Cenozolc Fossil Mollusks FromWestern Pacific Islandu;Gastropods (Etlimidae andVolutid ae Through Terebridae)87 HARRY S. LADD

GEOLOGICAL SURVEY PROFESSIONAL PAPER II71

Descriptions 0r citations of 271 representatiues ofgastropod Jamilies fro* seuen island groups

11ÀtAcfltoGlEmlsail, 9aRls

U N I'f I-l) s f'\'I ilS (;()\' IìR\\{ tNl PRINTING OI"FICE, WASHT)iGT()N: 19tì2

Measurements of the figured specimen, USNM175133: iength 44.8 mm, diameter 24.1 mm.

The Fijian fossil seems most closely related toC. ngat:i,artrrs described by Martin (1895, p. 23, pi.4, figs. 57-61) from the upper Tertiary of Java buthas a iou'er spire and is more constricted anteriorly.

Contts sp. A resembles C. monila Hlr'ass, a Hoìo-cene species from the tropical Indian and westernPacific Oceans but has a more angular shoulder, ismore constricted anteriorly, and has more stronglydeveìoped basal ribs. Il also resembles C. fctsteriCìench and Arguay (Clench, 7942, p.34, pl. 72, frg.5), a species living in the rvestern Atlantic, but hasa lor,l'er spire and a more strongly concave profile. Ifadditionaì specimens are collected, the Fijian fossilmay prove to be an unnamed species.

Q6g.11vvsn6s.-[ single specimen from stationK700 on \ranua Levu, Fiji; age, probably Pliocene(Tertiary /r).

Conus sp.

Conus sp. Ladd, 19?0, U. S. C'tological Survey Prof. Paper640-C, p. C8, pì. 5, frg. 2, late Eocene (Tertiary Ó);Eua, Tonga.

Genus KENYONIÀ Brazier

Brazíer. 1896, Linnean Soc. New South Wales Proc., v. 21,p. 346.

T -upe (b1, monotypy) .-Kenyonia pttlcherrimnntErazìer. Holocene. Nerv Hebrides.

Kenycnia cf. K. chiangi (Azuna)

.. Plate- 19. figures 17-19

Kengoniu cf. K. chiangi (Azuma), Ladd, 1976. Nautilus. v.70, no. 4, p. 135, fiSs. 21, 22.

A single erample of a highly decorated cone .was

coiiecied at USGS loc.alitli 25737 on the NavakaRiver. Sanfo, Ner.v Hebrides. The shell measures:ìength 10.4 mm, dian-reter 5.9 mm. USNI\{ 214297.It ma.'- be closely related to the Holocene sheìl de-

-qcribed by Erazier in 1896 as probabll' from Neu'Hehljrles. but 'rhat shell, unfortunatel-v, has beenÌosl. The fossil from Santo ma1' be conspecìfic rvithTai'aiiít:cotttts cltiangÌ clescribed b1' Azuma (1972, p.

59, flgs. 5, 6) and ìater figured b-v Okutani (19?5, pl.10, fig. 27 ) , but the fossil is more slender anteriorì1'and malr have stronger spiral -qculpture. Azuma'stype of T. chi,artgi u'as collected in the South ChinaSea at a depth of 200 fathoms (365 m). Okutanir't'unrl ìir-ìng specinrens fairll- conlrnon on banks atdeplhs of 150-i90 m in Japanese \raters.

Family TEREBRIDAE

Genus HASTULA H. and A. Adams

-{i:,il:-. H. and A., 1853, Genera oí Recent mollu-.ks, v. 1, p'22i.

PALEONTOLOGY 81

T17pe (b-v subsequent designation, Cossmann,1896, Essais de paléoconchoìogie comparée, pt' 2' p.

53).-Buccirtum striqíIotum Linnaeus. Holocene,Indo-Pacific.

Hastula sp.

Hastttla sp. Ladd, 19?0, U.S. Geo)ogical Survey, Prof. Paper6.10-C, p. C-8, pl. 5, fig. 3. Late Eoc'enc (Tertiary b);Eua, Tonga.

Genus TEREBRA Bruguiere

Rruguière, 1789, En:1'cÌope Cie méthodique. Histoire nat. vers,v. 1, p. 15 (trenus without sPecies).

Subgenus TEREBRA sensu stricto

Type (b-,- monoil'py) Lamarck, 7799, Soc- His-toire Nat., Paris, Mém, p. 79) '-fi71s6'ì7711m str'btt-

lrLttLm Linnaeus. Holocene, rvestern Pacific'

Te:ebra (Terebra) subulata (Linnaeus)

]lle'.t' 30, figu rc Ii

Bur:cinum subtrlotttnt Linnaeus. 1?67, -s1'stema naturae, 12thed., p. 1205.

