A catalog of Nearctic Chironomidaeesc-sec.ca/wp/wp-content/uploads/2017/03/AAFC_catalog_of_nearcti… · Nanocladius Kieffer, 1913d: 31. Type-species, vitellinus Kieffer (orig. des.)

AgricultureCanadaI*l

A catalog ofNearcticChironomidae

A catalog of Catalogue des

Nearctic ChironomidaeChironomidae delardgion

ndarctiqueD.R. Oliver and M.E. Dillon D.R. Oliver et M.E. DillonBiosystematics Research Centre Centre de recherches biosyst6matiquesOttawa, Ontario Ottawa (Ontario)K1A 0C6 K1A 0C6and etP.S. Cranston P.S. CranstonCommonwealth Scientific and Organisation de la recherche

Industrial Research scientifique et industrielle duOrganization, Entomology Commonwealth, Entomologie

Canberra ACT 2601 Canberra ACT 2601Australia Australie

Research BranchAgriculture Canada

Publication 1857181 990

Direction g6n6rale de la rechercheAgriculture Canada

Publication 1857181 990

@Minister of Supply and Services Canada 1990

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Oliver, D.R.

A mtalog of Nearctic Chironomidae

(Publication ; 1857/8)

Text in English and French-Includes bibliographiel referenes.Issued by Research Branch, Agriculture Canada.Cat No. A43-1857/19908ISBN 0-660-55839-4

1. Chironomidae--Ameria.2. Chironomidae--Nomenclature. I. Dillon, M.EII. Cranston, P.S. III. Canada. AgricultureCanada. IV. Title. V. Title: Catalogue desChironomidae de la r6gion n6arctique. VI.Title: Nearctic Chironomidae. VII. Series:PubUcation (Canada. Agriculture Canada).English ; 1857,18.

QL537.C45604 1990 595.77'7 C90-099ttt-98

CoverAdult male of Chirnmw pfumm (Linnaeus)

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Oliver, D.R.

A atalog of Nearctic Chironomidae

(Publiation ; 18578)

Texle en anglais et en frangais.Comprend des r6f6rences bibliogr.Publi6 par Direction g6n6rale de la recherche,Agriculture CanadaN' de cal A43-1857/1990FISBN 0-660-55839-4

1. Chironomidae--Am6rique. 2. Chironomidae--Nonenclature. I. Dillon, M.E. II. Cranston,P.S. III. Canada. Agriculture Canada. IV.Titre. V. Titre: C:talogue des Chironomidae dela r6gion n6arctique. VI. Titre: Chironomidaede la rEgion n6arctique. VII. Clllection:Publiation (Canada. Agriculture Canada).Franqais ; 1857,8.

QL537.C45604 1990 595.77',1 C90-099111-9F

CouvertureM6le adulte du Chironomw plumm (Linnaeus)

Production du Seruie aux programmes derecherches

TABLE OF CONTENTS

INTRODUCTION.. ....1

FORMAT ,,...1

ABBREVIATIONS .....2

ACKNOWLEDGMENTS .......3

FamilyCHIRONOMIDAE .......7

Subfamily TELMATOGETONINAE . . . . . . . . . 7

SubfamilyPODONOMINAE ....... 7

TTibeBOREOCHLINI ........ 7

TTibePODONOMINI ......... 8

SubfamilyTANYPODINAE ........ 8

Tribe COELOTANYPODINI . . .. 8

TTibeMACROPELOPIINI ...... 8

TTibeNATARSIINI ...10

TTibePENTANEURINI ....... 10

TTibePROCLADIINI ........ 74

Tribe TANYPODINI .. ....... 16

SubfamilyDIAMESINAE ...16

Tribe BOREOHEPTAGYIINL . . . . . . . .'16

TTibeDIAMESINI .. ........ 16

TTibePROTANYPINI . ....... 19

SubfamiIyPRODLAMESINAE ..... 19

SubfamilyORTIIOCLADIINAE . ... 20

SubfamilyCHIRONOMINAE ... ... 40

TTibeCHIRONOMINI ....... 40

Tribe PSEUDOCHIRONOMINI . . . . . . . 55

TTibeTANYIARSINI .. ...... 55

REFERENCES ,.., ... 60

rNDEX .... 76

TABLE DES MATIERES

INTRODUCTION . . ....4

MODEDEPRESENTATION .. ........ 4

ABREVIATIONS ....,.5

REMERCIEMENTS ....6

Famille des CHIRONOMIDAE ......... 7

Sous-famille des TELMATOGETONINAE . . . 7

Sous-famille des PODONOMINAE . . . . . . . . 7

Tribu des BOREOCHLINI .. . . ...... 7

Tribu desPODONOMINI ........... 8

Sous-famille des TANYPODINAE . . . . . . . . 8

Tribu des COELOTANYPODINI . . . . . . 8

Tribu des MACROPELOPIINI . . . . . . . . 8

TTibUdesNATARSIINI ..... 10

Tribu des PENTANEURINI . . . . . . . . . 10

Tribu des PROCLADIINI . . . . .. .... 14

Tribu desTANYPODINI ..... 16

Sous-famille des DIAMESINAE . . . . . . . . . 16

Tribu des BOREOHEPTAGYIINI . . . . . 16

Tribu desDIAMESINI ...... 16

Tribu des PROTANYPINI . . . . . . . .. . 19

Sous-famille des PRODIAMESINAE . . . . . . 19

Sous-famille des ORTHOCLADIINAE . . . . . 20

Sous-famille des CHIRONOMINAE . . . . . . 40

Tribu des CHIRONOMINI . . . . . .... 40

Tribu des PSEUDOCHIRONOMINL . . 55

Tribu des TANYTARSINI . . . . . . . . . . 55

BIBLIOGRAPHIE .. ... 60

INDEX ... 76

INTRODUCTIONAdults of the family Chironomidae are

commonly called chironomids, midges,nonbiting midges, or chironomid midges, andthe red larvae of some groups are calledbloodworms. Chironomid larvae occur in alltypes of aquatic and semiaquatic habitats andto a lesser extent in brackish and marinewaters, and semiterrestrial and terrestrialhabitats. Aquatic ecosystems are usuallydominated by larval chironomids, in terms ofnumber of species, biomass, or both. Theyare important in the breakdown of organicmaterial and recycling nutrients, as food forfish, birds, and both terrestrial and aquaticinvertebrates, and as predators of aquaticinvertebrates. Some species can tolerate themost polluted conditions, whereas others havevery special requirements and quicklydisappear when their habitat is stressed.Thus the Chironomidae have exceptionalpotential to act as biological indicators ofaquatic environmental health.

The family is distributioned around theworld, occurring in all zoogeographicalregions, including Antarctica. Currently 10subfamilies and 24 tribes (Ashe 1983) arerecognized, of which 7 subfamilies and 14tribes occur in the Nearctic region. Aminimum of 5000 species are found aroundthe world and at least 2000 of these in theNearctic region.

The nonspecialist was faced, until recently,with two generic classifications. This dualsystem arose during the first half of thiscentury, because one group of workersprincipally emphasized the immature stagesand the other emphasized adults. Pinder(1983) gives a brief history of thedevelopment of the two systems and theirreconciliation. Reconciliation of the twosystems has culminated in the classificationused in the three volumes of theChironomidae of the Holarctic region editedby Wiederholm (1983, 1986, 1989) and thatclassification is used herein.

The last catalog of Nearctic Chironomidae(Sublette and Sublette 1965) was publishedabout 25 years ago. The intervening yearshas been a period of intense chironomidresearch, including much taxonomic andecological study. The following table showsthe increase in the number of valid generic-and species-level names between 1965 and1989.

Subtette 0[ iver, Di [ [onand and

Subtette Cranston't965 1990

G ene Fa 205

Spec i es 601 1 051

The need for an updated catalog of NearcticChironomidae is evident.

Accordingly we have prepared a catalog ofthe taxa of Chironomidae recorded from theNearctic region, with information on theirdistribution and revelent literature. ThiScatalog is based only on published data to theend of 1989; it does not include unpublisheddata in collections, such as the CanadianNational Collection of Insects. We haveattempted to list all published names, valid ornot, that have been used in the literaturepertaining to the Nearctic region. A fewolder faunal lists (Leonard 1928; Johnson1925; Brimley 1938; Petch and Maltais 1932)are not included because they contain manyobviously incorrect determinations. As thetaxa are presented uncritically in their mostrecently published taxonomic position, itshould not be construed that we are inagreement with all the taxonomic placements.

This catalog will be continuously updatedwith periodic publication of additions andcorrections. To assist us in keeping anupdated list, we request any errors oromissions, and copies of publications withinformation on taxonomy and distribution ofNearctic Chironomidae. be sent to either ofthe first two authors.

FORMATThe arrangement of names is alphabetical,

except for the subfamily-group names whichfollow the phylogenetic arrangement of theHolarctic Chironomidae volumes. Species-group names follow the subfamily, tribe,genus, and subgenus headings (not all groupshave been assigned tribes or subgenera). Thefirst occurrence of all valid names are printedin boldface type along the left margin of eachcolumn. Valid taxa that have beenincorrectly reported from the Nearctic regionare also listed along the left margin but theyare printed in italics as are all other listing ofnames, valid or invalid.

Citations for genus-group names arearranged as in the following example.

NEARCTIC CHIRONOMIDAE

Genus NANOCLADIUS KiefferNanocladius Kieffer, 1913d: 31.

Type-species, vitellinus Kieffer(orig. des.).

Microcric otopas Thienemann &Ilarnisch, 1.932: 1.37.Type-species, Cricotopusal b icornis Goetghebuer(Goetghebuer in Goetghebuer &Lenz, 1944: 114) = Chironomusbicolor Zetterstedt.

Subgenus Nanocladius Kieffer

The original citation is given in the secondline, indented, followed by synonyms (if any),also indented. The genus name is followedby the original author, date, page, name oftype-species, and nature of type fixation. Theoriginat type-species combination is cited, butthe generic name is omitted if it is the sameas the genus for which it is the type. If theoriginal type-species is now a junior synonym,it is given at the end following an equalsymbol (=). Only the authors of subsequenttype fixation are cited. Much of theinformation about type fixation and synonymshas been obtained from Ashe (1983). Thispublication has detailed information ongeneric synonymy, type-species, andnomenclatorial problems. Subgeneric-namecitation format follows that of the genericcitations. Citations for nominate subgeneraare omitted when they are the same as thegeneric citations; otherwise they follow theformat of generic citations.

Species-group names are arranged asfollows:

triannulatus. Sask., Ont. to N.Y., s. toKy. and Del.; PA.

Chironomus triannulatus Macquart,7826: 202.-Germany.

Cricotopus exjlis Johannsen, 1905:25.5.-N.Y.

The species name is followed by a synopticaccount of the Nearctic region distributionstated by territory, province, or state in anorthwest to northeast and southwest tosoutheast pattern, except for Greenland.Occurrence in other zoogeographical regionsare listed next, separated by a semicolon.When a Mexican locality record occurs nearthe uncertain division between Nearctic andNeotropical regions, it is listed as occurring inboth Mexico and the Neotropical region. The

original binomial citation is next, followed bysubspecies or varieties and then synonyms (ifany), which are cited in their original formwith author, date, page, and type-locatity.Type-localities for Canada and the UnitedStates are listed by province, territory orstate; all others are listed by country orlarger geographical regions where country isnot known.

ABBREVIATIONSThe abbreviations used are as follows:

General&- ando., E.- east, easternemend.- emendedI., Is.- Island, IslandsICZN- International Commission

on Zoological Nomenclaturemisident.- misidentificationmon.- monotypynam.- namen. nam.- new namenom. nud.- nomen nudumorig. des.- original designationpreocc.- preoccupieds., S.- south, southernse., Se.- southeast, southeasternsub. mon.- subsequent monotypysubg.- subgenussyn.- synonym, synonymyvar.- varietyw.- west, western

Geographical NamesAF.- Afrotropical RegionAla.- AlabamaAlas.- AlaskaAlta.- AlbertaAmer.- AmericaAriz.- ArizonaArk.- ArkansasAU.- Australasian RegionBaja Calif.- Baja CaliforniaB.C.- British ColumbiaCalif.- CaliforniaCan.- CanadaColo.- ColoradoConn.- ConnecticutD.C.- District of ColumbiaDel.- DelawareFla.- FloridaGa.- GeorgiaGrld.- GreenlandIll.- IllinoisInd.- IndianaKans.- KansasKy.- Kentucky

CHIRONOMIDAE DE LA REGION NEARCNQUE

Ky.- KentuckyLa.- LouisianaLabr.- LabradorMan.- ManitobaMass.- MassachusettsMd.- MarylandMex.- MexicoMich.- MichiganMinn.- MinnesotaMiss.- MississippiMo.- MissouriMont.- MontanaN.B.- New BrunswickN.C.- North CarolinaN. Dak.- North DakotaNE.- Nearctic RegionNebr.- NebraskaNev.- NevadaNfld.- NewfoundlandN.H.- New HampshireN.J.- New JerseyN. Mex.- New MexicoN.S.- Nova ScotiaNT.- Neotropical RegionN.W.T.- Northwest TerritoriesN.Y.- New YorkOkla.- OklahomaOnt.- OntarioOR.- Oriental RegionOreg.- OregonPA.- Palaearctic RegionPa.- PennsylvaniaP.E.L- Prince Edward IslandQu€.- QudbecR.I.- Rhode IslandSask.- SaskatchewanS.C.- South CarolinaS. Dak.- South DakotaTenn.- TennesseeTex.- TexasU.S.A.- United States of AmericaU.S.S.R.- Union of Soviet Socialist

RepublicsVa.- VirginiaVt.- VermontWash.- WashingtonWis.- WisconsinW. Va.- West VirginiaWyo.- WyomingY.T.- Yukon Territory

ACKNOWLEDGMENTSWe gratefully acknowledge Mr. B.

