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1 Curriculum Vitae, Eugene W. Schupp Curriculum Vitae Eugene W. Schupp ResearcherID: F-1834-2010 Address: Department of Wildland Resources, and The Ecology Center 5230 Old Main Hill Utah State University Logan, UT 84322-5230 Phone: 435-797-2475 FAX: 435-797-3796 e-mail: [email protected] Education: 1987 Ph.D., Biology University of Iowa 1981 M.A., Zoology University of South Florida 1977 B.A., Biology University of South Florida Professional Experience (Post-Ph.D.): 2012-2013 College of Natural Resources Assistant Dean for Research & Graduate Education, Utah State University 2011-present Professor, Utah State University 1998-2011 Associate Professor, Utah State University 1995-present Adjunct Researcher, Integrated Ecology Group, Estación Biológica de Doñana, Sevilla, Spain 1992-1998 Assistant Professor, Utah State University 1989-1992 DOE Alexander Hollaender Distinguished Postdoctoral Fellow, Savannah River Ecology Laboratory 1988-1989 NSF–NATO Postdoctoral Fellow, Estación Biológica de Doñana, Spain 1987-1988 Postdoctoral Research Associate, University of Iowa 1987 Co-coordinator, Organization for Tropical Studies field course, “Tropical biology: an ecological approach,” Costa Rica

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1 Curriculum Vitae, Eugene W. Schupp

Curriculum Vitae Eugene W. Schupp

ResearcherID: F-1834-2010 Address: Department of Wildland Resources, and The Ecology Center

5230 Old Main Hill Utah State University Logan, UT 84322-5230 Phone: 435-797-2475 FAX: 435-797-3796 e-mail: [email protected]

Education: 1987 Ph.D., Biology University of Iowa 1981 M.A., Zoology University of South Florida 1977 B.A., Biology University of South Florida Professional Experience (Post-Ph.D.):

2012-2013 College of Natural Resources Assistant Dean for Research & Graduate Education, Utah State University

2011-present Professor, Utah State University 1998-2011 Associate Professor, Utah State University

1995-present Adjunct Researcher, Integrated Ecology Group, Estación Biológica de Doñana, Sevilla, Spain

1992-1998 Assistant Professor, Utah State University 1989-1992 DOE Alexander Hollaender Distinguished Postdoctoral Fellow, Savannah River

Ecology Laboratory 1988-1989 NSF–NATO Postdoctoral Fellow, Estación Biológica de Doñana, Spain 1987-1988 Postdoctoral Research Associate, University of Iowa 1987 Co-coordinator, Organization for Tropical Studies field course, “Tropical biology:

an ecological approach,” Costa Rica

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Grants and Contracts Funded: External, at USU:

2017-2019 USDI Bureau of Indian Affairs, “Range Inventory of the Uintah and Ouray Reservation,” Co-PI, $254,000

2015-2020 USDA Forest Service/USDI Bureau of Land Management, “Ongoing funding for SageSTEP regional experiment,” PI, $427,225

2015-2016 Utah Division of Wildlife Resources, “Interactive effects of soils and browsing on sagebrush: implications for restoration success,” K. Veblen, PI, $137,000; (co-investigator with K. Nehring, J. Boettinger, E. Schupp, E. Thacker, J. Villalba, C. Brungard)

2013-2015 National Fish and Wildlife Foundation, “Sclerocactus wetlandicus habitat characterization and seed germination,” Co-PI, $98,999

2011-2015 State of Colorado, Department of Natural Resources, Division of Parks and Outdoor Recreation, “Conservation of a candidate species: breeding system and seed pool dynamics of the endemic DeBeque Phacelia (Phacelia submutica),” PI, $50,000.

2011-2015 USDA Forest Service/USDI Bureau of Land Management, “Ongoing funding for SageSTEP regional experiment,” PI, $510,823

2011-2013 The Nature Conservancy, “North Elk Ridge/Cottonwood Ecosystem Condition Assessment,” PI, $135,000

2009-2014 US Fish & Wildlife Service, “Shrubby Reed-Mustard reproductive success and roads,” PI, $72,363

2007-2012 US National Park Service, “Restoring natural fire regimes at Golden Spike National Historic Site by developing a healthy sagebrush/grassland vegetation community to prevent the cheatgrass-wildfire cycle,” PI, $157,375

2005 NSF, workshop support for the “4th International Symposium–Workshop on Frugivores and Seed Dispersal,” PI, $33,000

2005-2011 Joint Fire Science Program, “A regional experiment to evaluate effects of fire and fire surrogate treatments in the sagebrush biome,” Co-PI $12,900,000 multi-institution project, USU PI (USU portion $1,404,745, my portion $869,884)

2004-2008 NSF, Research Coordination Networks, “Drought impacts on regional ecosystems network (DIREnet): Coordinating studies on southwest forests and woodlands,” as collaborator, $0, but opportunities for USU students

2001-2005 USDA CSREES IFAFS, “Integrated restoration strategies towards weed control on western rangelands,” Co-PI, $3,241,847 multi-institution project, USU PI (USU portion, $779,238)

2000-2001 NSF, “Overlaps in tropical tree fruiting phenology: implications for patterns of seed dispersal and seedling establishment,” $5,466 (Dissertation Improvement Grant of Tarek Milleron)

1999-2002 USDA NRI, “Nitrogen immobilization for restoration of cheatgrass-infested range,” PI, $100,959

1997-1999 USDA NRI, “Competition from native grasses for restoration of cheatgrass-infested range,” PI, $100,000

1994-1996 U.S. Department of Defense Legacy Resource Management Program, “Postfire restoration for ecosystem management at arid installations,” PI, $78,200

1993-1996 Oak Ridge Institute for Science and Education Travel Grant, $1,500 1993 Junta de Andalucía Travel Grant, for cooperative research with Estación

Biológica de Doñana, Spain, $3,000

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External, as Graduate Student and Postdoc:

1989-1992 Department of Energy Alexander Hollaender Postdoctoral Fellowship, “Seed predation and swamp forest dynamics,” Savannah River Ecology Laboratory

1988-1989 NSF–NATO Postdoctoral Fellowship, “The evolution of seed dispersal by animals: quantity versus quality,” Estación Biológica de Doñana, Spain

1986-1989 Smithsonian Institution Environmental Science Program Grant, “Long-term seedling dynamics,” Panama

1986 Smithsonian Tropical Research Institute Short-term Fellowship, “Tropical tree seedling recruitment,” Panama

1984-1985 Smithsonian Institution Predoctoral Fellowship, “Ecological consequences of masting in Faramea occidentalis,” Panama

1983 Smithsonian Tropical Research Institute Short-term Fellowship, “Ecological consequences of masting in Faramea occidentalis,” Panama

USU Internal:

2012-2014 Utah Agricultural Experiment Station Grant, “Restoration of healthy sagebrush ecosystems on the Colorado Plateau”

2010-2015 Utah Agricultural Experiment Station Project, “Reproductive ecology and recruitment of Utah rangeland plants”

2004-2010 Utah Agricultural Experiment Station Project, “Climate and Soil Impacts on Utah Piñon-Juniper Woodlands”

1998-2004 Utah Agricultural Experiment Station Project, “Effects of woody vegetation on plant recruitment in Utah rangelands”

1995-1997 USU Vice President for Research Grant, “Seed accumulation and natural regeneration across a juniper invasion front”

1995-1997 USU Vice President for Research Grant, “Bottlebrush squirreltail competition with cheatgrass: implications for restoration of native rangelands”

1994-1995 USU New Faculty Research Grant, “Seed production, dispersal and loss as factors limiting recruitment of Cercocarpus ledifolius”

1993-1998 Utah Agricultural Experiment Station Project, “Seed and seedling ecology of Utah juniper (Juniperus osteosperma)”

1993-1994 USU New Faculty Research Grant, “Seed dispersal ecology of Juniperus osteosperma”

Funded, declined:

2008-2010 Schupp, E.W., official PI and mentor; T.A.S. Newbold, primary author. USDA NRI Postdoctoral Proposal. "The role of harvester ant foraging behavior in the restoration of cheatgrass-degraded sagebrush-steppe rangelands." March 2008. $100,000, recommended for full funding, declined because accepted a job with USNPS.

Other:

2007 Part of 16-person writing team that submitted a successful proposal for a National Environmental Observatory Network (NEON) Wildland Core Site and Gradient Site to be located in our region.