Tcrebra. subu!.ata (Linnaeus), Reeve, 1860, ConchoìogicaIconica, t. 72, Terebra, pl. 6, no. 22; Tryon, 1885'l\{anual conchology, v.7, p. 10, pl. 1, fig- 3; Demond,1957, Pacific Sci., v. 11. no. 3, p. 334; Cernohorsky,196?, llarine shells of the Pacific, r'. 1, P. 205, pl. 52,

fig. 385.

A single fossil from the Ner.v Hebrides, thoughworn and incontplete, sho\4's traces of the subsuturalgroove on the early \4'horls and the double row ofsquarish brol-n spots that are unjque.

Ileasurements: heieht 82.5 mm. diameter" 15-9

run.Occut't't'nce.-Station SM242, Santo, Neu' Heb-

rides; aB€, Pleistocene. The species lives toda-v:

throughout the Indo-Pacific region. Not preriouslyreported as a fossil.

S'bgenus STRIOTEREBRUM Sacco

Sacco. 1891, I- molluschi dei t.err:ni terziarii del Piemontee delia Liguria, pt. 10, p. 33.

T'Ape (by original designation).-Terebra bas-t er ctt í Nyst. Miocene, If editerranean.

Terebra (Strioterebrum) torquata Adams and Reeve

Plale 30. figure iTertbra. t.orqtLota Adan'rs and Reeve, 1850, Zooìog1' of the voy-

age of H.M.S. -\tnna.ro:tg, p. 30, pì. 10. fig. 13; Mac-Neil. 1960, Ll.S. Geol. Survev Prof. Paper 339, p. 125,pl. li. figs. .l8,

19.

Cinguloterebra lorqiLat.a (Adams and Reeve), Habe, 1964,

Shells of the *-e-s-:e;-n Par.'jfic in coior. v.2, p. 128. pl.40, fig. 19..

Each u'horl bears two .strong, beadecl, subsuturaìriciges, the rirlge next to the suture being thest.r'onger; belou-the nrajor spirals are f()ur se{ondar-Yspiraìs and faini tlaces of tertiarl'spiraìs.

lleasuremenls of the figurecl specinten: height(incomplete) 35.7 mm, diameter 8.3 mm.

Occtn'r'ence.-One incomplete shell from stationSX2-12, Sarnto, Nelv Hebrides; age. Pleistocene. The.sp-:.- ies ìir-es torlay in Japrrn rntt olher par':s ,it' '.h:wes:ern Pacific; IfacNeil reportecl it from the Plio-cene of Okinau'a.

Terebra (Strioterebrum) nebulosa Sowerby

Plate 30, figure 2

Ter..bru, ncbLtlosa. Sowelb,v. 1825. Tanken'ille ca-,:rlogue. -\p-pendix, p. 25; Reeve. 1860, Conchologica lconic:r, v. 12,T,:rebru, pl. 1, fig. 1; Cemohorski,, 196î, ìf arine sheiÌsof the Pacific, v.1, p.204, pÌ.51, fig.380.

A medium-sized species with a swoilet subsuturalbanri thaL bears lìne spirals ancl is bounded anterior-ly by an open punctafe g-roove ; remaincier of sheilrei;culated by coarser spirals and strong cun'eclaxral ribs.

]Ieasurements of the figured specimen: height(aper incomplete) 50.9 mm, diameter' 11.4 mm.

()cczn't'ettce.-A single specimen from the Pleisto-cene marls on Santo (station SNÍ242), Neu. Heb-riLles. The species lives today in parts of the tropi-eai Indo-Pacific. Not previously' reporteci as a fossii.

Terebra (Strioterebrum) sp. A

Plale 30, figure 3

Shell large, sides neariy straight. Sculpture con-sisiing of a wide coarsely noded subsutural bandthat occupies more than one-third of each rvhorland spiral cords. The nodes ar€ regularly spacecl,each one pinched to a sharp ridge that parallels theaxis of the shell; nodes overridden by fine spiraìcords, traces of seconclary corcls intervening. Re-mainder of n'horl has three or four spiral cords thatare coarser than those of the subsutural band but,like them, have traces of fìner cords befween. Sutureinconspicuous ; coìumella has a heavy clouble fold;anterior canal extended ancj reflered; outer lip notpreservect.

lleasurements of the single specimen, USNM175084; height (incompiete) 33.3 HH, diameter12.8 mm.