Caldwell for checking several drafts. The lateDr. Sam Roback checked an early draft ofthe subfamily Tanypodinae. We thank Mr. L.Speers for formatting the manuscript forprinting, and Ms. L. Robillard for typingsome of the references.


INTRODUCTIONLes adultes de la famille des chironomidds

sont communement appeles chironomes oumoucherons et les larves d'un rouge sang decertains groupes sont les vers de vase utilisdspar les pOcheurs. On trouve des larves dechironomidds dans tous les types d'habitatsaquatiques et semi-aquatiques et, dans unemoindre mesure, dans les eaux saumAtres oumarines de m€me que dans les habitats semr-terrestres et terrestres. Les 6cosystdmesaquatiques sont habituellement domin6s parles formes larvaires, en ce qui touche lenombre d'espdces, la biomasse ou les deux.Ces insectes jouent un r0le important dans lad6gradation de la matidre organique et lerecyclage des 6l6ments nutritifs ou commenourriture pour les poissons, les oiseaux ainsique les invertdbr6s tant terrestresqu'aquatiques ou encore comme pr€dateursdes invertdbrds aquatiques. Certaines espdcespeuvent toldrer des conditions de pollutionextreme, alors que d'autres, trds sp6cialisdes,disparaissent rapidement quand leur habitatest menac6. Les chironomidds possddentdonc un potentiel exceptionnel commeindicateurs biologiques de la santd du milieuaquatlque.

Cette famille, de rdpartition mondiale, estprdsente dans toutes les rdgionszoog6ographiques, y compris l'Antarctique.On reconnait actuellement 10 sous-familles et24 tribus (Ashe 1983), dont 7 sous-familles et14 tribus dans la rdgion nearctique. Laplanete compte un minimum de5 000 espOces dont au moins 2 000 dans cetterdgion-ld.

Jusqu'A recemment, le profane seretrouvait devant deux classificationsgdndriques. Ce double systdme a pris formeau cours de la premidre moitid du sidcle, ungroupe de chercheurs ayant mis I'accent surles stades immatures et I'autre sur lesadultes. Pinder (1983) trace un brefhistorique du ddveloppement des deuxsystemes et de leur rapprochement. Lareconciliation des deux systemes a atteint unpoint culminant dans la classification utilis€edans les trois volumes de Chironomidae ofthe Holarctic region 6,dit€ par Wiederholm(1983, 1986, 1989), et c'est cette classificationque I'on utilise dans le pr6sent ouvrage.

Le dernier inventaire des chironomid6snearctiques (Sublette et Sublette 1965) a 6t6publid il y environ 25 ans. Les anndes quisuivirent furent une pdriode de recherchesintenses sur les chironomid6s, surtout leurtaxinomie et leur €coloeie. Le tableau

suivant montre la multiplication des nomsvalide au niveau des genres et des espCcesentre 1965 et 1989.

Subtette 0[iver, Dittonet et

Subtette Cranston1965 1990

Genres 205

EspCces 601 1 051

La n€cessitd d'un inventaire i jour deschironomid6s est dvidente.

C'est pourquoi nous avons dressd unrdpertoire des taxons des chironomid6s dontla prdsence a 6t6 signal6e dans la rdgionndarctique, assorti d'informations sur leurrepartition et la documentation pertinente.Ce catalogue n'est fond6 que sur des donndespublides jusqu'd la fin de 1989 et necomprend pas les donndes non publides quel'on trouve dans les collections commel'Insectarium national canadien. Nous avonsessayd d'€num6rer tous les noms publies,valides ou pas, qui ont ete mentionn6s dansla documentation relative d la rdgionndarctique. Quelques vieilles listes fauniques(Leonard 1928; Johnson 1925; Brimely 1938;Petch et Maltais 1932) n'ont pas 6td prises enconsideration parce qu'elles contiennent denombreuses appellations qui sont hors de toutdoute incorrectes. Comme les taxons sanscommentaires dans leur dernidre positiontaxinomique publi6e, il ne faudrait pas enconclure que nous y souscrivons dans tous lescas.

Le pr6sent inventaire sera continuellementtenu d jour par la publication pdriodiqued'ajouts et de corrections. Pour nous aiderdans cette teche, nous vous saurions grd defaire part d l'un ou I'autre des deux premiersauteurs nomm6s des erreurs ou des omissionsqui auraient pu s'y glisser et de leur envoyerdes exemplaires de publications contenant deI'information sur la taxinomie et la 16oartitiondes chironomidds.

MODE DE PRESENTATIONLes noms apparaissent en ordre

alphab€tique, sauf que ceux des groupes desous-familles suivent I'organisationphylog6nique des volumes sur leschironomid€s holarctiques. Les noms desgroupes d'espdces suivent les titres des sous-familles, tribus, genres et sous-genres (a

CHIRONOMIDAE DE I-A REGION NEARCNQUE

noter que ce ne sont pas tous les groupes quise sont fait assigner des tribus et des sous-genres). La premidre fois qu'un nom valideapparait, il est imprim6 en caractires grasprds de la marge de gauche de chaquecolonne. Les taxons valides qui ont 6t€signalds a tort comme de la rdgion n6arctiquesont aussi €numdr€s le long de la marge degauche, mais en italique, comme le sontd'ailleurs toutes les autres listes de noms,valides ou non.

Les citations pour les noms de groupes degenres se pr€sentent comme suit :

Genus NANOCLADIUS KiefferNanocladius Kieffer. 1913d: 31.

Type-species, vitellinus Kieffer(orig. des.).

Microcricotopas Thienemann &Harnisch, 7932: 737.Type-species, Cricotopusalbicornis Goetghebuer(Goetghebuer in Goetghebuer &Lenz,1944; 114) = Chironomusbicolor Zetterstedt.

Subgenus Nanocladius Kieffer

La citation originale se trouve sur ladeuxi0me ligne, en retrait, suivie dessynonymes (s'il y a lieu), eux aussi en retrait.Le nom du genre est suivi du nom deI'auteur original, de la date, de la page, dunom de I'espdce type et de la nature de lafixation du type. L'esp0ce type originale estmentionnee, mais le genre est omis s'il estidentique d celui dont elle est le type. Si lenom de l'espOce type originale est par contreun synonyme mineur, il apparait i la finaprds un symbole d'6galit6 (=). Seuls lesauteurs de fixation de type ultdrieure sontmentionnds. Une grande partie desinformations sur la fixation des types et lessynonymes sont tirdes de Ashe (1983). Cettepublication contient des renseignementsd6taill6s sur la synonymie g6n6rique, lesespoces types et les probldmes denomenclature. Les citations des noms dessous-genres suivent celles des genres, sauf s'ils'agit de rdpdtition (dans ce cas, elles sontalors omises).

Les noms des groupes d'espdces sepresentent comme suit :

triannulatus. Sask., Ont. to N.Y., s. toKy. and Del.; PA.

Chironomus ffiannulatus Macquart,1826: 202.-Germany.

Cricotopus exilis Johannsen, 1905:255.-N.Y.

Le nom de I'espdce est suivi d'un compterendu synoptique de la rdpartition dans lardgion ndarctique indiqude par le territoire, laprovince ou I'Etat, dans le sens nord-ouest anord-est et sud-ouest d sud-est, sauf pour leGroenland. L'apparition de l'espdce dansd'autres rdgions zoog€,ographiques est6num€r6e ensuite et sdpar6e par un point-virgule. Quand l'apparition d'une espdce estindiqude dans une r6gion du Mexique situdeprds de la division impr6cise entre les rdgionsn€arctique et n€otropicale, elle est indiqudecomme observable tant au Mexique que dansla rdgion n6otropicale. La citation binomialeoriginale apparait ensuite, suivie des sous-especes ou varidtds, puis des synonymes (s'il ya lieu), qui sont mentionnds sous leur formeoriginale accompagn€s du nom de I'auteur, dela date, de la page et de l'emplacement type.Les emplacements types pour le Canada etles Etats-Unis sont indiquds selon lesfrontidres des provinces, des territoires oudes Etats; tous les autres suivent lesfrontidres des pays ou encore des grandes169ions g6ographiques si celui-ci n'est pasconnu.

ABREVIATIONSLes abrdviations utilis€es sont les

suivantes :G6n6ralitds

&- ete., E.- est, de l'est, orientaleemend.- corrigdI., Is.- ile, ilesICZN.- Commission internationale de

nomenclature zoologiquemisident.- erreur d'identificationmon.- monotypienam.- nomn. nam.- nouveau nomnom. nud.- nom seulorig. des.- d€signation originalepreocc.- d6jd appliquds., S.- sud, du sud, mdridionalse., Se.- sud-est, du sud-estsub. mon.- monotypie ulterieuresubg.- sous-genresyn.- synonyme, synonymievar.- varidtdw.- ouest, de I'ouest, occidental


Noms gdographiquesAF.- r€gion afrotropicaleAla.- AlabamaAlas.- AlaskaAlta.- AlbertaAmer.- AmdriqueAriz.- ArizonaArk.- ArkansasAU.- r6gion australasienneBaja Calif.- Basse-CalifornieB.C.- Colombie-BritanniqueCalif.- CalifornieCan.- CanadaColo.- ColoradoConn.- ConnecticutD.C.- District de ColumbiaDel.- DelawareFla.- FlorideGa.- GdorgieGrld.- GroenlandIll.- IllinoisInd.- IndianaKans.- KansasKy.- KentuckyLa.- LouisianeLabr.- LabradorMan.- ManitobaMass.- MassachusettsMd.- MarylandMex.- MexiqueMich.- MichiganMinn.- MinnesotaMiss.- MississippiMo.- MissouriMont.- MontanaN.B.- Nouveau-BrunswickN.C.- Caroline du NordN. Dak.- Dakota du NordNE.- r€gion ndarctiqueNebr.- NebraskaNev.- NevadaNfld.- Terre-NeuveN.H.- New HampshireN.J.- New JerseyN. Mex.- Nouveau-MexiqueN.S.- Nouvelle-EcosseNT.- rdgion n€otropicaleN.W.T.- Territoires du Nord-

OuestN.Y.- New YorkOkla.- OklahomaOnt.- OntarioOR.- r6gion orientaleOreg.- OregonPA.- rdgion paldarctiquePa.- PennsylvanieP.E.I.- lle-du-Prince-EdouardQu6.- Qudbec

R.L- Rhode IslandSask.- SaskatchewanS.C.- Caroline du SudS. Dak.- Dakota du SudTenn.- TennesseeTex.- TexasU.S.A.- Etats-Unis d'Am6riqueU.S.S.R.- Union des Rdpubliques

Socialistes Sovi6tiquesVa.- VirginieVt.- VermontWash.- WashingtonWis.- WisconsinW. Va.- Virginie de I'OuestWyo.- WyomingY.T.- Territoire du Yukon

REMERCIEMENTSNous tenons d remercier M. B. Caldwell

d'avoir vdrifid plusieurs 6bauches. Feu D'Sam Roback a eu la bont6 de vdrifier une6bauche antdrieure de la sous-famille destanypodin6s. Nous remercions 6galementM. L. Speers d'avoir prdpard le manuscritpour I'impression et Mme L. Robitailled'avoir dactylographi6 une partie de labibliographie.

CHIRONOMIDAE DE LA REGION NEARCTIQUE

FAMILY CIIIRONOMIDAE

Subfamily TELMATOGETONINAE

Genus TELMATOGETON SchinerTelmatogeton Schiner, 1866: 931.

Type-species, sanctipauli Schiner (orig.des.).

Paraclunio Kieffer, 1911b: 103.Type-species, trilobatus Kieffer (orig.des.).

alaskensis. Alas. s. to Calif.; Mex.Telmatogeton alaskensis Coquillett, 1900:

395.-Alas.japonicus. Coastal, N.Y., s. to Miss.; PA.;

NT.; OR.; AU.Telmatogeton japonicus Tokunaga, 1933:

95.-Japan.macswaini. Calif.

Telmatogeton macswaini Wirth, 1949:170.-Calif.

spinosus. Calif.Paraclunio spinosus Hashimoto, 1.973:

266.-Calit.trilobatus. Oreg., Calif.

Paraclunio trilobatus Kieffer, 191 1b:105.-Calif.

Genus THALASSOMYA SchinerThalassomya Schiner, 1856: 219.

