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Wiens-C.V. page 1 of 37 15 May, 2019 Curriculum Vitae John J. Wiens Present address: Department of Ecology and Evolutionary Biology University of Arizona Tucson, AZ 85721 Web-page: http://wienslab.com/Home.html Research interests: (1) Phylogenetic approaches to evolution and ecology (2) Systematic theory and methodology (3) Phylogeny, evolution, and ecology of reptiles and amphibians Education: The University of Kansas, B.S. in Systematics and Ecology, 1991 Senior Honors Thesis: “Phylogenetic position and relationships of the tree lizards (genus Urosaurus)” Supervisor: William E. Duellman The University of Texas at Austin, Ph.D. in Zoology, 1995 Dissertation Title: “Phylogeny of the lizard genus Sceloporus: Systematics, evolutionary morphology, and evaluation of phylogenetic approaches.” Supervisors: David C. Cannatella and David M. Hillis Professional experience: 2013–present, Professor, Department of Ecology and Evolutionary Biology, University of Arizona 2006–2012, Associate Professor, Department of Ecology and Evolution, Stony Brook University 2003–2006, Assistant Professor, Department of Ecology and Evolution, Stony Brook University 2001–2002, Associate Curator, Section of Amphibians and Reptiles, Carnegie Museum of Natural History 2000–2002, Supervisor, Molecular Systematics Laboratory, Carnegie Museum of Natural History 1996–2001, Adjunct Professor, Department of Biology, University of Pittsburgh 1995–2001, Assistant Curator, Section of Amphibians and Reptiles, Carnegie Museum of Natural History 1991–95, NSF Graduate Fellow, University of Texas at Austin. 1993–94, Graduate Teaching Assistant, University of Texas at Austin. 1987–91, Undergraduate Research Assistant, Division of Herpetology, University of Kansas Museum of Natural History.

Curriculum Vitae John J. Wiens · Museum of Natural History 2000–2002, Supervisor, Molecular Systematics Laboratory, Carnegie Museum of Natural History 1996–2001, Adjunct Professor,

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Page 1: Curriculum Vitae John J. Wiens · Museum of Natural History 2000–2002, Supervisor, Molecular Systematics Laboratory, Carnegie Museum of Natural History 1996–2001, Adjunct Professor,

Wiens-C.V. page 1 of 37

15 May, 2019

Curriculum Vitae John J. Wiens

Present address: Department of Ecology and Evolutionary Biology University of Arizona Tucson, AZ 85721 Web-page: http://wienslab.com/Home.html Research interests:

(1) Phylogenetic approaches to evolution and ecology (2) Systematic theory and methodology (3) Phylogeny, evolution, and ecology of reptiles and amphibians

Education:

The University of Kansas, B.S. in Systematics and Ecology, 1991 Senior Honors Thesis: “Phylogenetic position and relationships of the tree lizards (genus Urosaurus)”

Supervisor: William E. Duellman The University of Texas at Austin, Ph.D. in Zoology, 1995

Dissertation Title: “Phylogeny of the lizard genus Sceloporus: Systematics, evolutionary morphology, and evaluation of phylogenetic approaches.”

Supervisors: David C. Cannatella and David M. Hillis Professional experience:

2013–present, Professor, Department of Ecology and Evolutionary Biology, University of Arizona

2006–2012, Associate Professor, Department of Ecology and Evolution, Stony Brook University

2003–2006, Assistant Professor, Department of Ecology and Evolution, Stony Brook University

2001–2002, Associate Curator, Section of Amphibians and Reptiles, Carnegie Museum of Natural History

2000–2002, Supervisor, Molecular Systematics Laboratory, Carnegie Museum of Natural History

1996–2001, Adjunct Professor, Department of Biology, University of Pittsburgh

1995–2001, Assistant Curator, Section of Amphibians and Reptiles, Carnegie Museum of Natural History

1991–95, NSF Graduate Fellow, University of Texas at Austin. 1993–94, Graduate Teaching Assistant, University of Texas at Austin. 1987–91, Undergraduate Research Assistant, Division of Herpetology,

University of Kansas Museum of Natural History.

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Teaching experience: Diversity of Life (Ecol 170C), University of Arizona, spring 2017, 2019 Herpetology (Ecol483), University of Arizona, spring 2013, 2014, 2015, 2016,

2018. Graduate seminar (Ecol586) “Macroecology, macroevoution, and speciation”

University of Arizona, fall 2014, 2015, 2016, 2017 Herpetology (Bio 348; Undergraduate; “Diversity and evolution of reptiles

and amphibians”), Stony Brook University, spring 2006, 2007, 2008, 2009, 2011

Evolution (BEE 551; Graduate level), Stony Brook University, fall 2004, 2005, 2007, 2008, 2009

Evolution (BIO 354; Undergraduate), Stony Brook University, fall 2003, 2005, 2007, 2009

Graduate seminar on evolutionary and ecological responses to climate change, Stony Brook University, fall 2011

Graduate seminar on evolution and species interactions, Stony Brook University, spring 2007

Graduate seminar on species richness, Stony Brook University, spring 2003 Phylogenetics, University of Pittsburgh, fall 1996, 2000 Herpetology, Pymatuning Laboratory of Ecology, University of Pittsburgh,

summer 1996 Teaching Assistant for: Herpetology, Comparative Anatomy, Structure and

Function of Organisms (Univ. Texas; 1993-1994) Graduate student direction (major advisor):

Emma Tuschhoff (Ph.D. program, Univ. Arizona, started fall 2018) Cristian Roman-Palacios (Ph.D. program, Univ. Arizona, started fall 2016) Elizabeth Miller (Ph.D. program, Univ. Arizona, started fall 2014; NSF

Graduate Fellow) Shea Lambert (Ph.D., Univ. Arizona, 2018) Xia Hua (Ph.D., Stony Brook, 2012); currently Assistant Professor at Australian

National University M.-Caitlin Fisher-Reid (Ph.D., Stony Brook, 2012); currently Assistant

Professor, Bridgewater State University Dan Moen (Ph.D., Stony Brook, 2012; NSF Graduate Fellow); currently

Assistant Professor at Oklahoma State University Patrick Stephens (Ph.D., Stony Brook, 2005); currently Research Assistant

Professor at Univ. Georgia Carl Hutter (M.A., Stony Brook, 2012); currently Ph.D. program at Univ.

Kansas Antonin Machac (M.A., Stony Brook, 2012); currently Ph.D. program at Stony

Brook Univ. Scott David (M.A., Stony Brook, 2012) Zachary Chejanovski (M.A., Stony Brook, 2012); currently Ph.D. program at

Univ. Rhode Island Nathan Butler (M.A., Stony Brook, 2011) Allison Onstine (M.A., Stony Brook, 2011) Carolina Ulloa (M.A., Stony Brook, 2011) Christina Wolfe (M.A., Stony Brook, 2004)

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Graduate committee: Arizona: Allison Harrington: Ph.D., EEB; Alice Cang: Ph.D., EEB; Joseph

Charboneau: Ph.D., EEB; Zheng Li, Ph.D., EEB. Antonio Gomez (Master’s, Dept. Entomology; 2014); Chris Gurguis, Master’s, EEB (2014); Tim O’Connor: Master’s, EEB (2015); Samantha Anderson, Master’s, EEB (2016);

Stony Brook: Mireya Mayor, Ph.D. 2008 (Dept. Anthropology); Shu-Dan Yeh, Ph.D. 2009 (Dept. Ecology & Evolution).

Pittsburgh: Charles Welsh, Univ. of Pittsburgh, Ph.D. 1997 External: James Austin, Queens Univ., Ontario, Ph.D. 2003 (now Assistant

Professor, Univ. Florida); Frank Fontanella, City Univ. New York, Ph.D. 2009; Alex Pyron, City Univ. New York, Ph.D. 2009; Allyson Fenwick-Modra, Univ. Central Florida, Ph.D. 2012; Daniel Scantlebury, Rochester Univ., Ph.D. 2014

External examiner (Ph.D.): S. P. Vijayakumar, Centre for Ecological Studies, Indian Institute of Science (2013)

External examiner (Ph.D.): Julie-Lee Yaw, University of British Columbia (2013)

External examiner (Master’s): Elkin Tenorio, Universidad de Los Andes, Colombia (2013)

Postdoctoral direction:

Daniel Portik (2017): current Zhou Chen (2017): current Tania Hernandez (2015–2017): currenty researcher at LANGEBIO (Laboratorio

Nacional de Genómica para la Biodiversidad) in Irapuato, Mexico Tereza Jezkova (2015–2017): currently Assistant Professor at Miami

University, Ohio Jeff Streicher (2013–2014); currently Assistant Curator at British Museum of

Natural History Alex Pyron (2009–2010); currently Assistant Professor, George Washington

University Windsor Aguirre (2007–2009); currently Associate Professor, DePaul

University Kenneth Kozak (2005–2007); currently Associate Professor, University of

Minnesota Sarah Smith (2004–2006); currently living in Australia Tag Engstrom (2003–2004); currently Associate Professor, California State

University at Chico James Fetzner (2001–2002); currently Assistant Curator, Invertebrate Zoology,

Carnegie Museum of Natural History Robert Espinoza (2000–2001); currently Professor, California State University

at Northridge Christopher Parkinson (2000–2001); currently Professor, University of Central

Florida Visiting foreign students and scientists (partial list):

Pablo Guerrero (Chile; Fullbright Fellow; Univ. Chile)–2010 Serkan Gul (Turkey)–2010–2011

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Thomas Blankers (Netherlands; Amsterdam Univ.)–2011 Greta Carrete-Vega (Spain)–2011 Rubi Nelsi Meza-Lazaro (Mexico; Ph.D. student; UNAM)–2011 Ignacio Quintero (Colombia; postgraduate: Univ. de los Andes)–2011 Virginia Valcarcel (Spain; postdoctoral researcher)–2012 Cendy Zeng (China; undergraduate Fudan Univ.)–2012 Nelsy Rocio Pinto Sanchez (Colombia; Univ. de los Andes)–2012 Maria Fernanda Bonetti (Brazil: Ph.D. student, Univ. Federal do Paraná)–2013 Yuchi Zheng (China; Chengdu Institute Biology; sabbatical visitor)–2014 Lucas Barrientos (Colombia: Ph.D. student: Univ. de los Andes)–2014 Victoria Souza (Brazil: Ph.D. student: Univ. Federal de Goiás)–2014–2015 Uri Omar Garcia (Mexico: Ph.D. student; UNAM)–2015 Adrian Nieto Montes de Oca (Mexico: Professor: UNAM)–2015 Andreas Meyer (Brazil: Ph.D. student: Univ. Federal do Parana)–2015–2016 Longhui Lin (China: Hangzhou University; sabbatical visitor)–2015 Melissa Bars-Closel (Brazil; Ph.D. student; Universidade de Sao Paulo)–2015–

2016 Yan Haifei (China: South China Botanical Garden; sabbatical visitor)–2015–

2016 Tiangang Gao (China: State Key Laboratory of Systematic and Evolutionary

Botany, Beijing; sabbatical visitor)–2016 Dongyuan Song (China, Fudan University, undergraduate)–2016 Kenji Hayashi (Brown University, undergraduate)–2016 Lian Chen (China, Jiangsu Second Normal University; sabbatical visitor)–

2016–2017 Hong Li (China, Nanjing Normal University; sabbatical visitor)–2016–2017 Weiwei Zhou (China, Kunming Institute of Zoology; visiting scientist)–2017-

2018 Peng Li (China, Nanjing Normal University; visiting scientist)–2017–2018 Yan-Fu Qu (China, Nanjing Normal University; visiting scientist)–2018–2019 Hui Liu (China, South China Botanical Garden; visiting scientist)–2018–2019 Adriana Morales (Brazil: Sao Paulo University; Ph.D. student)–2019 Matthew Moreira (Portugal: University of Areiro; Ph.D. student)–2019

Workshops given

Eco-evolutionary dynamics in biogeography (1 week, October 2018; Katalapi Park, Puerto Montt, Chile; co-taught with several researchers).