The major sculpfural features-the sLronglynoded subsutural band and the well-developed spiralcords-suggesL Terebru nelsotLi, Hanna and Israel,sk!-, an early Miocene species from Peru clescribeclby Nelson (1870, p. 10, reprínt) ancl by Spieker(1922, p.36, pl. 1, fig.2), but the nodes and spiralsof the Enirvetok fossil are much sharper than arethose of T. ttelsottt. The Enirvetok shell probablyrepresents an undescribecl form, but the single ex-ample is incomplete anri a nrìrÌle is withhelcl.

Occut't'ertct,.-A single spt:cinren from drill hole

F-1, Eni',,;etok, ata cleoth of 680 690 feet (210 m);age, late lf iocene (Tertiary g).

Terebra (Strioterebrum ?) sp.

Trrehrq t.-\-irir;Ér,r'ebr'zrin) species Ladd, 1g3,1, B. p. Bishcipll:rs Buil. i 19. p. 2:33, pl. 11, fig. 9.

No additional shells collected. The species is un-usual in that it shorvs no trace of spiral scuptureother than the groove marking the base of the lub-sntur:rl band. As noted by Cernohorsky (1966, p.6;), it rioes not resemble any of the living FijianTerebricl:re.

]Iezrsurements of the incomplete flgurecì sllecimen,B. P. Bishop ùIris., Geol. No. 1223: height 26.6 mm,diameler 5.8 mm.

OcctLrretzce.-station 59, Viti Levu, Fiji; age, late]liocene (Tertiary g).

Subgenus DEC0RIHASTULA Oyama

Oyama, 1961, Venus, v.21, no. 2, p. 1gb.

Type {by original designation) .-Tet.ebr.rt afi.nisGra;". Holocene, Inclo-Pacifi c.

Tercbra lDecorihastula) affinis Gray

Pla,ue 10, figur.es -1, íTert,bra uljinis Gray, 1814, Zooì_ Soc. Lonclon proc., pt. 2, p.

60.Der:orih.astultt a.ffinis (Cray), Kira, 1g62, Shells of the west_

e'r-n Pacific in color, v. 1, p.20g, pl. T1, fig.2: Cerno-horsky. 196íi, Veliger, v.9, no. 1, p. -15, pI.5, fig.26.

Fossil i'epresentatives are small and incomnlete.but they shou. major sculptLrral features, inclucilngfhe punctate grooves, that characterize this species.

Measurements of the figured specimen from Sa,i-pan, USNM 175081: height 23.8 mm, diameter 6.1mm.

Occtr.t't ettce.-Figured specimen from the Tana_pag Limestone of Saipan (USGS loc. 12g91) ; age,Pleistocene and Holocene. A second specimen fromdrill hole I'-1, Enirvetok, at depth of 180-140 feet(40 m); age, Quaternary. The species is commontoday throughoub most of the tropical Inclo-pacific.Not previor-rsly reported as a fossil.

Subgenus IVIICROTRYPETES Pilsbry and Lowe

Pilsbry antl Lowe, 1932, Acad. Nat. Sci. philadelphia proc..v. 8-1, o. .{8.

Ty 1te ( by original designation ) .-Terebra iolaPilsbr_v ;rncl Lowe. Holocene, west coast of Mexico.

The subgenus inclucìes slender forms rvith un-broken ribs that lack a disfinct subsutural groove.

82 CENOZOiC FOSSIL IÍOLLUSKS FROM WESTERN PACIFIC ISLANDS; GASTROPODS

Tcrebra

Sheìì smzrll,convex rvhorls

(Microtrypetes) vanualevensis Ladd, n. sp.

P.late i0, figures 6, 7

slencler, macle up of 10 or more gently; ilpex nof preserved. Sculpture con-

P.{LEONTOLOGY 83

sisting of strong, narrow-crestecì, curved axial ribs;11 presenb on body rvhorl; concave area betlveenribs scored by spiral grooves of variable size; upper-mosL two or three are faint, the next one n'ìoreprominent, suggesting the boundary of a subsuturalband. Suture impressed; anterior canal extended;columelìa rvjth a single narrow fold.

Measurements of the holotype, USNM 175085;height (incomplete) 15.6 mm, diameter 3.3 mm;paratype, USNM 175086:height 14.9 mm, diameter4.0 mm.

One smail nearly complete specimen and tu'o in-complete larger sheìls from \riti Levu closel.v- re-sembìe the \ranua Levu fornr, but their u'horls areless convex, their sutures shallower, and they havea more discernible subsutural band. The nearly com-pìete Viti Levu shell is from stzrtion C742 ar-rd prob-abìy is Pliocene (Tertiary h ) ; the trvo fragments oflarger sheils from station 165 are probabÌ1' older-late Miocene (Tertiary g). Final decision as to theidentity of the Viti Levu shells musl au'aif the col-lection of better material.