Type-species, frauenfeld i Schiner(mon.).

bureni. Ky., N.C., s. to Miss. & Fla.; NT.Thalassomya bureni Wirth, 1949:

167.-Fla.

Subfamily PODONOMINAE

Tribe BOREOCHLINI

Genus BOREOCHLUS EdwardsBoreochlus Edwards in Edwards &

Thienemann, t938: I52. Type-species,thienemanni Edwards (orig. des.).

gracilistylus. Alas., Alta., Wash., Wyo.Boreochlus gracilistylus Brundin, 1966:

304.-Alta.persimilis. Alas. to Qu€., Calif. to Va.

Trichotanypus persimilis Johannsen, 1926:99.-N.Y.

sinuaticornis. Wash., Mont., Calif.Boreochlus sinuaticornis Brundin, 1966:

30-5.-Calif.

Genus LASIODIAMESA KiefferLasiodiamesa Kieffer, I924b: 48 (as subg.

of Syndiamesa Kieffer, 1918a).Type-species, Syndiamesa gracilisKieffer (orig. des.).

arietina. Ont., Qu6., Wis., N.Y.Tanypus arietinus Coquillett, 1908:

144.-N.Y.brusti. N.W.T., Wyo., Minn.

Lasiodiamesa brusti Sather. 1969:15.-N.W.T.

rawsoni. Sask.Lasiodiamesa rawsoni Brundin, 1966:

325.-Sask.tenebrosa. Alas., N.W.T., B.C., Minn., N.H.

Tanypus tenebrosus Coquillett, 1905:66.-N.I{.

Linacerus piloala Garrett, 1925a: 9.-B.C.Lasiodiamesa nearctica Brundin, 1966:

326.-N.W.T.sphagnicola. Ont.; PA.

Isoplastus sphagnicola Kieffer, 7925c:16.-U.S.S.R.

Genus PARABOREOCHLUS ThienemannParaboreochlas Thienemann. 1939: 166.

Type-species, Tanypus minutissimusStrobl (orig. des.).

stahli. Pa., Kans.Paraboreochlus stahli Coffman in

Coffman, Ferrington & Seward, 1.988:192.-Pa.

Genus TRICIIOTANYPUS KiefferTrichotanypus Kieffer, 1906a:. 42.

Type-species, Tanypus posticalisLundbeck (mon.).

alaskensis. Alas.Trichotanypus alaskensis Brundin, 1966:

315.-Alas.foliaceus. Alas.

Trichotanypus foliaceus Wirth & Sublette,1970: 343.-Alas.

hanseni. Wyo.Trichotanypus hanseni Wirth & Sublette,

1970: 347.-Wyo.


mariae. Alas.Trichotanypus mariae Wirth & Sublette,

1970: 345.-Alas.posticalis. N.W.T., Grld.; PA.

Tanypus posticalis Lundbeck, 1898:295.-Grld.

Tribe PODONOMINI

Genus PAROCI{LUS EnderleinParochlus Enderlein, l9I2: 109.

Type-species, fuegianus Enderlein (orig.des.).

Paratanypus Garrett, 1925a: 8.Type-species kiefferi carrett (orig.des.).

kiefferi. Alas. to N.8., s. to Calif. & N. MexGrld.; PA.

Paratanypus kiefferi Garrett, 7925a:8.-B.C.

Tanypus tibialis Staeger, 1845: 354(preocc. Say, 1823).-Grld.

Podonomus peregrinus Edwards, 1929:296.-Scotland.

Subfamily TANYPODINAE

Tribe COELOTANYPODINI

Genus CLINOTANYPUS KiefferClinotanypus Kieffer, 1973c: 157.

Type-species, Procladius fuscosignatusKieffer (Sublette & Sublette, 1965:14-5).

Subgenus APONTEUS RobackAponteus Roback, I97I: 25. Type-species,

Clinotanypus sabensis Roback (orig.des.).

sabensis. Tex.Clinotanypus sabensis Roback, 1971:

25.-Tex.

Subgenus CLINOTANYPUS Kieffer

aureus. Kans., La., Fla.Clinotanypus aureus Roback, 1971:

25.-La.pinguis. Man., Ont., widespread U.S.A.

Tanypus pinguis Loew, 1861a: 308.-N.Y.Tanypus flavicinctus Loew, 1861a:

309.-Pa.Tanypus thoracicus Loew, 1866: 4.-D.C.

Procladius caliginosus Johannsen, 190-5:131.-N.Y.

planus. Fla.Clinotanypus planus Roback, 1971:

24.-Fla.wirthi. Fla.

Clinotanypus wirthi Roback, 7977:24.-Fla.

Genus COELOTANYPUS KiefferCoelotanypus Kieffer, l9L3c: t54.

Type-species, Tanypus humeralis Loew(orig. des.).

atus. Kans., La., Tex., Mex.; NT.Coelotanypus a/us Roback, 1977: 37.-Tex.

concinnus. Ilt., Neb. to Pa., s. to N. Mex. &Fla.

Tanypus concinnus Coquillett, 1895:308.-Fla.

Coelotanypus flavus Kieffer, 1923a:296.-Tex.

naelis. Tex.; NT.Coelotanypus naelis Roback, 1.963b:

170.-Surinam.scapularis. Ont., Ohio, e. U.S.A., s. to N.

Mex., Kans., Tex., Mex.; NT.Tanypus scapularis Loew, 1866: 2.-D.C.

tricolor. Kans., Ill., N. Mex. to N.C., s. toFla.

Tanypus tricolor Loew, 1861a: 309.-N.Y.

Tribe MACROPELOPIINI

Genus ALOTANYPUS RobackAlotanypus Roback, l97I: 95.

Type-species, Tanypus venustusCoquillett (orig. des.).

aris. N.H., W. Va., N.C., S.C., Ala., Ga.Alotanypus aris Roback, 7971: 97.-N.C.

venusta. Wash., Wyo., Utah, Calif. to N.Mex.

Tanypus venustus Coquillett, 1902b:91.-N. Mex.

Genus APSECTROTANYPUS FittkauApsectrotanypr.rs Fittkau, 7962: l4l.

Type-species, Tanypus trifascipennisZetterstedt (orig. des.).

algens. Alas., Alta., Wash., Oreg.Tanypus algens Coqttillett, 1902b:

90.-Alas.


florens. Alta., Man., Wash., Wyo., Calif., Genus FITTKAUIMYIA KarunakaranNev., Colo., Ariz. Fittkauimyia Karunakaran, 1969: 75.

Tanypus florens Johannsen, 1908: Type-species, disparipes (orig. des.).272.-Colo., Wash., N.Y. Parapelopia Roback, l97l: 91.

johnsoni. Sask., Qu6., N.H., N.J., N.C., S.C., Type-species, serta Roback (mon).Fla.

Tanypus johnsoni Coquillett, 1901b: serta. Fla.609.-N.J. Parapelopia serta Roback, t97l: 91.-Fla.

Genus BETHBILBECKIA Fittkau & Murray Genus MACROPELOPIA ThienemannBethbilbeckia Fittkau & Murray, 1988: Macropelopia Thienemann in Thienemann

253. Type-species, floridensis (orig. & Kieffer, 1916: 497. Type-species,des.). Isoplastus bimaculatus Kieffer (orig.

des.) = Tanypus nebulosus Meigen.floridensis. Fla.

Bethbilbeckia floridensis Fittkau & decedens. B.C. to Qu6. & R.I., Utah, Wyo.,Murray, 1988: 2-54.-Fla. Colo., Ky., N.C., s. to Miss. & Fla.

Tanypus decedens Walker, 7848: 22.-Ont.Tanypus hirtipennis Loew, 1866:

Genus BRUNDINIELLA Roback -5.-Maine.Brundiniella Roback, 1978b: 141 (n. nam. Anatopynia dena Roback in Sublette &

for Brundinia Roback). Sublette, 1965: 145 (n. nam. forBrundinia Roback, 1978a: 168 (preocc. Anatopynia brunnea Roback, 1955: 2,

Tottenham, 1949). Type-species, preocc. Edwards, 1931).-Mass.Anatopynia eumorpha Sublette (orig.des.).

Genus PSECTROTANYPUS Kieffereumorpha. Ont., Wash. to Pa., s. to Calif. & Psectrotanyplrs Kieffer, 1,909: 42.

W. Va. Type-species, brevicalcar KiefferAnatopynia eumorpha Sublette, I964a: (Fittkau, 1962: 129) = Tipula varici

107.-Calif. Fabricius.

discolor. Wash., Wis., N.H., S.C.Genus DEROTANYPUS Roback Tanypus discolor Coquillett, 1902b:

Derotanypus Roback, l97l:- 91. 89.-N.H.Type-species,Anatopynia aclines dyari. B.C. to Ont., s. to Calif. & FIa.Sublette (orig. des.). Tanypus dyari Coquillett,1902a:

Merotanypus Roback, 1971: 94" 85.-Mich.Type-species, Tanypus alaskensis Tanypus guttularis Coquillett, I902b:Malloch (orig. des.). 92.-Wash.

Tanypus garretti Walley, 1925: 275.-B-C.aclines. Alas., Wash., Oreg., Idaho, Utah, pictipennis. Grld.; PA.

Wyo., Calif., Nev. Tanypus pictipennis Zetterstedt, 1838:Anatopynia aclines Sublette, 1,964a: 818.-Grld.

104.-Calif.Derotanypus aclines var. aclines

Roback, L971,: 93.-Utah, Wyo. Genus RADOTANYPUS Fittkau & Murrayalaskensis. AIas. to N.W.T., B.C. to Man. Radotanypus Fittkau & Murray, 1985: 209.

Tanypus alaskensis Malloch, 1919: Type-species, Anatopynia submarginella35.-Alas. Sublette (orig. des.).

Anatopynia centralis Malloch, 1934:13.-N.W.T. submarginella. Oreg., Calif., Colo., Wyo.

Anatopynia submarginelld Sublette, 7964a:103.-Calif.

10 NEARCTIC CHIRONOMIDAE

Tribe NATARSIINI

Genus NATARSIA FittKauNatarsia Fittkau, 1962: l5l. Type-species,

Chironomus punctatus Fabricius (orig.des.).

miripes. Calif.Tanypus miripes Coquillett, 1905:

6-5.-Calif.baltimoreus. N.W.T., e. of Rockies to Kans.

& Fla.Tanypus baltimoreus Macquart, 1855:

35.-Md.Ablabesmyia fastuosa Johannsen, 190-5:

1.53.-N.Y.

Tribe PENTANEURINI

Genus ABLABESMYIA JohannsenAblabesmyia Johannsen, 1905: 135.

Type-species, Tipula monilis Linnaeus(Johannsen, 1,907a: 400).

Subgenus ABLABESMYIA Johannsen

aspera. B.C. to Maine, s. to Calif. to Fla.Pentaneura aspera Roback, 1959a:

'i.24.-Maine.

Pentaneura gera Roback, L959a:126.-N.H.

hauberi. Man., Ont., N.Y., Pa., N.J., Ky., s.to Miss. & Fla.

Ablabesmyia hauberi Beck & Beck, 1966:329.-Fla.

janta. N.W.T., Md., Pa., Tenn., S.C., Okla.,Tex., Ga., Fla.

Pentaneura janta Roback, 19-59a:131.-Md.

johannseni. Ill.Pentaneura j ohannseni Roback, 19-59a:

131.-Ill.mallochi. Widespread NE.

Tanypus mallochi Walley, 1925: 273.-Ont.Pentaneura auriensis Roback, 7957 a:

39.-Pa.Pentaneura aequifasciata Dendy &

Sublette, 1959: 507.-Ala.Ablabesmyia ornata Beck & Beck, 1966:

328.-Fla.Ablabesmyia tarella Roback, 1971:

368.-Fla.monilis. Widespread NE.; PA.; OR.

Tipula monilis Linnaeus, 1758:587.-'Europe'.

Ablabesmyia monilis var. nigra Roback,l97l;378.-Ariz.

Tanypus basalis Walley, 1925:. 273--Ont-Ablabesmyia americana Fittkau, 1.962: 430

(n. nam. for Tipula monilis, Johannsen,1905: 142, not Linnaeus 1758,misident.).

parajanta. Ont., N.Y., Neb. to Pa., s. toMiss. & Fla.

Ablabesmyia parajanta Roback, 1971:373.-S.C.

rasha. N.H.Ablabesmyia rasha Roback, I97I:.

371.-N.H.rhamphe. Minn. to Ohio, s. to N. Mex. &

Fla.Ablabesmyia rhamphe Sublette, 1964b:.

ll4.-La.simpsoni. N.Y., Pa., S.C.

Ablabesmyia sintpsoni Roback 1985:188.-N.Y.

Subgenus ASAYIA RobackAsayia Roback, 7985: 177- Type-species

Tanypus annulatus (orig. des.).

annulata. Sask. to Qu6., s. to Calif. & Fla.Tanypus annulatus Say, 1823: 15.-Pa.