Macroecology and macroevolution winter course (1 week, December 2014; École Normale Supérieure, Paris; co-taught with Brian Enquist and Neo Martinez)

Phylogenetic and comparative methods (~2 weeks; Santiago, Chile, October 2011; with Pablo Guerrero)

Comparative methods in R (1 day; Leiden, The Netherlands, fall 2010) Phylogenetic and comparative methods (2 weeks; Mysore, India, summer

2006; with R. Geeta) Field experience:

Approximately 14 months over various trips to Mexico, Ecuador, Peru, Argentina, Madagascar, and Cameroon, with more limited experience in

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Brazil, China, France, India, Japan, Kenya, Malaysia, Panama, Spain, and Taiwan. Extensive experience in the U.S.

Professional service:

Editor-in-Chief, Quarterly Review of Biology, 2007–2010 Editor, Quarterly Review of Biology, 2006–2007, 2010–present Associate Editor, Ecology Letters, 2009–present Associate Editor, BMC Evolutionary Biology, 2011–2014 Associate Editor, Ecography, 2007–2009 Associate Editor, American Naturalist, 2005–2009 Associate Editor, Evolution, 2001–2003 Associate Editor, Systematic Biology, 1996–2000 Associate Editor, Herpetologica, 1996–2000 Editorial Board, Systematic Biology, 1995–1996 Editorial Board, Copeia, 1999–2008 Editorial Board, Journal of Herpetology, 2001–present Editorial Advisory Board, Revista Mexicana de Biodiversidad, 2005–present Academic Editor, PLoS Biology (2014) Guest Editor, Proceedings Natl. Academy of Sciences, U.S.A. (2010, 2015, 2017) Ad Hoc Associate Editor, Conservation Biology (2007) Web-based journals: Editorial Board, The International Journal of Ecology, 2008–2013 Council, Society of Systematic Biologists (2003–2005) Long-Range Planning Committee, Society of Systematic Biologists (2007–2009) Board of Governors, American Society of Ichthyologists and Herpetologists

(2003-2008) Rosemary Grant Award Committee, Society for the Study of Evolution (2011) Scientific Committee, World Congress of Herpetology, 2016 Invited Participant, NSF Workshop on Integrative Biogeography, Spring 2007 Invited Participant, NSF Workshop on the Tree of Life, Spring 2008 Invited Participant, NCEAS workshop on “Niche conservatism and diversity

gradients,” Fall 2008, 2009 Invited Participant, workshop on niche width, University of Toronto,

Scarborough, Summer 2015 German Research Foundation, Grant Review Panel 2017 Swedish Research Council, Grant Review Panel 2014, 2015 NSF Systematic Biology Panel, Fall 2000 NSF Biotic Surveys and Inventory Panel, Spring 2002 NSF Doctoral Dissertation Improvement Grants, Spring 2003 NSF Evolutionary and Population Ecology Panel, Fall 2004 NSF Evolutionary and Population Ecology Panel, Fall 2005 NSF Systematic Biology Panel, Fall 2007 NSF Evolutionary Genetics Panel, Spring 2009

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NSF Evolutionary Genetics Pre-proposal Panel, Spring 2012 Declined (Evolutionary Genetics, Fall 2006; Evolutionary Ecology, Spring

2010, 2016) Symposium organizer (“Morphological data in phylogenetic analysis: Recent

progress and unresolved problems”), Society of Systematic Biologists, 1996

Symposium organizer (“Species delimitation: new approaches for discovering diversity”), Society of Systematic Biologists, 2006

Symposium co-organizer (“How lizards turn into snakes: combining fossils, phylogeny, function, genetics, development, morphology, and ecology”), co-organized with Agustin Camacho, 8th World Congress of Herpetology, 2016

Reviewer for 106 journals: Acta Biotheoretica, Acta Oecologica, American

Naturalist, American Journal of Physical Anthropology, Animal Behavior, Annals of the Carnegie Museum, Bioinformatics, Biological Conservation, Biological Journal of the Linnean Society, Biological Reviews (Cambridge Philosophical Society), Biology Letters, Bioscience, BMC Biology, BMC Evolutionary Biology, BMC Genetics, BMC Genomics, Canadian Journal of Zoology, Cladistics, Comparative Biochemistry and Physiology, Conservation Biology, Conservation Genetics, Copeia, Current Biology, Ecography, Ecology, Ecology Letters, Ecosphere, Evolution, Evolution Letters, Evolution and Development, Evolutionary Biology, Evolutionary Ecology, Frontiers in Zoology, Functional Ecology, Gene, Genome Biology and Evolution, Global Change Biology, Global Ecology and Biogeography, Heredity, Herpetologica, Herpetological Journal, Herpetological Monographs, Herpetological Review, Human Mutation, Integrative and Comparative Biology, International Journal of Evolutionary Biology, International Journal of Zoology, Journal of Anatomy, Journal of Animal Ecology, Journal of Biogeography, Journal of Ethology, Journal of Evolutionary Biology, Journal of Heredity, Journal of Herpetology, Journal of Human Evolution, Journal of Molecular Evolution, Journal of Morphology, Journal of Natural History, Journal of Paleontology, Journal of the Royal Society Interface, Journal of Theoretical Biology, Journal of Vertebrate Paleontology, Journal for Zoological Systematics and Evolutionary Research, Marine Ecology Progress Series, Methods in Ecology and Evolution, Molecular Biology and Evolution, Molecular Ecology, Molecular Phylogenetics and Evolution, Nature, Nature Climate Change, Nature Communications, Nature Ecology and Evolution, Nature Reviews Genetics, New Phytologist, Oecologia, Oikos, The Open Conservation Biology Journal, The Open Evolution Journal, Organisms Diversity and Evolution, Paleobiology, PeerJ, Philosophical Transactions of the Royal Society of London, Population Ecology, PNAS, Proceedings of the Royal Society of London Series B, PLoS Biology, PLoS One, PLoS Currents: Tree of Life, Quarterly Review of Biology, Royal Society Open Science, Science, Science Advances, Scientific Correspondence (Nature), Scientific Reports (Nature), Scientific Publications of the University of Kansas Museum of Natural History, Systematic Biology, Systematic Botany, Toxins, Trends in Ecology and Evolution, Trends in Genetics, Tropical Zoology, Zoologica Scripta, Zoological Journal of the Linnean Society, Zoology, ZooKeys, Zootaxa

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Reviewer for (book chapters): "Phylogenies and the Comparative Method in Animal Behavior " (1996; E. Martins, ed.), "Biology of Plethodontid Salamanders" (2000; Bruce et al., eds.), "Temperature Dependent Sex Determination in Vertebrates" (2003; Valenzuela and Lance, eds.), “Evolution since Darwin: the first 150 years” (2010; Bell et al., eds.)

Reviewer for (book proposals): Cambridge University Press, Oxford University Press, Princeton University Press, Roberts and Company, Sinauer, University of Chicago Press

Reviewer for (funding agencies): U.S. National Science Foundation (Programs: Systematic Biology, Evolutionary Ecology, Evolutionary Genetics, Population Biology, Animal Behavior, Long-Term Research in Environmental Biology, Paleontology, Biotic Surveys and Inventory), National Geographic Society, Austrian Science Fund, Comision Nacional de Investigacion Cientifica y Tecnologica de Chile, French National Research Agency, German Research Foundation, Leverhulme Trust (United Kingdom), Marie Curie Co-Fund (Belgium), Natural Environment Research Council (United Kingdom), Netherlands Organisation for Scientific Research, Polish National Science Center, Portugese Foundation for Science and Technology, Royal Society of New Zealand (Marsden Fund, Centre of Research Excellence Fund), Research Committee of the University of Crete (Greece), Research Council of Norway, South Africa’s National Research Foundation, Swiss National Science Foundation, Swedish Research Council, U.S.-Israeli Binational Science Foundation

Reviewer for (promotion/tenure): American Museum of Natural History, Arizona State University, Boston University, Bucknell University, City University of New York, Indian Institute for Science, Iowa State University, National Museum of Natural History, Peking University, Rochester University, Rutgers University, San Diego State University, Tulane University, University of California (Los Angeles, Merced, Riverside), University of Colorado, University of Connecticut, University of Florida, University of Hawaii, University of Minnesota, University of Nebraska, University of Nevada (Reno), University of New Orleans, University of Oklahoma, University of Tennessee, University of Texas (Arlington), University of Toronto

Chair, Henri Seibert Award Committee (Best Student Paper Competition), Society for the Study of Amphibians and Reptiles, 1998

Judge, Ernst Mayr Award (Best Student Paper Competition, Society of Systematic Biologists), 1999

Judge, Stoye Award (Best Student Paper Competition, American Society of Ichthyologists and Herpetologists), 2000

Judge, Storer Award (Best Student Poster Competition, American Society of Ichthyologists and Herpetologists), 2001

Judge, Award for Graduate Student Research, Society of Systematic Biologists, 2009

Judge, Rosemary Grant Award for Graduate Student Research, Society for the Study of Evolution, 2010

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Departmental Service University of Arizona Graduate Admissions Committee 2016–2018 Monday Seminar Committee, 2016 Promotion and Tenure Committee, 2014 Peer Review Committee, 2014, 2015 Retreat Committee, 2013–2014 Stony Brook Undergraduate Program Director, 2009–2012 Preliminary Exams Committee, 2007, 2010 Departmental Retreat Coordinator, 2006–2007 Evolutionary Genomics Search Committee, 2006–2007 Departmental Colloquium Coordinator, 2006–2007, 2010–2011 Entering Students Advisory Committee (ESAC), 2004–2006 Graduate Admissions Committee, 2003–2004

University Service (Stony Brook)

Student and Faculty Awards Committee, 2008 Delegate for OTS (Organization for Tropical Studies), 2008–2012 Undergraduate Biology Executive Committee, 2010–2012

Professional organizations:

American Society of Ichthyologists and Herpetologists American Society of Naturalists Herpetologist's League Society of Systematic Biologists Society for the Study of Amphibians and Reptiles Society for the Study of Evolution

Grants:

National Science Foundation, 2017–2020, “Collaborative Research: Understanding large-scale patterns of ecomorph evolution” ($278,905 to PI Wiens). Collaborative project with fellow PI Dan Moen

Chinese Academy of Sciences President’s International Fellowship Initiative (PIFI) for one-month stay at the Kunming Institute of Botany, summer 2017 (49,474 Yuan, ~$7,574 USD)

National Science Foundation, 2011–2012, “Dissertation Research: The role of history in adaptation to novel environments: the role of morphology, performance, and phylogenetic history in frogs” ($12,821; for Dan Moen).

Temminck Fellowship, 2010, Netherlands Centre for Biodiversity Naturalis (4,000 Euros).

National Science Foundation, Systematic Biology Program, 2004–2008 (no-cost extension, 2009), “Collaborative Research: ATOL: The Deep Scaly Project: Resolving squamate phylogeny using genomic and morphological approaches” ($2,426,390 total; $645,700 to PI Wiens). Collaborative project with fellow PIs Tod Reeder, Jack Sites, and Maureen Kearney (NSF EF 0334923).

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National Science Foundation, Systematic Biology Program, 2004-2005, “Dissertation Research: A phylogenetic perspective on community ecology and ecological diversification in emydid turtles” ($11,956; for Patrick Stephens).

National Science Foundation, Systematic Biology Program, 2002–2005, "Collaborative Research: Phylogeny and montane species richness in plethodontid salamanders" ($275,000 total; $125,000 to PI Wiens). Collaborative project with with fellow PI Paul Chippindale.

Netting Fund Grant, Carnegie Museum, 2002, “Phylogenetic relationships of hylid frogs in Middle America” ($6,695).

