Chapter 54-55 The difference between a fundamental niche and a realized niche. The role of competitive exclusion in interspecific competition

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  • Chapter 54-55
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  • The difference between a fundamental niche and a realized niche. The role of competitive exclusion in interspecific competition. The symbiotic relationships of parasitism, mutualism, and commensalism The impact of keystone species on community structure The difference between primary and secondary succession
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  • Community: A collection of populations that interact with one another in a given area. Within a community, different populations will play different roles. What do we call the role a population plays in its community? Its Niche
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  • Inter- means between different groups. Intra- means within the same group. Intraspecific competition for resources are examples of density dependent limiting factors for a single population.
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  • Community interactions can be classified by whether they help, harm, or have no effect on the species involved relationships between species interspecific interactions Ecologists call relationships between species in a community interspecific interactions Examples include: competition predation herbivory symbiosis (parasitism, mutualism, and commensalism) facilitation
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  • Interspecific interactions Interspecific interactions can affect the survival and reproduction of each species, and the effects can be summarized as positive (+), negative (), or no effect (0)
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  • the carrier crab carries a sea urchin on its back for protection against predators
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  • Interspecific competition (/ interaction) Occurs when species compete for a resource in short supply Two different species compete for the same limited resource Squirrels and black bears compete for acorns
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  • Strong competition can lead to competitive exclusion, local elimination of a competing species two species competing for the same limiting resources cannot coexist in the same place The competitive exclusion principle states that two species competing for the same limiting resources cannot coexist in the same place What will decide which species will win out?
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  • Ecological niche (ecological role) Sum of an organisms use of biotic and abiotic resources Interspecific competition occurs when the niches of two populations overlap Competition lowers the carrying capacity of competing populations
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  • FundamentalRealized The potential niche a species can occupy. The actual niche a species occupies
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  • Ecologically similar species can coexist in a community if there are one or more significant differences in their niches Resource partitioning Resource partitioning is differentiation of ecological niches, enabling similar species to coexist in a community
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  • A. distichus perches on fence posts and other sunny surfaces. A. insolitus usually perches on shady branches. A. ricordii A. aliniger A. insolitus A. distichus A. christophei A. cybotes A. etheridgei
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  • When two organisms interact closely in a way that benefits both species. Honeybees have a mutualistic relationship with flowers. How is this so?
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  • Reef-building corals require mutualism Photosynthetic dinoflagellates Live in the cells of each coral polyp Produce sugars used by the polyps Provide at least half of the energy used by the coral animals
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  • Predation (+/ interaction) refers to interaction where one species, the predator, kills and eats the other, the prey Some feeding adaptations of predators are claws, teeth, fangs, stingers, and poison
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  • Predation benefits the predator but kills the prey Prey adapt using protective strategies Camouflage Mechanical defenses Chemical defenses
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  • (a)Cryptic coloration (b) Aposematic coloration Canyon tree frog Poison dart frog (c) Batesian mimicry : A harmless species mimics a harmful one. (d) Mllerian mimicry: Two unpalatable species mimic each other. Hawkmoth larva Cuckoo bee Yellow jacket Green parrot snake
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  • Camouflage
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  • Poison
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  • Herbivory (+/) (type of predation) refers to an interaction in which an herbivore eats parts of a plant or alga It has led to evolution of plant mechanical and chemical defenses and adaptations by herbivores
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  • Herbivory is not usually fatal to the plant Plants must expend energy to replace the loss Plants have numerous defenses against herbivores Spines and thorns Chemical toxins Herbivores must adapt to the defenses created by their food.
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  • Coevolution: A series of reciprocal evolutionary adaptations in two species. A change in one species acts as a new selective force on another Poison-resistant caterpillars seem to be a strong selective force for Passiflora plants
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  • Eggs Sugardeposits Heliconius, caterpillar has enzymes to break down poison of Passiflora (plant). The Plant has since adapted sugar-deposits to mimic butterfly eggs. The Heliconius butterfly will not lay eggs on a leaf with another butterflies eggs already attached.