Subgenus KARELIA RobackKarelia l{oback, l97l: 357. Type-species,

Tanypus illinoensis Malloch (orig. des.).

alaskensis. Alas.Ablabesmyia alaskensis Roback, 1971:

360.-Alas.cinctipes. Calif., Nev., Ariz., N. Mex., Ky., s.

to Miss. & Fla.Pentaneura cinctipes Johannsen, 1946:

271..-Fla.idei. Ont., N.Y., Pa., S.C.

Tanypus idei tNalley, 7925: 272.-Ont.illinoensis. N.W.T., Alta. to Qu6., N.Y.,

Maine, Colo., N. Mex., Kans., W. Va.Tanypus illinoensis Malloch, 191-5b:

376.-\t.peleensis. Alas., widespread Can. & U.S.A.

Tanypus peleensis Walley, 1926a: 64.-Ont.philosphagnos. B.C., Ont., N.B., Pa. to Miss.

& FIa.Ablabesmyia philosphagnos Beck & Beck,

1966: 330.-Fla.pulchripennis. N.W.T., B.C. to Qu6., Wash.,

Wyo., S. Dak., D.C., Col., Kans., Grld.Tanypus pulchripennis Lundbeck, L898:

293.-Grld.Tanypus prudens Walley, L925: 275.-Man.

CHIRONOMIDAE DE LA REGION NEARCTIQUE 11

Genus ARCTOPELOPIA FittKauArctopelopia Fittkau, 1962: 194.

Type-species, Tanypus barbitarsisZetterstedt (orig. des.).

cana. N.W.T.Arctopelopia cana Roback, 1,977:

253.-N.W.T.melanosoma. Grld.; PA.

Ablabesmyia melanosoma Goetghebuer,1933: 20.-Grld.

Genus CANTOPELOPIA RobackCantopelopia Roback, l97l: 270.

Type-species, Cantopelopia gestaRoback (orig. des.).

gesta. Qu6., Ill., N.Y., Fla.Cantopelopia gesta Roback, 197I:

270.-Qu6..

Genus CONCHAPELOPIA FittkauConchapelopia Fittkau, L957: 317-

Type-species, Tanypus pallidus Meigen(orig. des.).

Mesopelopia Roback, 1977: 252. Typespecies, aleta (mon).

aleta. Widespread in e. Can. & U.S.A.Conchapelopia aleta Roback, 1971:

252.-N.Y.bruna. Alas. to N.W.T.

Conchapelopia bruna Roback, 1971:249.-N.W.T.

currani. Man. to Qu€., Wyo., Iowa, N.Y.,Ohio. s. to Miss. & Fla..

Tanypus currani Walley, 1925: 276.-Qu€.fasciata. Alta. to Ont., s. to Kans. & Fla.

Conchapelopia fasciata Beck & Beck,1966: 346.-Fla.

Conchapelopia dusena Roback, 1971:247.-N.Y.

melanops. Not NE.Tanypus melanops Meigen, 1818:

65.- Not stated.pallens. Alas. to Qu6., s. to Calif. & Fla.

Tanypus pallens Coquillett, I902b: 91.-N.Mex.

Pentaneura goniodes Sublette, 1964a:89.-Calif.

Conchapelopia goniodes var. trifidaRoback. I97I: 246.-Calif.

mera. Wash., Ariz.Conchapelopia mera Roback, 1971:

246.-Wash.paramelanops. N.Y., Va.

Conchapelopia paramelanops Roback,1971: 249.-Ya.

rurika. Man., Qu6., s. to Kans. & Ga.Pentaneura rurika Roback. 1957a: 34--Pa.

telema. N.W.T. to Ont., Mont., N.Y., Ill.,Kans.. N.C.

Conchapelopia telema Roback, 1971:247.-Alta.

Yarna. W. Va.Conchapelopia varna Roback, 1981:

93.-W. Va.

Genus DENOPELOPIA Roback & RutterDenopelopia Roback & Rutter, 1988: 118.

Type-species, atria (orig. des.).

atria. Fla.Denopelopia atria Roback & Rutter, 1988:

120.-Fla.

Genus GUTTIPELOPIA FittkauGuttipelopia Fittkau, 1962: 251.

Type-species, Tanypus guttipennis Wulp(orig. des.).

guttipennis. N.W.T., Man., Ont., s. to Ala. &Fla.; PA.

Tanypus guttipennis Wulp, 1861:16.-Netherlands.

Tanypus multipunctatas Curran, 1930:29.-N.Y.

Guttipelopia curuani Beck & Beck, 1966:332.-Fla.

rosenbergi. Man., Ont.Guttipelopia rose nbergi Bilyj, 1984:

6-59.-Man.

Genus IIAYESOMYIA Murray & FittkauHayesomyia Murray & Fittkau, 1985: 196.

Type-species Tanypus lripunctatusGoetghebuer (orig. des.).

senata. Alta. to N.S., s. to Calif., Kans. &Fla.

Tanypus senatus Walley, L925: 276: -Ont.' Pentaneura chrysos Sublette, 1964a:90.-Calif.

?Chironomus unicolor Walker.1848: 19.-N.S.

t2 NEARCTIC CIIIRONOMIDAE

Genus HELOPELOPIA Roback neopilosella. Neb., Ky., N.Y., N.C., s. toHelopelopia Roback, l97l: 250 (as subg. Miss. & Fla.

of Conchapelopia Fittkau, 1957). Labrundinia neopilosella Beck & Beck,Type-species, Tanypus cornuticaudatus 1966: 341.-Fla.Walley (orig. des.). pilosella. Alas. to Qu6., s. to Ariz. & Fla.;

NT.cornuticaudata. Alta. to Qu6., N.Y., Ohio, Tanypus pilosellus Loew, 1866: 5.-D.C.

Pa., Kans., N.C., S.C., Fla. Labrundinia floridana Beck & Beck, 1966:Tanypus cornuticaudatus Walley, 1,925: 339.-Fla.

277--Qu6. virescens. Pa. s. to Tex. & Fla.pilicaudata. N.W.T. to N.B., N.Y., N.C. Labrundinia virescens Beck & Beck, 1966:

Tanypus pilicaudatus Walley, 1925: 341.-Fla.277.-Ont.

Genus LARSIA FittkauGenus HUDSONIMYIA Roback. Larsia Fittkat, 1962.339. Type-species,

Hudsonimyia Roback, 1979: I. Ablabesmyia atrocincta GoetghebuerType-species karelena Roback (orig. (orig. des.).des. ).

berneri. Minn., Fla.karelena. N.Y., N.C., S.C., Ga. Larsia berneri Beck & Beck, 1966:

Hudsonimyia karelena Roback, 1979: 3-50.-Fla.3.-S.C. canadensis. Can. e. of Rockies.

parrishi. Ga. Larsia canadensis Bilyj, 1984: 663.-Man.Hudsonimyia parrishi Caldwell & Soponis, decolorata. Wis., Mich., N.Y., Kans., Ill.,

1982: 507.-Ga. Ohio. La., Fla.Tanypus decoloratus Malloch, 191-5b:

370.-Ill.Genus KRENOPELOPIA Fittkau Larsia lurida Beck & Beck, 1966:

Krenopelopia Fittkau, 1962: 262. 3-51.-Fla.Type-species, Chironomus binotatus Larsia indistincta Beck & Beck, 1966:Wiedemann (orig. des.). 3-52.-Fla.

Iyra. Calif.hudsoni. S.C. Pentaneura /yra Sublette, 1964a.

Krenopelopia hudsoni Roback, 1983: 96.-Calif.254.-S.C. marginella. Qu6., Ill., Kans.

narda. Alas., ?Oreg. Tanypus marginellus Malloch, 1915b:Krenopelopia narda Roback, l97l: 374--Ill.

274.-Alas. planensis. Ont., Qu€., Calif., Tex., Ohio,N.Y., Va., Mex.

Pentaneura planensis Johannsen, 1946:Genus LABRUNDINIA Fittkau 284.-Tex.

Labrundinia Fittkau, 1962: 372. sequoiaensis. Calif.Type-species, Tanypus longipalpis Pentaneura sequoiaensis Sublette, 1964a:Goetghebuer (orig. des.). 99.-Calif.

becki. Okla., Ky., N.C., s. to Miss. & Fla.Labrundinia becki Roback, I97I: Genus MEROPELOPIA Roback

279.-Fla- Meropelopia Roback, l97l: 253 (as subg.johannseni. Ky., N.C., s. to Miss. & Fla. of Arctopelopia Fittkau, 7962).

Labrundinia johannseni Beck & Beck, Type-species, Ablabesmyia flavifrons1966: 342.-Fla. Johannsen (orig. des.).

maculata. Calif., Kans., N. Mex., Tex., N.C.,Mex.; NT.

Labrundinia maculata Roback. 1971:278.-Calif.

CHIRONOMIDAE DE I-A REGION NEARCTIQUE l-t

americana. N.Y., Conn., N.J., Ohio, N.C.,S.C., Kans.

Conchapelopia americana Fittkau, 19-57:320 (n. nam. for Pentaneura vitellinaJohannsen, t946: 281, not Kieffer,1916, misident.).-Conn.

flavifrons. Alas., Alta. to Qu6., s. to Calif.,Fla.

Ablabesmyia flavifrons Johannsen, 1905:150.-N.Y.

Pentaneura gesra Roback in Sublette &Sublette, 1965: I47 (n. nam. forPentaneura alba Roback, 1957a: 32,preocc. Tokunaga, 1937).-Pa.

Arctopelopia fittkaui Beck & Beck, 1966:348.-Fla.

Genus MONOPELOPIA FittkauMonopelopia Fittkau, 1962: 394.

Type-species, Tanypus tenuic al c arKieffer (orig. des.).

s. to Miss. &Fla.

Monopelopia boliekae Beck & Beck, 1966:335.-Fla.

tenuicalcar. 8.C., Ont., N.8., Nfld., Calif.,Fla.: PA.

Tanypus tenuicalcar Kieffer, 1918a:182.-Czechoslovakia.

tillandsia. Ky., N.C., s. to Miss. & Fla.Monopelopia tillandsia Beck & Beck,

t966: 336.-Fla.

Genus NILOTANYPUS KiefferNilotanypus Kieffer, 1923b: 191.

Type-species, r emot is s imus Kieffer(mon.).

americanus. Ga., N.C., S.C., Fla.Nilotanypus americanus Beck & Beck,

1966: 345.-Fla.du,bius. Not NE.

Tanypus dubius Meigen, 1804: 25.-notstated.

fimbriatus. Widespread NE.Chironomus fimbriatus Walker, 1848:

20.-Ont.ka.nsensis. Kans.

Nilotanypus kansensis Roback, 1986a:458.-Kans.

Genus PARAMERINA FittkauParamerina Fittkau, 1962: 317.

Type-species, Chironomus cingulatusWalker (orig. des.).

anomala. Ky., N.C., s. to Miss. & Fla.Paramerina anomala Beck & Beck, 1966:

344.-Fla.fragilis. N.W.T., B.C. to Qud., Oreg., Va.

Tanypus fragilis Walley, 1926b. 205.-Ont.hanseni, Wash.

Paramerina hanseni Roback, 1971:274.-Wash.

smithae. Wash., N. Dak., S. Dak., Calif.,IItah, Wyo., N. Mex., Kans.

Pentaneura smithae Sublette, 1964a:100.-Calif.

testa. Okla., Tex.Paramerina testa Roback. l97 l: 272.-Tex.

Genus PENTANEURA PhilippiPentaneura Philippi, 18651. 629.

Type-species, grisea Philippi (mon.).

inconspicua. Ont., Wash., s. to Ariz., Kans.,Okla., Ill. to Pa., s. to Fla.

Tanypus inconspicuus Malloch, 191-5b:371.-Ill.

Pentaneura comosa Sublette, 1,964a:93.-Calif.

Pentaneura inculta Beck & Beck, 1966:334.-Fla.

inyoensis. Calif., N.Y., Ill., Ark.Pentaneura inyoensis Sublette, 1964a:

98.-Calif.

Genus REOMYIA RobackReomyia Roback, 1986b: 283. Type-

species, Zavrelimyia wartinbei Roback(orig. des.).

wartinbei. Alas., Oreg.Zavrelimyia wartinbei Roback, 1984:

17.-Alas.

Genus RHEOPELOPIA FittkauRheopelopia Fittkau, 1962: 209.

Type-species, Tanypus ornatus Meigen(orig. des.).

acra. Qu6., Maine, Mich., s. to W. Va., Ga.Thienemannimyia acra Roback, 1971:

243.- Mich.

L4 NEARCTIC CIIIRONOMIDAE

paramaculipennis. Qu6., Ill., Va.Thie n emannimy i a p ar amacul ipe nnis

Roback, I97l: 243.-Ya.perda. Alta., Idaho, Ont., Minn., N.Y., Pa.

Thienemannimyia perda Roback, 1971:244.-N.Y.

Genus TELOPELOPIA RobackTelopelopia Roback, l97l: 255.

Type-species, Tanypus okoboji Walley(orig. des.).

okoboji. Man., Minn., Iowa, Ohio, Md., Va.,Kans.. N. Mex.