T. ranttolr:t;ensis al)pears to be most closell- le-lated to T. iokL Pilsbry and Lorve, the t1'pe of XIicro-tt'ypetes, but has fell'er axial ribs than does thatspecies, a Holocene shell obtained front 20 fathomsoff Mazatlan. The Fijian species is larger and moreslender than is T. (M.) ntcLriatct Pilsltrl' attd Lou'e,another Holocene species occurring off the Pacificcoast of Pananra, and ii has slronger spiraìs butfev'er axjals than does that species.

T. (X[.) sTrcr Br"orvn ar-rd Pilsbr-v (1913, 1r. 497).a Pleistocene species from the Isthmus of Panama,has a more prominent subsutural groove and moreaxial ribs than does the Fijian species.

Occut't'ence.-Three incomplete specimens fromstation 817, \ranua Levu, Fìji; age, Pliocene (Ter-tiary h ) .

Terebra (Microtrypetes) kercnsis Ladd

Plate 2.1. figuI e-s 20. 21

Tcrcbra (llict'otrufietc.s) À-rrozsis Ladd. 1976. Nautiius, v.90, no. 4, p. 137, figs. 3i-33.

Holotype, USNM 274340, measures: heíght 10.0mm, diameter 2.0 mm. Many sheils collected fromUSGS locality 25775 on the Kere River, Santo, Neu'Hebrides ; age, Pleistocene.

. Subgenus 0XYMERIS Dall

Dall, 1903, tl.S. Natì. llus. Ploc., r-.26, p. fì51. ]ierl'nanrefor ,4czrs Gray.

Tylte (Gray, 1847, ZooI. Soc. London Pi'oc., p.

139).-BuccittLLnt nracttl.otitnt Linnaeus. Holocette.Indo-Pacific.

Terebra (Oxymeris) maculala (Linnaeus)

Platc 30, figure [ì

Iiuct:itrtutt macula.tunt Linn:reus, 1?58, S1'stcnra Naturae, 1OLh

cd., p. 741.Tercbro maòulata (Linnaeus) , I)e mond, 1f)57, Pacific Sci.,

v. 1i, 334; Cernohorsky, 196?, Marine shelìs of thePacific, p.204, pl. 51, fig.376.

A single example of this u'idely distributed livingIndo-Pacific species rvas collected from the Pìeisto-cene and Holocene Tanapag Limestone of Saipan(USGS locaìitl' 17891). The shell, USNM 1750?9,measures: height 165 mm, diameter 45 mm. Theearlv u'horls of the shell retain traces of the originalspiral lines of brot'n spots. Xtartin (1890, p. 280)reported the specìes from the Quaternary of Timor.

Terebra (0x;-meris) interlineata Deshayes

Plalc 30. fie'ure g

Ttt't'Itro ìrtttrlitteala Di'sha1'es. 185!), Zool. Soc. London Proc.,p. 277; Reeve, 18C0. Conchologica Iconica, v. 12 Ttre-bro plr.12. no. 51.

Tercbra (O:runeris) 1:renulata (Linnaeus) variant, Cernehorskv and Jennings. 1966. \religer, v. I, no. 1, p.47*42.

Recognized by man1. authors as a sl'non]'m or asa varianl of the rvidespread living lndo-Pacificspecies T erebt a ct enlLlats (Linnaeus ) , characterizedparticularìy by numerous axial wrinkles that per-sist over the entire shell. Such lvrinkled sheìls haveheletofore been founcl onl-v* in Harvaii. The Net'Hebrides Pleistocene shell from station SM242 onSanto is the fir'st fcissii o3currence. It presen-estt"aces of reddish-brou-n markings on the subsuturalband l:ut none of the bronn dots that mark thervhorls belorv the band.

l'Ieasulements: heisht 56.0 mm, cliameter 13.6mm.

. Terebra (0xymeris) cf. T. feìina (Dillwyn)

Plate 30, figures 10-12

An unusuall-v 1arge, nearly compiete shell (pl. 30,fig. l0) was reco\rerecl from the Pleistocene marls ofSanto in the Neu' Hebricles (station SM242). Theiììplinran mèrqrrlp{' heioht I înos miqqìnrrl 110

mm. tliameter 25 mm. 7. f clitta live-q today in manyparts of the tlopical Pacific but rareì1' exceeds 80mm in height. The fossil shell, u-hen con'rplete, prob-abì-v exceecìed 150 mm in height. Its suture and sub-sutural gì'ocve ar"e nloÌ'e plonríttent than on Holo-cene shells. as ar'e llres of glou-th on later u-borls.