Netting Fund Grant, Carnegie Museum, 2001, “A phylogenetic analysis of the coevolution of ecology, morphology, and locomotor performance in emydid turtles” ($5,879) with Patrick Stephens (PI).

National Geographic Society, 2000–2001, “The evolution of herbivory in liolaemine lizards” ($9,400) co-PI with Robert Espinoza (PI).

Carnegie Institute, 2000–2002, "Establishing a molecular phylogenetics laboratory at Carnegie Museum of Natural History" ($275,000).

National Science Foundation, Research Collections in Systematics and Ecology, 1997, “Renovation of herpetological collections at Carnegie Museum of Natural History” ($49,727), co-PI with Ellen Censky (PI).

Provost’s Resarch Instrumentation Program (Univerity of Pittsburgh), 1997. “Molecular equipment for novel ecological and evolutionary research” ($14,908), with Susan Kalisz (PI) and Tia-Lynn Ashman (Co-PI).

Netting/O”Neill Fund Grants, Carnegie Museum, 1997-1998 “Speciation, phylogeny, and gene flow in the spiny lizards of the Sceloporus jarrovi complex inferred from mtDNA sequences and morphology” ($12,808).

National Science Foundation Graduate Fellowship, 1991–95. Ernst Mayr Grant, Harvard University, 1992 ($1,100). Gaige Fund Award, American Society of Ichthyologists and Herpetologists,

1992 ($500). Theodore Roosevelt Award, American Museum of Natural History, 1992

($500). 4 Research Fellowships, Department of Zoology, Univ. Texas, 1992-94 ($1,900). Research Fellowship, Texas Memorial Museum, 1994 ($300). Collection Study Grant, American Museum of Natural History, 1994 ($400). Sigma Xi Research Grant, 1992 ($300). Thomas J. Dee Award, Field Museum of Natural History, 1992 ($250). 3 Undergraduate Research Awards, Univ. Kansas, 1988-89 ($2,500). Panorama Society Grant, Univ. Kansas Museum of Natural History, 1989

($400). Research Grant, Dept. Systematics and Ecology, Univ. Kansas, 1989 ($500).

Awards and Honors:

ISI Highly Cited Researcher, 2014, 2015, 2018 American Society of Naturalists President's Award 2011 (for outstanding

paper published in the American Naturalist in 2010) for Kozak and Wiens 2010

BIOS Distinguished Lecturer, Univ. Nevada-Las Vegas, April 2009

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Kirschner Lecture, Department of Zoology, Washington State University, March 2008

Research Highlight in Nature (for Wiens, 2016; PLoS Biology) Research Highlight in Nature (for Jezkova and Wiens, 2016; Proc. R. Soc. Lond.) Research Highlight in Nature (for Scholl and Wiens, 2016; Proc. R. Soc. Lond.) Research Highlight in Nature Climate Change (for Quintero and Wiens, 2013;

Ecol. Lett.) Research Highlight in Nature (for Carrete-Vega and Wiens, 2012; Proc. R. Soc.

Lond.) Editor’s Choice in Science (for Wiens et al., 2011; Ecol. Lett.) Editor’s Choice in Science (for Wiens and Servedio, 2000; Proc. R. Soc. Lond.) “Recommended” paper in Faculty of 1000 Prime, December 2016 (Wiens,

2016, PLoS Biology) “Recommended” paper in Ecology, Faculty of 1000, July 2010 (Kozak and

Wiens 2010; Am. Nat.) “Recommended” paper in Ecology, Faculty of 1000, June 2009 (Wiens et al.

2009; Evolution) “Recommended” paper in Ecology, Faculty of 1000, November 2006 (Wiens et

al. 2006; Am. Nat.) “Must Read” paper in Ecology, Faculty of 1000, November 2004 (Espinoza et

al., 2004; PNAS) “Recommended” paper in Evolution, Faculty of 1000, August 2004 (Wiens,

2004; Am. Nat.) Outstanding Paper Award from Journal of Systematics and Evolution (for Wiens

and Moen 2008), for 2008–2013 Citation for top cited paper in Molecular Phylogenetics and Evolution 2008–2010

(for Townsend et al. 2008)—November 2010 Subject of “Inspirations” interview for the Systematics Association Newsletter,

2011 “Top Referee” for Proceedings of the Royal Society of London for 2011 Nominated for Treasurer, American Society of Naturalists, 2002 Honorable Mention for Ernst Mayr Award (Best Student Paper, Society of

Systematic Biologists), 1993. Honorable Mention for Best Student Paper Presentation, Herpetologist's

Leage, 1991. National Science Foundation Graduate Fellowship, 1991–1995. Outstanding Senior Honors Thesis in Biology, Univ. Kansas, 1991. Graduated Summa Cum Laude, with Honors from Division of Biological

Sciences, Univ. Kansas, 1991.

Invited seminars: 1990. Universidad Catolica, Quito, Ecuador 1992. National Museum of Natural History (Smithsonian) 1994. Carnegie Museum of Natural History, Washington University (St. Louis;

herpetology group) 1995. American Museum of Natural History, University of Pittsburgh,

University of Alabama 1996. University of Texas at Arlington, Slippery Rock University, Edinboro

University of Pennsylvania

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1997. Cornell University, San Diego State University 1998. George Washington University, Drexel University 1999. Ohio University, State University of New York at Stony Brook

(Department of Anatomical Sciences), University of Kansas (Natural History Museum), University of Nevada at Reno (two talks), University of Oregon

2000. American Museum of Natural History, Pennsylvania State University-Erie, Field Museum of Natural History (two talks), Rutgers University

2001. University of Nebraska (two talks), University of Texas at Arlington (student-invited speaker), University of Kansas (Natural History Museum), IV Congreso Argentino de Herpetologia (plenary lecture)

2002. University of California at Berkeley (two talks), State University of New York at Stony Brook (Department of Ecology and Evolution), University of Akron, University of Minnesota (Bell Museum), Brigham Young University

2003. Queens University (Ontario), Yale University (phylogenetics group) 2004. Hofstra University, University of Connecticut (two talks), University of

Iowa (Dept. Geosciences), Yale University (lecture in biogeography) 2005. Yale University, Smithsonian Tropical Research Institute-Panama

(Tupper lecture, Bambi talk), City University of New York–New York Consortium in Evolutionary Primatology

2006. Ohio State University, Center for Ecological Studies (Indian Institute of Science, Bangalore, India), Iowa State University, Harvard University, IX Reunion Nacional de Herpetologia, Mexico (plenary lecture), Stony Brook University (Living World public lecture series)

2007. University of Calgary, University of Rochester, University of Minnesota, Biodiversity Research Center-Academica Sinica (Taiwan), Hangzhou University (China), Louisiana State University, University of North Carolina-Chapel Hill

2008. University of Toronto-Mississauga, Washington State University (Kirschner Lecture; annual student-invited speaker), University of Montana, University of Oklahoma, Cornell University, American Museum of Natural History (niche modeling group), Fudan University (Shanghai, China; 4 talks), University of Connecticut (annual student-invited speaker in Ecology and Evolutionary Biology Dept.)

2009. University of Nevada, Las Vegas (BIOS Distinguished Lecturer; annual student-invited speaker), Institute of Advanced Studies (Jerusalem, Israel; Darwin 09 celebration), Instituto de Biologia (Universidad Nacional Autonoma de Mexico; Darwin 09 celebration); IV Congresso Brasileiro de Herpetologia (2 plenary lectures); Universidade de Goiania (Brazil); Instituto de Ecologia (Xalapa, Mexico; Darwin 09 Colloquium; student-invited plenary speaker); Dona Ana Biological Station (Sevilla, Spain); Berlin Museum (Germany)

2010. University of Chicago; Wageningen University (The Netherlands; student-invited speaker); National Museum of Natural History (Leiden, The Netherlands); Plenary lecture, XV Spanish Herpetological Congress (Sevilla); Plenary lecture, XI Reunion Nacional de Herpetologia, Mexico (Toluca); University of Arizona; Keynote lecture, Colombian

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Herpetology Association meeting (Medellin); Universidad Nacional de Colombia (Bogota)

2011. University of Toronto (Scarborough); City College of New York; Keynote Lecture, Munich Graduate Programs for Evolution, Ecology and Systematics Summer School, Chiemsee, Germany; Rice University; Universidad de Chile (Santiago); Opening Plenary Lecture, meeting of the Sociedad de Biología de Chile (Puerto Varas, Chile); McGill University

2012. New York Botanical Garden; Washington University (annual student-invited speaker for Ecology and Evolution Program); Fudan University Advanced Ecology Lecture; University of Central Florida; Columbia University

2013. Opening Keynote Lecture, BioSystEU (Vienna, Austria); University of Rennes (France); College of France (Paris); Porto Alegre Biological Evolution Workshop (Brazil); University of South Florida; American Museum of Natural History

2014. Keynote Lecture, 3rd Network for Neotropical Biogeography Meeting (Bogota, Colombia); U.S. Department of the Interior (webinar); University of Rochester; Qionzhou University (Sanya, China); Colorado State University; University of Colorado; University of California, Berkeley (Global Change Biology seminar series); École Normale Supérieure, Paris

2015. University of Zurich 2016. Sun-Yat Sen University (Guangzhou, China) 2017. State Key Laboratory of Systematic and Evolutionary Botany (Beijing,

China); South China Botanical Garden (Guangzhou, China); Kunming Institute of Botany (China); Nanjing Normal University (China); University of Arizona (School of Natural Resources and the Environment); Public Keynote lecture: Chiricahua-Peloncillo Heritage Days (Rodeo, New Mexico).

2018. Oklahoma State University, University of Florida (graduate student invited speaker)

2019. University of Georgia, University of New Mexico Invited symposium papers:

(1) “Phylogeny of Sceloporus I: Analysis of diverse data sets.” Symposium

entitled “Sceloporus. A model system.” American Society of Ichthyologists and Herpetologists and Herpetologist's League, Austin, TX, May, 1993 (with Tod W. Reeder).

(2) “Phylogeny of Sceloporus II: Corroboration and classification.” Symposium

entitled “Sceloporus. A model system.” American Society of Ichthyologists and Herpetologists and Herpetologist's League, Austin, TX, May, 1993 (with Tod W. Reeder).

(3) “Phylogenetic analysis of polymorphic morphological characters:

Inferences from empirical and simulated data sets.” Symposium

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entitled “Morphological data in phylogenetic analysis: Recent progress and unresolved problems.” Society of Systematic Biologists, Saint Louis, MO, June, 1996.

(4) “Evolution of display morphology and display behavior in spiny lizards

(genus Sceloporus): A statistical, phylogenetic analysis.” Symposium entitled “A tropical natural history” (in honor of William E. Duellman). Society for the Study of Amphibians and Reptiles, Lawrence, KS, August, 1996.

(5) “Phylogeny and taxonomy of hoplocercid lizards.” Symposium entitled

“Lizard systematics and evolutionary biology for the next millenium: A symposium honoring the life work of Richard Etheridge.” American Society of Ichthyologists and Herpetologists, Herpetologists’ League, Society for the Study of Amphibians and Reptiles, State College, PA, June, 1999 (wth Richard Etheridge).

(6) "Character selection and the methodology of morphological phylogenetics." Etheridge Symposium, State College, PA, June, 1999 (with Steve Poe).

(7) "War of the iguanas: Conflicting phylogenies, long-branch attraction, and disparate rates of molecular and morphological evolution in iguanid lizards." Etheridge Symposium, State College, PA, June, 1999 (wth Brad Hollingsworth).

(8) "Effects of including incomplete taxa and characters on phylogenetic

analysis" Symposium on missing data and phylogeny reconstruction. Society of Vertebrate Paleontology. Mexico City, October 2000.