Tanypus okoboji Walley, I928b:582.-Iowa.

Genus THIENEMANNIMYIA FittkauThienemannimyia Fittkau, 1.9 57 : 375.

Type-species, Ablabesmyia geij skesiGoetghebuer (orig. des.).

barberi. Wash. s. to Calif., Nev., Ariz., Kans.,N. Mex.

Tanypus barberi Coquillett, 1902b: 90.-N.Mex.

fusciceps. Alas., N.W.T., B.C., Man., Calif.,Wyo.; PA.

Pentaneura fusciceps Edwards, 1929:290.-England.

norena. Man., Ont., Mich., Maine s. to Pa.,Oreg., Kans.

Pentaneura norena Roback, 7957a:38.-Pa.

Thie nemannimyia norena var.quadrata Roback, 197L:242.-Ill.

Genus TRISSOPELOPIA KiefferTrissopelopia Kieffer, 1923d: L78.

Type-species, flavida Kieffer (orig.des.).

ogemawi. Alta., Ont., Qud., Mich., N.Y., Pa.,Ga.

Trissopelopia ogemawi Roback, 1971:268.-Mich.

Genus XENOPELOPIA FittkauXenopelopia Fittkau, 1962: 381.

Type-species, Pelopia falcigera Kieffer(orig. des.).

tincta. Calif.Xenopelopia tincta Roback, l97I:

256.-Calif .

Genus ZAVRELIMYIA FittkauZavrelimyia Fittkau, 1962: 285-

Type-species, Tanypus melanurusMeigen (orig. des.).

bifasciata. Man. to Nfld., U.S.A. s. to S.C.Tanypus bifasciatus Coquillett, 1901b:

609.-N.J.sinuosa. Alta. to Qu6., s. to Kans. & Fla.,

Calif.Tanypus sinuosus Coquillett, 1905:

6-5.-N.H.Zavrelimyia carneosa Fittkau, 1,962: 375

(n. nam. for Ablabesmyia carneaJohannsen, 190-5: 140, not Fabricius,180-5, misident.).-N.Y.

Pentaneura fluminalis Sublette, 1964a:88.-Calif.

thryptica. Alas., Alta., Oreg., Ont., N.Y., s.to Kans. & S.C., Calif.; ?PA.

Pentaneura thryptica Sublette, L964a:9-5.-Calif.

Incerta sedis in PentaneuriniTanypus futilis Wulp, 1867:

130.-Wis.

Nominen dubium in PentaneuriniAblabesmyia aurea J ohannsen, 1907b:

110.-Kans.

Tribe PROCLADIINI

Genus DJALMABATISTA FittkauDj almab atis ta Fittkau, I968a: 328.

Type-species, director Fittkau (orig.des.).

Calotanypus Roback, 7977: I52 (as subg.of Procladius Skuse, 1889).Type-species, Protenthes pulcherJohannsen (orig. des.).

pulcher. Ont., Qu6., N.Y., Mich., Ohio, Pa.,N.C., S.C., Fla., Ga.

Protenthes pulcher Johannsen, 1908:273.-N.Y.

Tanypus apicalis Walley, L926a: 64.-Qu6.Protenthes fasciger Curran, 1930:

29.-N.Y.Procladius maculatus Roback. 1971:

152.-Fla.

CHIRONOMIDAE DE LA REGION NEARCTIQUE 1-5

Genus PROCLADIUS SkuseProcladius Skuse. 1889: 283.

Type-species, paludic ola Skuse(coquillett, 1910: 594).

Subgenus HOLOTANYPUS RobackHolotanypus Roback, 1982: 99 (n. nam.

for subg. Procladius sensu Roback1971, not Skuse 1.889, misident.).Type-species, Tipul a culicifor misLinnaeus (orig. des.).

abetus. Alta., Ont.Procladius abetus Roback. 1971:

185.-Alta.barbatulus. Calif.

Procladius b arb atulus Sublette. 1964a;l2l.-Calif.

clavus. N.W.T., B.C.Procladius clavus Roback, 1971:

187.-N.W.T.culiciformis. N.W.T., B.C. to Ont., s. to

Kans. & Fla., Calif., Nev., Ariz.; PA.Tanypus culiciformis Linnaeus, L767:

978.-Sweden.Tanypus crassinervis Zetterstedt, 1,838:

817.Protenthes claripennis Malloch, 1915b:

387.-Mich.curtus. Fla.

Procladius curlas Roback, l97l: 181.-Fla.denticulatus. Widespread NE.

Procladius denticulatus Sublette. 1964a;I24.-Calif.

dentus. Alas., N.W.T., B.C. to Labr.Procladius dentus Roback. 1971:

186.-Man.desis. Alas.. N.W.T.

Procladius desis Roback, l97l: 178.-Alas.freemani. Alas., N.W.T., B.C. to Qu€., s. to

Calif. & Ga.Procladius freemani Sublette, 7964a:

122.-Calif.Procladius freemani var. asetus Roback,

l97l: 172.-Ga., N.C., S.C.Procladius freemani var. brunneus Roback,

1971: 17l.-Nfld.Procladius freemani var. cretis Roback,

l97l: 172.-8.C.Procladius freemani var. elongatus Roback,

l97t: L7Z.-Fla., N.C., S.C.gretis. N.W.T.

Procladius Sretis Roback, 1971:184.-N.W.T.

jeris. Alas.. N.W.T.Procladius jeris Roback, 197 l: 187.-Alas.

johnsoni. Ont., N.Y.Procladius johnsoni Roback, 1980a:

43.-Ont.paragretis. ?Alas., Grld.

Procladius paragretis Roback, 1971:185.-Grld.

prolongatus. Alas., N.W.T.Procladius prolongatus Roback, 1971:

188.-N.W.T.ruris. N.W.T., B.C. to Labr., Minn.

Procladius ruris Roback, 1971.: 182.-Alta.Procladius ruris var. grandis Roback,

1977: 183.-N.w.T., 8.C., Nfld.sublettei. Widespread NE.

Procladius sublettei Roback, 1971:191.-Calif.

Procladius sublettei var. grandis Roback,l9'71: 793.-N.W.T., Sask., Ont.

Procladius sublettei var. minuta Roback,1971.: 194.-N.C., S.C., Fla.

vesus. Alas., N.W.T.Procladius vesas Roback, 1971:

181.-N.W.T.wilhmi. Tenn.

Procladius wilhmi Roback, 1966:177.-Tenn.

Subgenus PSILOTANYPUS KiefferPsilotanypus Kieffer, 1906a: 39.

Type-species, Tanypus bellus Loew(Coquillett, 1910: -597).

bellus. N.W.T., B.C., Sask. to N.B., s. toCalif. & Fla.

Tanypus bellus Loew, 1866;4.-D.C.Procladius adumbratus Johannsen, 190-5:

132.-N.Y.Tanypus pusillus Loew, 1866: 5.-D.C.Procladius flavidus Kieffer, 1923a:

297.-Tex.Procladius malifero Garrett, 1925:

10.-8.C.deltaensis. Man.

Procladius deltaensis Roback in Wrubleski& Roback. 1987: 208.-Man.

macrotrichus. Minn.Procladius macrotrichus Roback, 1971:

168.-Minn.nietus. B.C., Alta., Sask., Man.

Procladius nietus Roback. 1,971:.168.-B.C.

riparius. Kans., Ill.Protenthes riparius Malloch, 1915b:

389.-Ill.

t6 NEARCTIC CHIRONOMIDAE

Incerta sedis in ProcladiusProcladius lundstromi Goetghebuer in

Goetghebuer & Lenz, 1936: 11.-notstated.

Tanypus abrupta Garrett, I925a: 8.-B.C.Tanypus arcuata Garrett, 7925a: 7.-I|.C.Tanypus bifida Garrett, 1925a: 8.-B.C.Tanypus trifolia Garrett, 1925a: 7.-B.C.Trichotanypus vestitipennis Kieffer, 1917:

339.-N.Y.

Tribe TANYPODINI

Genus TANYPUS MeigenTanypus Meigen, 1,803: 261. Type-species,

Tipula cincta Fabricius (Latreille, 1810:442; misident.). = Tanypuspunctipennis Meigen.

Pelopia Meigen, 1800: 18. Type-species,Tipula cincta Fabricius (Coquillett,1910: -586; misident.) = Tanypuspunctipennis Meigen. Supressed byICZN, 1963: 339.

Protenthes Johannsen, 7907a: 400.Type-species, Tanypus punc tipennisMeigen (orig. des.).

Subgenus APELOPIA RobackApelopia Roback, l97L: 55. Type-species,

Tanypus neopunctipennls Sublette (orig.des).

clavatus. Ky. to N.C., s. to Tex. & Fla.Tanypus clavatus Beck, 1962: 92.-Fla.

grodhausi. Calif.Tanypus grodhausi Sublette, I964a:

116.-Calif.imperialis. Calif., N. Mex.

Tanypus imperialis Sublette, 1964a:II2.-Calif.

neopunctipennis. Widespread U.S.A. e. ofRockies, Mex.; NT.

Tanypus neopunctipennis Sublette, 1964a:118.-Ill.

nubifer. Man., Calif., Nev., Utah, N. Mex.,Neb., Kans.

Tanypus nubifer Coquillett, 190-5:66.-Utah.

Subgenus TANYPUS Meigen

carinatus. Calif., Miss., Pa., S.C.Tanypus carinatus Sublette, 1964a:

110.-Calif.

concavus. Iowa, Kans., Ark., Mich., Ohio,N.Y., Va., N.C., S.C., Tex.

Tanypus concavus Roback, 1971:63.-Mich.

parastellatus. Calif., Nev., Ariz., Tex.Tanypus parastellatus Sublette, 1964a:

113.-Calif.punctipennis. B.C., Sask. to Qu6., Calif. to

Md., Ala., S.C., Fla.; PA.; OR.Tanypus punctipennis Meigen, 1818:

61.-Not stated.stellatus. Alta., Ont., Utah to Pa., s. to Tex.

& Fla.Tanypus stellatus Coquillett, 1902b:

89.-Tex.telus. Fla.

Tanypus relus Roback, l97l. 68.-Fla.vilipennis. Man., Wis., Mich.; PA.

Protenthes vilipennis Kieffer, 1918a:164.-Hungary.

Nomen dubium in TanypodinaeTanypus pubitarsis Zetterstedt, 1850:

3603.-Grld.; Sweden.

Subfamily DIAMESINAE

Tribe B OREOHEPTAGYIINI

Genus BOREOHEPTAGYIA BrundinBoreoheptagyla Brundin, L966: 420.

Type-species, Heptagia rugosa Saunders(orig. des.).

lurida. Alas., B.C., N.Y., Wash., Wyo.Diamesa lurida Garrett. t925a: 6.-8.C.

Tribe DIAMESINI

Genus ARCTODIAMESA MakarchenkoArctodiamesa Makarchenko, 1983; 264.

Type-species, Diamesa appendiculataLundstrOm (orig. des.).

appendiculata. Alas.; PA.Diamesa appendiculata LundstrOm, 1915:

23.-Siberia.

Genus DIAMESA MeigenDiamesa Meigen in Gistl, 1835: 66.

Type-species, cinerella Meigen in Gistl(orig. des.).


Adiamesa Kieffer, 1906a: 36. geminata. N.W.T., Grld.; PA.Type-species, Chironomus tonsus Diamesa geminata Kieffer, 1926:Walker, 19-56a: 195 (mon.). 79.-Grld.

Eutanypus Coquillett, 1899: 341.. Diamesa biappendiculata Goetghebuer inType-species, borealis Coquillett (orig. Remy, 1928: 52.-Gtld-des.). Diamesa furcata Edwards, 1933:

Psilodiamesa Kieffer, 1918a: 104. 616.-N.W.T.Type-species, Psilodiamesa gregsoni. N.W.T., N.B.; PA.spitzbergensis Kieffer (orig. des.). Diamesa gregsoni Edwards, 1.933:

618.-N.W.T.aberrata. Alas., Wyo., Grld.; PA.; OR. haydaki. Alas., Wyo., Minn., Col., Ariz.

Diamesa abetata Lundbeck, 1898: Diamesa haydaki Hansen in Hansen &289.-Grld. Cook, 1976: 93.-WYo.

alpina. Alas.; PA. heteropus. Alas. s. to N. Mex., to Neb. &Diamesa alpina Tokunaga, 1936: Minn.

539.-Japan. 'Tanypus heteropus Coquillett, 1905:amplexivirilia. B.C., Alta., Wash., Mont.; PA. 66.-?N.H., N. Mex., Wash.

Diamesa amplexivirilia Ilansen in I{ansen Adiamesa confusa Garrett, 192-5: 5.-B.C.& Cook, 19?6: -53.-Wash. Diamesa banana Garrett, 1925: 6.-8.C.

ancysta. Alas., Wash., Idaho, Mont., Nev., Diamesa onteona Roback, 1957b:Utah. Colo. 6.-Utah.