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  • parasitism parasitenourishment host harmed In parasitism (+/ interaction), one organism, the parasite, derives nourishment from another organism, its host, which is harmed in the process endoparasites Parasites that live within the body of their host are called endoparasites Such as nematodes and tapeworms ectoparasites Parasites that live on the external surface of a host are ectoparasites Such as mosquitoes and ticks
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  • Ticks Tapeworms Liver flukes
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  • Pathogensdisease-causing parasites Pathogens are disease-causing parasites Pathogens can be bacteria, viruses (non-living), fungi, or protists Non-native pathogens Non-native pathogens can have rapid and dramatic impacts American chestnut devastated by chestnut blight protist A fungus-like pathogen currently causing sudden oak death on the West Coast Non-native pathogens can cause a decline of the ecosystem
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  • What adaptations have predators & prey evolved in order to help survive? Explain coevolution. Explain using examples why non-native parasitism, or pathogens, are typically dramatic for an ecosystem. Explain and give an example of an autotrophic organism. Explain and give an example of an heterotrophic organism. What do you think the suffix trophic refers to?
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  • Create a food chain that spans 4 trophic levels. Be sure to start with a producer.
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  • Trophic structure A pattern of feeding relationships consisting of several different levels Food chain Sequence of food transfer up the trophic levels
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  • Producers Support all other trophic levels Autotrophs Photosynthetic producers Plants on land Cyanobacteria in water
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  • Where does the energy for life processes come from?
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  • Producers Without a constant input of energy, living systems cannot function. What do you think is the main energy source for life on Earth? Sunlight
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  • Only plants, some algae, and certain bacteria can capture energy from sunlight and use that energy to produce food. Autotrophs What are these organisms called?
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  • In a few ecosystems, some organisms obtain energy from a source other than sunlight.
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  • Life Without Light Some autotrophs can produce food in the absence of light. Chemosynthetic bacteria are important in deep sea ecosystems.
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  • Some chemosynthetic bacteria live in very remote places on Earth, such as volcanic vents on the deep-ocean floor and hot springs. Others live in more common places, such as tidal marshes along the coast.
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  • Consumers Organisms that rely on other organisms for their energy and food supply are called ________________. Heterotrophs are also called consumers. heterotrophs
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  • There are many different types of heterotrophs. ____________ eat plants. ____________ eat animals. ____________ eat both plants and animals. ____________ feed on plant and animal remains and other dead matter. _____________, like bacteria and fungi, break down organic matter. Herbivores Carnivores Omnivores Scavengers Decomposers
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  • Consumers Heterotrophs Primary consumers: Eat Producers Secondary consumers Tertiary consumers Quaternary consumers Detritivores and decomposers: Derive energy from dead matter and wastes
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  • Plant A terrestrial food chain Producers Phytoplankton An aquatic food chain
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  • Plant A terrestrial food chain Producers Phytoplankton An aquatic food chain Primary consumers Grasshopper Zooplankton
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  • Plant A terrestrial food chain Producers Phytoplankton An aquatic food chain Primary consumers Grasshopper Zooplankton Secondary consumers Mouse Herring
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  • Plant A terrestrial food chain Producers Phytoplankton An aquatic food chain Primary consumers Grasshopper Zooplankton Secondary consumers Mouse Herring Snake Tuna Tertiary consumers
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  • Plant A terrestrial food chain Producers Phytoplankton An aquatic food chain Primary consumers Grasshopper Zooplankton Secondary consumers Mouse Herring Snake Tuna Tertiary consumers Hawk Killer whale Quaternary consumers Trophic level
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  • Detritivores, such as scavengers, eat detritus, or dead organic material. Decomposers, mainly fungi & prokaryotes, secrete enzymes to digest molecules in organic material and convert them to inorganic forms. Great Circle Of Life Think of the Great Circle Of Life.
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  • Copyright 2009 Pearson Education, Inc. Food web A network of interconnecting food chains
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  • Producers (plants) Primary consumers Secondary and primary consumers Tertiary and secondary consumers Quaternary, tertiary, and secondary consumers
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  • This food web shows some of the feeding relationships in a salt-marsh community.
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  • Species diversity is the variety of organisms that make up the community It has two components: Species richness Species richness is the total number of different species in the community Relative abundance is the proportion each species represents of the total individuals in the community Plant species diversity in a community affects the animals Species diversity has consequences for pathogens
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  • Community 1 A: 25%B: 25%C: 25%D: 25% Community 2 A: 80%B: 5%C: 5%D: 10% ABCD Compare and contrast the species richness and species abundance for communities 1 & 2.
