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New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21 Photosynthesis
21.1 The basic concepts of photosynthesis (Book 3, p. 21-3)
� Plants and algae are autotrophs. They make their own food by (1) _______________.
� Photosynthesis is the process by which (2) _______________ substances (such as
carbohydrates) are made from simple (3) _______________ substances (carbon dioxide and
(4) _______________) using (5) _______________ energy. During the process, light energy
is captured by (6) _______________ in plants and algae. Chlorophyll is found in
(7) _______________ in most plants.
� The light energy captured is converted to (8) _______________ energy that is stored in the
carbohydrates produced. (9) _______________ is released as a by-product.
� Photosynthesis is a / an (10) _______________ (catabolic / anabolic) process.
� (11) _______________ and (12) _______________ are made from intermediates of
photosynthetic reactions. Their production requires (13) _______________ absorbed from the
soil.
oxygen (released to the air)
carbon dioxide (from air)
water (from soil)
light
chlorophyll
photosynthesis
from
in
term
ed
iate
s
carbohydrates
lipids
proteins
plant materials, e.g. enzymes, chlorophyll, cell membrane and cell wall
▲ How plants make the materials they need
sun
�
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.2 The site of photosynthesis (Book 3, p. 21-5)
� In plants, (1) _______________ are the main site of photosynthesis.
A What are the adaptive features of leaves for photosynthesis? (Book 3, p. 21-5)
Structure Feature Adaptation for photosynthesis
� Broad and flat � Provides a large (2) _______________
_____________ for absorbing sunlight Leaf blade
� Thin � Gases and light can reach the
photosynthetic cells easily
Palisade
mesophyll
� Consists of tightly packed cells that
contain many (3) _______________
� Located on the (4) ______________
(upper / lower) side of the leaf
� Allows effective absorption of sunlight
Spongy
mesophyll
� Consists of loosely packed cells with
many (5) _______________
_______________
� The cells contain chloroplasts (fewer
than palisade mesophyll)
� Allows gases to (6) _______________
freely
� Allows absorption of sunlight
� Covered by a thin layer of waxy
(7) _______________
� Reduce (8) _______________ loss
from the leaf, so that water is kept
inside for photosynthesis and keeping
the cells turgid Upper
epidermis
and lower
epidermis
� Contain numerous
(9) _____________ (more on lower
epidermis)
� Each stoma is surrounded by a pair
of (10) _______________
_______________
� Stomata open when the conditions are
favourable for photosynthesis. They
close when the conditions are less
favourable. This regulates the passage
of gases and water vapour into and out
of the leaf
Midrib and
veins
� Contain vascular bundles which are
made up of (11) _______________
and (12) _______________
� Allow efficient transport of materials
into and away from the leaf
� Support and spread the leaf blade
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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B What are the adaptive features of chloroplasts for
photosynthesis? (Book 3, p. 21-7)
� A chloroplast is bounded by a (13) _______________ membrane and is filled with a
jelly-like fluid called (14) ______________ (基質). The stroma contains (15) ______________.
It also holds (16) _______________ grains.
� A network of disc-like membranous sacs called (17) _______________ (類囊體) is suspended
in the stroma. The thylakoids are arranged in stacks called (18) _______________ (基粒).
(19) _______________ is located on the thylakoid membranes.
� A chloroplast has about 50 grana, each consisting of about 50 thylakoids. This provides a large
(20) _______________ _______________ to pack more chlorophyll for absorbing light. The
grana are (21) _______________ by extensions from thylakoids. This allows efficient
(22) _______________ of photosynthetic products within the chloroplast.
21.3 The requirements for photosynthesis (Book 3, p. 21-9)
A How do we know if photosynthesis has occurred? (Book 3, p. 21-9)
1 Detecting the presence of starch
� We can carry out (1) _______________ test to test for the presence of starch in a plant.
2 Detecting the release of oxygen
� We can test for oxygen in the gas produced by a plant using a (2) _______________ splint.
