoral Fellow, Melissa Collier
1998 California Predoctoral Fellow, Carlos Navarrete
Symposium on University Research and Creative Expression (SOURCE) Presentations (many of these also presented at other venues; student co-authors underlined)
2016 Sheeran LK; Wagner RS; Sun LX; Li JH; Xia DP. Human and Tibetan macaque (Macaca
thibetana) interactions in the Valley of the Wild Monkeys, Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 18-19 May.
2015 Bell MC; Sheeran LK; Skollár G. Javan gibbons (Hylobates moloch) vary gesture use by
recipient’s attentional state at the Gibbon Conservation Center, CA. SOURCE, CWU,
Ellensburg, WA, 21 May. [poster]
2015 Clifton CJ; Sheeran LK; Wagner RS; Li JH. A Comparison of Female-Female Bridging to Male-
Male Bridging in Tibetan Macaques (Macaca thibetana). SOURCE, CWU, Ellensburg, WA,
21 May.
2015 Clifton CJ; Sheeran LK; Wagner RS; Li JH. Tibetan Macaque Bridging Behavior. SOURCE,
CWU, Ellensburg, WA, 21 May. [video]
2015 Fratellone GP; Sun L; Sheeran LK; Wagner RS; Li JH. Leadership in the collective movements
of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 21 May. [poster]
2014 Price E; Sheeran LK; Li JH. Developmental changes in vocalizations of Tibetan
macaques (Macaca thibetana). SOURCE, CWU, Ellensburg, WA, 15 May.
2014 Staven A; Sheeran LK; Li JH; Wagner RS. Interactions Between tourists and juvenile Tibetan
macaques at Mt. Huangshan, China. SOURCE, CWU, Ellensburg, WA, 15 May.
2013 Murphy A; Link J; Li JH; Sheeran LK Maternal investment in free-ranging Tibetan
macaques (Macaca thibetana). SOURCE, CWU, Ellensburg, WA, 16 May.
2013 Pritchard A; Sheeran LK; Gabriel K; Li JH; Wagner RS. Measuring personality traits in
provisioned Tibetan macaques (Macaca thibetana), Mt. Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 16 May.
2013 Usui R; Sheeran LK; Pritchard A; DuVall-Lash A; Li JH; Wagner RS. Impact of park ranger
quality on Tibetan macaque (Macaca thibetana) behaviors at Mt. Huangshan, China. SOURCE, CWU, Ellensburg, WA, 16 May.
2012 Batts CB; Sheeran L; Li J; Wagner R. Infant and juvenile play patterns in Tibetan macaques
(Macaca thibetana). SOURCE, CWU, Ellensburg, WA, 17 May.
Updated May 2017
19
2012 Dunayer E; Matheson M; Sheeran L; Beck D; Li J; Wagner R. Does tourism have a market effect
on the grooming for tolerance interchange in Tibetan macaques (Macaca thibetana) at Mt.
Huangshan, China? SOURCE, CWU, Ellensburg, WA, 17 May.
2012 Faulkner A; Sheeran L. (faculty mentor). Tourist knowledge and perceptions of Tibetan
macaques at Mt. Huangshan, China. SOURCE, CWU, Ellensburg, WA, 17 May.
2011 Simons N; Lorenz J; Sheeran L; Matheson M; Li J; Wagner S. Methods and implications from
the noninvasive collection of saliva from nonhuman primates. SOURCE, CWU, Ellensburg, WA,
21 May.
2010 Ginn L; Sheeran LK; Matheson MD; Li JH; Wagner RS. The influence of affiliative relations on
copulation strategies of Tibetan macaques (Macaca thibetana). SOURCE, CWU, Ellensburg,
WA, 20 May.
2010 Hall B; Sheeran LK; Matheson MD; Li JH; Wagner RS. Maternal stress and infant attachment in
free-living Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 20 May.
2010 Rinear J; Sheeran LK; Cheyne S. Effects of tree characteristics in sleeping tree choice in
Bornean agile gibbons (Hylobates albibarbis). SOURCE, CWU, Ellensburg, WA, 20 May.
2010 Self S; Sheeran LK; Matheson MD; Li JH; Wagner RS; Harding S; Pelton O. Tourist density and
infant-directed aggression in Macaca thibetana at Mt. Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 20 May.
2010 Sheeran LK; Matheson MD; Li JH; Wagner RS. Sociability and aging in free living Tibetan
macaques (Macaca thibetana). SOURCE, CWU, Ellensburg, WA, 20 May.
2009 Sheeran LK; Matheson M; Li JH; Wagner RS. Age and potential social partners in Tibetan
macaques (Macaca thibetana). SOURCE, CWU, Ellensburg, WA, 21 May.
2008 Jones A; Matheson M; Sheeran LK; Li JH; Wagner RS. Human-macaque (Macaca thibetana)
agonistic encounters in Mt. Huangshan, China. SOURCE, CWU, Ellensburg, WA, 15 May.