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Publications: = undergraduate student, = graduate student, ‡ = postdoctoral researcher

Peer-Reviewed: Lewis, M.B., E.W. Schupp, and T.A. Monaco. 2017. Road dust correlated with decreased reproduction of the endangered Utah shrub Hesperidanthus suffrutescens. Western North American Naturalist 77: 430-439. Schupp, E.W., P. Jordano, and J.M. Gómez. 2017. A general framework for effectiveness concepts in mutualisms. Ecology Letters 20: 577-590 (open access: http://onlinelibrary.wiley.com/doi/10.1111/ele.12764/full) Homet-Gutierrez, P., E.W. Schupp, and J.M. Gómez. 2015. Naturalization of almond trees (Prunus dulcis) in semi-arid regions of the western Mediterranean. Journal of Arid Environments 113: 108-113. McIver, J., M. Brunson, S. Bunting, J. Chambers, P. Doescher, J. Grace, A. Hulet, D. Johnson, S. Knick, R. Miller, M. Pellant, F. Pierson, D. Pyke, B. Rau, K. Rollins, B. Roundy, E. Schupp, R. Tausch, J. Williams. 2014. A synopsis of short-term response to alternative restoration treatments in sagebrush-steppe: the SageSTEP project. Rangeland Ecology and Management 67: 584-598. Rau, B.M., J.C. Chambers, D.A. Pyke, B.A. Roundy, E.W. Schupp, P. Doescher and T.G. Caldwell. 2014. Soil resources influence vegetation and response to fire and fire-surrogate treatments in sagebrush-steppe ecosystems. Rangeland Ecology and Management 67: 506-521. Pyke, D.A., S.E. Shaff, A.I. Lindgren, E.W. Schupp, P.S. Doescher, J.C. Chambers, J.S. Burnham, and M.M. Huso. 2014. Region-wide ecological responses of arid Wyoming big sagebrush communities to fuel treatments. Rangeland Ecology and Management 67: 455-467. Chambers, J.C., R.F. Miller, D.I. Board, D.A. Pyke, B.A. Roundy, J.B. Grace, E.W. Schupp, R.J. Tausch. 2014. Resilience and resistance of sagebrush ecosystems: Implications for state and transition models and management treatments. Rangeland Ecology and Management 67: 440-454. Lewis, M.B. and E.W. Schupp. 2014. Reproductive ecology of the endangered Utah endemic Hesperidanthus suffrutescens with implications for conservation. The American Midland Naturalist 172: 236-251. Poulos, J.M., A.P. Rayburn, and E.W. Schupp. 2014. Simultaneous, independent, and additive effects of shrub facilitation and understory competition on the survival of a native forb (Penstemon palmeri). Plant Ecology 215: 417-426. Rayburn, A.P., E.W. Schupp, and S. Kay. 2014. Effects of perennial semi-arid bunchgrass spatial patterns on performance of the invasive annual cheatgrass (Bromus tectorum L.) . Plant Ecology 215: 247-251.

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Rayburn, A.P. and E.W. Schupp. 2013. Effects of community- and neighborhood-scale spatial patterns on semi-arid perennial grassland community dynamics. Oecologia 172: 1137-1145.

Ostoja, S.M., E.W. Schupp, S. Durham, and R. Klinger. 2013. Seed harvesting is influenced by associational effects in mixed seed neighbourhoods, not just by seed density. Functional Ecology 27: 775-785.

Rayburn, A.P., J.B. Davidson, and E.W. Schupp. 2013. Effect of storage time, site, and floral morph on seed germination of the threatened distylous primrose Primula cusickiana var. maguirei. Plant Species Biology 28: 101-108.

Ostoja, S.M., E.W. Schupp, and R. Klinger. 2013. Seed harvesting by a generalist consumer is context dependent: Interactive effects across multiple spatial scales. Oikos 122: 563-574.

Pekas, K.M. and E.W. Schupp. 2013. Influence of aboveground vegetation on seed bank composition and distribution in a Great Basin Desert sagebrush community. Journal of Arid Environments 88: 113-120. Schupp, E.W. and P. Jordano. 2011. The full path of Janzen-Connell effects: genetic tracking of seeds to adult plant recruitment. Molecular Ecology 20: 3953–3955. (Invited Perspective).

Sivy, K.J., S.M. Ostoja, and E.W. Schupp. 2011. Effects of rodent species, seed species, and predator cues on seed fate. Acta Oecologica 37: 321-328

Rau, B.M., D.W. Johnson, R.R. Blank, A. Lucchesi, T.G. Caldwell, and E.W. Schupp. 2011. Transition from sagebrush steppe to annual grass (Bromus tectorum): influence on belowground carbon and nitrogen. Rangeland Ecology and Management 64: 139-147.

Rayburn, A.P., K.J. Schiffers, E.W. Schupp. 2011. Use of precise spatial data for describing spatial patterns and plant interactions in a diverse Great Basin shrub community. Plant Ecology 212: 585-594.

Mazzola, M.B., J.C. Chambers, R.R. Blank, D.A. Pyke, E.W. Schupp, K.G. Allcock‡, P.S. Doescher, and R.S. Nowak. 2011. Effects of resource availability and propagule supply on native species recruitment in sagebrush ecosystems invaded by Bromus tectorum. Biological Invasions 13: 513-526.

Schupp, E.W., P. Jordano and J.M. Gómez. 2010. Tansley Review: Seed dispersal effectiveness revisited: a conceptual review. New Phytologist 188: 333-353. Puerta-Piñero, C., J.M. Gómez, and E.W. Schupp. 2010. Spatial patterns of acorn dispersal by rodents: does the environment matter? Oikos 119: 179-187.

Morris, C. and E.W. Schupp. 2009. Comparison of emergence speed and sterility in two sterile annual hybrid cereal grasses developed for use in restoration. Restoration Ecology 17: 678-685.

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Ostoja, S.M. and E.W. Schupp. 2009. Conversion of sagebrush shrublands to exotic annual grasslands negatively impacts small mammal communities. Diversity and Distributions 15: 863-870. Ostoja, S.M., E.W. Schupp, and K.J. Sivy. 2009. Ant assemblages in intact big sagebrush and converted cheatgrass-dominated habitats in Rush Valley, Tooele County, Utah, USA. Western North American Naturalist 54: 223-234.

Mazzola, M.B., K.G. Allcock‡, J.C. Chambers, R.R. Blank, E.W. Schupp, P.S. Doescher, and R.S. Nowak. 2008. Effects of nitrogen availability and cheatgrass competition on the establishment of Vavilov Siberan wheatgrass. Rangeland Ecology and Management 61: 475-484.

Hampe, A.‡, J.L. García-Castaño, E.W. Schupp, and P. Jordano. 2008. Spatio-temporal dynamics and local hotspots of initial recruitment in vertebrate-dispersed trees. Journal of Ecology 96: 668-678.

Gómez, J.M., C. Puerta-Piñero, and E.W. Schupp. 2008. Effectiveness of rodents as local seed dispersers of Holm oak. Oecologia 155: 529-537.

Humphrey, L.D.‡ and E.W. Schupp. 2004. Competition as a barrier to establishment of a native perennial grass (Elymus elymoides) in alien annual grass (Bromus tectorum) communities. Journal of Arid Environments 58: 405–422.

Ibáñez, I. and E.W. Schupp. 2002. Effects of litter, soil surface conditions, and microhabitat on Cercocarpus ledifolius Nutt. seedling emergence and establishment. Journal of Arid Environments 52: 209–221.

Humphrey, L.D.‡ and E.W. Schupp. 2002. Seedling survival from local and commercially obtained seeds on two semiarid sites. Restoration Ecology 10: 88–95.

Ibáñez, I. and E.W. Schupp. 2001. Positive and negative interactions between environmental conditions affecting Cercocarpus ledifolius seedling survival. Oecologia 129: 543–550.

Humphrey, L.D.‡ and E.W. Schupp. 2001. Seedbanks of Bromus tectorum-dominated communities in the Great Basin. Western North American Naturalist 61: 85–92.

Jordano, P. and E.W. Schupp. 2000. Seed disperser effectiveness: the quantity component and patterns of seed rain for Prunus mahaleb. Ecological Monographs 70: 591–615.

Ahmad, S., C.A. Call, and E.W. Schupp. 2000. Regeneration ecology of Chrysopogon aucheri and Cymbopogon jwarancusa in upland Balochistan: I. Morphology, viability and movement of seeds (spikelets). Pakistan Journal of Biological Sciences 3: 1583–1587.

Ahmad, S., C.A. Call, and E.W. Schupp. 2000. Regeneration ecology of Chrysopogon aucheri and Cymbopogon jwarancusa in upland Balochistan: II. Dispersal, predation and soil reserves of seeds (spikelets). Pakistan Journal of Biological Sciences 3: 1880–1883.

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Ahmad, S., C.A. Call, and E.W. Schupp. 2000. Regeneration ecology of Chrysopogon aucheri and Cymbopogon jwarancusa in upland Balochistan: III. Effects of precipitation and seedbed microhabitat on seedling recruitment. Pakistan Journal of Biological Sciences 3: 2041–2047.

Chambers, J.C., S.B. Vander Wall, and E.W. Schupp. 1999. Seed and seedling ecology of piñon and juniper species in the pygmy woodlands of western North America. Botanical Review 65: 1–38.