Diamesa ancysta Roback, 1959b: hyperborea. Not NE.l.-Mont. Diamesa hyperborea Holmgren, 1869:

arctica. Alas., N.W.T.; PA. 48.-Bear Island.Chironomus arcticus Boheman, 1865: Adiamesa ursus Kieffer, 1918a: 104.-Bear

574.-Spitzbergen. Island.bertrami. Grld.; PA. incallida. Alas., Y.T., Man., Ont., Wash.,

Diamesa bertrami Edwards, 1935: Utah, Wyo.; PA.470.-Grld. Chironomus incallidus Walker, L956a:

bohemani. N.W.T., Grld.; PA. 183.-England.Diamesa bohemani Goetghebuer, 1932a: Diamesa fonticola Sether, 1969:

181.-Spitzbergen. 24.-Man.cheimatophila. N.Y. insignipes. Wyo., Utah; PA.

Diamesa cheimatophila Hansen in Hansen Diamesa insignipes Kieffer in Kieffer && Cook, 1976: 68.-N.Y. Thienemann, 1908: 3.-Germany.

chiobates. Minn., Wis. japonica. Wash., Mont., Wyo., Utah, Calif.;Diamesa chiobates Hansen in Hansen & PA.

Cook, 1976: 70.-Wis. Diamesa japonica Tokunaga, 1936:chorea. Alas., Calif., Wyo., Grld. 542.-Japan.

Diamesa chorea Lundbeck. 1898: Diamesa leoniella Hansen in Hansen &291.-Grld. Cook, 1976: 111.-Wyo.

clavata. N.W.T. leona. Wash. to N. Mex., to Minn., Colo.,Diamesa clavata Edwards, 1933: Qu6., N.B.; PA.

61-5.-N.W.T. Diamesa leona Roback, 1957b: 7.-Utah.colenae. Alas., Y.T., Wyo. Diamesa caena Roback, 1957b: 9.-Utah.

Diarnesa colenae Hansen in Hansen & Diamesa pieta Roback, 1957b: 8.-Utah.Cook, 1976: 76.-Wyo. lindrothi. Grld.; PA.

davisi. N.W.T., Utah, Wyo., N.H., Grld. Diamesa lindrothi Goetghebuer inDiamesa davisi Edwards, 1933: Goetghebuer & Lindroth, 1931:

614.-N.W.T. 281.-Iceland.garretti. B.C., Wash., Idaho, Mont., Wyo. lupus. Alas., Alta.

Diamesa garretti Sublette & Sublette, Diamesa /apas Willassen, 1985:1965: 151 (n. nam. for Diamesa 127.-Alas.borealis Garrett, 1925: 6, preocc.Kieffer, 1915b).-B.C. mendotae. S. Dak., Minn., Wis.

Diamesa mendotae Muttkowski. 1915:116.-Wis.

l7


nivicavernicola. Alas.. Wash. Genus POTTHASTIA KiefferDiamesa nivicavernicola Hansen in Hansen Potthastia Kieffer. 7922c: 362.

& Cook, 1976: 123.-Wash. Type-species,longimana Kieffer (orig.nivoriunda. Minn. to Nfld., Ky., N.C., s. to des.).

Miss. & Fla.Chironomus nivoriundus Fitch, 1847: gaedii. E. U.S.A.; PA.

282.-N.Y. Diamesa gaedii Meigen, 1838:simplex. Alas., N.W.T., B.C., Qu6., Wyo., 13.-Belgium.

Grld. iberica. Idaho, Wyo., Ga.; PA.Diamesa simplex Kieffer, 1926: Potthastia iberica Serra-Tosio, 1971:

81.-N.W.T. 160.--Spain.sommermani. Alas. longimana. Ont., Qu6., N.Y., N.C., S.C.; PA.

Diamesa sommermani Hansen in Hansen Potthastia longimanus Kieffer, 1922c:& Cook, 7976: 134.-Alas. 362.-Germany.

sonorae. Calif. montium. Se. U.S.A.; PA.Diamesa sonorae Willassen, 1985: Diamesa montium Edwards, 1929:

125.-Calif. 307.-Great Britain.spinacies. Alas., Alta., Mont., Idaho, Wyo.,

utah, col., calif.Diamesa spinacies Sether, 1969: Genus PSEUDODIAMESA Goetgbebuer

21.-Alta. Pseudodiamesa Goetghebuer insubletti. Alta. Goetghebuer & Lenz, 1939a: 9 (as

Diamesa subletti Makarchenko, 1986: subg. of Syndiamesa Kieffer, 1918a).155.-Alta. Type-species, Syndiamesa pilosa Kieffer

vockerothi. Ont., Qu6. (orig. des.) = Diamesa branickiiDiamesa vockerothi Hansen in Hansen & Nowicki.

Cook, 1976: 139.-Ont.Subgenus PACIIYDIAMESA Oliver

Pachydiamesa Oliver, 1959: 54.Genus LAPPODIAMESA Serra-Tosio Type-species, Diamesa arctica Malloch

Lappodiamesa Serra-Tosio, 1968: 140. (orig. des.).Type-species, b rundini Serra-Tosio(orig. des.). arctica. N.W.T.

Diamesa arctica Malloch, 1919:boltoni. Ohio. 37.-N.W.T.

Lappodiamesa boltoni Sether & Willassen,1988: 78.-Ohio.

Subgenus PSEUDODIAMESA Goetghebuer

Genus PAGASTIA Oliver branickii. B.C., Alta., Calif., N. Mex., Grld.;Pagastia Oliver, 1959: 49. Type-species, PA.; OR.

orthogonia Oliver (orig. des.). Diamesa branickii Nowicki, 1873:Hesperodiamesa Sublette, I967a: 305. 3.-Poland.

Type-species, Prodiamesa sequax pertinax. N.W.T., B.C., Utah, Wyo., Colo.Garrett (orig. des.). Prodiamesa pertinax Garrett, 1925:

6.-8.C.orthogonia. Alas., Mich.

Pagastia orthogonia Oliver, 1959:51.-Alas. Genus PSEUDOKIEFFERIELLA Zavrel

partica. Alas., Y.T., Utah, Wyo. Pseudokiefferiella Zavrel, 1947: 1.1.Syndiamesa partica Roback, I95'7b: Type-species, Diplomesa

4.-Utah. lapponica Pagast = DiamesaSyndiamesa artisia Roback, 19-57b: parva Edwards (orig. des.). (see

5.-Utah. Cranston, 1975: 88; Ashe, 1983:sequax. B.C., Alta. 46- ICZN ruling required).

Prodiamesa sequax Garrett, 1925: 7.-8.C. Diplomesa Pagast, 1947: 539.Type-species, lapponica Pagast (mon.).

CHIRONOMIDAE DE LA REGION NEARCTIQUE

parva. N.W.T.; PA. ramosus. N.W.T., Ont.Diamesa parva Edwards, 1.932a: Protanypus ramosus Sether, 1975c:

4-5.-Scotland. 368.-Ont.saetheri, Alas.

Protanypus saetheri Wiederholm, 797 5:Genus SYMPOTTHASTIA Pagast 224.-Alas.

Sympotthastia Pagast, 1947: 457.Type-species, zavreli Pagast (mon.).

diastena. Oreg., Colo., Calif. Subfamily PRODIAMESINAEPseudodiamesa diastena Sublette. I964a:

128.-Calif. Genus COMPTEROMESA Setherfulva. N.Y., N.C., S.C. Compteromesa Sather, 1987c: 794.

Diamesa fulva Johannsen, 192L: Type-species, oconeensis Srether (orig.229.-N.Y. des.).

zavreli. N.C.Sympotthastia zavreli Pagast, 1947: oconeensis. S.C.

458.-Germany. Compteromesa oconeensis Saether, 1981c:195.-S.C.

Genus SYNDIAMESA KiefferSyndiamesa Kieffer, 1918a: 103. Genus MONODIAMESA Kieffer

Type-species, Diamesa hygropetrica Monodiamesa Kieffer, 1922a: 287.Kieffer (Edwards, 1929: 303). Type-species, Prodiamesa bathyphila

Kieffer, Goetghebuer in GoetghebuerUnnamed species.-N.W.T., Ont. & Lenz, 7939a: 3).

Nomina dubia in Diamesinae bathyphila. Alas., N.W.T., Minn., Ill. to Ohio,Diamesa waltlii Meigen, 1838: Grld.; PA.

l3.-Germany. Prodiamesa bathyphila Kieffer, 1918a:Syndiamesa polaris Kieffer, 1926: 102.-Germany.

81.-N.W.T. depectinata. Man., Ont., Pa.Monodiamesa depectinata Srther, I973b:

666.-Man.Incerta sedis in Diamesinae prolilobata, B.C.

Diamesa polaris Kieffer, 1926: Monodiamesa prolilobata Sether, 1,973b:79.-N.W.T. 673.-B.C.

tuberculata. N.W.T., 8.C., Ont., Mich., Ind.Monodiamesa tuberculata Srther. 1,973b:.

Tribe PROTANYPINI 671.-Mich.

Genus PROTANYPUS KiefferProtanypus Kieffer, I9O6a: 40. Genus ODONTOMESA Pagast

Type-species, Tanypus morio Odontomesa Pagast, 1947: 502.Zetterstedt (Edwards, 1924a: 719). Type-species, Prodiamesa fulva Kietfer

(orig. des.).caudatus. N.W.T.: PA.

Protanypus caudatus Edwards, 1,924a: ferringtoni. Colo.122.-Norway. Odontomesa ferringtoni S&ther, 1985e:

hamiltoni. N.W.T., B.C. 25.-Colo.Protanypus hamiltoni Saether, 197-5c: fulva. Mont. s. to N. Mex., Wis., II[., Pa. s. to

375.-8.C. Miss. & Fla.: PA.pseudomorio. Alas.; PA. Prodiamesa fulva Kief.fer, 1919a:

Protanypus pseudomorio Makarchenko, 132.-Hungary.1982: L33.-U.S.S.R. Odontomesa fulva nearctica Srther,

1985e: 20.-Ga.

19


lutosopra. B.C. Chironomus lucens lucensProdiamesa lutosopra Garrett, 7925: Zetterstedt.-Not NE.

7.-B.C. Chironomus lucens var. baueriGoetghebuer, I955. 167.Unrecognized.

Genus PRODIAMESA Kieffer nitidellus. B.C. to Man., S. Dak., Ill., N.8.,Prodiamesa Kieffer, 1906a: 37. N.Y., Pa.

Type-species, Diamesa praecox Kieffer Orthocladius nitidellus Malloch, 1915b:(orig. des.) = Chironomus olivaceus 515.-Ill.Meigen. Spaniotoma senex Johannsen, 1937a:

63.-N.Y.cubita. B.C.

Prodiamesa cubita Garrett, 1925: 7.-8.C.olivacea. Widespread NE.; PA. Genus ANTILLOCLADIUS Sether

Chironomus olivaceus Meigen, 1818: Antillocladius Srether, 1981b: 4.29.-Not stated, ?Germany. Type-species, antecalvus Sether (orig.

Chironomus turpis Zetterstedt, 1838: des.).81 1.-Sweden.

Tanypus occidentalis Coquillett, 1902b: arcuatus. Kans., S.C.92.-Colo. Antillocladius arcuatus Srether, 1982:

474.-S.C.pluspilalus. Kans., S.C.

Antillocladius pluspilalus Sether, 1982:Subfamily ORTHOCLADIINAE 474.-S.C.

Genus ABISKOMYIA EdwardsAbiskomyia Edwards, 1937: I4O. Genus APOMETRIOCNEMUS Sether

Type-species, virgo Edwards (orig. Apometriocnemus Snther, 1985c: 536.des.). Type-species, fontinalis Sether (orig.

des.).virgo. Alas., N.W.T.; PA.

Abiskomyia virgo Edwards, 1937: beringensis. Y.T.l41.-Sweden. Apometriocnemus beringensis Cranston &

Oliver. 1988a: 428.-Y.T.fontinalis. Tenn.

Genus ACAMPTOCLADIUS Brundin Apometiocnemus fontinalis Srether, 1985c:Acamptocladlas Brundin, 1956: 162. 537.-Tenn.

Type-species, Spaniotoma submontanaEdwards (orig. des.).

Phycoidella Srether, 1971: 1810. Genus BAEOCTENUS SretherType-species, dentolatens Srether (orig. Baeoctenus Srether, \976:. 4I.des.). Type-species, bicolor Sether (orig.

des.).dentolatens. Ont.

Phycoidella dentolatens Sether, 1971: bicolor. Man., N.B.1814.-Ont. Baeoctenus bicolor Sether, 1976:

43.-Man.

Genus ACRICOTOPUS KiefferAcricotopus Kieffer, l92la: 90. Genus BRILLIA Kieffer

Type-species, grandis Kieffer (orig. Brillia Kieffer, 1913b: 34. Type-species,des.) - Chironomus lucens Zetterstedt. Metriocnemus bifidus Kieffer (orig.

des.) = Chironomus modestus Meigen.Iucens. N.W.T., Alta.; PA.

Chironomus lucens Zetterstedt. 1850:3574.-Denmark.