� Go to
Practical 21.1 Detection of starch produced in photosynthesis (the iodine test)
(Book 3, p. 21-9; Practical Workbook for SBA 3 • 4, p. 21-1)
Practical 21.2 Detection of oxygen produced in photosynthesis
(Book 3, p. 21-10; Practical Workbook for SBA 3 • 4, p. 21-5)
thylakoid
stroma
granum
outer membrane
inner membrane
▲ 3-D drawing of a chloroplast showing grana and thylakoids
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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B Investigation of the requirements for photosynthesis(Book 3, p. 21-11)
� If we plan to use the presence of starch in leaves as the evidence that photosynthesis has
occurred, any starch that is present in the leaves originally must be removed before the
investigation.
� Plants used in such investigations must first be put in the (3) _______________ for at least
(4) _______________ hours to remove all the starch. This process of removing starch is called
(5) _______________ (脫澱粉).
1 Chlorophyll
� We can carry out the investigation on plants with (6) _______________
leaves (斑葉). Chlorophyll is (7) _______________ (present / absent) in
the non-green parts of the leaves.
� The part of a leaf with chlorophyll is the (8) _______________
(experimental / control) set-up.
� The part of a leaf without chlorophyll is the (9) _______________
(experimental / control) set-up.
▲ Variegated leaf of
Coleus
2 Carbon dioxide
� We can use (10) _______________ _______________ (氫氧化鉀) to remove carbon dioxide
from the air inside the control set-up.
� The leaf exposed to normal air is the (11) _______________ (experimental / control) set-up.
� The leaf exposed to air without carbon dioxide is the (12) _______________ (experimental /
control) set-up.
3 Light
� We can use a piece of black paper or aluminium foil to mask part of a leaf.
� The unmasked part of the leaf is the (13) _______________ (experimental / control) set-up.
� The masked part of the leaf is the (14) _______________ (experimental / control) set-up.
� Go to
Practical 21.3 Investigation of the need for chlorophyll in photosynthesis
(Book 3, p. 21-12; Practical Workbook for SBA 3 • 4, p. 21-7)
Practical 21.4 Investigation of the need for carbon dioxide in photosynthesis
(Book 3, p. 21-13; Practical Workbook for SBA 3 • 4, p. 21-11)
Practical 21.5 Investigation of the need for light in photosynthesis
(Book 3, p. 21-14; Practical Workbook for SBA 3 • 4, p. 21-14)
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.4 The process of photosynthesis (Book 3, p. 21-16)
� Photosynthesis involves many reactions, each of which is controlled by a different enzyme.
The reactions occur in two main stages: (1) _______________ _______________ (光反學反應)
and (2) _______________ _______________ (碳固定).
A Photochemical reactions (Book 3, p. 21-16)
� Photochemical reactions occur in the (3) _______________ of chloroplasts. The reactions
require (4) _______________. Therefore, they are also called (5) _______________
_______________ (光反應).
1 Light absorption
� When the chlorophyll molecule in the thylakoid captures (6) _______________ energy, some
(7) _______________ of the chlorophyll molecule are raised to a higher energy level and are
emitted from the chlorophyll molecule. These excited electrons then pass through a series of
electron carriers of (8) _______________ (increasing / decreasing) energy levels in the
thylakoid membrane. (9) _______________ is released in a step-wise manner. The series of
electron carriers is called the (10) _______________ _______________ _______________
(電子傳遞鏈).
▲ Absorption of light energy by a chlorophyll molecule
initial energy level
higher energy level
� Light energy is
absorbed by chlorophyll.
� An electron becomes ‘excited’.
� The ‘excited electron’
releases energy
when it passes
through an electron transport chain.
energy
chlorophyll molecule in thylakoid membrane
thylakoid
chloroplast
electron transport chain
sun
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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2 Synthesis of ATP
� Some of the energy released by the electrons in the electron transport chain is used to form
(11) _______________ (adenosine triphosphate 腺苷三磷酸) by combining a phosphate with
(12) _______________ (adenosine diphosphate 腺苷二磷酸). This process is called
(13) _______________ (光磷酸化).