Humphrey, L.D.‡ and E.W. Schupp. 1999. Temporal patterns of seedling emergence and early survival of Great Basin perennial plant species. Great Basin Naturalist 59: 35–49. Fuentes, M.‡ and E.W. Schupp. 1998. Empty seeds reduce seed predation by birds in Juniperus osteosperma. Evolutionary Ecology 12: 823-827.

Feener, D.H. and E.W. Schupp. 1998. Effect of treefall gaps on the patchiness and species richness of Neotropical ant assemblages. Oecologia 116: 191–201. Russell, S.K., E.W. Schupp, and V.J. Tepedino. 1998. Reproductive biology of curlleaf mountain mahogany, Cercocarpus ledifolius (Rosaceae): self-compatibility, pollen limitation, and wind pollination. Plant Species Biology 13: 7–12.

Russell, S.K. and E.W. Schupp. 1998. Effects of microhabitat patchiness on patterns of seed dispersal and seed predation of Cercocarpus ledifolius (Rosaceae). Oikos 81: 434–443.

Alzérreca-Angelo, H., E.W. Schupp, and S.G. Kitchen. 1998. Sheep grazing and plant cover dynamics of a shadscale community. Journal of Range Management 51: 214–222. Clark, J., C. Fastie, G. Hurtt, S. Jackson, C. Johnson, G. King, M. Lewis, J. Lynch, S. Pacala, C. Prentice, E.W. Schupp, T. Webb III and P. Wyckoff. 1998. Dispersal theory offers solutions to Reid's paradox of rapid plant migration. BioScience 48: 13–24.

Boettinger, J.L., J.A. Doolittle, N.E. West, E.W. Bork, and E.W. Schupp. 1997. Non-destructive assessment of rangeland soil depth to petrocalcic horizon using electromagnetic induction. Arid Soil Research and Rehabilitation 11: 375–390.

Schupp, E.W., H.J. Heaton, and J.M. Gómez‡. 1997. Lagomorphs and the dispersal of seeds into communities dominated by exotic annual weeds. Great Basin Naturalist 57: 253–258.

Schupp, E.W., J.M. Gómez‡, J.E. Jiménez, and M. Fuentes‡. 1997. Dispersal of Juniperus occidentalis (western juniper) seeds by mammals on Juniper Mountain, southeastern Oregon. Great Basin Naturalist 57: 74–78.

Schupp, E.W. and M. Fuentes‡. 1995. Spatial patterns of seed dispersal and the unification of plant population ecology. ÉcoScience 2: 267–275.

Schupp, E.W. 1995. Seed-seedling conflicts, habitat choice and patterns of plant recruitment. American Journal of Botany 82: 399–409.

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Schupp, E.W. 1993. Quantity, quality and the effectiveness of seed dispersal by animals. Vegetatio 107/108: 15–29.

Schupp, E.W. 1992. The Janzen-Connell Model for tropical tree diversity: population implications and the importance of spatial scale. American Naturalist 140: 525–530.

Schupp, E.W. 1990. Annual variation in seedfall, post-dispersal predation and recruitment of a neotropical tree. Ecology 71: 504–515. Schupp, E.W., H.F. Howe, C.K. Augspurger and D.J. Levey. 1989. Arrival and survival in tropical treefall gaps. Ecology 70: 562–564 (invited Special Feature contribution).

Schupp, E.W. and E.J. Frost. 1989. Differential predation of Welfia georgii seeds in treefall gaps and the forest understory. Biotropica 21: 200–203. Schupp, E.W. 1988. Factors affecting postdispersal seed survival in a tropical forest. Oecologia 76: 525–530.

Schupp, E.W. 1988. Seed and early seedling predation in the forest understory and in treefall gaps. Oikos 51: 71–78.

Schupp, E.W. 1986. Azteca protection of Cecropia: ant occupation benefits juvenile trees. Oecologia 70: 379–386. Howe, H.F., E.W. Schupp and L.C. Westley. 1985. Early consequences of seed dispersal for a neotropical tree (Virola surinamensis). Ecology 66: 781–791.

Schupp, E.W. 1976. Cattle egrets feeding at a carcass. Florida Field Naturalist 4: 37-38.

Rochow, T.F. and E.W. Schupp. 1976. Lake fluctuation and drawdown and their effects on the lakeside vegetation of Lake Tarpon, Pinellas County. Florida Scientist 39.

Books:

Monaco, T.A., E.W. Schupp, R.L. Pendleton, S.G. Kitchen, and P.K. Palacios, editors. 2011. Proceedings – Threats to Shrubland Ecosystem Integrity; 2010 May 18-20; Logan, UT. Natural Resources and Environmental Issues, Volume XVII. S.J. and Jessie E. Quinney Natural Resources Research Library, Logan Utah, USA.

Dennis, A.J., E.W. Schupp, R.J. Green, and D.A. Westcott, editors. 2007. Seed Dispersal: Theory and Its Application in a Changing World. CAB International, Wallingford, UK, 684 pp.

Invited Book Chapters: Chambers, J.C., M.J. Germino, J. Belnap, C.S. Brown, E.W. Schupp, and S.B. St. Clair. 2016. Plant community resistance to invasion by Bromus species: the roles of community attributes, Bromus interactions with plant communities, and Bromus traits. In: Germino, M.J., J.C. Chambers, and C.S. Brown, eds., Exotic Brome-Grasses in Arid and Semiarid Ecosystems of the Western US.

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Causes, Consequences, and Management Implications. Springer International Publishing Switzerland.

Schupp, E.W. 2007. Part III. Seed fate and establishment. In: Dennis, A.J., E.W. Schupp, R.J. Green, and D.A. Westcott, eds., Seed Dispersal: Theory and Its Application in a Changing World. CAB International, Wallingford, UK, pp. 363-367.

Schupp, E.W. 2007. Suitable sites for dispersal are context dependent. In: Dennis, A.J., E.W. Schupp, R.J. Green, and D.A. Westcott, eds., Seed Dispersal: Theory and Its Application in a Changing World. CAB International, Wallingford, UK, pp. 445-462. Schupp, E.W., T. Milleron, and S.E. Russo. 2002. Dissemination limitation and the origin and maintenance of species–rich tropical forests. In: D.J. Levey, W.R. Silva, and M. Galetti, eds. Seed Dispersal and Frugivory: Ecology, Evolution and Conservation. CABI Publishing, Wallingford, Oxfordshire, UK, pp. 19-33.

Schupp, E.W. and D.H. Feener. 1991. Phylogeny, lifeform, and habitat-dependence of ant-defended plants in a Panamanian forest. In, C. R. Huxley and D. K. Culver, eds. Ant-Plant Interactions, pp. 175–197. Oxford University Press, Oxford.

Encyclopedia Entries:

Schupp, E.W. 2011. Dispersal ability, plant. In: Simberloff D. and M. Rejmánek, Editors. Encyclopedia of Biological Invasions, pp. 159-165. University of California Press, Berkeley, CA, USA.

Proceedings and Other Non-Peer-Reviewed: Pyke, D.A., Chambers, J.C., Pellant, M., Miller, R.F., Beck, J.L., Doescher, P.S., Roundy, B.A., Schupp, E.W., Knick, S.T., Brunson, M., and McIver, J.D., 2017, Restoration handbook for sagebrush steppe ecosystems with emphasis on greater sage-grouse habitat—Part 3. Site level restoration decisions: U.S. Geological Survey Circular 1426, 62 p., https://doi.org/10.3133/cir1426. Pyke, D.A., J.C. Chambers, M. Pellant, S.T. Knick, R.F. Miller, J.L. Beck, P.S. Doescher, E.W. Schupp, B.A. Roundy, M. Brunson, M., and J.D. McIver. 2015. Restoration handbook for sagebrush steppe ecosystems with emphasis on greater sage-grouse habitat—Part 1. Concepts for understanding and applying restoration: U.S. Geological Survey Circular 1416, 44 p., http://dx.doi.org/10.3133/cir1416. Pyke, D.A., S.T. Knick, J.C. Chambers, M. Pellant, R.F. Miller, J.L. Beck, P.S. Doescher, E.W. Schupp, B.A. Roundy, M. Brunson, and J.D. McIver. 2015. Restoration handbook for sagebrush steppe ecosystems with emphasis on greater sage-grouse habitat—Part 2. Landscape level restoration decisions: U.S. Geological Survey Circular 1418, 21 p., http://dx.doi.org/10.3133/cir1418. Schupp, E.W., C.S. Boyd, and S. Green. 2015. Woody Fuels Reduction in Wyoming Big Sagebrush Communities. Great Basin Factsheet Series no. 13, 6 p.