(9) "Consensus, combination, and supertrees: integrating diverse data to

reconstruct evolutionary trees" DIMACS (Center for Discrete Mathematics and Theoretical Computer Science) Tutorial and Workshop on Bioconsensus II, Rutgers University, NJ, October 2001.

(10) "Phylogeny and paedomorphosis in salamanders" Symposium entitled

"Contemporary herpetological research: a tribute to Robert Jaeger" Herpetologists League, Kansas City, MO, July 2002.

(11) "Phylogenetic incongruence, combining data, and large-scale phylogenies"

Symposium entitled "Incongruence, data partitions, and phylogenetic signal" ICSEB VI, Sixth International Congress of Systematic and Evolutionary Biology, Patras, Greece, September, 2002.

(12) “Paedomorphosis and evolutionary morphology of salamanders”

Symposium entitled “Urodele phylogeny and reproduction” sponsored by the Society for the Study of Amphibians and Reptiles, Manaus, Brazil, June 2003.

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(13) “Hidden phylogenetic diversity of Neotropical treefrogs” Symposium entitled “Diversity and ecology of Amazonian frogs” sponsored by the Herpetologist’s League, Manaus, Brazil, June 2003.

(14) “The impact of missing data and incomplete taxa on phylogenetic

accuracy” Symposium entitled “Missing data in phylogenetic analysis” for NSF-funded Deep Time workshop, held at George Washington University, Washington, D.C., March 2004.

(15) “A phylogenetic perspective on global biodiversity patterns” Symposim

entitled “Phylogeny and biodiversity science” held at the “First DIVERSITAS Open Science Conference: Integrating biodiversity for human well-being” in Oaxaca, Mexico, November 2005 (with Michael Donoghue).

(16) “Evolutionary and ecological causes of the latitudinal diversity gradient in amphibians” American Society of Naturalist’s Vice Presidential Symposium entitled “Merging ecological and evolutionary approaches to interpreting geographic gradients in species richness.” June 24, 2006, Stony Brook, New York

(17) “Molecules, morphology, and species delimitation” Introduction for

symposium entitled “Species delimitation: new approaches for discovering diversity” sponsored by the Society of Systematic Biologists. June 23, 2006, Stony Brook, New York.

(18) “Paleontology, genomics, and combined-data phylogenetics: can

molecular data improve phylogeny estimation for fossil taxa?” Symposium entitled “Bringing together the living and dead: integrating extant and fossil biodiversity in evolutionary studies” sponsored by the Botanical Society of America. August 1, 2006, Chico, California.

(19) “Combining phylogenies and GIS-based methods to study evolutionary

and ecological patterns” symposium “Evoluionary biology in the 21st Century–tracing patterns of evolution through the Tree of Life” sponsored by the National Natural Science Foundation of China, Chinese Academy of Sciences, and DIVERSITAS. June 6, 2007, Beijing China.

(20) “Using GIS and biogeographic data to help understand character

evolution” colloquium “Integration of spatial and ecological data in evolutionary studies” sponsored by the Botanical Society of America, July 11, 2007, Chicago, Illinois.

(21) “Empowering herpetological phylogenetics with genomic resources”

symposium “Herpetology in the age of genomics“ sponsored by the Society for the Study of Amphibians and Reptiles. July 13, 2007, Saint Louis, Missouri.

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(22) “Why are there so many species of tropical bolitoglossine salamanders?” Symposium honoring David Wake at Fifth Conference on Biology of Plethodontid Salamanders, San Cristobal de Las Casas, Mexico, August 4, 2007.

(23) “Global biodiversity patterns: evolutionary and ecological causes” Buffon

International Symposium “Natural history museums and institutions in the 21st century.” October 18, 2007, Paris, France.

(24) “Phylogenetic perspectives on community assembly and the competition-

divergence-coexistence conundrum” Symposium on “Phylogenetic approaches to community ecology” sponsored by the Ecological Society of Japan, March 17, 2008. Fukuoka, Japan.

(25) “Evolutionary ecology and the Tree of Life” NSF-sponsored workshop

“Where to next with the Tree of Life?” April 8, 2008. Washington D.C. (26) “The evolution of snake-like body form in squamate reptiles: patterns,

rates, and levels of explanation” Symposium entitled: “Evolutionary transitions of body shape in extant amphibians and reptiles: call for integrative approaches” sponsored by the 6th World Congress of Herpetology, August 20, 2008. Manaus, Brazil.

(27) “The unexpected ecology of allopatric speciation” Keynote lecture for

symposium on “Mechanisms of speciation” at XXth International Congress of Zoology, August 29, 2008. Paris, France.

(28) “Phylogenies and paradigm shifts in evolutionary biology” Keynote

lecture for symposium “Recent advances in macroevolutionary approaches to evolutionary studies” European Society for Evolutionary Biology, August 24–29, 2009. Torino, Italy.

(29) “Using phylogenies in evolutionary biology and ecology” for workshop

“Phylogenetic methods and applications” August 31, 2009. Asti, Italy. (30) “The niche, large-scale biogeography, and species interactions ” for

symposium “Biogeography and ecology: two lenses in one telescope” at Vth International Biogeography Meetings, January 9, 2011. Heraklion, Crete, Greece.

(31) “Using phylogenies to study niche evolution” for symposium on trait

evolution at the Willi Hennig Society meetings, June 2012, Riverside, California.

(32) "Niche conservatism, ecology, and speciation" plenary lecture for

symposium "The role of behaviour in non-adaptive and non-ecological speciation" at the International Behavioral Ecology Congress, August 2012, Lund, Sweden.

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(33) "Speciation, niche conservatism, and niche divergence in theory and in amphibians" for symposium "Speciation in amphibians and reptiles: from patterns to processes and mechanisms" at the 7th World Congress of Herpetology, August 2012, Vancouver, British Colombia.

(34) "Understanding biodiversity patterns by integrating phylogenetic and

ecological information" plenary lecture for symposium "Mathematics for an evolving biodiversity", September 2013, Montreal, Canada.

(35) "A twisted view of ecology and speciation" for American Society of

Naturalists symposium "Beyond reproductive isolation: microevolutionary controls on macroevolutionary speciation dynamics", June 2014, Raleigh, North Carolina.

(36) “Why morphology-based phylogenies (sometimes) lie, and why it still

matters” for symposium on heterochrony and phylogenetics at Willi Hennig Society meeting, July 2014, Trento, Italy.

(37) “Phylogenies, niche evolution, and the origin and future of tropical

biodiversity” for symposium “The origin and future of tropical biodiversity: an evolutionary perspective”for the 51st meeting of the Association for Tropical Biology and Conservation, Cairns, Australia, July 2014.

(38) “The phylogeny of lizard families: resolving relationships, overcoming

controversy” for symposium “Reproductive biology and phylogeny of lizards” for the Joint Meeting of Herpetologists and Ichthyologists, August 2014, Chattanooga, Tennessee.

(39) “The ecology of speciation: a phylogenetic and biogeographic perspective”

Keynote lecture for symposium “Genomics of the speciation continuum” in Fribourg, Switzerland, September 2014.

(40) “Using phylogeny to understand the origins and future of biodiversity”

for symposium “Frontiers in integrative organismal biology: herpetological horizons” at the annual meetings of the Society for the Study of Amphibians and Reptiles in Lawrence, Kansas, July 2015.

(41) “Niche evolution versus climate change in diversity hotspots: phylogenetic

analyses in plants and animals” for symposium “Ecological implications of endemism hotspots as climate change refugia” at the annual meeting of the Ecological Society of America in Baltimore, Maryland, August 2015.

(42) “Lizard diversity: origins and future” plenary lecture for “9th Symposium

on the Lacertids of the Mediterranean Basin” in Limassol, Cyprus, June 2016.

(43) “Macroevolutionary patterns in the evolution of snake-like reptiles” for

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symposium “How lizards turn into snakes: Combining fossils, phylogeny, function, genetics, development, morphology, and ecology” at 8th World Congress of Herpetology, in Hangzhou, China, August 2016.

(44) “Using phylogenetic databases for large-scale herpetological research” for

symposium “Herpetological information in a networked world” at 8th World Congress of Herpetology, in Hangzhou, China, August 2016.

(45) “Climate change and the biogeography of a montane lizard in Arizona’s

Sky islands” for symposium on the biogeography of southwestern North America at Interional Biogeography Meetings, Tucson, Arizona, January 2017.

(46) “The evolution of biodiversity patterns” plenary lecture for the VI

meetings of the Colombian Assocation of Evolutionary Biology, Cali, Colombia, August 2017.

(47) “What to do with the new Tree of Life?” at the 59th Phylogenetic

Symposium “Phylogeny in the post-genomic era” in Berlin, Germany, 19 November 2017.

(48) “Ecological niche dynamics in biogeography” and “Evolutionary

approaches to diversity gradients” for workshop on “Eco-evolutionary dynamics in biogeograph” in Katalapi (Puerto Montt) Chile, October 2018

(49) “Making and using big trees” for workshop on “Trees in the Desert” in

Oracle, Arizona, April 2019

Publications:

(1) Wiens, J. J. 1989. Ontogeny of the skeleton of Spea bombifrons (Anura: Pelobatidae). Journal of Morphology 202:29–51.

(2) Wiens, J. J., and T. A. Titus. 1991. A phylogenetic analysis of Spea (Anura:

Pelobatidae). Herpetologica 47:21–28. (3) Wiens, J. J., and L. A. Coloma. 1992. A new species of the Eleutherodactylus

myersi (Anura: Leptodactylidae) assembly from Ecuador. Journal of Herpetology 26:196–207.

(4) Duellman, W. E., and J. J. Wiens. 1992. The status of the hylid frog genus

Ololygon and the recognition of Scinax Wagler, 1830. Occasional Papers of the Museum of Natural History University of Kansas 151:1–23.

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(5) Duellman, W. E., and J. J. Wiens. 1993. Hylid frogs of the genus Scinax Wagler 1830, in Amazonian Ecuador and Peru. Occasional Papers of the Museum of Natural History University of Kansas 153:1–57.

(6) Wiens, J. J. 1993. Phylogenetic relationships of phrynosomatid lizards and

monophyly of the Sceloporus group. Copeia 1993:287–299. (7) Wiens, J. J. 1993. Systematics of the leptodactylid frog genus Telmatobius

in the Andes of northern Peru. Occasional Papers of the Museum of Natural History University of Kansas 161:1–76.

(8) Wiens, J. J. 1993. Phylogenetic systematics of the tree lizards (genus

Urosaurus). Herpetologica 49:399–420. (9) Wiens, J. J. 1993. Review of "Herpetology" by George Zug. Systematic

Biology 42:592–596. (10) Chippindale, P. T., and J. J. Wiens. 1994. Weighting, partitioning, and

combining characters in phylogenetic analysis. Systematic Biology 43:278–287 (order of authorship alphabetical).

(11) Wiens, J. J., and P. T. Chippindale. 1994. Combining and weighting

characters and the prior agreement approach revisited. Systematic Biology 43:564–566.

(12) Wiens, J. J. 1995. Polymorphic characters in phylogenetic systematics.

Systematic Biology 44:482–500. (13) Wiens, J. J., and T. W. Reeder. 1995. Combining data sets with different

numbers of taxa for phylogenetic analysis. Systematic Biology 44:548–558.

(14) Wiens, J. J., and M. R. Morris. 1996. Character definitions, sexual

selection, and the evolution of swordtails. American Naturalist 147:866–869.

(15) Wiens, J. J., and D. M. Hillis. 1996. Accuracy of parsimony analysis using

morphological data: A reappraisal. Systematic Botany 21:237–243. (16) Reeder, T. W., and J. J. Wiens. 1996. Evolution of the lizard family

Phrynosomatidae as inferred from diverse types of data. Herpetological Monographs 10:43–84.

(17) Wiens, J. J., and M. R. Servedio. 1997. Accuracy of phylogenetic analysis

including and excluding polymorphic characters. Systematic Biology 46:332–345.