CHIRONOMIDAE DE LA REGION NEARCTIQUE 2l

flavifrons. B.C. to N.B., s. to Oreg., N. Mex.to La., N.C., S.C., Ga.

Metriocnemus flavifrons Johannsen, L905:301.-N.Y.

laculata. Wash., Oreg., Wyo.Brillia laculata Oliver & Roussel, 1983:

268.-Oreg.longifurca. Not NE.

Brillia longifurca Kieffer, I92la:.86.-Germany.

parva. Ont. to N.B., s. to S.C.Brillia parva Johannsen, 1934: 3-51.-N.Y.

retifinis. Alas., B.C. to Man., s. to Calif. &Colo.

Brillia retifinis Srether, t969: 37.-8.C.sera. N.If., Pa., S.C., Ga.

Brillia sera Roback. 1951a:. 64.-Pa.

Genus BRYOPHAENOCLADIUSThiennemann

Bryophaenocladius Thienemann, 1934: 36.Type-species, Orthocladius mus cic olaKieffer (orig. des.).

Clinocladius Sublette. 1970: -50Type-species, Orthocl adiussubparallelus Malloch (orig. des.).

Platycladius Sublette, t97O: 82.Type-species, Orthocladius pleuralisMalloch (orig. des.).

aestivus. Alas.; PA.Eudacrylocladius aestivus Brundin, 1947:

30.-Sweden.astis. 8.C., Utah.

Hydrobaenus astis Roback, 1957b:IZ.-Utah.

Hydrobaenus traenis Roback, 1957b;12.-Utah.

digitatus. S. Dak., N.C., S.C.Bryophaenocladius digitatrrs Sether, 1973a:

55.-S. Dak.flavoscutellatus. Ont., N.8., Ill.

Orthocladius flavoscutellalas Malloch,1915b: 523.-lll.

fumosinus. N.Y.Camptocladius fumosinus Curran, 1930:

33.-N.Y.furcatus. N.Y.; PA.

Dactylocladius furcatus Kieffer inThienemann & Kieffer. 1916:535.-Sweden.

impectinus. N.C., S.C.Bryophaenocladius impeuinus Sather,

1976: 273.-5.C.

laticaudus. S. Dak.Bryophaenocladius I atic audus Sether,

1973a: 55.-S. Dak.pleuralis. Ill.

Orthocladius pleuralis Malloch, 1915b:527.-rll.

psilacrus. S.C.Bryophaenocladius psilacrus Sether, 1982:

500.-s.c.subparallelus. S. Dak., Ill., Ohio.

Orthocladius subparallelus Malloch, 1915b:522.-ilL.

Genus CAMPTOCLADIUS WulPCamptocladirzs WulP, 1874: 133.

Type-species, Tipula byssina Schrank(Coquillett, 1910: 518) = Tiqulastercorarius De Geer.

stercorarius. Widespread NE.; PA.; AU.Tipula stercorarius De Geet, I776:

388.-Sweden.Tipula byssina Schrank, 1803:

76.-Germany.

Genus CARDIOCLADIUS KiefferCardiocladius Kieffer, L9l2a: 22.

Type-species, ceylanicus Kieffer (orig.des.).

albiplumus. E|.C., Ont., Qu6., s. to N.C-Cardiocladius albiplumus Saether, 1969:

34.-Ont.fulvus. Mich., N.Y., Ill., Miss'

Thalassomyia fulva Johannsen, 1908:275.-N.Y.

obscurus. Ont., Qu6., Wy., Utah to N.Y-' s.to Miss. & Fla.

Thalassomyia obscura Johannsen,1903: 437.-N.Y.

Thalassomyia fusca Johannsen, \903:. 437(nom. nud.).-N.Y.

platypus. Calif., Nev., Ariz., N. Mex., ?Qu6.Orthocladius platypus Coquillett, L902b'-

93.-Ariz.

Genus CHAETOCLADIUS KiefferChaetocladius Kieffer, 191lc: 182 (as

subg. of Dacrylocladias Kieffer, 1906a).Type-species, Dacrylocladius setigerKieffer (Goetghebuer in Goetghebuer& Lenz, 1942: 57) = Chironomusperennis Meigen.

zz NEARCTTC CHIRONOMIDAE

adsimilis. Grld.Orthocladius adsimilis Goetghebuer, 1933:

25.-Grld.grandilobus. N.W.T.; PA.

Chaetocladius grandilobus Brundin, 1956:126.-Sweden.

holmgreni. N.W.T.; PA.Chironomus holmgreni Jacobson, 1898:

204 (n. nam. for Chironomus festivusHolmgren, 1869: 43, preocc. Say,1823).-Spitsbergen & Bear I.

ligni. Oreg.Chaetocladius ligni Cranston & Oliver,

1988b: 144.-Oreg.perennis. Alas., N.W.T.; PA.

Chironomus perennis Meigen, 1830:249.-Europe.

stamfordi. Qu6., Ont., S. Dak., Ohio, Conn.Hydrob aenus stamfordi Johannsen, 1947:

171.-Conn.Chaetocladitts oliveri Sether, 1969:

93'-Que'

Genus CHASMATONOTUS LoewChasmatonotas Loew, 1864: -51. Type-

species, unimaculatus Loew (mon.).

atripes. Ont. to N.B., s. to Ill., Ohio, Md.Chasmatonotus atripes Rempel, 1937 a:

2_53.-N.Y.bicolor. N.Y., Pa., Tenn.

Chasmatonotus bicolor Rempel, 1937 a:254.-Tenn.

bimaculatus. Qu€. s. to Pa., Tenn.Chasmatonotus bimaculalas Osten Sacken,

1877: 19l.-N.Y.fascipennis. B.C., Idaho, Mont., Wyo.

Chas matonotus fasc ipe nnis Coqu illett,1905: 66.-8.C.

hyalinus. Calif.Chasmatonotus hyalinus Coquillett, 190-5:

67.-Calit.maculipennis. B.C. s. to Calif.

Chasmatonotus maculipennls Rempel,1937a:. 254.-Wash.

unimaculatus. Ont. to N.S., s. to Mich., N.C.,S.C.

Chasmatonotus unimaculatus Loew, 1864:50.-N.H.

univittatus. Alas. s. to Wash., Calif.Chasmatonotus univittatus Coquillett,

1900: 395.-Alas.

Genus CLUNIO HalidayClunio Haliday, 1855: 62. Type-species,

marinus Haliday (mon.).

cal iforniensis. Calif.Clunio californiensis Hashimoto, 1.97 4:

108.-Calif.marshalli. Ky., N.C., s. to Fla.

Clunio marshalli Stone & Wirth. 1947:214.-Fla.

Genus COMPTEROSMITTIA SretherCompterosmittia Se,ther, 1981b: 20.

Type-species, dentispina Sather (orig.des.).

nerius. N.Y., S.C.Camptocladius nerius Curran, 1930:

34.-N.Y.Compterosmittia clavigera Sather, 1982:

491.-S.C.

Genus CORYNONEURA WinnertzCorynoneura Winnertz, 1846: 12.

Type-species, s cutell ata Winnertz(Coquillett, 1910: 528).

arctica. N.W.T.; PA.Corynoneura arctica Kieffer, 1923c:

4.-U.S.S.R.celeripes, Ont., Maine, N.Y., Ill., Ind., Ohio,

Grld.: PA.Corynoneura celeripes Winnertz, 18-52:

-50.-Germany.Corynoneura atra Winnertz, 1852:

50.-Germany.diara. Utah.

Corynoneura diara Roback, 1957b:10.-Utah.

fittkaui. N.C., S.C.; PA.Corynoneura littkaui Schlee, 1968:

19.-Germany.lacustris, Man., Ont.; PA.

Corynoneura lacustris Edwards,7924c: I87.-Great Britain.

lobata. Alta., Ont., N.B., N.C., S.C.;PA.

Corynoneura lobata Edwards, I924c:186.-England.

minuscula. Ont.; PA.Corynoneura minuscula Brundin,

1949: 833.-Sweden.oxfordana. Ohio.

Corynoneura oxfordana Boesel & Winner1980: -50.5.-0hio.

CHIRONOMIDAE DE I-A REGION NEARCTIQUE z-)

scutellata. N.W.T. s. to Fla., Grld.; PA.; NT;AU.

Corynoneura scutellata Winnertz, 1846:13.-Germany.

taris. Ohio, N.Y., s. to Miss. & Fla.Corynoneura /aris Roback, 1957a:

61.-N.J.

Genus CRICOTOPUS WulpCricotopus Wulp 1874: 132. Type-species,

Chironomus tibialis Meigen (Coquillett1910b: 528).

Subgenus CRICOTOPUS Wulp

abanus. B.C., Man.Cricotopus abana Curran, 1929: 1.-Man.

absurdus. N.Y., Ohio, N. Mex.Orthocladius absurdus Johannsen. 190-5:

277.-N.v.annulator. Ont., Calif., N. Mex.; PA.

Cricotopus annulator Goetghebuer, 1927:.52.-Belgium.

Cricotopus irwini Sublette & Sublette,l97l: 97.-Calif.

baptistensis. Ont.Cricotopus baptistensis Boesel, 1983:

87.-Ont.beringensis. Alas.

Cricotopus beringensis Oliver & Dillon,1988: 475.-Alas.

bicinctus. Widespread NE.; PA.; NT.; AF.Chironomus bicinctus Meigen, 1818:

41.-Austria.coronatus. N.W.T., Man., Qu6.; PA.

Cricotopus coronatus Hirvenoja, 1973:22Z.-Finland.

cumulatus. N.W.T., Grld.; PA.Cricotopus cumulatus Hirvenoja, 1973:

146.-Grld.curtus. Sask., N.Y.; PA.

Cric otopus curtus Hirvenoja, 1973:199.-Germany.

cylindraceus. Man., N.Y.; PA.Trichocladius cylindraceus Kieffer in

Kieffer & Thienemann. 1908:8.-Germany.

elegans. Mich., Pa., Ohio.Cricotopus elegans Johannsen, 1943:

78.-Mich.ephippium. Alas.; PA.

Chironomus ephippium Zetterstedt, 1838:814.-Sweden.

Ceratopogon humeralis Zetterstedt, 1838:820.-Sweden.

festivellus. Man., Ont.; PA.Chironomus festivellus Kieffer, 1906a: 18

(n. nam. for Chironomus festivusMeigen, 1830:. 252, Preocc. SaY,1823).-Europe.

flavocinctus. Man.; PA.Tricho cl adius fl av o c inc tus Kieffer,

1924b: 92.-Germany.fugax. Ont., N.Y., Ky., N.C., s. to Miss. &

Fla.Orthocladius fugax Johannsen, 1905:

279.-N.Y.fuscus. Ont.; PA.

Trichocladius fuscus Kieffer, 1909:46.-Germany.

gelidus. N.W.T., Grld.; PA.Trichocladius gelidus Kieffer, 1922b:

18.-Novaya Zemlya.infuscatus. Ont., S. Dak., Ill., Ohio, Pa., N.

Mex.Orthocladius infuscatus Malloch, 1915b:

517.-Ill.Cricotopus aratus Roback, 1957a: 73.-Pa.Cricotopus ceris Roback, 7957a: 72.-Pa.

lestralis. Alas., N.W.T., Wash., Grld.; PA.Trichocladius lestralis Edwards, I924b:

172.-Spitsbergen.luciae. Ont.

Cricotopus luciae LeSage in LeSage &Harrison, 1980: 91.-Ont.

mackenziensis. N.W.T.Cricotopus mackenziensis Oliver, 1977 :

99.-N.W.T.macraei. Man.

Cricotopus macraei Srether, 1971:1799.-Man.

Cricotopus flannagani Srether, 1971:1809.-Man.

patens. Y.T., N.W.T.; PA.Cricotoptts patens Hirvenoja, 1973:

218.-Finland.pedatus. Mont., N. Mex.

Cricotopus pedatus Sublette, 1967b:.558.-Mont.

pilosellus. Y.T., N.W.T., Grld.; PA.Cricotopus pilosellus Brundin, 1956:

114.-Sweden.politus. Sask., Colo:, Ill., Ohio, N.Y., D.C.,

Md., N. Mex. to Fla.Orthocladius politus Coquillett, 1902b:

93.-D.C.pulchripes. N.Y.; PA.

Cric otopus pulchripe s Y errall, 19 12:22--Wales.

aA NEARCTIC CHIRONOMIDAE

slossonae. Sask. to Ont., s. to Ill., N.Y.,Ohio, Ga.

Cricotopus slossonae Malloch, 1915b:506.-Iil.

tibialis. Y.T., N.W.T., Man., N.B., Grld.; PA.Chironomus tibialis Meigen, 1804:

16.-Europe.Chironomus basalis Staeger, 184-5:

351.-Grld.Chironomus pavidus Holmgren, 1869:

42.-Spitsbergen.Cricotopus polaris Kieffer, 1926:

84.-Grld.Trichocladius b acilliger Kieffer, 1926:

86.-Grld.tremulus. Alta. to N.S., Oreg., Ohio, N.Y.,

N'J'; Pa'Tipula tremula Linnaeus, 17-58:

587.-Sweden.triannulatus. Sask., Ont. to N.Y., s. to Miss.