Tii-c' ir.ruirrrrplele sheììs (rrl. 30, figs. 11, 12) frcnrstation 165 on \-iti Ler-u. Fiji, are comparabìe insize u'ith Holocetre shelÌs of T. f eli.rta, but theirslul-Ìr:ule. lil,e tha'L of the \-ounger Neu' Hebrìdesíossil. is co:ri'sei' iharr on Holc'cene shells. The Fijian

8.! cENozotc Fossrl ùIoLLLrsKs FRo-1r wESTERN pAcrFrc TSLANDS; G-{STRopoDSfos.sils are late Miocene (Teriiary {l) in age. Boilra* frgur.eri. The larger, LrS.\-lI iZiirsi, meiìsures:he:ght (incomplete) 85.6 mm. dianreter. 15..? mm.The .smaller fragmentary fossii, USNII 1Z50gg, hasthe coarse axials of the early.- whorls ; it mea.:r.rres :hlrfht 15.8 ffift, diametei. l{1.1 mm. T. it:!ittit(D,ìirv,v'n) lives tocì:ry throughout the fropical Inclo_Pacdc (Cernohorsky, 1g62, p. 1g9).

Terebra (Oxymeris) argus (Hinds)plate 30, {ìgure 13

Ter. ir;,:L rtrgus Hincls, Lg43, Zc:oI. Soc. Lonclon pioc.. p. 160;Reeve, 1860, Conchologic.r lconica, v. 12. Tet.ebra, pl.6, rrg. 11; Tn,on, 1gg5, Manual conchoicgX,, ". r, O. ii,pl. 2, fig. 21; Cei.nohor.sky-, 1967, ]Ialine sheìls paciflc,

1, p. 196, pt. .19, fig. 35.1.Subitl,tt. ar'gris { Hinds) , Kira, 1g62, SheÌls of w-e ster-n pacific

in color, v. 1, p. 209, pl. 71, fig. 15.

-{ small, near.ly complete fossil shell from Saipanreplesents this livrng species. The fos:sil lacks thepale color pattern founcl on fresh shells. The rvhorlsare smocth excepf for traces of axials on earlyrvhoris and a punctate subsutural groove.lfeasui'ements of the figurecl specimen, USNII

175rr9.3; height (apex incomplete ) :32.ó mm, cliam_eter 6.8 mm.Occurrence.-UsGS locaìity 1?ggi on Saipan inthe Pleistocene and Holocené Tanapag Limestone.

The species lives today in many p*rt-"' of the rvest_ern Pacific, inclucling the Marianas.

Terebra (0xymeris) sp. B

Plate 30, figure 14

A juvenile .shell consisting of 10 strongly plicate,flaf-sided whorls form a sharp spire u,ith an apicalangle of 26". Both suture ancl subsutural groove arewell imples^secl; the coiumella ha^s a single narrowfold.

Measui.ements of the specimen, USNM 1250g9:height 3-1.4 mm, diameter 11.5 mm.The fossil resembles the abunclant anri rvidespread

Holocene Inclo-p:rcific species, T. nruttt[rtta ( Linn_aeus) but hns a slighil-v. greater apical angle ancl ismuch more strongly plicate. It may represent anundescribed form, but, as notecl, the only specimenrs J u\-entle.

Occttt't'ence._-Drill hole F_1,of 800-tì10 feet (245 m); ,,.ge,tiary' g).

Subgenus MyURELLA HindsHinds" i8.1.1, ln Sowe rby, The saurus Conchl,liorunt, f erehra,p.1?1.

Ty pe (by sLlbseqlrent cìesignation, I)all, 1g0gNautilus, v. 21, p. I2lt).-Tr:r.,,i'rrr ,rry,r,.o.s l,amiri.ck.Holocene, Indo-pacifìc.

Terebra (Myurella) myuros LamarckPlaie 30, figrire 16

Tere brrt rnlLLt"os Lamerck, 1g22, Systèms aninraux sans verte-bres, v. 7, p. 2g9; Reer.e, 1g6lt, Con.iroìogÌ"u Iconica, v.72. Terebra, pl. g,31,

Spire slencler ancl Ilat_sicled. SubsuLulal banclmiìde up of tu'o nearìy ilat but slighil-v beadeclspir-als, the one next to the suture lvider than theother; belorv the subsutural band u." ."u*J;;;É;secondar). spiral,*.

Measur-ement,s of the rncomplete flgurecl spe,cimen: height 56.1 mm, dianteterl 11.0 rnm.Occtlr'i.gi7.6s.-Station Sli2_i2 r)Ìl Santo, Nerv Heb_t'icles: age, Pleistocene. The .species lives toclay inser.eral parLs of Indonesia, ancl a vartety tvas re_

î:ll1 by.Marrin (1883, p. 2r7) from the upper-!ttocene of Javtì.