(18) Wiens, J. J. 1997. Review of “Homoplasy. The Recurrence of Similarity

in Evolution.” Copeia 1997:472–474.

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(19) Wiens, J. J., and T. W. Reeder. 1997. Phylogeny of the spiny lizards

(Sceloporus) based on molecular and morphological evidence. Herpetological Monographs 11:1–101.

**Cover article (20) Pianka, E. R., D. M. Hillis, M. R. Ryan, D. C. Cannatella, and J. J. Wiens.

1998. Teaching herpetology. Herpetologica 54:S3–S5. (21) Wiens, J. J., and M. R. Servedio. 1998. Phylogenetic analysis and

intraspecific variation: Performance of parsimony, likelihood, and distance methods. Systematic Biology 47:228–253.

(22) Wiens, J. J. 1998. The accuracy of methods for coding and sampling

higher-level taxa for phylogenetic analysis: A simulation study. Systematic Biology 47:381–397.

(23) Wiens, J. J. 1998. Testing phylogenetic methods with tree-congruence:

Phylogenetic analysis of polymorphic morphological characters in phrynosomatid lizards. Systematic Biology 47:411–428.

(24) Wiens, J. J. 1998. Combining data sets with different phylogenetic

histories. Systematic Biology 47:568–581. (25) Wiens, J. J. 1998. Does adding characters with missing data increase or

decrease phylogenetic accuracy? Systematic Biology 47:625–640. (26) Wiens, J. J. 1999. Phylogenetic evidence for multiple losses of a sexually

selected character in phrynosomatid lizards. Proceedings of the Royal Society of London, Series B 266:1529–1535.

(27) Wiens, J. J. 1999. Polymorphism in systematics and comparative biology.

Annual Review of Ecology and Systematics 30:327–362. (28) Wiens, J. J., T. W. Reeder, and A. Nieto Montes de Oca. 1999. Molecular

phylogenetics and evolution of sexual dichromatism among populations of the Yarrow’s spiny lizard (Sceloporus jarrovii ). Evolution 53:1884–1897.

(29) Wiens, J. J., and B. D. Hollingsworth. 2000. War of the iguanas:

conflicting molecular and morphological phylogenies and long-branch attraction in iguanid lizards. Systematic Biology 49:143–159.

(30) Wiens, J. J., and M. R. Servedio. 2000. Species delimitation in systematics:

inferring diagnostic differences between species. Proceedings of the Royal Society of London, Series B 267:631–636.

**Featured in "Editor's Choice" in Science (Vol. 288; p. 577)

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(31) Wiens, J. J. 2000. Decoupled evolution of display morphology and display behaviour in phrynosomatid lizards. Biological Journal of the Linnean Society 70:597–612.

(32) Wiens, J. J. 2000. Reconstructing phylogenies from allozyme data: testing

method performance with congruence. Biological Journal of the Linnean Society 70:613–632.

(33) Chippindale, P. T., A. H. Price, J. J. Wiens, and D. M. Hillis. 2000.

Phylogenetic relationships and systematic revision of central Texas hemidactyliine plethodontid salamanders. Herpetological Monographs 14:1–80.

(34) Wiens, J. J. (Editor). 2000. Phylogenetic Analysis of Morphological Data.

Smithsonian Institution Press, Comparative Evolutionary Biology Series.

(35) Hillis, D. M., and J. J. Wiens. 2000. Molecular versus morphological systematics: Conflicts, artifacts, and misconceptions. Pages 1–19 in Phylogenetic analysis of morphological data (J. J. Wiens, ed.). Smithsonian Institution Press, Washington, D.C.

(36) Poe, S., and J. J. Wiens. 2000. Character selection and the methodology of

morphological phylogenetics. Pages 20–36 in Phylogenetic analysis of morphological data (J. J. Wiens, ed.). Smithsonian Institution Press, Washington, D.C.

(37) Wiens, J. J. 2000. Coding morphological variation for phylogenetic

analysis: polymorphism and interspecific variation in higher taxa. Pages 115–145 in Phylogenetic analysis of morphological data (J. J. Wiens, ed.). Smithsonian Institution Press, Washington, D.C.

(38) Wiens, J. J. 2001. Widespread loss of sexually-selected traits: how the

peacock lost its spots. Trends in Ecology and Evolution 16:517–523. **Cover article (39) Wiens, J. J. 2001. Shape shifters. Time after time, lizards have dropped

their legs in favor of a snakelike body form. Natural History 110:70–75. (40) Wiens, J. J. 2001. Character analysis in morphological phylogenetics:

problems and solutions. Systematic Biology 50:689–699. (41) Wiens, J. J., and J. L. Slingluff. 2001. How lizards turn into snakes: a

phylogenetic analysis of body-form evolution in anguid lizards. Evolution 55:2303–2318.

(42) Wiens, J. J., and T. L. Penkrot. 2002. Delimiting species based on DNA

and morphological variation and discordant species limits in spiny lizards (Sceloporus). Systematic Biology 51:69–91.

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(43) Wiens, J. J. 2002. Review of “Patterns of Distribution of Amphibians” edited by William E. Duellman. Systematic Biology 51:980–981.

(44) Stephens, P. R., and J. J. Wiens. 2003. Explaining species richness from

continents to communities: the time-for-speciation effect in emydid turtles. American Naturalist 161:112–128.

(45) Wiens, J. J., and M. R. Morris. 2003. Review of "Sexual Selections" by

Marlene Zuk. Quarterly Review of Biology 78:127. (46) Wiens, J. J. 2003. Incomplete taxa, incomplete characters, and

phylogenetic accuracy: what is the missing data problem? Journal of Vertebrate Paleontology 23:297–310.

**Invited symposium paper (47) Wiens, J. J., P. T. Chippindale, and D. M. Hillis. 2003. When are

phylogenetic analyses misled by convergence? A case study in Texas cave salamanders. Systematic Biology 52:501–514.

**Cover article (48) Wiens, J. J. 2003. Missing data, incomplete taxa, and phylogenetic

accuracy. Systematic Biology 52:528–538. (49) Stephens, P. R., and J. J. Wiens. 2003. Ecological diversification and

phylogeny of emydid turtles. Biological Journal of the Linnean Society 79:577–610.

(50) Wiens, J. J., and R. E. Etheridge. 2003. Phylogenetic relationships of

hoplocercid lizards: coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica 59:375–398.

(51) Wiens, J. J. 2004. Speciation and ecology revisited: phylogenetic niche

conservatism and the origin of species. Evolution 58:193–197. (52) Wiens, J. J. 2004. What is speciation and how should we study it?

American Naturalist 163:914–923. **”Recommended” article in Evolution, Faculty of 1000 (53) Stephens, P. R., and J. J. Wiens. 2004. Convergence, divergence, and

homogenization in the ecological structure of emydid turtle communities: the effects of phylogeny and dispersal. American Naturalist 64:244–254.

(54) Wiens, J. J. 2004. The role of morphological data in phylogeny

reconstruction. Systematic Biology 53:653–661. (55) Wiens, J. J. 2004. Development and evolution of body form and limb

reduction in squamate reptiles: a response to Sanger and Gibson-Brown. Evolution 58:2107–2108.

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(56) Wiens, J. J., and M. J. Donoghue. 2004. Historical biogeography, ecology,

and species richness. Trends in Ecology and Evolution 19:639–644. **Featured on cover (57) Espinoza, R. E., J. J. Wiens, and C. R. Tracy. 2004. Recurrent evolution of

herbivory in small, cold-climate lizards: breaking the ecophysiological rules of reptilian herbivory. Proceedings of the National Academy of Sciences, U.S.A. 101:16819–16824.

**Featured on cover, with accompanying commentary article by Laurie Vitt (“Shifting paradigms: Herbivory and body size in lizards,” PNAS, 101:16713–16714)

**”Must Read”article in Ecology, Faculty of 1000 (58) Chippindale, P. T., R. M. Bonett, A. S. Baldwin, and J. J. Wiens. 2004.

Phylogenetic evidence for a major reversal in life history evolution in plethodontid salamanders. Evolution 58:2809–2822.

(59) Wiens, J. J., R. M. Bonett, and P. T. Chippindale. 2005. Ontogeny

discombobulates phylogeny: paedomorphosis and higher-level salamander phylogeny. Systematic Biology 54:91–110.

(60) Chippindale, P. T., and J. J. Wiens. 2005. Re-evolution of the larval stage

in the plethodontid salamander genus Desmognathus. Herpetological Review 36:113–117.

(61) Wiens, J. J. and C. H. Graham. 2005. Niche conservatism: integrating

evolution, ecology, and conservation biology. Annual Review of Ecology, Evolution, and Systematics 36:519–539.

(62) Wiens, J. J. 2005. Can incomplete taxa rescue phylogenetic analyses from

long-branch attraction? Systematic Biology 54:731–742. (63) Wiens, J. J., J. W. Fetzner, C. L. Parkinson, and T. W. Reeder. 2005. Hylid

frog phylogeny and sampling strategies for speciose clades. Systematic Biology 54:778–807.

(64) Smith, S. A., P. R. Stephens, and J. J. Wiens. 2005. Replicate patterns of

species richness, historical biogeography, and phylogeny in Holarctic treefrogs. Evolution 59:2433–2450.

(65) Wiens, J. J., M. C. Brandley, and T. W. Reeder. 2006. Why does a trait

evolve multiple times within a clade? Repeated evolution of snake-like body form in squamate reptiles. Evolution 61:123–141.

(66) Wiens, J. J. 2006. Missing data and the design of phylogenetic analyses.

Journal of Biomedical Informatics 39:34–42. **Invited review

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(67) Wiens, J. J., C. H. Graham, D. S. Moen, S. A. Smith, and T. W. Reeder. 2006. Evolutionary and ecological causes of the latitudinal diversity gradient in hylid frogs: treefrog trees unearth the roots of high tropical diversity. American Naturalist 168:579-596.

**Featured in “Science News” October 21, 2006 **Featured in “EurekAlert!” November 1, 2006 **Featured article for November 2006 on the American Naturalist web-site **Featured in “ScienceDaily” November 6, 2006 **Recommended article in Ecology, Faculty of 1000, November 9, 2006 **Featured (with illustration) in textbook “Evolution” by D. J. Futuyma (2009) (68) Wiens, J. J. 2006. Molecular evolution for the masses (review of “The

making of the fittest” by Sean Carroll). Bioscience 56:1014–1015. (69) Wiens, J. J., T. N. Engstrom, and P. T. Chippindale. 2006. Rapid

diversification, incomplete isolation, and the “speciation clock” in North American salamanders (genus Plethodon): testing the hybrid swarm hypothesis of rapid radiation. Evolution 60:2585–2603.

(70) Kozak, K. H., and J. J. Wiens. 2006. Does niche conservatism drive speciation? A case study in North American salamanders. Evolution 60:2604–2621.

**Featured (with illustration) in textbook “Evolution” by D. J. Futuyma (2009) (71) Wiens, J. J. 2007. Review of “The amphibian tree of life” by Frost et al.

Quarterly Review of Biology 82:55–56. (72) Wiens, J. J., G. Parra-Olea, M. Garcia-Paris, and D. B. Wake. 2007.

Phylogenetic history underlies elevational patterns of biodiversity in tropical salamanders. Proceedings of the Royal Society of London 274:919–928.

**Cover article (73) Smith, S. A., A. Nieto Montes de Oca, T. W. Reeder, and J. J. Wiens. 2007.

A phylogenetic perspective on elevational species richness patterns in Middle American treefrogs: why so few species in lowland tropical rainforests? Evolution 61:1188–1207.