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McIver, J.D., M. Brunson, S. Bunting, J. Chambers, N. Devoe, P. Doescher, J. Grace, D. Johnson, S. Knick, R. Miller, M. Pellant, F. Pierson, D. Pyke, K. Rollins, B. Roundy, E.W. Schupp, R. Tausch, D. Turner. 2010. The Sagebrush Steppe Treatment Evaluation Project (SageSTEP): a test of state-and-transition theory. RMRS-GTR-237. USDA Forest Service General Technical Report, Rocky Mountain Research Station, Ft. Collins, CO.

Allcock K.‡, R. Nowak, B. Blank, T. Jones, T. Monaco, P. Doescher, T. Tanaka, D. Ogle, L. St. John, M. Pellant, D. Pyke, V. Satyal, J. Tanaka, E.W. Schupp, and C. Call. 2006. Integrating weed management and restoration on western rangelands. Ecological Restoration 24:199-200.

Chambers, J.C., E.W. Schupp, and S.B. Vander Wall. 1999. Seed dispersal and seedling establishment of piñon and juniper species within the piñon-juniper woodland. Pp. 1–6 In, S.B. Monsen, R. Stevens, R.J. Tausch, R. Miller, and S. Goodrich, eds. Proceedings: Ecology and Management of Pinyon–Juniper Communities Within the Interior West. USDA Forest Service RMRS–Proceedings-0.

Ibáñez, I., E.W. Schupp, and J.L. Boettinger. 1999. Successional history of a curlleaf mountain mahogany stand: a hypothesis. In, E.D. McArthur, W.K. Ostler, and C.L. Wambolt, eds. Proceedings: Shrubland Ecotones. USDA Forest Service Proceedings RMRS–P–11: 102–107.

Schupp, E.W., J.C. Chambers, S.B. Vander Wall, J.M. Gómez‡, and M. Fuentes‡. 1999. Piñon and juniper seed dispersal and seedling recruitment at woodland ecotones. In, E.D. McArthur, W.K. Ostler, and C.L. Wambolt, eds. Proceedings: Shrubland Ecotones. USDA Forest Service Proceedings RMRS–P–11: 66–70. Schupp, E.W., M. Fuentes‡, and J.M. Gómez‡. 1996. Dispersal of Utah juniper (Juniperus osteosperma) seeds by lagomorphs. Proceedings Vth International Rangeland Congress, p. 496–497.

Book Reviews:

Schupp, E.W. 2012. The interplay between place-based research and ecological theory. (review of I. Billick and M.V. Price, eds. The ecology of place: contributions of place-based research to ecological understanding). Ecology 93: 690-691. Schupp, E.W. 2000. Process, scale, and wildland repair: a framework. (review of S. G. Whisenant. 1999. Repairing Damaged Wildlands: A Process-Oriented Landscape-Scale Approach). Ecology 81: 3264–3265.

Schupp, E.W. 1993. Applications of population biology: can we do it better? (review of S.K. Jain and L.W. Botsford, eds. 1992. Applied Population Biology). Ecology 74: 1293-1294.

Text Boxes:

Schupp, E.W. 2002. La eficacia del agente dispersor: un modelo conceptual jerárquico. (Disperser effectiveness: a hierarchical conceptual model) In: M. Guariguata and G. Kattan, eds. Écología y Conservación de Bosques Neotropicales. Pp. 357-358. Libro Universitario Regional, Cartago, Costa Rica.

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Schupp, E.W. 2002. La eficacia del dispersor: cómo determinarla (Disperser effectivenss: how we measure it) In: M. Guariguata and G. Kattan, eds. Écología y Conservación de Bosques Neotropicales. Pp. 359-360. Libro Universitario Regional, Cartago, Costa Rica.

Environmental Assessments: Rochow, T.F., L.F. Bartos and E.W. Schupp. 1976. Environmental Assessment of the Jay B. Starkey Wilderness Park. Southwest Florida Water Management District Report.

Courser, W.D., T.F. Rochow and E.W. Schupp. 1974. Cypress Creek wellfield baseline environmental data. Southwest Florida Water Management District Report.

Theses:

Schupp, E.W. 1987. Studies on Seed Predation of Faramea occidentalis, an Abundant Tropical Tree. Ph.D. dissertation, University of Iowa, Iowa City.

Schupp, E.W. 1981. Interactions between Ants and Plants: Azteca Protection of Cecropia. M.A. thesis, University of South Florida, Tampa.

Professional Service: Society Memberships:

American Society of Naturalists Association for Tropical Biology and Conservation Ecological Society of America Sigma Xi Society for Ecological Restoration Society for Range Management Utah Native Plant Society

Committee Memberships, etc.: 2009-2011 Member, NEON Domain 15 (Great Basin) Domain Science and Education

Coordination Committee 2008-present Uinta Basin Rare Plant Forum 2006-present Canyonlands Research Center Science Committee 2006-present USU representative, Great Basin Cooperative Ecosystems Studies Unit 2005-present Idaho-Nevada-Utah Interagency Plant Materials Committee

2000-2009 Utah representative, WERA-21, “Revegetation and stabilization of deteriorated and altered lands”

2004-2005, Secretary 2005-2007, Chairperson 2000-2006 Board of Directors, Utah Native Plant Society

1995-2012 Shrub Consortium Executive Committee, USU representative 1993 Botanical Society of America Corresponding Fellows Committee

Judging Student Presentations: 2017 Best Undergraduate Student Poster Judge, Student Showcase, USU 2014 Best Undergraduate Student Poster Judge, Student Showcase, USU 2013 Best Student Paper Judge, Ecological Society of America Annual Meeting 2013 Best Undergraduate Student Poster Judge, Student Showcase, USU

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2012 Best Student Paper Judge, Intermountain Graduate Student Symposium 2011 Best Student Paper Judge, Intermountain Graduate Student Symposium

2011 Best Student Paper Judge, Society for Range Management 2009 Best Student Paper Judge, Ecological Society of America Annual Meeting 2006 Best Student Paper Judge, Ecological Society of America Annual Meeting 2004 Best Student Paper Judge, Ecological Society of America Annual Meeting 2002 Best Student Paper Judge, Intermountain Graduate Student Symposium 1994 Best Student Paper Award Committee, Association for Tropical Biology

Manuscript Reviews (Journals since 1 January 2005): Acta Oecologica; African Journal of Range & Forage Science; American Journal of Physical Anthropology; American Midland Naturalist; Austral Ecology; Basic and Applied Ecology; Biological Conservation; Biological Invasions; Biological Reviews; Biotropica, Conservation Biology; Diversity & Distribution; Ecological Applications; Ecological Engineering; Ecological Research; Ecology; Ecology Letters; Ethology Ecology & Evolution; Evolutionary Ecology; Functional Ecology; Global Change Biology; Integrative Zoology; International Journal of Plant Sciences; International Journal of Primatology; Invasive Plant Science and Management; Journal of Applied Ecology; Journal of Arid Environments; Journal of Ecology; Journal of Natural History; Journal of Vegetation Science; Molecular Ecology; Naturwissenschaften; New Phytologist; Northwest Science; Oecologia; Oikos; Oxford Bibliographies in Ecology; Perspectives in Plant Ecology, Evolution & Systematics; Plant Biology; Plant Ecology; Plant Ecology & Diversity; Plant Species Biology; Rangeland Ecology and Management; Rangelands; Restoration Ecology; Western North American Naturalist; various book chapters

Proposal Reviews:

Fauna and Flora International, International Foundation for Science, Israel Science Foundation, National Aeronautics and Space Administration, National Fish & Wildlife Foundation, National Geographic Society, National Science Foundation, Nevada Agricultural Experiment Station, Organization for Tropical Studies, Smithsonian Institution, United States Department of Agriculture National Research Initiative, U.S.–Israel Bi-National Science Foundation, Utah Agricultural Experiment Station, USU Undergraduate Research and Creative Opportunities Grants; restoration ecology book proposals for Sinauer Press and for Island Press

Unpaid Consulting Activities:

1994 Educational Testing Service: usefulness and validity of new GRE tests 1994 Eastside Ecosystem Management Project: responses of rare plants to large-scale

forest disturbance 1994 Utah Department of Health: stratified sampling design for Hanta virus survey 1993-1995 Oxford Scientific Films: information on seed dispersal by monkeys and birds

Meeting Sessions Chaired: 2005 4th International Symposium/Workshop on Frugivory and Seed Dispersal, “Frugivore

community composition” 2000 3rd International Symposium/Workshop on Frugivory and Seed Dispersal, “Theoretical

aspects” 1998 10th Wildland Shrub Symposium, “Shrubland ecotones” 1997 Ecological Society of America, “Frugivory, seed dispersal and seed and seedling ecology” 1994 Association for Tropical Biology, “Plant population ecology”

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Seminars:

Estación Experimental de Zonas Áridas, Almería, Spain, 2015; University of Missouri-St. Louis/Missouri Botanical Gardens, 2009; Utah State University, Department of Wildland Resources, 2006, 2009; Universidad de Granada, Spain, 2003; Muséum National d’Histoire Naturelle, Brunoy, France, 1999; Universidad de Sevilla, Spain,1999; Idaho State University, 1997; Brigham Young University, 1997; University of South Florida, 1995; University of Nevada, Reno, 1995, 2002; Utah State University, Department of Plants, Soils and Biometeorology, 1995; University of Arkansas, Department of Biology, 1995; University of Arkansas, Department of Entomology, 1995; Utah State University, Department of Fisheries & Wildlife, 1994; Utah State University, Department of Forest Resources, 1993, 1998, 2000; Estación Biológica de Doñana, Sevilla, Spain,1993; University of Utah, 1992, 1997; Utah State University, Department of Rangeland Resources, 1992, 1996, 1999, 2000; Laval University, 1992; Savannah River Ecology Laboratory, 1990; University of Georgia, Department of Botany, 1990; University of Georgia, Institute of Ecology, 1990; Ohio University, 1989; Miami University (Ohio), 1989; Organization for Tropical Studies, 1987; University of Illinois, 1987; University of Puerto Rico, 1987; University of Miami, 1983, 1995; Smithsonian Tropical Research Institute, Panamá,1982, 1985, 1991

Meetings Organized:

2010 16th Wildland Shrub Symposium, “Threats to Shrubland Ecosystem Integrity. Linking Research & Management, Utah State University, Logan, UT (with Tom Monaco and Mark Brunson)

Organized Oral Sessions Organized:

2017 Ecological Society of America, Portland, OR (Co-organized Organized Oral Session with Beckman, N.G., R. Snell, B.A. Loiselle, and E. Fricke). “Consequences of Individual Variation in Dispersal for Recruitment, Populations, and Communities.”

Workshops:

Organized: 2010 5th International Symposium-Workshop on Frugivory and Seed Dispersal, Montpellier,

France (with Pierre-Michel Forget, Pedro Jordano, Joanna Lambert, Anna Traveset, Joe Wright, and others)

2005 4th International Symposium-Workshop on Frugivory and Seed Dispersal, Griffith University, Brisbane, Australia (with Andrew Dennis, Mauro Galetti, Ronda Green, Pedro Jordano, David Westcott)

1996 National Center for Ecological Analysis and Synthesis Workshop, “The role of seed dispersal in the Holocene expansion of trees,” (with J. Clark, Duke University [lead] and T. Webb, Brown University)

Invited:

2003 Workshop on Colorado Plateau Research. Salt Lake City, UT 2000 3rd International Symposium–Workshop on Frugivory and Seed Dispersal, Brazil 1997 University of Iowa's Oberman Center for Advanced Studies, workshop on modeling seed

dispersal in heterogeneous landscapes 1991 2nd International Symposium–Workshop on Frugivory and Seed Dispersal, Mexico

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Member of Roundtable Discussion Panels: 2008 member, concluding roundtable discussion panel, Spanish Association of Terrestrial

Ecology Symposium, “Evolutionary ecology of plant-animal interactions, Palma de Mallorca, Spain

2000 member, concluding roundtable discussion panel, 3rd International Symposium–Workshop on Frugivory and Seed Dispersal, Brazil

Oral Papers and Posters Presented: = undergraduate student, = graduate student, ‡ = postdoctoral researcher

Invited: 2017 Mediterranean Ecology Society/Spanish Terrestrial Ecology Association Meeting, Sevilla,

Spain. “How might climate change affect seed dispersal by animals?” 2016 Ecological Society of America, Ft. Lauderdale, FL (presented in symposium as first author

with M. Germino, J. Stark, E. Allen, J. Belnap, and B. Rau). “Ecosystem impacts of invasive annual brome grasses.”

2014 Association for Fire Ecology/International Association of Wildland Fires, “Large Wildland Fires: Social, Political and Ecological Effects,” “SageSTEP: Short-term vegetation responses to fuels treatments in sagebrush ecosystems.”

2008 Spanish Association of Terrestrial Ecology Symposium, “Evolutionary ecology of plant-animal interactions,” Palma de Mallorca, Spain. “The context dependence of seed dispersal effectiveness and constraints on the evolution of specialization.”

2008 The Association for Fire Ecology, Regional Meeting, Tucson, AZ (with Ostoja, J.M., first author, and R. Klinger). “Fire, weed invasion and associated modifications of plant-animal interactions: Examples from the Great Basin.”

2005 4th International Workshop/Symposium on Frugivory and Seed Dispersal, Brisbane, Australia. “What is a suitable site? Or, where does a seed really want to land?”

2005 Taller Iberoamericano sobre "Conservación de interacciones animal-planta: conceptos y transferencias" (IberoAmerican workshop on "conservation of plant-animal interactions: concepts and extension), Santiago de Chile, Chile (with Gómez, J. M., first author; and C. Puerta-Piñero). “Dispersión de Quercus ilex por roedores y arrendajos: consecuencias demográficas y genéticas a escala de paisaje (Dispersal of Quercus ilex by rodents and jays: demographic and genetic consequences at the landscape scale).”

2005 XXXV Annual Meeting of the Ecological Society of Germany, Austria and Switzerland, Regensburg, Germany (with Hampe, A.‡, first author; J. L. García-Castaño; and P. Jordano). “Initial recruitment of vertebrate-dispersed plants: dissecting spatial patterns through time.”

2005 XVII International Botanical Congress, Symposium “How much is seed dispersal limiting?”, Vienna, Austria (with Hampe, A.‡, first author; J. L. García-Castaño; and P. Jordano). “Spatial and temporal dynamics of seed rain and seedling emergence in animal-dispersed species: bridging dispersal and establishment.”

2005 8th Biennial Conference Integrating Science and Management on the Colorado Plateau, Symposium on “Weed Invasions,” Flagstaff, AZ (declined)

2003 7th Biennial Conference Integrating Science and Management on the Colorado Plateau, Symposium on “Evolution and Management of Invasive Weeds,” Flagstaff, AZ. “Nitrogen immobilization and competition between exotic annual and native perennial grasses.”

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2003 Coordinated Intermountain Restoration Project Meeting on Great Basin Restoration, Boise, ID. “Nitrogen immobilization and competition between exotic annual and native perennial grasses.”

2000 3rd International Symposium–Workshop on Frugivory and Seed Dispersal, Brazil (first author, with T. Milleron and S.E. Russo). “Dissemination limitation and the origin and maintenance of species–rich tropical forests.”

1998 VII International Congress of Ecology Symposium, Florence, Italy, Processes that structure tropical tree communities. (first author, with T. Milleron). “Dispersal limitation in tropical forests: consequences for species richness.”

1997 Conference on Ecology and Management of Pinyon–Juniper Woodlands Within the Interior West, Utah, (with J. Chambers, first author, and S. B. Vander Wall). "Seed dispersal and seedling establishment of piñon and juniper species within the piñon-juniper woodland."

1994 Association for Tropical Biology/Society for Conservation Biology Symposium, Mexico, “Plant–animal interactions in tropical dry forests,” (declined)

1993 XV International Botanical Congress Symposium, Japan, “Role of herbivores in controlling structure and dynamics of plant communities,” (declined)

1992 Botanical Society of America Symposium, Hawaii, Seed Fates: Importance for Structuring Plant Populations and Communities. “Seed-seedling conflicts, habitat choice and patterns of plant recruitment.”

1991 2nd International Symposium–Workshop on Frugivory and Seed Dispersal, Mexico. “Quantity, quality and the effectiveness of seed dispersal by animals.”

1989 Oxford Symposium, England, “Ant–Plant Interactions” (first author, with D.H. Feener). “Phylogeny, lifeform, and habitat-dependence of ant-defended plants in a Panamanian forest.”

Contributed, While at USU: 2017 Ecological Society of America, Portland, OR. “Sources of inter-individual variation in seed

dispersal: A plant’s perspective.” 2017 Ecological Society of America, Portland, OR. (presented with J.L. Boettinger, first author,

K.T. Harding, J.D. Armentrout, and J.M. Norton). "Habitat associations of the threatened Sclerocactus wetlandicus: vegetation and soil characteristics."

2017 14th Biennial Conference of Science & Management on the Colorado Plateau & Southwest Region, Flagstaff, AZ. (presented as first author with J.L. Boettinger, K.T. Harding, J.D. Armentrout, and J.M. Norton). “Habitat associations of the threatened

Sclerocactus wetlandicus: vegetation and soil characteristics.” 2016 Association for Tropical Biology and Conservation, Montpellier, France (presented as

sole author). “The influence of seed size and environment on seedling growth of the neotropical tree Faramea occidentalis.”

2016 Western Regional Soil Survey Conference, Fairbanks, AK (presented with Boettinger, J.L., first author, J. Armentrout, K. Harding, and J. Norton). “Habitat associations of the threatened Sclerocactus wetlandicus: soil characteristics.”