Terebra (Myurella) pretiosa Recveplate 30, figures 1?, 1g

7't're lt'ra pretíoso Reeve. 1g42, Zool. Soc. Lontlon proc., p. 200;Reeve, 1860, Conchologica Iconica, v. 12, Tr:rcbra, pl.8. fig. .10; Tryon, 1gg5, ]lanual conchology, v.7, p.27,pl. 8, fig 33; llacl,icil. 1960, U.S. Geoi. Surl_gy p1ei.,Paper .3iì9, p. 12i, pl. 10, fig. 18.-'lt1r'i'a'i111

p.i-etíosu.s (Reeve), Kira. 1g62, Sheils of the rvesternPacific in color, v. 1, p. 110, pl. J'S, iig. S.

Shell large rvith rnanrv g:ently convex and slighilylulretecl w,horls. Sculpture consisting of a clistinctlight-coloreci subsutural banci ancl close_set curvedaxial ricìges. Belorv the sub-qutural band, fìne spirals,some of rvhich overricle the axials, give rise to alatticecl appearance. Reclclish_brown" axial bandscover much of the area below the subsutural banclon one shell; on the other shell, cliscontinuouspatches of brorvn occur along axial lines.

Measurements of the figureci specimen: height116 mm, diameter 15 mm.

Ocan.r't:nr:e..-T\\,o sllecimens fr.om station SMZ.12,S:rnto, Nelv Hebricles ; àge, pleistocene. MacNeil(1960) reporteci on shells from the upper .Iertiarirof Okinau'a. Described originally lrom Chine,se,,r'aters; founcl aìso near Japan at depths of 50_g0I:Ì tnom.s.

Enirvetok, at depthìate Miocene (Ter.-

Terebra (Myurella) alveolata Hindsplaie ilO, fi.gures 1g, 20

Ttrcbra a!ueo[atu Hínrls, 1g.11], Zool. Soc. Lonclon proc., p.151 ; Reevc, lg(i0, Conchologica Iconic a, v. 12, Terebra,pl. 19, fig. 89; Tryon, 1gg5, lfanual Conchology, v. T,p. tj. pl. 5, fig. s7.

Ilutt.stttlopsís ulut,olcttrt ( Ilintls), Habe. 1g6.1, Shells ,rf thewestern P:rcific in cr;lor, v.2, p. 12g, pl..1l, fig. 1g.

Sm:rll, u'horls near.l1. flat, slighily turretecl to pro_cìuce an impressecl ..uture. 1 moderately rleep

groove on the upper parf of each $'horl sets off asubsulural band, ancl there are trvo or more incon-spicuous spiral grooves belou'; rvhorls covered byclose-set, curved, roundecl axial ribs. The suturalband carries a series of regularly spaced brownspots, and a second series is present on the bodyu'horl near the base. The fossil sheils retain cleartraces of this unusual color pattern.

Measurements of the figured specimen, USNM1?5082 from stalion SM242 on the Kere River,Santo, Nerv Hebrides: hei.sht 52.5 mm, diameter 9.0

mm.Ocr:rrt't'cn.ce.-Nine specimens from Pleistocene

beds on the I{ere River, Santo, Neu' Hebrides. Thespecie.s lives today in Indonesia anci Japan.

Subgenus TRIPLOSTEPHANUS Datl

Dall. 1ft0E, Nautilus. v.2L. p. 125-125.

Tupc (b,v original designatiott').-Terebra trí-seriata Gray. Holocene, Pacifrc.

Terebra (Triplostephanus) triseriata Gray

Pìate 30. fisure 21

7-t'rtln'u tt istt'iuta Grar', 1834, Zool. Soc. Lcndon Proc., p- 62;

Ccrnohorskl',1967. Ilarinc sbclls of the Pacific, v' 1,

p. 206, pl. ir2. fig. 389-Triplost,'phanus triscriata (Gra-v), Kira, 1962, Shells of the

weslern Pacìfic in color, v.1, p.110, p1.39' fig.9.T ercbro (T r ip! <tst.cplt o ntts) trist r ict lo ( Gral' ) . Cer-nohorskl',

1{ì66, \-elig-er, v.9, p.57, pÌ.4. fig. 14.