2008 Mack H; Matheson M; Sheeran LK; Li JH; Wagner S. Grooming behavior of Tibetan macaques
(Macaca thibetana) in the presence of tourists at Mt. Huangshan, China. SOURCE, CWU,
Ellensburg, WA, 15 May.
2008 Yenter TA; Matheson MD; Sheeran LK; Li JH; Wagner RS. Self-directed behaviors in Tibetan
macaques as a function of tourist density and proximity at the Valley of the Wild Monkeys.
SOURCE, CWU, Ellensburg, WA, 15 May (poster presentation).
2007 Halberg RH; Sheeran LK; Jensvold ML. Laughter, number of play partners, age and play bout
duration in chimpanzees (Pan troglodytes) living in an African sanctuary. SOURCE, CWU,
Ellensburg, WA, 17 May.
2007 Lester JD; McCarthy M; Matheson M; Sheeran LK; Li JH; Wagner RS. Green Cascade Frog
(Rana livida) Detection in The Valley of the Wild Monkeys, Mt Huangshan, China. SOURCE,
CWU, Ellensburg, WA, 17 May.
2007 McCarthy M; Matheson MD; Sheeran LK; Lester JD; Li JH; Wagner RS. Sequences of Tibetan
macaque behaviors and tourist behaviors at Mt. Huangshan, China. SOURCE, CWU, Ellensburg,
WA, 17 May.
2007 Wallin J; Jensvold ML; Sheeran LK. Play, Laughter, and Humor in Captive Chimpanzees (Pan
troglodytes). SOURCE, CWU, Ellensburg, WA, 17 May.
Updated May 2017
20
2006 Hartel J; Matheson M; Sheeran LK; Li JH; Wagner S. Post-conflict and self-directed behaviors
in Tibetan macaques. SOURCE, CWU, Ellensburg, WA, 19 May.
2006 Jensvold ML; Sheeran LK; Halberg R; Keyser J. Laughter, Number of Play Partners, & Play
Bout Duration in Captive Chimpanzees (Pan troglodytes). SOURCE, CWU, Ellensburg, WA, 19
May.
2006 Ruesto L; Sheeran LK; Matheson M; Li JH; Wagner S. Investigation of possible impacts of
tourist density, behavior, and decibel level on Tibetan macaque aggression. SOURCE, CWU,
Ellensburg, WA, 19 May.
2006 Smith C; Wagner S; Matheson M; Sheeran LK; Li JH. Population monitoring of the mountain
stream frog (Rana spinosa) in Huangshan, China. SOURCE, CWU, Ellensburg, WA, 19 May.
2005 Matheson MD; Sheeran LK; Wagner RS. Preliminary report of tourist impact on aggression and
social behavior in two Tibetan macaque groups (Macaca thibetana). SOURCE, Ellensburg, WA,
19 May.
Service to the Profession
Editing/Reviewing
2015-present Associate Editor for Scientific Reports, Nature Publishing Group
Ecology and Evolutionary Biology
http://www.nature.com/srep/about/editorial-board#ecologyevolutionary
2014 Impact factor for Scientific Reports is 5.578
(http://www.nature.com/srep/about)
2016-17 Panel Reviewer for National Science Foundation
2015-16 Panel Reviewer for National Science Foundation
2014-present Reviewer for Choice: Current Reviews for Academic Libraries, a publication of
the Association of College & Research Libraries, a division of the American
Library Association
2007-11 Annual abstract reviewer for Biomedical Research Conference for Minority
Students Ad hoc reviewer Oryx, Folia Primatologica, Biodiversity & Conservation, Zoo Biology,
Anthropology: Bachelors to Doctorates, Wadsworth Publishing, McGraw Hill,
Rowman & Littlefield, U of Missouri Press, Graduate Women in Science, National
Geographic Society, National Science Foundation, Earthwatch, Primate
Conservation, Inc., Lincoln Park Zoo Asia/Africa Fund
2001-2,
2000-1
Member, Program Committee, American Society of Primatologists
2001 Ad hoc peer reviewer of the World Conservation Union’s Guidelines for
Nonhuman Primate Reintroductions
1996-2008 Annual proposal reviewer (technical panel) for Charles A. Lindbergh Fund, Inc.