� In this way, (14) _______________ energy captured by the chlorophyll molecule is converted
to (15) _______________ energy stored in ATP. The ATP will provide energy to drive the
reactions in carbon fixation.
3 Photolysis of water
� The light energy captured by chlorophyll molecules is also used to split (16) ______________
molecules to form (17) _______________ and (18) _______________. This process is called
(19) _______________ (光解) of water.
- Hydrogen is accepted by (20) _______________ (nicotinamide adenine dinucleotide
phosphate 菸酰胺腺嘌呤二核苷酸磷酸) to form (21) _______________. NADPH is the
(22) _______________ (oxidized / reduced) form of NADP. It will act as a
(23) _______________ agent by donating (24) _______________ in carbon fixation.
- Oxygen is a (25) _______________ in the reaction. It is released as a
(26) _______________ to the atmosphere.
Light energy captured by chlorophyll
Synthesis of ATP
Photolysis of water
to carbon fixation
stroma thylakoid
space thylakoid
membrane
▲ Photochemical reactions
(photophosphorylation) sun
oxygen released to atmosphere
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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B Carbon fixation: Calvin cycle (Book 3, p. 21-19)
� ATP and NADPH formed in the photochemical reactions are used to fix (27) ______________
into organic compounds. This stage is called (28) _______________ _______________. It
occurs in the (29) _______________ of chloroplasts.
� This stage is also called (30) ____________ _______________ (暗反應) because it does not
require (31) _______________.
� This stage is also known as the (32) _______________ _______________ (卡爾文循環). It
involves a cyclic series of reactions and occurs in three main steps.
1 Carbon dioxide fixation and formation of a 3-C compound
� Under the action of enzymes, a molecule of (33) _______________ _______________
combines with a molecule of a 5-C compound to form (34) _______________ molecules of a
3-C compound. The 5-C compound acts as a carbon dioxide (35) _______________ to ‘fix’
carbon dioxide into an organic compound which starts the reactions in the cycle.
2 Reduction of the 3-C compound and formation of glucose
� Using energy from (36) _______________ and hydrogen from (37) _______________
produced in photochemical reactions, the 3-C compound is (38) ______________ (oxidized /
reduced) to another 3-C compound called (39) ______________ _______________ (丙糖磷酸).
ADP and NADP are formed in the process. They will be reused in (40) _______________
reactions.
� Triose phosphate is a sugar. It is the primary product of the Calvin cycle. Two molecules of
triose phosphate combine to form one molecule of (41) _______________ (6-C).
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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3 Regeneration of carbon dioxide acceptor
� Most triose phosphate molecules are used to (42) _______________ the original 5-C carbon
dioxide acceptor, so that the cycle can repeat itself to form more glucose. The regeneration of
carbon dioxide acceptor requires (43) _______________ from ATP.
Carbon dioxide fixation
Reduction of 3-C compound
Regeneration of carbon dioxide acceptor
3-C compound 5-C compound
triose phosphate (3-C)
glucose (6-C)
from photochemical reactions
from photochemical reactions
▲ The Calvin cycle
Calvin cycle
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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C Overall process of photosynthesis (Book 3, p. 21-22)
� Photochemical reactions and the Calvin cycle are linked by (44) _______________ and
(45) _______________.
� The equation of the overall process of photosynthesis:
It shows that (46) _______________ molecules of carbon dioxide and (47) _______________
molecules of water are required to form one molecule of (48) _______________.
� The hydrogen and oxygen atoms in water molecules are found in the (49) _______________
and (50) _______________ _______________ produced respectively.
6 CO2 + 6 H2O
carbon dioxide water
C6H12O6 + 6 O2
glucose oxygen
light captured
by chlorophyll
ATP
NADPH
NADP
ADP + P
light
Photochemical reactions
thylakoid (with chlorophyll)
Calvin cycle
chloroplast
glucose
▲ Summary of the process of photosynthesis
stroma
sun
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.5 Factors affecting the rate of photosynthesis (Book 3, p. 21-24)
A Effect of light intensity (Book 3, p. 21-24)
� Initially, the rate of photosynthesis
(1) _______________ proportionately with light
intensity as more (2) _______________ is
supplied to the photochemical reactions.