2016 Western Regional Soil Survey Conference, Fairbanks, AK (presented as first author with K. Harding, J.L. Boettinger, J. Armentrout, and J. Norton). “Habitat associations of the threatened Sclerocactus wetlandicus: the vegetative community.”

2016 Utah Rare Plant Meeting, Salt Lake City, UT (presented with Sodja, E., first author). “Pediocactus sileri, a threatened cactus in southwest Utah.”

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2016 Utah Rare Plant Meeting, Salt Lake City, UT (presented as first author with J.L. Boettinger, K. Harding, J. Armentrout, and J. Norton). “Sclerocactus wetlandicus habitat associations.”

2016 Washington County Rare Plant Meeting, St. George, UT (presented with Sodja, E., first author). “Pediocactus sileri (Siler’s pincushion cactus) 2015 monitoring results.”

2015 13th Biennial Conference of Science & Management on the Colorado Plateau & Southwest Region, Flagstaff, AZ (presented with Darger, A.M., first author, and J.L. Boettinger). “Sagebrush decline on the Colorado Plateau: A look at sagebrush and soils.”

2015 Society for Ecological Restoration International Meeting, Manchester, UK (presented with D.A. Pyke, first author, S.T. Knick, J.C. Chambers, M. Pellant, and J.L. Beck). “Restoration of Greater Sage-Grouse habitat: A framework for landscape and site-specific decisions.”

2015 6th International Symposium-Workshop on Frugivory & Seed Dispersal. Drakensberg, South Africa (first author, presented with L.D. Bennion, J.M. Gómez, J.M. Arroyo, P. Jordano, and S.G. Kitchen). “Preliminary observations of dispersal and recruitment dynamics of Prunus fasciculata, a dry-fruited native almond of the SW United States.”

2014 IUFRO World Congress, Salt Lake City, UT (presented with J.C. Chambers, first author, R.F. Miller, D.I. Board, D.A. Pyke, B.A. Roundy, J.B. Grace, R.J. Tausch). “Resilience to Disturbance and Management Treatments and Resistance to Invasive Annual Plants in Sagebrush and Piñon Pine/Juniper Ecosystems.”

2014 Ecological Society of America, Sacramento, CA (presented with M. Lee Davis, first author, J. Burnham, J.C. Chambers, P.S. Doescher, D.A. Pyke, and S.E. Shaff). “Imazapic interacts with perennial grass cover to mediate cheatgrass cover after fuels reduction treatments in arid Wyoming big sagebrush sites.”

2013 Ecological Society of America, Minneapolis, MN (presented with Kristen Pekas, first author). “Effects of sagebrush fire and fire surrogate treatments on a Great Basin seed bank community.”

2013 Ecological Society of America, Minneapolis, MN (presented with Alicia Langton, first author). “Reproductive strategy of Phacelia submutica (Boraginaceae; formerly in Hydrophyllaceae), a threatened plant species in western Colorado.”

2013 Ecological Society of America, Minneapolis, MN (presented with Matthew Van Scoyoc, first author). “Assessing ecosystem health using the ecological site framework in the absence of ecological site descriptions.”

2012 Fifth International Fire Ecology and Management Congress, Portland, OR (presented with K.M. Pekas, first author). “Effects of Sagebrush Fire and Fire Surrogate Treatments on a Great Basin Seed Bank Community.”

2012 Fifth International Fire Ecology and Management Congress, Portland, OR (presented with S.E. Shaff, first author, D.A. Pyke, A.I. Lindgren, J. Burnham, P.S. Doescher, and J.C. Chambers). “Short-term results from arid SageSTEP (Sagebrush Steppe Treatment Evaluation Project) fuel treatments.”

2012 Ecological Society of America, Portland, OR (presented with S.E. Shaff, first author, D.A. Pyke, A.I. Lindgren, J. Burnham, P.S. Doescher, and J.C. Chambers). “Inter-perennial gaps may indicate invisibility of sagebrush systems.”

2012 Ecological Society of America, Portland, OR (presented with A.D. Reinwald, first author, J.R. Summerhays, S.M. Ostoja, and M.L. Brooks). “Evaluating restoration techniques using native perennial grasses in cheatgrass (Bromus tectorum L.) invaded sagebrush-steppe ecosystems.”

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2012 Ecological Society of America, Portland, OR (presented with M.B. Lewis, first author, and T.A. Monaco). “Dust and proximity to unpaved roads correlated with decreased reproduction of an endangered Utah endemic shrub.”

2012 Ecological Society of America, Portland, OR (presented with J. Burnham, first author, S.E. Shaff, D.A. Pyke, A.I. Lindgren, J.C. Chambers, and P.S. Doescher). “Effects of tebuthiuron and imazapic on sagebrush and herbaceous understory.”

2012 17th Wildland Shrub Symposium, Las Cruces, NM (presented with J. Burnham, first author, S.E. Shaff, D.A. Pyke, A.I. Lindgren, J.C. Chambers, and P.S. Doescher). “Effects of tebuthiuron and imazapic on sagebrush and herbaceous understory.”

2011 Ecological Society of America, Austin, TX (with J.R. Summerhays, first author, and A.D. Reinwald). “Changes to nutrient availabilities and cheatgrass (Bromus tectorum L.) metrics following non-surface disturbing restoration treatments in a sagebrush-steppe ecosystem.”

2011 Ecological Society of America, Austin, TX (with J.M. Poulos, first author, and S.M. Ostoja). “Life-stage conflicts and the shifting balance between interference and facilitation: A case study with Penstemon palmeri and shrubs.”

2011 Ecological Society of America, Austin, TX (with T.A.S. Newbold, first author). “The role of harvester ant foraging behavior in the restoration of cheatgrass-degraded sagebrush-steppe rangelands.”

2010 5th International Symposium-Workshop on Frugivory and Seed Dispersal, Montpellier, France (with J.M. Gómez, first author, and M. Verdú). “How evolutionarily conserved are the dispersal systems of Ficus and Quercus?”

2010 5th International Symposium-Workshop on Frugivory and Seed Dispersal, Montpellier, France (first author, with J.M. Gómez and M. Fuentes). “Seed dispersal and microhabitat patterns of Juniperus osteosperma recruitment.”

2010 16th Wildland Shrub Symposium, Logan, UT (with Jesse Poulos, first author). “Effects of Artemisia tridentata on spatial patterns, seedling survival, and water stress of Penstemon palmeri.”

2009 Ecological Society of America (with D.A. Pyke, first author, and S.E. Shaff, A. Lindgren, M.D. Reisner, E.W. Schupp, J. Burnham, P.S. Doescher, and J.C. Chambers). “Gaps among perennial plants: spatial relationships, root biomass and cheatgrass cover.”

2009 Ecological Society of America (with S.E. Schaff, first author; and D.A. Pyke, A. Lindgren, J. Burnham, E.W. Schupp, J.C. Chambers, and P.S. Doescher). “Pretreatment vegetation similarities and differences among arid Sagebrush Steppe Treatment Evaluation Project (SageSTEP) sites.”

2009 Ecological Society of America (with A.P. Rayburn, first author). “Spatial patterns and species interactions in a diverse Great Basin (USA) shrub community.”

2009 International Association of Landscape Ecology Conference (with A.P. Rayburn, first author). “Spatial patterns and species interactions in a diverse Great Basin (USA) shrub community.”

2009 Society for Range Management (with J.S. Burnham, first author). “Growth in mixtures of bluebunch wheatgrass, cheatgrass, and squarrose knapweed under factorial nitrogen and phosphorus additions to a low nutrient soil.”

2008 Ecological Society of America (with K.M. Pekas, first author). “Seed pools, aboveground vegetation, and prescribed fire in the context of sagebrush restoration.”

2008 15th Wildland Shrub Symposium (with K.M. Pekas, first author). “Seed pools, aboveground vegetation, and prescribed fire in the context of sagebrush restoration.”

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2008 Annual Winter Meeting of the Utah Section of the Society for Range Management (with K.M. Pekas, first author). “Seed pools, aboveground vegetation, and prescribed fire in the context of sagebrush restoration.”

2008 Annual Winter Meeting of the Utah Section of the Society for Range Management (with J.S. Burnham*, first author). “Bluebunch wheatgrass competition with invasive weeds.”

2007 Ecological Society of America (with K. Sivy, first author; and S.M. Ostoja). “Tales from the sandbox: Do predator cues influence patch selection and seed-handling behaviors of two Heteromyid species?”

2007 Ecological Society of America (S.M. Ostoja, first author). “Man versus rat: A comparison of two traditional seeding methods to seeding done by Dipodomys ordii (Ord's kangaroo rat).”

2007 Ecological Society of America (with L.D. Humphrey‡, first author). “Reproductive and growth traits related to life histories of two rangeland grasses of western North America.”

2007 Ecological Society of America (with C. Morris, first author). “Comparison of emergence speed and sterility in two sterile annual hybrid cereal grasses developed for use in restoration.”