Shell long and ver)' nalrl'o\l-' compo-sed of manyu'horis (20 on the incomplete fossil segment).Scuìpfurecl uniformll', each g"horl having three con-spicuousl)' beaded spiral ccrds ; col:d immediateiybelor,r'the suture is the largest, the cord immediateiybeìos' it í.r nexf in size, and the one immerliatelvabove the suture is the smallest; the lou- zrea oc-

rrrnr.'inq thp middle of each rvhorl bears four beadedLsy.r arrò

secondar)' spiral corrls, the lorvest -"lightly largerthan the other three. Apex and aperture notpreserrrerì.

I'feasr-iler-r.ients of the flgur"ecl --"pecimen : height(incomplete) 34.3 mm, dian.ieter 4.1 mm.

An examination of Holocene shells shorvs that thegpiral cords vary con-siderabli' in number and rela-tive size. Tu'o other fossils from SI 242 exceed thevariabiìit1'shos'n b1- most Ilolorene shells. Thel' ai"e

less slender than the t1'pìcaì sperilì'ìen alleadl- de-

scriÌ,red bul are assigned to T. lii;riíata' One is

nearl.u- c:ompleie, consisting of a pt"oiocotlch of aboutfir'e glass5' rvhorls follorvecl bi' 24 scuìplured u-horlsthal have a. large stìbsutural cord, ctlt.trparativell'sri;rll coi'ds above attcì beìou', pìus tt-o or ihl'ee stillsnr:'llel cords in the cenier of each llhot']. The thil"d

LOCALITIES 85

specimen, not figuled, is likewise dominated by a

sincle subsutural cord.Or:ctrt't en.ct:.-station 5M242 on Santo, Nerv

Hebridès; age, Pleistocene' Living exampies have

been collected u'idely ín the tropical Pacific.

LOCALITI[,S

Some of the fossils described or cited in the pres-

ent. report $,ere collectecl from localities (see fig. 2)not listecl or sho\l'n on maps in the previous reporLs

Lacìd 1966, 1972,1977). A few fossils t'ere obtainedflom Rotuma and islands nol previously report'edupon. The lcicaiity dafa given belorv suppÌementthose given pi-eviouslv by Ladd (1966, p. 81-89;1972, p. 63-69; 7977, p. 68-73). Ontline maps ofmost of the islands u'ere rlrovided in the reports justcited.

I aland.

U.S. GcolosicalSurt:cu Ccnozoic

lot:ality orSldtiml

Localittt ortd collcctq

Guanr

llariana Islands

------About 2 miles (3 km) N\Wof Fadian Point, rr,'aisi ofjsland. Pacific Island Engi-neels.

Do ,------20997 ----- Cascajo pit near Agat. close

to west coast of islani- H. T'Sleaitrs,

Nerr Hebrides

Santo --------Slf 298 -----Lat 15'33.81' S., long 166-'-16.7'l'

E., altitude 80 rn 1260 ft).D. I. J. trlallick.

Do - -----,-Sl{307 --,--trIiddle Pakalolo River. a iribu-far] to ti'c Wambu. D- I' J.IlaÌlick.

Do -,, *,--SM308

Do ---------SM309

----- Upper \À-ambu River nearSM309. D. I. J. Maliick-

--,-- South bank of Wambu Rirerand as much as 50 n {165ft) dorvnstream frorn r:outhof Pakalolo. a tribr:ta:]--D. L J. Ifallick'pper \\:anbu River neelSM309. D. I. J. Ifallich-

------Lat 15'26'30" S., ìong167'05'48" E., aiiitud: 55 m(t 80 ft).

---- Sarahala Rive-r debris-

-- Lat 15'26'40" S., long16?'5'10" E., altitude :5 m(180 ft).

----- Lat 15'25'30" S., long -ri;'56'E., altitude 360 nl í1.:S0 ft)David Greenbaum'

Do --- --- -SG78

I-t o

Do

SG;9

- __SGE2

L)o , - SG9.1

-tt lcl:.; F 1. Terebt'a (Stríoterrbt u.rn) tot-qtLatoHeìght (incomplete) lì5.? mm (

T erebra \Stt-io tere brunt) nebuloso,H:ight ( apex incomplete) 50.9

308 151.

PLATE 30

Adams and Reeve (p. 81).X 1). Station Sil12{2, Santo,Sowertry (p. 82).

ò,

Nerv Hebricles. Pleistocene. USNùÍ 808150.

mm (X 1). Station SN{242. Santo. Nerl,

'I'ercbrt {StriotcrebrtLrz) sp. A (p. g2).Height (incoùrpiebe) ii3'3 mm (x 2). D|iii hole F-1. Enìrvetol<, depth 6b0-tju0 ft (210 nr). Late Miocene

( Tertiai'y g) . USNII 175084.Tertbra 1De corihastttla) u./ùnis Glay (p. g2).