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  • Communities with higher diversity are: More productive and more stable in their productivity Better able to withstand and recover from environmental stresses More resistant to invasive species, organisms that become established outside their native range
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  • Which would you expect to have higher species diversity, a well-maintained lawn, or one that is poorly maintained? Explain. The poorly maintained lawn would have higher species diversity. A well-maintained lawn should have low species diversity. While a lawn that is cared for may not be a perfect monoculture, any weeds that are present would have low relative abundance.
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  • Explain competitive exclusion Difference between a fundamental and realized niche. When 2 species are competing for a resource, the species with a slight advantage will eliminate the other. Fundamental niche is the niche potentially occupied by the species. The realized niche is the portion of the fundamental niche the species actually occupies.
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  • What is symbiosis? A +/- symbiotic interaction in which one organism derives nourishment from a host. An interspecific interaction that benefits both species. Symbiotic relationship that benefits one of the species but neither harms nor helps the other. A fern growing in the shade of another plant is an example. When individuals of 2 or more species live in direct contact with one another. Parasitism Mutualism Commenalism
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  • Dominant Species Population in a community that has the highest biomass Biomass is the sum weight of all the members of a population.
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  • Keystone species exert strong control on a community by their ecological roles, or niches A keystone species has a much larger impact on its community than its biomass would suggest. Field studies of sea stars illustrate their role as a keystone species in intertidal communities Keystone absent
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  • Pisaster ochraceus may prey on sea urchins & mussels with no other natural predators. If the sea star is removed from the ecosystem, the mussel population explodes, driving out most other species The urchin population annihilates coral reefs
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  • EXPERIMENT RESULTS With Pisaster (control) Without Pisaster (experimental) Year 737271706968676665641963 0 5 10 15 20 Number of species present
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  • RESULTS With Pisaster (control) Without Pisaster (experimental) Year 1963 0 5 10 15 20 Number of species present 64656667686970717273
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  • The beaver is an ecosystem engineer transforms its territory from a stream to a pond or swamp. Beavers affect the environment by cutting down older trees to use for their dams. allows younger trees to take their place. Beaver dams alter the riparian area (where water and land interact) Dams change the edges of streams and rivers into wetlands, meadows, or riverine forests. Beneficial to groups of species such as amphibians, salmon, and song birds
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  • Disturbances Events that damage biological communities Storms, fire, floods, droughts, overgrazing, or human activity The types, frequency, and severity of disturbances vary from community to community
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  • Communities change drastically following a severe disturbance Ecological succession Colonization by a variety of species A success of change gradually replaces other species
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  • Primary succession Begins in a virtually lifeless area with no soil Secondary succession When a disturbance destroyed an existing community but left the soil intact
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  • Time Shrubs Annual plants Perennial plants and grasses Softwood trees such as pines Hardwood trees Primary or secondary succession?
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  • After what type of natural disasters would one expect to see primary succession?
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  • Primary succession could occur on the barren slopes of a recently erupted volcano.
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  • A large-scale fire struck Yellowstone National Park in 1988. It quickly responded. Would this be primary or secondary succession? (a) Soon after fire(b) One year after fire
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  • Ecosystem All the organisms in a community as well as the abiotic environment Components of ecosystems Energy flow Passage of energy through the ecosystem Chemical cycling Transfer of materials within the ecosystem Why is this important to us? What are the four major macromolecules, what do they look like? Where do we get the material to make these molecules?
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  • Energy flow Light energy Chemical energy Chemical elements Heat energy Bacteria and fungi Chemical cycling A terrarium has the components of an ecosystem
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  • Primary production The amount of solar energy converted to chemical energy Carried out by producers (autotrophs) Produces biomass Amount of living organic material in an ecosystem
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  • gross primary production (GPP). The total primary production in an ecosystem is known as that system gross primary production (GPP). The amount of energy available to consumers is the net primary production (NPP). What is the difference? NPP = GPP-R R is the energy used in respiration by the producers.
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  • Open ocean Estuary Algal beds and coral reefs Desert and semidesert scrub Tundra Temperate grassland Cultivated land Boreal forest (taiga) Savanna Temperate deciduous forest Tropical rain forest 0 500 Average net primary productivity (g/m 2 /yr) 1,000 1,500 2,5002,000
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  • Trophic Levels Each step in a food chain or food web is called a trophic level. Producers make up the first trophic level. Consumers make up the second, third, or higher trophic levels. Each consumer depends on the trophic level below it for energy.
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  • So energy is created by producers and then consumed by consumers. Is there a limit to the trophic levels this energy can travel?