2007 Society for Range Management (first author, with P.S. Doescher and D.A. Pyke). “Plant functional groups and soil N: secondary weed responses.”

2007 Society for Range Management (with S.M. Ostoja, first author). “Rodent granivory and ecological restoration.”

2007 Society for Range Management (with M. Mazzola, first author, K. Alcock‡, J. Chambers and D.A. Pyke). “Plant functional groups and soil N: cheatgrass and native plant responses.”

2006 Ecological Society of America (with S.M. Ostoja, first author). “Total seed densities and relative proportions affect seed preferences by granivorous rodents.”

2006 Ecological Society of America (with L.D. Humphrey‡, first author). “Competition between an invasive annual grass, and mature individuals of a perennial grass species.”

2006 Fourteenth Wildland Shrub Symposium. “SageSTEP: Sagebrush steppe treatment evaluation project.”

2006 Fourteenth Wildland Shrub Symposium (with J.S. Burnham, first author; and T.A. Monaco). “Competition between bluebunch wheatgrass, cheatgrass and squarrose knapweed under factorial nitrogen-phosphorous regimes.”

2006 Annual Meeting of the Spanish Association of Terrestrial Ecology (with J.M. Gómez, first author; C. Puerta-Piñero; and M.L. Huertas Martínez). “Dispersión de Quercus ilex por roedores y arrendajos: consecuencias demográficas a escala de paisaje.”

2005 Ecological Society of America (with J. Landmesser, first author; T. Jones; T. Monaco; R. Nowak; D. Pyke). “Success of cheatgrass (Bromus tectorum) control and native plant restoration techniques on western rangelands.”

2005 Ecological Society of America (with S.M. Ostoja, first author). “Evidence for indirect effects in seed removal by granivorous rodents in the Great Basin of Utah.”

2005 Ecological Society of America (with J. Burnham, first author). “Competition between bluebunch wheatgrass, cheatgrass, and squarrose knapweed under different nutrient regimes.”

2005 Ecological Society of America (with L.D. Humphrey‡, first author). “Grasshopper herbivory in Bromus tectorum-dominated annual grasslands and establishment of perennial-plant seedlings.”

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2005 Society for Range Management (with S.M. Ostoja, first author). “Evidence for indirect effects in seed removal by granivorous rodents in the Great Basin of Utah.”

2005 Ecology & Management of Pinyon-Juniper & Sagebrush Communities Conference. “A regional experiment to evaluate effects of fire and fire surrogate treatments in the sagebrush biome.”

2004 Ecological Society of America (first author, with J.M. Gómez and M. Fuentes). “Spatial and microhabitat patterns of tree recruitment in three juniper woodlands in west-central Utah, USA.”

2003 7th International Conference on the Ecology and Management of Alien Plant Invasions (with R.S. Nowak, first author, K. Allcock‡, R. Blank, J.C. Chambers, P.S. Doescher, D. Ogle, M. Pellant, D.A. Pyke, and C. Wilson). “Integrating weed control and restoration on western rangelands.”

2003 Ecological Society of America (first author, with R. Sharitz). “Effects of burial, litter, and canopy openness on Aesculus pavia seedling emergence.”

2003 Ecological Society of America (with L.D. Humphrey‡, first author). “A test of a long-lived and a short-lived perennial grass for use in restoring Bromus tectorum communities.”

2003 Society for Range Management (with M. Myklebust, first author). “Nitrate and the germination of 10 common annual and perennial species.”

2002 World Congress of Soil Science (with J.L. Boettinger, first author, J.M. Gómez, J. Castro, and R. Zamora) “Human-induced soil changes: pine plantations above apparent natural tree line, Sierra Nevada, Spain.”

2002 Ecological Society of America (first author, with J.L. Boettinger and P. Jordano). “Prunus mahaleb regeneration: does the balance between interference and facilitation depend on the environment?”

2002 Ecological Society of America (with L.D. Humphrey‡, first author). “Life histories of some perennial grasses in semiarid environments.”

2001 Ecological Society of America (with L.D. Humphrey‡, first author). “Competition and facilitation during the establishment of perennial grasses.”

2001 Ecological Society of America (with R. Luster, first author). “Evaluating nitrogen immobilization for the restoration of cheatgrass (Bromus tectorum L.) infested rangeland.”

2001 Ecological Society of America (with J.L. Boettinger, first author; J.M. Gómez; J. Castro; and R. Zamora). “Soil acidification following burning of pine plantations, Sierra Nevada, Spain.”

2000 Ecological Society of America (with L.D. Humphrey‡, first author). “Model comparisons for evaluating competitive relationships in mixed-density plots.”

2000 Ecological Society of America (with R. Luster, first author). “Nitrogen immobilization for the restoration of cheatgrass-infested range.”

2000 Soil Science Society of America (with J.L. Boettinger, first author, J.M. Gómez, J. Castro, and R. Zamora). “Influence of pine plantations on native shrub soils, Sierra Nevada, Spain.”

1999 Ecological Society of America (with L.D. Humphrey‡, first author). "Effects of Bromus tectorum on seedling and second-year Elymus elymoides plants."

1998 Ecological Society of America (with I. Ibañez, first author, and J.L. Boettinger). "Environmental conditions affecting emergence and seedling establishment of the tree Cercocarpus ledifolius during the first growing season."

1998 Ecological Society of America (with L.D. Humphrey‡, first author). "Seedling survival from local and commercially-obtained seeds on two semiarid sites."

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1998 10th Wildland Shrub Symposium (with I. Ibañez, first author; and J.L. Boettinger). "Successional history of a curlleaf mountain mahogany stand."

1998 10th Wildland Shrub Symposium (first author, with J.C. Chambers, S.B. Vander Wall, J.M. Gómez‡, M. Fuentes‡, and T. Milleron). "Piñon and juniper seed dispersal and seedling recruitment at woodland ecotones."

1998 Society for Range Management (first author, with J.M. Gómez‡ and M. Fuentes‡). “Seed dispersal and recruitment of Utah juniper in the Great Basin, USA.”

1998 Society for Range Management (with J. McMillan, first author; and J.L. Boettinger). “Soil properties influence western harvester ant nest site density in the upper Green River Basin.”

1997 Ecological Society of America (first author, with M. Fuentes‡ and J.M. Gómez‡). "Microhabitat patterns of seed dispersal and seedling recruitment of Utah juniper (Juniperus osteosperma)."

1997 Ecological Society of America (with L. D. Humphrey‡, first author). "Can squirreltail (Elymus elymoides) withstand competition from cheatgrass (Bromus tectorum)?"

1996 Ecological Society of America (with S. Russell, first author). "The effects of habitat patchiness on patterns of seed dispersal and survival of Cercocarpus ledifolius (Rosaceae).”

1995 7th European Congress of Ecology (with M. Fuentes‡, first author, and J.M. Gómez‡). "Spatio-temporal variability in the seed dispersal of Juniperus osteosperma."

1995 International Rangeland Congress (first author, with M. Fuentes‡ and J.M. Gómez‡). "Dispersal of Utah juniper (Juniperus osteosperma) seeds by lagomorphs."

1994 Ecological Society of America. (first author, with R. Sharitz). "Propagule characteristics associated with potential seed dispersal by water in a South Carolina bottomland community."

1994 Association for Tropical Biology. "Pollen and resource limitation of reproduction in the abundant tropical tree Faramea occidentalis."

1993 Ecological Society of America. "Seed size, light environment and seedling vigor of a tropical tree."

1993 Association for Tropical Biology. "Seed size, light environment and seedling vigor of a tropical tree."

Webinars Presented:

2012 “Patterns of Understory Growth during the First Three Years Following Prescribed Burn,” for the Great Basin Science Delivery Project.

2014 “Effects of imazapic over four years post-treatment,” for the Great Basin Science Delivery Project.

Honors and Awards as Student: 1978 University of South Florida 1-yr Graduate Fellowship Honors and Awards at USU: 2014 USU Quinney College of Natural Resources Faculty University Service Awardee 2009 USU College of Natural Resources Undergraduate Research Mentor of the Year 2006 USU College of Natural Resources Undergraduate Advisor of the Year Academic and Research Advising:

Ph.D. Students:

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Andrew P. Rayburn, S.J. and Jesse E. Quinney Ph.D. Fellow (graduated 2011), presently, Restoration Ecologist & Science Director, River Partners, Sacramento, CA. Dissertation: “Causes and consequences of plant spatial patterns in natural and experimental Great Basin plant Communities.”

Steven M. Ostoja, Ph.D. (graduated 2008), presently, Director, USDA ARS California Climate Hub, UC Davis, CA. Dissertation: “Granivores and restoration: implications of weed invasion and context-dependent seed removal.”

Tarek Milleron, Ph.D. (graduated 2005), presently, Executive Director of Caura Futures, an environmental/cultural non-profit. Dissertation: “Context-dependent dispersal and survival of tropical tree seeds.”