� As the light intensity increases, the rate levels off
when the light intensity reaches a
(3) _______________ ______________ (飽和點).
The rate is now (4) _______________ by other
factors. The light intensity at the saturation point
is called (5) _______________ light intensity.
B Effect of carbon dioxide concentration (Book 3, p. 21-25)
� Initially, the rate of photosynthesis (6) _______________
proportionately with carbon dioxide concentration
because more (7) _______________ are supplied to the
Calvin cycle.
� As the carbon dioxide concentration further increases, the
rate (8) _______________ _______________ because the
rate becomes limited by other factors.
� Go to
Practical 21.6 Investigation of the effect of light intensity on the rate of photosynthesis
(Book 3, p. 21-24; Practical Workbook for SBA 3 • 4, p. 21-17)
Practical 21.7 Design an investigation of the effect of carbon dioxide concentration on the rate of
photosynthesis
(Book 3, p. 21-26; Practical Workbook for SBA 3 • 4, p. 21-22)
rate limited by light intensity
rate limited by other factors
rate
of
pho
tosynth
esis
saturation point
optimum light intensity
light intensity
▲ Effect of light intensity on the rate of photosynthesis
CO2 concentration
rate
of
pho
tosynth
esis
▲ Effect of CO2 concentration on the rate of photosynthesis
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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How can we know which factor is limiting?
� Without light, no photosynthesis takes place.
� At low light intensities, the rate (9) _______________ (increases / decreases) with increasing
light intensity. (10) _______________ _______________ is the limiting factor.
� At high light intensities, the rate in curve A (11) _______________ _______________. The
rate is limited by a factor other than light intensity.
� The rate (12) _______________ (increases / decreases) with higher CO2 concentration. This
shows that (13) _______________ _______________ _______________ is the limiting
factor when curve A levels off.
� The rate in curve B levels off again. It is limited by a factor other than
(14) _______________ _______________ _______________.
C Regulating the rate of photosynthesis (Book 3, p. 21-28)
� Farmers can build (15) _______________ (溫室) to monitor
environmental factors such as temperature, light intensity
and carbon dioxide concentration, so as to regulate the rate
of photosynthesis of their crops.
� In this way, the productivity (生產力) and the quality of the
crops can be (16) _______________.
▲ A greenhouse
rate
of
pho
tosynth
esis
light intensity
low CO2 concentration (curve A)
high CO2 concentration (curve B)
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.6 The fate of photosynthetic products (Book 3, p. 21-31)
� (1) _______________ _______________ is the primary product of photosynthesis. This
molecule and other intermediates of photosynthesis are synthesized into biomolecules essential
for plant growth.
1 Carbohydrates
� Triose phosphate is quickly synthesized into (2) _______________. Some of the glucose is
used up in photosynthesizing cells as an (3) _______________ source.
� Most of the glucose formed is quickly built into (4) ______________ and is stored in cells. It
is a suitable form for storage because it is (5) _______________. It can be stored in leaves
without affecting the (6) _______________ _______________ of the cells.
� Starch can be broken down to form (7) _______________, which is transported by the
(8) _______________ to other parts of the plant. Sucrose is a suitable form for transport
because it is soluble but relatively (9) _______________.
� Some glucose molecules are built into (10) _______________, which is a major component of
(11) _______________ _______________.
2 Lipids
� The intermediates of photosynthesis can also be used to synthesize (12) _______________
and (13) ______________ ______________. Glycerol and fatty acids then combine to form
lipids.
� Lipids are used to form (14) _______________ _______________. They are also stored as
energy reserve in some plants.
3 Proteins
� The intermediates of photosynthesis can also combine with (15) _______________ to form
(16) _______________ _______________, which are the building blocks of proteins.
� Proteins are important for (17) _______________ and repair, and for the synthesis of
enzymes. They are also the components of (18) _______________ _______________.