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  • At each stage of consumption, or trophic level, some energy is lost as heat to the environment. This is why most producers can only support 4-5 consumer levels.
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  • Explain the difference between populations, communities, and ecosystems.
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  • Ecological Pyramids An ecological pyramid is a diagram that shows the relative amounts of energy or matter contained within each trophic level in a food chain or food web.
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  • 0.1% Third-level consumers 1% Second-level consumers 10% First-level consumers 100% Producers Energy Pyramid: Shows the relative amount of energy available at each trophic level.
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  • What do you notice about the amount of stored energy as it passes from one trophic level to the next? What does this mean about the amount of living tissue (biomass) able to be supported at each level? Because each trophic level harvests only about one tenth of the energy from the level below, it can support only about one tenth the amount of living tissue.
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  • 1,000,000 kcal of sunlight 10 kcal 100 kcal 1,000 kcal 10,000 kcal Producers Primary consumers Secondary consumers Tertiary consumers
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  • The more levels that exist between a producer and a top-level consumer in an ecosystem, the less energy that remains from the original amount. Only about 10 percent of the energy available within one trophic level is transferred to organisms at the next trophic level.
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  • Ecosystems are supplied with a continual influx of energy Sun Earths interior Life also depends on the recycling of chemicals Organisms acquire chemicals as nutrients and lose chemicals as waste products
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  • Biogeochemical cycles Nutrient cycles that contain both biotic and abiotic components. Cycle chemicals between organisms and the Earth Can be local or global Decomposers play a central role in biogeochemical cycles
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  • Consumers Geologic processes Producers Decomposers Nutrients available to producers Abiotic reservoir 4 1 2 3
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  • Carbon is the major ingredient of all organic molecules respiration balances photosynthesis The return of CO 2 to the atmosphere by respiration closely balances its removal by photosynthesis The carbon cycle is affected by burning wood and fossil fuels (these increase CO 2 in the atmosphere)
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  • Photosynthesis Decomposers (soil microbes) Cellular respiration Detritus 4 1 2 3 5 Plants, algae, cyanobacteria Primary consumers Higher-level consumers Burning CO 2 in atmosphere Plant litter; death Wastes; death Decomposition Wood and fossil fuels
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  • Moves nitrogen from the atmosphere through the living world. Nitrogen is a common limiting factor of plant growth.
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  • Nitrogen is an essential component of proteins and nucleic acids Nitrogen has two abiotic reservoirs Air Soil Nitrogen fixation converts N 2 to nitrogen used by plants Carried out by some bacteria and cyanobacteria
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  • The Nitrogen Cycle Certain types of bacteria that live in the soil and on the roots of plants can convert nitrogen gas (N 2 ) into ammonia in a process known as nitrogen fixation.
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  • nitrification Other bacteria in the soil convert ammonia into nitrates and nitrites in a process known as nitrification. Once these products are available, producers can use them to make proteins. Consumers then eat the producers and reuse the nitrogen to make their own proteins.
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  • Nitrogen (N 2 ) in atmosphere 8 Plant Animal Assimilation by plants Organic compounds Organic compounds Death; wastes Denitrifiers Nitrates in soil (NO 3 ) Detritus Decomposers Decomposition Nitrifying bacteria Ammonium (NH 4 + ) in soil Nitrogen fixation Nitrogen fixation Nitrogen-fixing bacteria in root nodules Free-living nitrogen-fixing bacteria and cyanobacteria 6 1 2 7 4 3 5
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  • Water moves between the ocean, atmosphere, and land.
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  • Water molecules enter the atmosphere as water vapor, a gas, when they evaporate from the ocean or other bodies of water.
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  • Water can also enter the atmosphere by evaporating from the leaves of plants in the process of transpiration.
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  • Water vapor condenses into tiny droplets that form clouds. The water returns to Earths surface in the form of precipitation. Water enters streams or seeps into soil where it enters plants through their roots. Water can also infiltrate the ground and travel as groundwater.groundwater
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  • Copyright 2009 Pearson Education, Inc. 1. Describe the characteristics of a community 2. Explain how interspecific interactions affect the dynamics of populations 3. Describe the trophic structure of a community 4. Explain how species diversity is measured 5. Describe the role of environmental disturbance on ecological succession 6. Explain energy and nutrient cycling in ecosystems
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  • Why is it important to study communities? What is a community? Give both an aquatic and land organism examples of predation. Give both an aquatic and land organism examples of mutualism. Give both an aquatic and land organism examples of competition. Give an example of commensalism. Give an example of parastism.