Height 2'3'3 mm (X 3). USGS locality 17891, Szripan. -lanapag Lims,stone, pleistocene and Holocene.usNM 175081.Ttrebra (JIicrotryp:;tes) ranuale,-ezsrs Laclcl, n. sp. (p. g2).

6' Holotype. Height (incompletè) 15.6 mm (x 21,i). Station 817, Vanua Levu, Fiji. pliocene (Tertiarlr/,). LrsN}{ l7;08r,.

Hebricles. Pleistocene. USNIÍ Z0g7 40.

IHebrides. Pleistocene. USNM

Hebriries. Pleistocene. USNM 1?b0g2.

Santo, Nerv Hebrides. pleistocene. USNM B0g74I.

| ,,;.

f 6,7-

as holotl-pe. USNII 175086.

. Tanapag Limsstone, pleistocene ancl Holocene.

9. Te rebra (O:L:unerts) int,rliil,:ata Desiral.s5 ip. g:l).Height ó{i.0 mm ( )< l). StaLion S}I212. Santo, Ne$-Hebrides. Pleistocene. USNII g0g152.

t0-12. Terebra (O.tymeris) cf.7'. ír:lina (Diihv5.n) (p. g:ì).10' Height (incomplete) 110 mm (x t). Station sM242, Santo, New Hebrides. pleistocene. usÀIlI iog158.11' 12' fwo fra.glrentarv fossils from the late llioc,ne of Viti Levu, Fiji (station ltib) ; larger is 15.6mm in height, smaller 15.g mm (both x 1r/2, rJiNM 1750g? ancl 17.r0gg).13. 7'erebra (OtynLet.ís) argrrs (Hintls) (p. g.1).Height (apex incomplete) 32..r mm (X 2). usGS Ìocality 17tg1, Saipan. Tanapag Limeston:, pleisto-

cene and Holocene. USNIÍ 1?5093.l-1. Torehru (Otyme.rís) sp. B (p. 84).

Height 34.4 mm (x z;. Drill hole F-1, Eniwetok, cìepth E00-E10 ft (J-t; m). Lare Miocene (Tertiarl-r). usNùI 175089.7'erebru (Tcrebra) sttbulata (Linnaeus) (p. g1).

Height 82.5 mm (x t). Station SM242, Santo, New Hebricles. Pleistocene. USNM 30g149.Tet'ehra (ùIyu.rella) rn.lJtL.t.os Lamarck (p. 34).Height i6.1 mm. (X 1). Station SM24Z, Santo, Ne*, Hebi.icles. Pleistocene. USNM 3081b4.

1?, 18. Terehrrt (1'[yurella) .pretiosa Reeve (p. g4).Height lt6 mm (X 1). StaLion SM24Z, Santo. Ne;v

19,20. Terr:bra (JIuurella) al.t-eolata Hinrls (p. 8.1).

8.

7. Paratype. He,ight 1-1.9 mm. (X 21:). Same localit1.Te rebra (Otyneris) tttaculatu (Linnaeus) (p. BJ).

Height 165 mm ()( 1). USGS locality 1?g91, Sa panusNlI 1130i9.

Height 52.5 mm (X 1). Station SM242. Santo. New2L. 7'r:r'cbro (TriplsslpOTnrits) ii.isrnafo Gray (p. g;).

Height ( inconrplete) 3.1.3 mm ( >< 2). Station SMJ.12,

I

I

iII

i!

t'

tjla.

lo.

{t\

d

{

i

ij

i;ì1Ol.o( ;l( .{ J. Sl'lì\'!llPII(u-'t:sst()NAt, I,AI,t-)i lt;t i,t..1Tt_ jtì

.:.tij..!!i1

jr{"-tr"'I

Ì{i' 'liil

{'1+1. r.Ir*?a:.,*

l, tl- {-b.'u'^.ti .:"';ii

".: I

.:jet"'

i,!

,'#

G't7'*,{

"*r,$

ú,rji$

&lt'.?-t_. "ú. \€s

€Tig

:.

.#*

É*Bt:'q

;irt,,i tii

:t.:

%L

-ì{{{.,'Nl*

,*',-d,tffis&iil-Xtry

11

,r_,,,.

Ttr}IEIJR]DAE

&' ,o

' 'i.

I4

Fî; Ig :

,, 't,. .î,1 .ú

ya';; jr'ljÉttilt

f r-{g Y' 'tE'':

':d Àt;.l

lrti '/.l/':t

'EtL rl''a '

*É !r$'4ér -'\ ]RE

- ln

,.ti

.iì?t-;it

I-t;A

tt *;..i.4

.*'i*#rc

jì -.t-..t*:.:' ,:.,

----<a:

'"".t-

l- I

I