Humberto Alzérreca-Angelo, Ph.D. (graduated 1996), presently, private range consultant, Bolivia. Dissertation: “Spatial and temporal dynamics of plant populations in salt-desert shrub vegetation grazed by sheep.”

M.S. Students:

Scott Zimmer, M.S. candidate (2017-present; Co-Advised by Janis L. Boettinger). Thesis topic: Range Inventory of the Uintah and Ouray Reservation. Jeremiah Armentrout, M.S. candidate (2013-present; Primary Advisor: Janis L. Boettinger). Thesis topic: Abiotic habitat relationships of Sclerocactus wetlandicus, a threatened cactus. April Darger, M.S. candidate (2013-present; Co-Advisor: Janis L. Boettinger). Thesis topic: Soil-vegetation relationships in Beef Basin, Canyonlands Research Center, Utah R. Chesson Colburn, M.S. candidate (2012-present). Thesis topic: Developing a new release of blue flax for restoration. (Tom Jones, USDA ARS, is funder and co-research advisor). Kourtney Harding, M.S. (graduated 2017; Primary Advisor: Janis L. Boettinger). Thesis topic: Biotic habitat relationships of Sclerocactus wetlandicus, a threatened cactus. Alicia Langton, M.S. candidate (graduated 2014), presently Biological Consultant. Thesis topic: Reproductive ecology of the annual Debeque phacelia, a candidate endangered species. Leah Lewis, M.S. (graduated 2014). Thesis: “Habitat characteristics of Mexican Spotted Owls (Strix occidentalis lucida) in the Canyonlands of southern Utah.” Matt Van Scoyoc, M.S. (graduated 2014), presently Ecologist, National Park Service, Moab, UT. Thesis: “A quantitative approach to the development of ecological sites and state-and-transition models.” Pablo Homet Gutiérrez, University of Granada, Spain (co-advised with José María Gómez; graduated 2013). Thesis: “Estudio de la dispersión natural del almendro (Prunus dulcis) D.A. Webb en un ambiente árido del sur de España” (“Study of the natural dispersal of the almond (Prunus dulcis) D.A. Webb in an arid environment of the south of Spain.”)

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Alexandra Reinwald, USU Vice President for Research Fellow, M.S. (graduated 2013), presently Manager of Native Plant Nursery Program for the Confederated Tribes of Warm Springs, Prairie City, Oregon. Thesis: “Effects of non-surface-disturbing restoration treatments for native grass revegetation and soil seedbank composition.”

Matt Lewis, M.S. (graduated 2013), presently Botanist, Bureau of Land Management, Vernal, UT. Thesis: “Roads and the reproductive ecology of Hesperidanthus suffrutescens, an endangered shrub.”

Jesse Poulos, M.S. (graduated 2013). Thesis: “Interspecific interactions between Penstemon palmeri and shrubs in the arid shrublands of the Spring Mountains, Nevada.”

Jan Summerhays, USU Vice President for Research Fellow, M.S. (graduated 2011), presently consultant with URS/AECOM, Boise, ID. Thesis: “Effects of non-surface-disturbing treatments for native grass revegetation on cheatgrass (Bromus tectorum L.) metrics and soil ion availabilities.”

Jeff Burnham, M.S. (graduated 2010), presently Range Ecologist, Washington Department of Fish and Wildlife, Yakima, WA. Thesis: “Native bunchgrass and invasive weed establishment in low nutrient rangeland soils with nitrogen and phosphorous manipulation.” Kristen Pekas, M.S. (graduated 2010), presently, Botanist, Idaho Natural Heritage Program, Idaho Department of Fish & Game, Lewiston, ID. Thesis: “Seed pool dynamics of a Great Basin sagebrush community in the context of restoration.”

Dara Scherpenisse, M.S. (graduated 2009), presently, Restoration Ecologist, Lake Mead National Recreation Area, NV. Thesis: “Mycorrhizae in sagebrush-steppe community restoration: mycorrhizal dependency of invasive and native grasses with intraspecific and interspecific competition.”

Matt Parsons, M.S. (graduated 2008), presently, USDA Agricultural Research Service technician, Ft. Collins, CO. Thesis: “Ecotypic Variation in Elymus elymoides ssp. brevifolius race C in the Intermountain West. (Tom Jones, USDA ARS, was research advisor).

Ryan Luster, M.S. (graduated 2002), presently, Restoration Coordinator, The Nature Conservancy, Chico, CA. Thesis: “Nitrogen manipulation and the competitive balance between Bromus tectorum, an annual weedy grass, and Elymus multisetus, a native perennial grass: Implications for restoration of native rangelands.”

Inés Ibáñez, Fulbright Fellow, M.S. (graduated 1998), Ph.D. Duke University; presently, Associate Professor, University of Michigan. Thesis: “Early recruitment in curlleaf mountain mahogany (Cercocarpus ledifolius Nutt.).”

Susan Russell, M.S. (graduated 1997), presently, Science Department Chair and Biology & Biotechnology Teacher, Lynnwood High School, Seattle, WA. Thesis: “Reproduction and recruitment of curlleaf mountain mahogany (Cercocarpus ledifolius): pollination and seed ecology.”

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M.N.R. Students: Eli Angus, professional degree program (graduated 2014). Project topic: organizing a bio-blitz for endangered plant species in the Uintah Basin, Utah.

Failed to Finish:

Dusty Carpenter, M.S. candidate (2002–2007; left USU without finishing). Thesis topic: Mycorrhizae and restoration of cheatgrass–infested rangelands.

May Myklebust, M.S. candidate (2000–2003; changed major advisor). Thesis topic: Nitrogen manipulation and the germination, emergence, and establishment of exotic weeds and native perennials.

Yvonne Wilder, M.S. candidate (1993-1994; left USU without finishing). Thesis topic: The role of rodents in juniper seed dispersal.

Undergraduate Independent Researchers: Eric Sodja (2015), “Shrodinger’s cactus: revisiting Pediocactus sileri.” Lee Bennion (2014-15), “Germination ecology of the desert almond, Prunus fasciculata.” Kourtney Blanc (2011-13), “Seed neighborhood species richness and seed harvesting by western harvester ants.”

Kelly Sivy (with Steve Ostoja, 2006), “Seed Preferences of Dipodomys ordi in the presence of predator cues.”

Tyler Logan (2006), “Comparing effects of restoration treatments on the foraging behavior of harvester ants in the eastern Great Basin sagebrush desert.”

Ryan Woodland (2002), “The effects of aeration on irrigated meadows at Deseret Land and Livestock, Woodruff, Utah.”

Hoyt Heaton (1996), “Lagomorphs and the dispersal of seeds into communities dominated by exotic annual weeds.”

Postdoctoral Associates:

Scott Newbold, Utah State University (2006-2008), presently Assistant Professor, Sheridan College, Sheridan, WY. Research topic: The effects of cheatgrass on harvester ant foraging: implications for restoration.

L. David Humphrey, Utah State University (1996-1999), presently, private biological consultant. Research topic: Rangeland restoration.

José María Gómez, Universidad de Granada, Spain, Spanish Ministry of Science Fellowship (1994-1995), presently Professor, Universidad de Granada, Spain. Research topic: Seed and seedling ecology of Juniperus osteosperma.

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Marcelino Fuentes, Universidad de Santiago de Compostela, Spain, Fulbright Fellowship (1993-1994), presently Associate Professor, Universidade da Coruña, Spain. Research topic: Seed and seedling ecology of Juniperus osteosperma.

Visiting Researchers:

José María Gómez, Universidad de Granada, Spain (September 1998, September 2003, October 2005, October 2007, October 2009, July-September 2011). Research topics: Balance between interference and facilitation during establishment of arid land plants, Drought impacts on piñon-juniper woodlands, and long-term dynamics of piñon-juniper woodlands.

Marcelino Fuentes, Universidade da Coruña, Spain (January 1997). Research topic: Juniperus osteosperma seed dispersal.

Maria Teresa Sebastià, Universidad de Lleida, Spain (July 1994). Research topic: Community structure of alpine grasslands.

Research Associates: Maggie Gray, M.S. Southern Illinois University (2016-present), manages restoration projects M. Lee Davis, M.S. Syracuse University (2012–2015), manages restoration projects. Christo Morris, B.S. University of California Santa Cruz, M.S. USU (2005–2006), managed USDA restoration project. Presently, Instructor, Eastern Oregon University, La Grande, OR. Jeff Burnham, B.S. Brigham Young University, M.S. USU (2004-2012), managed restoration projects, presently Range Ecologist, Washington Department of Fish and Wildlife, Yakima, WA.

Laura Blonski, B.S. & M.S. University of Alberta (2002-2004), managed cheatgrass restoration projects. Presently, Range Ecologist, Ministry of Forests and Range, British Columbia, CA.