(74) Wiens, J. J. 2007. Pissing away the eons. Review of “Phylogeny and

conservation” by Purvis et al. 2005. Evolution 61:1788–1790. (75) Wiens, J. J. 2007. Global patterns of species richness and diversification

in amphibians. American Naturalist 170:S86–S106. (76) Wiens, J. J., C. Kuczynski, W. E. Duellman, and T. W. Reeder. 2007. Loss

and re-evolution of complex life cycles in marsupial frogs: can ancestral trait reconstruction mislead? Evolution 61:1886–1899.

**Cover article.

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(77) Smith, S. A., S. Arif, A. Nieto Montes de Oca, and J. J. Wiens. 2007. A phylogenetic hotspot for evolutionary novelty in Middle American treefrogs. Evolution 61:2075–2085.

(78) Kozak, K. H., and J. J. Wiens. 2007. Climatic zonation drives latitudinal

variation in speciation mechanisms. Proceedings of the Royal Society of London 274:2995–3003.

(79) Wiens, J. J. 2007. Species delimitation: new approaches for discovering

diversity. Systematic Biology 56:875–878. (80) Stephens, P. R., and J. J. Wiens. 2008. Testing for evolutionary trade-offs

in a phylogenetic context: ecological diversification and locomotor performance in emydid turtles. Journal of Evolutionary Biology 21:77–87.

(81) Kozak, K. H., C. H. Graham, and J. J. Wiens. 2008. Integrating GIS data

into evolutionary studies. Trends in Ecology and Evolution 23:141–148. **Invited review **Featured on cover (82) Townsend, T. M., E. R. Alegre, S. T. Kelley, J. J. Wiens, and T. W. Reeder.

2008. Rapid development of multiple nuclear loci for phylogenetic analysis using genomic resources. Molecular Phylogenetics and Evolution 47:129–142

(83) Wiens, J. J. 2008. Systematics and herpetology in the age of genomics.

Bioscience 58:297–307. **Invited review **Cover article (84) Gomez-Mestre, I., J. J. Wiens, and K. M. Warkentin. 2008. Evolution of

adaptive plasticity: risk-sensitive hatching in Neotropical leaf-breeding treefrogs (Agalychnis: Hylidae). Ecological Monographs 78:205–224.

(85) Wiens, J. J., and D. S. Moen. 2008. Missing data and the accuracy of

Bayesian phylogenetics. Journal of Systematics and Evolution 46:307–314. **Invited symposium paper **Award from journal for “Outstanding Paper” for 2008–2013 (86) Wiens, J. J., and J. T. Hoverman. 2008. Digit reduction, body size, and

paedomorphosis in salamanders. Evolution and Development 10:449–463. (87) Wiens, J. J., C. A. Kuczynski, S. A. Smith, D. Mulcahy, J. W. Sites, Jr., T.

M. Townsend, and T. W. Reeder. 2008. Branch lengths, support, and congruence: testing the phylogenomic approach with 20 nuclear loci in snakes. Systematic Biology 57:420–431.

(88) Brandley, M. C., J. P. Huelsenbeck, and J. J. Wiens. 2008. Rates and

patterns in the evolution of snake-like body form in squamate reptiles:

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evidence for repeated re-evolution of lost digits and long-term persistence of intermediate body forms. Evolution 62:2042–2064.

(89) Wiens, J. J. 2008. Commentary on Losos (2008): niche conservatism déjà

vu. Ecology Letters 11:1004–1005. (90) Moen, D. S., and J. J. Wiens. 2009. Phylogenetic evidence for

competitively-driven divergence: body-size evolution in Caribbean treefrogs (Hylidae: Osteopilus). Evolution 63:195–214.

(91) Wiens, J. J. 2009. Estimating rates and patterns of morphological

evolution from phylogenies: lessons in limb lability from Australian Lerista lizards. Journal of Biology 8:19.

**Invited review (92) Stephens, P. R., and J. J. Wiens. 2009. Evolution of sexual size

dimorphisms in emydid turtles: ecological dimorphism, Rensch's rule, and sympatric divergence. Evolution 63:910–925.

(93) Wiens, J. J., J. Sukumaran, R. A. Pyron, and R. M. Brown. 2009.

Evolutionary and biogeographic origins of high tropical diversity in Old World frogs (Ranidae). Evolution 63:1217–1231.

**”Recommended” article in Ecology, Faculty of 1000 **Featured on cover (94) Kozak, K. H., R. W. Mendyk, and J. J. Wiens. 2009. Can parallel

diversification occur in sympatry? Repeated patterns of body-size evolution in co-existing clades of North American salamanders. Evolution 63:1769–1784.

(95) Wiens, J. J. 2009. Paleontology, genomics, and combined-data

phylogenetics: can molecular data improve phylogeny estimation for fossil taxa? Systematic Biology 58:87–99.

(96) Adams, D. C., C. M. Berns, K.H. Kozak, and J. J. Wiens. 2009. Are rates of

species diversification correlated with rates of morphological evolution? Proceedings of the Royal Society of London, Series B 276:2729–2738.

(97) Moen, D. S., S. A. Smith, and J. J. Wiens. 2009. Community assembly

through evolutionary diversification and dispersal in Middle American treefrogs. Evolution 63:3228–3247.

(98) Stephens, P. R., and J. J. Wiens. 2009. Bridging the gap between

biogeography and community ecology: niche conservatism and community structure in emydid turtles. Molecular Ecology 65:246–259.

(99) Hua, X., C. Fu, J. Li, A. Nieto-Montes de Oca, and J. J. Wiens. 2009. A

revised phylogeny of Holarctic treefrogs (genus Hyla) based on nuclear and mitochondrial DNA sequences. Herpetologica 65:246–259.

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(100) Wiens, J. J., and M. C. Brandley. 2009. The evolution of limblessness. The

Grzimek Encyclopedia of Animal Evolution. (101) Wiens, J. J., C. A. Kuczynski, S. Arif, and T. W. Reeder. 2010.

Phylogenetic relationships of phrynosomatid lizards based on nuclear and mitochondrial data, and a revised phylogeny for Sceloporus. Molecular Phylogenetics and Evolution 54:150–161.

(102) Hua, X., and J. J. Wiens. 2010. Latitutudinal variation in speciation

mechanisms in frogs. Evolution 64:429–443. (103) Wiens, J. J., C. A. Kuczynski, and P. R. Stephens. 2010. Discordant

mitochondrial and nuclear gene phylogenies in emydid turtles: implications for speciation and conservation. Biological Journal of the Linnean Society 99:445–461.

(104) Kozak, K. H., and J. J. Wiens. 2010. Niche conservatism drives

elevational diversity patterns in Appalachian salamanders. American Naturalist 176:40–54.

**”Recommended” article in Ecology, Faculty of 1000 **Winner of the 2011 President's Award by the American Society of Naturalists

(for an outstanding paper published in the American Naturalist in 2010). (105) Wiens, J. J., C. A. Kuczynski,X. Hua, and D. S. Moen. 2010. An expanded

phylogeny of treefrogs (Hylidae) based on nuclear and mitochondrial sequence data. Molecular Phylogenetics and Evolution 55:871–882.

(106) Buckley, L. B., T. J. Davies, D. D. Ackerly, N. J. B. Kraft, S. P. Harrison, B.

L. Anacker, H. V. Cornell, E. I. Damschen, J. A. Grytnes, B. A. Hawkins, C. M. McCain, P. R. Stephens, and J. J. Wiens. 2010. Phylogeny, niche conservatism, and the latitudinal diversity gradient in mammals. Proceedings of the Royal Society of London, Series B 277:2131–2138.

(107) Wiens, J. J., D. D. Ackerly, A. P. Allen, B. L. Anacker, L. B. Buckley, H. V.

Cornell, E. I. Damschen, T. J. Davies, J. A. Grytnes, S. P. Harrison, B. A. Hawkins, R. D. Holt, C. M. McCain, and P. R. Stephens. 2010. Niche conservatism as an emerging principle in ecology and conservation biology. Ecology Letters 13:1310–1324.

**Invited review (108) Kozak, K. H., and J. J. Wiens. 2010. Accelerated rates of climatic niche

evolution underlie rapid species diversification. Ecology Letters 13:1378–1389.

(109) Wiens, J. J., C. A. Kuczynski,T. Townsend, T. W. Reeder, D. G. Mulcahy,

and J. W. Sites, Jr. 2010. Combining phylogenomics and fossils in higher level squamate reptile phylogeny: molecular data change the placement of fossil taxa. Systematic Biology 59:674–688.

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(110) Pyron, R. A., F. T. Burbrink, G. R. Colli, A. Nieto Montes de Oca, L. J.

Vitt, C. A. Kuczynski, and J. J. Wiens. 2011. The phylogeny of advanced snakes (Colubroidea), with discovery of a new subfamily and comparison of support methods for likelihood trees. Molecular Phylogenetics and Evolution 58:329–342.

(111) Wiens, J. J. 2011. Re-evolution of lost mandibular teeth in frogs after

more than 200 million years, and re-evaluating of Dollo’s law. Evolution 65:1283–1296.

**Featured in The New York Times, BBC News, National Geographic News, EarthSky, The Scientist (under Naturally Selected, “Biology’s Personal Best”), and in news outlets from 23 countries including the U.S., U.K., Turkey, Iran, Russia, Netherlands, Hungary, Norway, Spain, Italy, Poland, Belgium, Colombia, Cuba, Ecuador, Mexico, Brazil, Indonesia, India, Bangladesh, Pakistan, China, Thailand

(112) Wiens, J. J., R. A. Pyron, and D. S. Moen. 2011. Phylogenetic origins of local-scale diversity patterns and the causes of Amazonian megadiversity. Ecology Letters 14:643–652.

**Editor's Choice in Science (May 2011) **Featured on msnbc.com, TODAY.com, BBC news, Yahoo News, LiveScience, ScienceDaily, Top Science News, and others, with interviews on EarthSky and OurAmazingPlanet

(113) Wiens, J. J. 2011. The causes of species richness patterns across space, time, and clades and the role of “ecological limits.” Quarterly Review of Biology 86:75–96.

(114) Wiens, J. J., and M. C. Morrill. 2011. Missing data in phylogenetic

analysis: reconciling results from simulations and empirical data. Systematic Biology 60:719–731.

(115) Wiens, J. J. 2011. The niche, biogeography, and species interactions.

Philosophical Transactions of the Royal Society of London, Series B 366:2336–2350.

***Invited paper for symposium on "Ecology and Biogeography" (116) Wiens, J. J., M. Sparreboom, and J. W. Arntzen. 2011. Crest evolution in

newts: implications for reconstruction methods, sexual selection, phenotypic plasticity, and the origin of novelties. Journal of Evolutionary Biology 24:2073–2086.

***Cover article. (117) Townsend, T., D. G. Mulcahy, J. W. Sites Jr., C. A. Kuczynski, J. J. Wiens,

and T. W. Reeder. 2011. Phylogeny of iguanian lizards inferred from 29 nuclear loci, and a comparison of concatenated and species-tree approaches for an ancient, rapid radiation. Molecular Phylogenetics and Evolution 61:363–380.

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(118) Pyron, R. A., and J. J. Wiens. 2011. A large-scale phylogeny of Amphibia

including over 2,800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61:543–583.

(119) Fisher-Reid, M. C., and J. J. Wiens. 2011. What are the consequences of

combining nuclear and mitochondrial data for phylogenetic analysis? Lessons from Plethodon salamanders and 13 other vertebrate clades. BMC Evolutionary Biology 11:300

***Highly accessed (120) Sites, J. W., Jr., T. W. Reeder, and J. J. Wiens. 2011. Phylogenetic insights

on evolutionary novelties in lizards and snakes: sex, birth, bodies, niches, and venom. Annual Review of Ecology, Evolution and Systematics 42:227–244.

**Invited review (121) Carrete-Vega, G., and J. J. Wiens*. 2012. Why are there so few fish in the

sea? Proceedings of the Royal Society of London 279:2323–2329. (published on line 8 Feb.)