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.7 The significance of photosynthesis (Book 3, p. 21-33)
1 Providing the basic food source in ecosystems
� Most organisms depend on plants for food. Plants are (1) _______________ in ecosystems.
They serve as the basic food source for other organisms.
2 Maintaining energy flow in ecosystems
� During photosynthesis, light energy from the (2) _______________ is captured by plants. The
light energy is converted to (3) _______________ energy that is stored in plant bodies. The
chemical energy is transferred along (4) _______________ _______________ and is
eventually lost as (5) _______________ to the surroundings.
3 Maintaining oxygen and carbon dioxide balance
� (6) _______________ of organisms consumes oxygen and releases carbon dioxide. The
oxygen consumed is replaced by the oxygen released by plants during (7) _______________.
Therefore, photosynthesis is important in maintaining the oxygen balance in the atmosphere.
� Plants take in carbon dioxide from the environment for (8) _______________. This balances
the increase in carbon dioxide concentration in the atmosphere due to (9) _______________.
photosynthesis
respiration
carbon dioxide in atmosphere
oxygen in atmosphere
taken up by plants during
releases
releases consumed
during
▲ Balance of oxygen and carbon dioxide in the atmosphere
light energy from sun
producers (e.g. plants)
decomposers
heat heat
heat
consumers
▲ Energy flow in an ecosystem
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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Answers
Ch 21 Photosynthesis
21.1 1 photosynthesis 2 organic 3 inorganic 4 water 5 light
6 chlorophyll 7 chloroplasts 8 chemical 9 Oxygen 10 anabolic
11 Proteins / Lipids 12 lipids / proteins 13 minerals
21.2 1 leaves 2 surface area 3 chloroplasts 4 upper 5 air spaces
6 diffuse 7 cuticle 8 water 9 stomata 10 guard cells
11 xylem / phloem 12 phloem / xylem 13 double 14 stroma 15 enzymes
16 starch 17 thylakoids 18 grana 19 Chlorophyll 20 surface area
21 interconnected 22 transport
21.3 1 iodine 2 glowing 3 dark 4 24 5 destarching
6 variegated 7 absent 8 experimental 9 control 10 Potassium hydroxide
11 experimental 12 control 13 experimental 14 control
21.4 1 photochemical reactions 2 carbon fixation 3 thylakoids 4 light
5 light reactions 6 light 7 electrons 8 decreasing 9 Energy
10 electron transport chain 11 ATP 12 ADP
13 photophosphorylation 14 light 15 chemical 16 water
17 hydrogen / oxygen 18 oxygen / hydrogen 19 photolysis 20 NADP 21 NADPH
22 reduced 23 reducing 24 hydrogen 25 by-product 26 gas
27 carbon 28 carbon fixation 29 stroma 30 dark reactions 31 light
32 Calvin cycle 33 carbon dioxide 34 two 35 acceptor 36 ATP
37 NADPH 38 reduced 39 triose phosphate 40 photochemical 41 glucose
42 regenerate 43 energy 44 ATP / NADPH 45 NADPH / ATP 46 six
47 six 48 glucose 49 glucose 50 oxygen gas
New Senior Secondary Mastering Biology Oxford University Press 2014 (Second Edition)
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21.5 1 increases 2 energy 3 saturation point 4 limited 5 optimum
6 increases 7 substrates 8 levels off 9 increases 10 Light intensity
11 levels off 12 increases 13 carbon dioxide concentration
14 carbon dioxide concentration 15 greenhouses 16 improved
21.6 1 Triose phosphate 2 glucose 3 energy 4 starch 5 insoluble
6 water potential 7 sucrose 8 phloem 9 inactive 10 cellulose
11 cell walls 12 glycerol 13 fatty acids 14 cell membranes 15 minerals
16 amino acids 17 growth 18 cell membranes
21.7 1 producers 2 sun 3 chemical 4 food chains 5 heat
6 Respiration 7 photosynthesis 8 photosynthesis 9 respiration