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  • Name two adaptations that may enable predators to capture prey. Name two adaptations that may enable prey to evade predators. Write a brief (fiction or nonfictional) scenario about two organisms coevolving in a predator-prey relationship. Write a brief (fiction or nonfictional) scenario about two organisms coevolving in a mutualism relationship.
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  • Imagine the fields behind the school as an ecosystem: Name a producer, primary consumer, and secondary consumer. How much of the producers potential energy will be consumed to the secondary consumer? From where does each trophic level obtain its carbon & nitrogen? Think back to musical chairs when would one expect interspecific competition occur?
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  • What are detritivores and what do they do for an ecosytem? Explain what is meant by species richness in an ecosytem. What are keystone species and what would happen if they were removed from an ecosystem? Explain and provide examples of primary & secondary succession. Explain what invasive species are and the impact they play on an ecosystem.
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  • Explain what this equation means: NPP = GPP R a Net Primary Production = Gross energy used in respiration
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  • Amount of light and the depth it can reach. Light decreases with depth. What is a limiting factor? Limits growth of populations Most common limiting factors in marine environments are nitrogen and phosphorous. A lake that is nutrient-rich and supports a vast array of algae is said to be eutrophic.
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  • Conservation Biology and Global Change Chapter 56 Chapter 56 (only section 1)
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  • Scientists have named and described 1.8 million species Biologists estimate 10200 million species exist on Earth Tropical forests contain some of the greatest concentrations of species and are being destroyed at an alarming rate Humans are rapidly pushing many species toward extinction
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  • Conservation biology, which seeks to preserve life, integrates several fields Ecology Physiology Molecular biology Genetics Evolutionary biology
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  • Bioremediation: Using organisms, like prokaryotes, plants and fungi to detoxify polluted ecosystems. Bioaugmentation: Is the intorduction of desirable species such as nitrogen fixers to add essential nutrients.
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  • Rates of species extinction are difficult to determine under natural conditions The high rate of species extinction is largely a result of ecosystem degradation by humans Humans are threatening Earths biodiversity
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  • Biodiversity has three main components Genetic diversity Species diversity Ecosystem diversity
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  • Genetic diversity in a vole population Species diversity in a coastal redwood ecosystem Community and ecosystem diversity across the landscape of an entire region
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  • Genetic diversity comprises genetic variation within a population and between populations
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  • Species diversity is the variety of species in an ecosystem or throughout the biosphere According to the U.S. Endangered Species Act An endangered species is in danger of becoming extinct throughout all or a significant portion of its range A threatened species is likely to become endangered in the foreseeable future
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  • Conservation biologists are concerned about species loss because of alarming statistics regarding extinction and biodiversity Globally, 12% of birds, 20% of mammals, and 32% of amphibians are threatened with extinction Extinction may be local or global
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  • Philippine eagle Javan rhinoceros Yangtze River dolphin
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  • Human activity is reducing ecosystem diversity, the variety of ecosystems in the biosphere More than 50% of wetlands in the contiguous United States have been drained and converted to other ecosystems
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  • The local extinction of one species can have a negative impact on other species in an ecosystem For example, flying foxes (bats) are important pollinators and seed dispersers in the Pacific Islands
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  • Human biophilia allows us to recognize the value of biodiversity for its own sake Species diversity brings humans practical benefits
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  • Species related to agricultural crops can have important genetic qualities For example, plant breeders bred virus- resistant commercial rice by crossing it with a wild population In the United States, 25% of prescriptions contain substances originally derived from plants For example, the rosy periwinkle contains alkaloids that inhibit cancer growth
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  • The loss of species also means loss of genes and genetic diversity The enormous genetic diversity of organisms has potential for great human benefit
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  • Ecosystem services encompass all the processes through which natural ecosystems and their species help sustain human life Some examples of ecosystem services Purification of air and water Detoxification and decomposition of wastes Cycling of nutrients Moderation of weather extremes
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  • Most species loss can be traced to four major threats Habitat destruction Introduced species Overharvesting Global change
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  • Human alteration of habitat is the greatest threat to biodiversity throughout the biosphere In almost all cases, habitat fragmentation and destruction lead to loss of biodiversity For example In Wisconsin, prairie occupies