*Corresponding author ***Featured as “Research Highlight” in Nature, 16 Feb. 2012 ***Phone interviews on CBS News Radio, Deutschlandfunk Radio (Germany), EarthSky ***Featured on many news web sites including CBS News, msnbc.com, EarthSky, LiveScience, NewScientist, DailyMail, Huffington Post, World News Australia, TheNewsWorld, WebIndia123, TopNews (India), SkyNews (Australia), NewsTrackIndia, LeFigaro (France), Spiegel Online (Germany), News Pano, PhysOrg.com, WorldNews.com, Yahoo News, Dallas Morning News, Australian Geographic, United Press International, Press Association, OnePakistan, IndiaVision, Zanost.com (Croatia), Dalje.com (Croatia), JornalCiencia (Brazil), ExpressoAmbiental (Brazil), TechMoney (Poland), Polska Agencja Prasowa (Poland), Technologijos (Lithuania), aePiot (Romania), Seasons (France), Stop (Hungary), Nu (Netherlands), News Center (China), China Daily (China), Mysterious Earth (China), and others for a total of 17 countries (U.S., U.K., Australia, New Zealand, France, Germany, Netherlands, Croatia, Lithuania, Poland, Hungary, Romania, Czech Republic, Brazil, China, India, Pakistan)

(122) Blankers, T., D. C. Adams, and J. J. Wiens*. 2012. Ecological radiation

with limited morphological diversification in salamanders. Journal of Evolutionary Biology 25:634–646.

*Corresponding author (123) Kozak, K. H., and J. J. Wiens. 2012. Phylogeny, ecology, and the origins of

climate-richness relationships. Ecology 93:S167–S181. **Invited paper for special issue on phylogeny and ecology.

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(124) Wiens, J.J., and J. Tiu. 2012. Highly incomplete taxa can rescue phylogenetic analyses from the negative impacts of limited taxon sampling. PLoS One 7:42925.

(125) Gomez-Mestre, I., R. A. Pyron, and J. J. Wiens. 2012. Phylogenetic

analyses reveal unexpected patterns in the evolution of reproductive modes in frogs. Evolution 66:3687–3700.

***Featured on cover ***Featured in ScienceDaily, daily-news.eu, innovations report (Germany), Sci

Guru-Science News, TendenciasCientificas (Spain), and other websites (126) Fisher-Reid, M.C., K. H. Kozak, and J. J. Wiens. 2012. How is the rate of

climatic-niche evolution related to climatic-niche breadth? Evolution 66:3836–3851.

(127) Wiens, J. J., C. R. Hutter, D. G. Mulcahy, B. P. Noonan, T. M. Townsend,

J. W. Sites, Jr., and T. W. Reeder. 2012. Resolving the phylogeny of lizards and snakes (Squamata) with extensive sampling of genes and species. Biology Letters 6:1043–1046. (published on-line 18 September 2012)

***Featured in ScienceNow, ScienceDaily, EarthTimes, United Press International (upi.com), PhysOrg.com, and others

***Featured in Science magazine (14 Dec. 2012) (128) Wiens, J.J. 2012. Why biogeography matters: historical biogeography

versus phylogeography and community phylogenetics for inferring evolutionary and ecological processes. Frontiers of Biogeography (published online).

***Invited perspectives piece (129) Mulcahy, D. G., B. P. Noonan, T. Moss, T. M. Townsend, T. W. Reeder, J.

W. Sites, Jr., and J. J. Wiens. 2012. Estimating divergence dates and evaluating dating methods using phylogenomic and mitochondrial data in squamate reptiles. Molecular Phylogenetics and Evolution 65:974–991.

(130) Cahill, A.E., M. E. Aiello-Lammens, M. C. Fisher-Reid, X. Hua, C. J.

Karanewsky, H. Y. Ryu, G. C. Sbeglia, F. Spagnolo, J. B. Waldron, O. Warsi, and J. J. Wiens*. 2013. How does climate change cause extinction? Proceedings of the Royal Society of London 280: 20121890.

*Corresponding author ***Cover article ***Most cited paper published in PRSL in 2013 ***Featured on many news websites (including The Guardian (UK), DailyMail (UK), MSN News (UK), YahooNews (UK), Business Insider), science websites (including ScienceDaily, physorg, Royal Society Website, io9, EarthTimes, and SciTechDaily), and environmental websites and blogs (including Conservation Magazine and SimpleClimate)

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(131) Blankers, T., T. M. Townsend, K. Pepe, T. W. Reeder, and J. J. Wiens. 2013. Contrasting global-scale evolutionary radiations: phylogeny, diversification, and morphological evolution in the major clades of iguanian lizards. Biological Journal of the Linnean Society 108:127–143.

*Corresponding author (132) Quintero, I., and J. J. Wiens*. 2013. What determines the climatic niche

width of species? The role of spatial and temporal climatic variation in three vertebrate clades. Global Ecology and Biogeography 22:422–432.

*Corresponding author

(133) Wiens, J. J., K. H. Kozak, and N. Silva. 2013. Diversity and niche evolution along aridity gradients in North American lizards (Phrynosomatidae). Evolution 67:1715–1728.

(134) Hua, X., and J. J. Wiens. 2013. How does climate influence speciation?

American Naturalist 182:1–12. (135) Pyron, R.A., F. T. Burbrink, and J. J. Wiens. 2013. A phylogeny and

revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 13:93

(136) Guerrero, P. C., M. Rosas, M. T. K. Arroyo, and J. J. Wiens. 2013.

Evolutionary lag times and recent origin of the biota of an ancient desert (Atacama-Sechura). Proceedings of the National Academy of Sciences, U.S.A. 110:11469–11474.

***Featured in newspaper “El Mercurio” (Chile) and on website “Climate News Network” (UK)

(137) Quintero, I., and J. J. Wiens*. 2013. Rates of projected climate change dramatically exceed past rates of climatic-niche evolution among vertebrate species. Ecology Letters 16:1095–1103.

*Corresponding author **Featured as “Research Highlight” in Nature Climate Change **TV interview: CNN-International (July 20, 2013) **Radio interview on: Minnesota Public Radio (July 31, 2013) **Interviews for: ScienceNOW, DiscoveryNews, MotherJones, UAnews, ArizonaDailyStar **Coverage on the web at many news, science, and environmental web sites, including: RollingStone (US), Huffington Post (US), The Guardian (UK), The Conversation (Australia), MotherJones (US), Examiner (UK), wired (UK), weather.com, ScienceNOW, DiscoveryNews, TheScientist, Science News, Smithsonian.com, futurity.org, Science Daily, upi.com, phys.org, io9, e!ScienceNews (Eureka), digitaljournal, dailytech.com, redorbit, natureworldnews, vr-zone, scitechdaily, silobreaker, worldnewsviews, newsvine, sciencenewsline, scinewsnetwork, inhabitat, ArsTechnica, policymic, OracleTalk, MotherNatureNetwork, envirobeat, thinkprogress, WorldNews.org, climateark, globalactionnow.org, earthaction, forests.org, molecularecologist, Kaleton, pollutionsolutions, insideclimatenews, televisa (Spanish), wiadmosci

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(Poland), Mongabay (Indonesia), Taipei Times (Taiwan), Cromo (Uruguay), El Once (Argentina), ZeeNews (India), TruthDive (India), YahooNews (India), sify.com (India), NewKerala (India), NeoFronteras, LaFlecha, Globedia, Taringa, diarioecologia (Spanish), acclimate, HypeScience (Portugese), Posta (Turkey), Vatan (Turkey), insanhaber (Turkey), Atlantico (France), Corriere (Italy), NotizarioItaliono (Italy), meteo (Italy), China Daily (China), Zimbabwe Star (Zimbabwe), Mail&Guardian (South Africa), SverigeRadio (Sweden), star.com (Toronto Star; Canada), stuff.org (New Zealand), uanews, ArizonaDailyStar

(138) Hutter, C. R., J. M. Guayasamin, and J. J. Wiens. 2013. Explaining

Andean megadiversity: the evolutionary and ecological causes of glassfrog elevational richness patterns. Ecology Letters 16:1135–1144.

(139) Machac, A., D. Storch, and J. J. Wiens. 2013. Ecological causes of

decelerating diversification in carnivoran mammals. Evolution 67:2423–2433.

(140) Lambert, S. M., and J. J. Wiens. 2013. Evolution of viviparity: a

phylogenetic test of the cold-climate hypothesis in phrynosomatid lizards. Evolution 67:2614–2630.

***Featured on ScienceNOW ***Cover article

(141) Fisher-Reid, M. C., T. N. Engstrom, C. A. Kuczynski, P. R. Stephens, and

J. J. Wiens. 2013. Parapatric divergence of sympatric morphs in a salamander: incipient speciation on Long Island? Molecular Ecology 22:4681–4694.

(142) Pyron, R.A., and J. J. Wiens. 2013. Large-scale phylogenetic analyses

reveal the causes of high tropical amphibian diversity. Proceedings of the Royal Society of London 280: 20131622

(143) Moen, D., D. J. Irschick, and J. J. Wiens. 2013. Evolutionary conservatism

and convergence both lead to striking similarity in ecology, morphology, and performance across continents in frogs. Proceedings of the Royal Society of London 280:20132156.

(144) Bonett, R., M. Steffen, S. Lambert, J. J. Wiens, and P. T. Chippindale. 2014.

Evolution of paedomorphosis in plethodontid salamanders: ecological correlates and re-evolution of metamorphosis. Evolution 68:466–482.

(145) Cahill, A.E., M. E. Aiello-Lammens, M. C. Fisher-Reid, X. Hua, C. J.

Karanewsky, H. Y. Ryu, G. C. Sbeglia, F. Spagnolo, J. B. Waldron, and J. J. Wiens*. 2014. Causes of warm-edge range limits: systematic review, proximate factors and implications for climate change. Journal of Biogeography 41:429–442.

*Corresponding author

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(146) Zeng, C., I. Gomez-Mestre, and J. J. Wiens*. 2014. Evolution of rapid development in spadefoot toads is unrelated to arid environments. PLoS One 9:e96673.

*Corresponding author (147) Pinto-Sánchez, N. R., A. J. Crawford, and J. J. Wiens*. 2014. Using

historical biogeography to test for community saturation. Ecology Letters 17:1077–1085.

*Corresponding author **Featured on cover (148) Jiang, W., S.-Y. Chen, H. Wang, and J. J. Wiens. 2014. Should genes with

missing data be excluded from phylogenetic analyses? Molecular Phylogenetics and Evolution 80:308–318.

(149) Bonetti, M. F., and J. J. Wiens*. 2014. Evolution of climatic niche

specialization: a phylogenetic analysis in amphibians. Proceedings of the Royal Society of London 281:2013229.

*Corresponding author (150) Chejanovski, Z., and J. J. Wiens. 2014. Climatic niche breadth and species

richness in temperate treefrogs. Journal of Biogeography 10:1936–1946.

(151) Lambert, S. M., T. W. Reeder, and J. J. Wiens. 2015. When do species-tree and concatenated estimates disagree? An empirical analysis with higher-level scincid phylogeny. Molecular Phylogenetics and Evolution 82:146–155.

(152) Reeder, T.W., T. M. Townsend, D. G. Mulcahy, B. P. Noonan, P. L. Wood,

Jr., J. W. Sites, Jr. and J. J. Wiens*. 2015. Integrated analyses resolve conflicts over squamate reptile phylogeny and reveal unexpected placements for fossil taxa. PLoS One 10:e0118199.

*Corresponding author (153) Qian, H., J. J. Wiens, J. Zhang, and Y. Zhang. 2015. Evolutionary and

ecological causes of species richness patterns in North American angiosperm trees. Ecography 38:241–250

(154) Gomez-Rodriguez, C., A. Baselga, and J. J. Wiens. 2015. Is diversification

rate related to climatic niche width? Global Ecology and Biogeography 24:383–395.