Board Membership/Elected Professional Position
2015-present Board Member, Pacific Primate Sanctuary, Maui, HI
2013-present,
1994-2011
Board Member, Gibbon Conservation Center, Santa Clarita, CA
1997-2000 President, Southwestern Anthropological Association
Updated May 2017
21
Panelist/Participant/Advisor/Commentator
2015 Nominated as foreign member of the China Primatological Society
2015 Comment on the Animal and Plant Health Inspection Service (APHIS) Proposed
Rule: Petition to Develop Specific Ethologically Appropriate Standards for
Nonhuman Primates in Research
2014-present Co-petitioner with Species Survival Network Primate Working Group to CITES to
request review of trade in long-tailed macaques in Laos and Vietnam
2012 Invited panelist, Conservation Workshop for China’s Gibbons, Jingdong, Yunnan
Province, China, 5 October
2012 Invited panelist, ASP education committee student workshop luncheon, 35th
Annual Meeting of the American Society of Primatologists, Sacramento, CA, 20-
23 June
2004 Scientific and technical advisor to Royal Forest Department of Thailand, Pileated
Gibbon Rehabilitation Project
2000-3 Advisor to the American Zoological Association Gibbon Species Survival Plan,
Conservation and Research Coordinator
2000 Member, Protection of Primate Health in the Wild, Resolution adopted by the
American Society of Primatologists
1999 Scientific Advisor Wuliangshan Gibbon Conservation Conference, 04-06
November
1998 Member, Private Ownership of Primates International Task Force, Resolution
adopted by the American Society of Primatologists
1996-2002 Member of the Organizing Committee of the Southern California Primate
Research Forum
Service to the University and College
2016-17 Member, Graduate Council Curriculum Subcommittee
2014-16 Member, Graduate Council Bylaws Subcommittee
2014-15,
2013-14
Member, Office of International Studies & Programs Executive Director Search
Committee
2012, 13, 14 SOURCE Session Chair
2012, 13, 14 Guest lecturer, Dr. Leo D’Acquisto’s Exercise Science classes
2011 Panelist, CWU Promotion and Tenure forum
2009, 10, 11,
12, 13, 14, 15
SOURCE Judge
2008-present Member, McNair Advisory Council
2007-8 PBE Program Review
2006-present Director, Primate Behavior and Ecology Program (MS and BS degrees)
2004-12 Member, Institutional Animal Care and Use Committee
2004 Panelist, New Faculty Orientation Luncheon
2003-present Member, Graduate Faculty
Updated May 2017
22
Service to the Department of Anthropology and Museum Studies
Search Committees
2017 Chair, Biological Anthropologist NTT Search Committee
2015-16 Chair, Primatologist Search Committee
2015-16 Member, Forensic Anthropologist Search Committee
2014-15 Member, Biological Anthropologist Search Committee
2012-13 Member, Cultural Anthropologist Search Committee
2010-11 Member, Cultural Anthropologist Search Committee
2008-09 Chair, Search Committee PBE Spheres of Distinction
2007-08 Member, Biological Anthropologist Search Committee
Personnel Committees
2005-6, 2013-14 Chair, Personnel Action Committee/Department Personnel
Committee
2006-7, 2007-8, 2010-11,
2011-12, 2014-15
Member, Departmental Personnel Committee
Faculty Advisor to Student Groups
2010-present Primate Awareness Network
2007-8 Wildlife Conservation Group
2003-4 Anthropology Students Association
Other Departmental Service
2016-17 Curriculum Committee
2014-15 Curriculum Committee
2006-7 Alternate Representative, Faculty Senate
2003-6 Department Library Representative
Service to the Community
2015 Presenter (with Professors Hope Amason, Lourdes deLeon, Karisa Terry, and Jessica Mayhew),
Expanding Your Horizons Conference at CWU
2012 Featured (with Primate Behavior graduate student Sofi Bernstein) in segment in CWU’s “Your
World” http://www.cwu.edu/international-programs/your-world-videos
2011 Featured in segment on the Tibetan macaques of Mt. Huangshan, CCTV China
http://news.cntv.cn/program/zoubianzhongguo/20120102/118895.shtml
2004-7 Annual Presenter (with Professor Mary Lee Jensvold), Expanding Your Horizons Conferences
at CWU
Updated May 2017
23
Awards and Honors
2018 Central Washington University Distinguished Professor of the University for Research Award
2017-18 Central Washington University Excellence in Advising Award
2015 Nominee, SOURCE Faculty Mentor of the Year
2014 Awardee, Honoring Our Professors’ Excellence, National Residence Hall Honorary
Nominee, SOURCE Graduate Faculty Mentor of the Year
2013 SOURCE Graduate Faculty Mentor of the Year
2001 California State University-Fullerton Outstanding Faculty Service Recognition: Scholarship
1992 Inductee Phi Kappa Phi Honor Society
Professional References
Dr. Kathleen Barlow
Professor
Department of Anthropology
Central Washington University
400 E University Way
Ellensburg, Washington 98926-
7544 (509)963-3212
Dr. Biruté Mary F. Galdikas
Orangutan Foundation International
8888 University Dr.
Burnaby, British Columbia V5A
1S6 CANADA
(604)291-3135 / (323)938-6046
Dr. JinHua Li
Anhui University
3 Feixi Road
Heifei, Anhui Province
CHINA 230039