(155) Zheng, Y., and J. J. Wiens*. 2015. Do missing data influence the accuracy

of divergence-time estimation with BEAST? Molecular Phylogenetics and Evolution 85:41–49.

*Corresponding author

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(156) Fisher-Reid, M. C., and J. J. Wiens. 2015. Is geographic variation within species related to macroevolutionary patterns between species? Journal of Evolutionary Biology 28:1502–1515.

(157) Wiens, J. J. 2015. Explaining large-scale patterns of vertebrate diversity.

Biology Letters 11:20150506.

(158) Wiens, J. J., R. T. Lapoint, and N. K. Whiteman. 2015. Herbivory increases diversification across insect clades. Nature Communications 6:8370.

(159) Wiens, J. J. 2015. Faster diversification on land than sea helps explain

global biodiversity patterns among habitats and animal phyla. Ecology Letters 18:1234–1241.

(160) Streicher, J. W., J. A. Schulte, and J. J. Wiens. 2016. How should genes and

taxa be sampled for phylogenomic analyses with missing data? An empirical study in iguanian lizards. Systematic Biology 65:128–145.

(161) Moen, D. S., H. Morlon, and J. J. Wiens. 2016. Testing convergence versus

history: convergence dominates phenotypic evolution for over 150 million years in frogs. Systematic Biology 65:146–160.

(162) Zheng, Y., and J. J. Wiens*. 2016. Combining phylogenomic and

supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4,162 species. Molecular Phylogenetics and Evolution 94:537–547.

*Corresponding author (163) Streicher, J. W., and J. J. Wiens*. 2016. Phylogenomic analyses reveal

novel relationships among snake families. Molecular Phylogenetics and Evolution 100:160–169.

*Corresponding author

(164) Kozak, K. H., and J. J. Wiens*. 2016. Testing the relationships between diversification, species richness, and trait evolution. Systematic Biology 65:975–988.

*Corresponding author (165) Kozak, K. H., and J. J. Wiens*. 2016. What explains patterns of species

richness? The relative importance of climatic niche evolution, morphological evolution, and ecological limits in salamanders. Ecology and Evolution 6:5940–5949.

*Corresponding author (166) Scholl, J.P., and J. J. Wiens*. 2016. Diversification rates and species

richness across the Tree of Life. Proceedings of the Royal Society of London 283: 20161335.

*Corresponding author **Featured in Nature’s “Research Highlights”

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(167) Cang, F.A., A. A. Wilson, and J. J. Wiens*. 2016. Climate change is

projected to outpace rates of niche change in grasses. Biology Letters 12:20160368.

*Corresponding author **Featured on news and science websites including BBC News (with

interview), MSN News (1 of 5 “Science Stories of the Week” U.S., South Africa), The Independent, Irish Examiner, New Zealand Herald, SBS (Australia), BreakingNews (Ireland), ThinkProgress, Daily Kos, NewScientist, phys.org, and others

(168) Jezkova, T., and J. J. Wiens*. 2016. Rates of change in climatic niches in

plant and animal populations are much slower than projected climate change. Proceedings of the Royal Society of London 283:20162104.

*Corresponding author **Featured in Nature’s “Research Highlights” **Featured in on various news, science, and environmental websites, including

BBC News (with interview), MSN, The Independent (UK), Gizmodo (Australia, Brasil), ScienceWorldReport, MarketBusinessNews, El Ciudadano (Chile), CDNews (Taiwan), Baomoi (Vietnam), Skanaa (Indonesia), GontorNews (Indonesia), Epoch News (China), BCharts (Brasil), Dunya (Turkey), Umhuriyet (Turkey), Aktualne-Spravy (Slovakia), Webnovinsky (Slovakia), Chronmy Klimat (Poland), Iran Daily, Pravo (Czech Republic)

(169) Wiens, J. J. 2016. Climate-related local extinctions are already

widespread among plant and animal species. PLoS Biology 11, e2001104. **Featured in Nature’s “Research Highlights” **Recommended by Faculty1000 Prime **Featured in on various news, science, and environmental websites and

magazines, including interviews with Los Angeles Times, CBC News, BBC Focus Magazine, Popular Science, Pacific Standard Magazine, Inside Climate News, The Independent, Seeker.com, Mongabay, and reported on at TIME, The Guardian (UK), National Geographic, Deutsche Welle (Germany), Science Daily, Phys.org, Iran Daily, El Ciudadano (Chile), AccuWeather, FoxNews

**Radio interview on Ireland’s “The Moncrief Show” on NewsTalk Radio **Radio interview on “The Show about Science” (170) Jezkova, T., and J. J. Wiens*. 2017. What explains patterns of

diversification and richness among animal phyla? American Naturalist 189:201–212.

*Corresponding author **Featured on various science-related websites, including ScienceDaily,

futurity.org, phys.org, earth.com, scienmag, Astrobiology Magazine, ScienceSprings, Hail Science, latesttechnology, technology.org, Emag, The Exception, LifeSciencesWorld, GaiaNews (Italy)

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**Featured on some news sites, including Express (UK), Pacific Standard Magazine (with interview), Novinky.cz (Czech Republic), Sozcu (Turkey)

**Radio interview on BYU News (171) Wiens, J. J. 2017. What explains patterns of biodiversity across the Tree of

Life? BioEssays 39:1600128. *Invited review

(172) Pontarp, M., and J. J. Wiens. 2017. The origin of species richness patterns

along environmental gradients: uniting explanations based on time, diversification rate, and carrying capacity. Journal of Biogeography 44:722–735.

*Designated as “Special Paper”

(173) Moen, D. S., and J. J. Wiens. 2017. Microhabitat and climatic-niche change explain patterns of diversification among frog families. American Naturalist 190:29–44.

(174) Miller, E. C., and J. J. Wiens. 2017. Extinction and time help drive the

marine-terrestrial biodiversity gradient: is the ocean a deathtrap? Ecology Letters 20:911–921.

(175) Anderson, S. R., and J. J. Wiens*. 2017. Out of the dark: 350 million years

of conservatism and evolution in diel activity patterns in vertebrates. Evolution 71:1944–1959.

*Corresponding author (176) Lin, L., and J. J. Wiens*. 2017. Comparing macroecological patterns across

continents: evolution of climatic niche breadth in varanid lizards. Ecography 40:960–970.

*Corresponding author (177) Larsen, B. B., E. C. Miller, M. K. Rhodes, and J. J. Wiens*. 2017.

Inordinate fondness multiplied and redistributed: the number of species on Earth and the new Pie of Life. Quarterly Review of Biology 92:229–265.

*Corresponding author ***Featured on some science news websites, including Science Daily, Science

Newsline, phys.org, Ensia.com (178) Bars-Closel, M., T. Kohlsdorf, D. S. Moen, and J. J. Wiens*. 2017.

Diversification rates are more strongly related to microhabitat than climate in squamate reptiles (lizards and snakes). Evolution 71:2243–2261.

*Corresponding author (179) Streicher, J. W., and J. J. Wiens. 2017. Phylogenomic analyses of more

than 4,000 nuclear loci resolve the origin of snakes among lizard families. Biology Letters 13:20170393.

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(180) Hutter, C. R., S. M. Lambert, and J. J. Wiens. 2017. Rapid diversification

and time explain amphibian species richness at different scales in the Tropical Andes, Earth's most biodiverse hotspot. American Naturalist 190:828–843.

(181) Meyer, A.L.S., and J. J. Wiens*. 2018. Estimating diversification rates for

higher taxa: BAMM can give problematic estimates of rates and rate shifts. Evolution 72:39–53.

*Corresponding author (182) Streicher, J. W., E. C. Miller, P. C. Guerrero, C. Correa, J. C. Ortiz, A. J.

Crawford, M. R. Pie, and J. J. Wiens. 2018. Evaluating methods for phylogenomic analyses, and a new phylogeny for a major frog clade (Hyloidea) based on 2,214 loci. Molecular Phylogenetics and Evolution 119:128–143.

(183) Hanschen, E.R., M.D. Herron, J. J. Wiens, H. Nozaki, and R. E. Michod.

2018. Repeated evolution and reversibility of self-fertilization in the volvocine green algae. Evolution 72:386–398.

(184) Wiens, J. J. 2018. Patterns of local community composition are linked to

large-scale diversification and dispersal of clades. American Naturalist 191:184–196.

(185) Yan, H.-F., Cai-Yun Zhang, A. A. Anderberg, G. Hao, X.-J. Ge, J. J.

Wiens*. 2018. What explains high plant richness in East Asia? Time and diversification in the tribe Lysimachieae (Primulaceae). New Phytologist 219:436–448

*Corresponding author (186) Miller, E. C., and J. J. Wiens. 2018. Demystifying the marine-terrestrial

biodiversity gradient: Response to Vermeij et al. Ecology Letters 21:940–941.

(187) Jezkova, T., and J. J. Wiens. 2018. Testing the role of climate in speciation:

new methods and applications to squamate reptiles (lizards and snakes). Molecular Ecology 27:2754–2769.

(188) Castro-Insua, A., C. Gómez-Rodríguez, J. J. Wiens, and A. Baselga. 2018.

Climatic niche divergence drives patterns of diversification and richness among mammal families. Scientific Reports 8:8781.

(189) Román-Palacios, C., and J. J. Wiens*. 2018. The Tortoise and the Finch:

testing for island effects on diversification using two iconic Galápagos radiations. Journal of Biogeography 45:1701–1712.

**Highlighted as “Editor’s Choice” in the Journal fo Biogeography *Corresponding author

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(190) Hanschen, E.R., M.D. Herron, J. J. Wiens, P. J. Ferris, H. Nozaki, and R. E. Michod. 2018. Multicellularity drives the evolution of sexual traits. American Naturalist (published online).

(191) Meyer, A. L. S., C. Román-Palacios, and J. J. Wiens*. 2018. BAMM gives

misleading rate estimates in simulated and empirical datasets. Evolution 72:2257–2266.

*Corresponding author (192) Miller, E. C., K. T. Hayashi, D. Song, and J. J. Wiens. 2018. Explaining the

ocean’s richest biodiversity hotspot and global patterns of fish diversity. Proceedings of the Royal Society of London, Series B 285:20181314.

***Featured in the New York Times, with interviews, and on news websites (WorldNews SBS, UA news, KJZZ) and science-related websites (phys.org, Science Daily, Futurity, Earth.com)

(193) Li, H., and J. J. Wiens*. 2019. Time explains regional richness patterns

within clades more often than diversification rates or area. American Naturalist 193:514–529.

*Corresponding author (194) Lambert, S.M., J. W. Streicher, M. C. Fisher-Reid, F. R. Méndez de la

Cruz, N. Martínez-Méndez, U. O. García-Vázquez, A. Nieto-Montes de Oca, and J. J. Wiens. 2019. Inferring introgression using RADseq and DFOIL: power and pitfalls revealed in a case study of spiny lizards (Sceloporus). Molecular Ecology Resources (published online).

(195) Lu, L., P. W. Fritsch, N. J. Matzke, H. Wang, K. A. Kron, D.-Z. Li, and J. J.

Wiens. 2019. Why is fruit color so variable? Phylogenetic analyses reveal relationships between fruit-color evolution, biogeography, and diversification. Global Ecology and Biogeography (in press).

(196) Wiens, J. J., A. Camacho, A. Goldberg, T. Jezkova, M. E. Kaplan, S. M.

Lambert, E. C. Miller, J. W. Streicher, and R. L. Walls. 2019. Climate change, extinction, and Sky Island biogeography in a montane lizard. Molecular Ecology (in press).

(197) Wiens, J. J., Y. Litvinenko, L. Harris, and T. Jezkova. 2019. Rapid niche

shifts in exotic species can be a million times faster than changes among native species and ten times faster than climate change. Journal of Biogeography (in press).