& Fla.; PA.Chironomus lriannulatus Macquart, 1826:

202.-Germany.Cricotopus exjlis Johannsen, 1905:

2.5-5.-N.Y.trifascia. Sask., Ont., Ohio, Pa., N.Y., N.C.;

PA.Cricotopus trifascia Edwards, 1929:

322.-England.Cricotopus ithacanensis Sublette, 1967b:

555.-N.Y.trilobus. Alas.

Cricotopus trilobus Oliver & Dillon, 1988:485.-Alas.

tristis. Y.T., N.W.T.; PA.Cricotopus /rislis Flirvenoja, 1973;

196.-Finland.varipes. N.W.T., Ont., Wash., Ohio, N.Y.,

Conn., Ga., Fla.Cricotopus varipes Coquillett, l9l2b:

93.-Md.vierriensis. Man., Ohio, Pa., N.Y., N. Mex.;

PA.Cricotopus vierriensis Goetghebuer, 1935:

417.-Belgium.

Subgenus ISOCLADIUS KiefferIsocladius Kieffer, I9O9: 44 (as genus).

Type-species, albipes Kieffer (mon.) =Tipula silvestris Fabricius.

flavibasis. B.C., Alta., S. Dak., Ill.Cricotopus flavibasis Malloch, 1915b:

502.-Iil.flavipes. Ohio, Mich., Conn.

Cricotopus flavipes Johannsen, 1942:73.-Mich.

glacialis. Not NE.Cricotopus glacialis Edwards, 7922:

209.-Spitsbergen.intersectus. Sask., Man., Ohio; PA.

Chironomus intersectus Staeger, 1839:574.-Denmark.

laetus. Ont.: PA.Cricotopus laetus Hirvenoja, 1,973:

264.-Finland.laricomalis. Alas., N.W.T., Qu6., Grld.; PA.

Cricotopus laricomalis Edwards, 1932a:46.-Scotland.

myriophylli. B.C., Ont.Cricotopus myriophylli OIiver, 1984:

L287.-8.C.ornatus. Alas. to N.W.T., Sask., Man., Ont.,

N.Y.; PA.Chironomus ornatus Meigen, 1818:

43.-Germany.pilitarsis. Sask., N.Y.; PA.

Chironomus pilitarsis Zetterstedt, 18-50:3-56-5.-Sweden.

sylvestris. Widespread NE.; PA.Tipula sylvestris Fabricius, 1794:

252.-Etrope.tricinctus. Utah to N.Y., s. to Tex.; PA.

Chironomus tricinctus Meigen, 1818:41.-Europe.

Cricotopus lebetis Sublette, t964b:118.-La.

trifasciatus. 8.C., Ont., Idaho to N.Y., s. toN. Mex. & Fla.; PA.

Chironomus trifasciatus Meigen in Panzer,1813: l8.-Germany.

Cricotopus remus Sublette, 1,964b:11-5.-La.

Subgenus NOSTOCOCLADIUS Ashe &Murray

Nostococladias Ashe & Murray, 1980:105. Type-species, lygropis Edwards(orig. des.).

nostocicola. Alta., Mont., N.Y., Ohio s. toGa., Oreg., Calif.

Cricotopus nostocicola Wirth, 1957:722.-Calif.

Subgenus PSEUDOCRICOTOPUS NishidaPseudocricoropus Nishida, 1987'. 460 (as

subg. of Cricotopus Wulp).Type-species, montanus Tokunaga(orig. des.).

bifurcatus, 8.C., Wash.Cricotopus bifurcatus Cranston & Oliver,

1988a: 430.-B.C.


Cricotopus unassigned to subgenus

belkini. N. Mex. to Ala.Cricotopus belkini Dendy & Sublette,

1959:51O.-Ala.brunnicans. Sask.

Cricotopus brunnicans Walley, L928a:21.-Sask.

diversus. Ont., Ohio., Mich., N.Y., Del.Cricotopus diversus Boesel, 1983:

85.-Ohio.eilurus. Calif.

Cricotopus edurus Sublette & Sublette,1971: 85.-Calif.

fuscatus. Calif.Cricotopus fuscatus Wirth, 1957:

t24.-Calif.furtivus. Calif.

Cricotopus furtivus Sublette & Sublette,L97L: 99.-Calif.

geminatus. Pa.Chironomus geminatus Say, 1823: 14.-Pa

globistylus. Utah, N. Mex.Cricotopus globistylus Roback, I957b:

10.-Utah.junus. Pa.

Cricotopus jznr.rs Roback, 1957a; 72.-Pa.obscurifuscus. Calif.

Cricotopus obscurifuscus Sublette &Sublette, I97 l: 96.-Calif.

oceanicus. Mass.Chironomus oceanicus Packard, 1869:

42.-Mass.olivetus. Ohio.

Cricotopus olivetus Boesel, 1983:88.-Ohio.

parafuscatus. Calif.Cricotopus parafuscatus Sublette &

Sublette, 1971: 9O.-Calif.subfuscus. Calif.

Cricotopus subfuscus Sublette & Sublette,1971: 98.-Calif.

Genus DIPLOCLADIUS KiefferDiplocladius Kieffer in Kieffer &

Thienemann, 1908: 6. Type-species,cultriger Kieffer (mon.).

cultriger. Widespread Can. to Nebr., Ohio,N.C., S.C., Grld.; PA.

Diplocladius cultiger Kieffer in Kieffer &Thienemann, 1908: 6.-Germany

Diplocladius bilobatus Brundin, L956:71.-Sweden.

Genus DIPLOSMITTIA SetherDiplosmittia Sather, I98lb: 29-

Type-species, hatisoni Srether (orig.des.).

carinata. Mich.Diplosmittia carinata Sether, 1985h:

55.-Mich.

Genus DOITHRIX Saether & SubletteDoithrix Srether & Sublette, 1983: 6.

Type-species, villosa Sether & Sublette(orig. des.).

barberi. Calif.Doithrix barberi Sether & Sublette, 1983:

2I.-Calif.dillonae. N.S.

Doithrix dillonae Cranston & Oliver,1988a: 432.-N.S.

ensifer. Calif.Doithrix ensifer Sether & Sublette, 1983:

15.-Calif.hamiltoni. B.C.

Doithrk hamiltoni Sather & Sublette,1983: 17.-B.C.

parcivillosa. Tenn.Doithrix parcivillosa Sether & Sublette,

1983: 12.-Tenn.vitlosa. Man., N.8., N.S., S.C., Tenn.

Doithrit villosa Sather & Sublette, 1983:9.-Tenn.

Genus DONCRICOTOPUS SatherDoncricotopas Srether, l98la:. 223.

Type-species, bicaudatus Saether (orig.des.).

bicaudatus. N.W.T.Doncricotopus bicaudatus Srether, 1981a:

225.-N.W.T.

Genus EPOICOCLADIUS ZavrelEpoicocladius Zavrel in Sulc & Zavrel,

1,924: 368 (as subg. of CamptocladiusWulp, 1874). Type-species,Camptocladius ephemerae Kieffer (orig.des.) - Camptocladius flavens Malloch.

flavens. Ont., Mont., N. Dak., s. to Tex.,Mich., Ill., s. to Fla.; PA.

Camptocladius f-lavens Malloch, 191-5b:511.-Ill.


Camptocladius ephemerae Kieffer in Sulc illoba. Calif.& Zavrel, 1924: 385.-Czechoslovakia. Euryhapsis illoba Oliver, 1981: 720.-Calif.

Genus ERETMOPTERA Kellogg Genus GEORTHOCLADIUS StrenzkeEretmoptera Kellogg, 1900: 82. Georthocladias Strenzke, 1941: 185.

Type-species, browni Kellogg (orig. Type-species, Orthocladius luteicornisdes.). Goetghebuer (orig. des.).

browni. Calif. Subgenus ATELOPODELLA SetherEretmoptera browni Kellogg, 1900: Atelopodella Sather, 7982: 488.

82.-Calif. Type-species, curlicornus Sether (orig.des.).

Genus EUKIEFFERIELLA Thienemann curticornus. S.C.Eukiefferiella Thienemann, 1926: 325. Georthocladius curticornrzs Sether, 1982:

Type-species, Dactylocladius longicalcar 488.-5.C.Kieffer, 1.9llc, sensu Thienemann(misident. = Spaniotoma gracei Subgenus GEORTIIOCLADIUS StrenzkeEdwards, 1929), (orig. des.) (ICZNruling required on the true type - fimbriosus. Tenn.species (ICZN 70a)). Georthocladius fimbriosus Sether &

Adactylocladizrs Sether, 1,968: 464. Sublette, 1983: 32.-Tenn.Type-species,finsensis Sether (mon.) platystylus. Calif.= Spaniotoma minor Edwards. Georthocladius platystylus Sether &

Sublette, 1983: 28.-Calif.brevinervis. Ill., Ohio, Pa., N. Mex. triquetrus. S.C.

Orthocladius brevinervis Malloch, 1915b: Georthocladiu.s tiquetus Seether &526.-Ill. Sublette, 1983: 25.-S.C.

claripennis. N.W.T., B.C., Alta., N. Mex., wirthi. Calif.Grld.; PA.; Hawaii. Georthocladius wirthi Sather & Sublette,

Chironomus claripennis Lundbeck, 1898: 1983: 30.-Calif.281.-Grld.

Spaniotoma hospita Edwards, 1929:351.-England. Genus GYMNOMETRIOCNEMUS

cyanea. Alas., ?Oreg.; PA. GoetghebuerEukiefferiella cyanea Thienemann, 1936b: Gymnometriocnemus Goetghebuer, 1932a:

61.-Germany. 23 (as subg. of Metriocnemus Wulp,devonica. Widespread Can., U.S.A.; PA. 1874). Type-species, Metriocnemus

Spaniotoma devonica Edwards, 1929: subnudus Edwards (Edwards, 7932b:349.-England. 141). (Technically invalid as no

unicalcar. Alta. type-species originally designatedAdactylocladius unicalcar Sather, 1969: (ICZN 13b), but treated as above by

59.-Alta. usage).

. Subgenus GYMNOMETRIOCNEMUSGenus EURYHAPSIS Oliver Goetghebuer

Euryhapsis Oliver, 7981: 7ll.Type-species, cilium Oliver (orig. des.). subnudus. B.C., Ont., N.S., Tenn., N.C.; PA.

Metiocnemus subnudus Edwards, 1929:annuliventris. Calif. 316.-England.

Metriocnemeus annulivenrrls Malloch, Gymnometriocnemus marionensis Sather,191-5a: 46.-Calif. 1969: 729.-8.C.

cilium. Y.T., N.W.T., Alta.; PA.Euryhapsis cilium Oliver, 1981:

714.-N.W.T.

CHIRONOMIDAE DE I-A REGION NEARCNQUE 27

Subgenus RAPIIIDOCLADIUS SetherRaphidocladizs Srether, 1983b: 211.

Type-species, acigus Sether (orig. des.)= Metriocnemus brumalis Edwards.

brumalis. 8.C., Man. to N.S., Tenn., S.C.;PA.

Metriocnemus brumalis Edwards,1929: 316.-England.

Gymnometriocnemus acigus Snther, 1983b:214.-S.C.

Genus HALOCLADIUS HirvenojaHalocladius Hirvenoja, 1973: 106.

Type-species, Chironomus variansStaeger (orig. des.).

Subgenus HALOCLADIUS Hirvenoja

variabilis. Man., Grld.; PA.Chironomus variabilis Staeger, 1839:

571.-Denmark.

Nomen dubium in IlalocladiusChironomus vitripennis Meigen, 1818:

32.-Europe.

Genus HELENIELLA GowinHeleniella Gowin. 1943: 116.

Type-species, thienemanni Gowin (orig.des.). = Spaniotoma ornaticollisE,dwards.

curtistila. B.C., Alta., Ont.Heleniella curtistila Sether, 1969:

130.-8.C.hirta. Ont., Qud., N.S.

Heleniella hirta Snther. 1969: 134.-Qu6.parva. S.C., Tenn.

Heleniella parva Sather, 1985b: -532.-5.C.

Genus HETEROTANYTARSUS SplirckHeterotanytarsas Spdrck, t923: 92.

Type-species, Metiocnemus apicalisKieffer (mon.).

nudalus. Ont.Heterotanytarsus nudalus Sether, 1975b:

263.-Ont.perennis. B.C.

Heterotanytarsus perennis Srether, I975b:260.-B.C.

Genus HETEROTRISSOCLADIUS SpdrckHeterotris socl adius SpAr ck,'1.923: 94.

Type-species, Metriocnemus cubitalisKieffer (Goetghebuer in Goetghebuer& Lenz,1940: 6) ?= Chironomusmarcidus Walker.

changi. N.W.T., Alta. to Ont., N.Y., s. to Ill.,Ind.

Heterotrissocladius changi Srether, 197 5a:47.-Man.

cooki. Minn.Heterotrissocladius cooki Sether, I97 5a:.

41.-Minn.hirtapex. Ont.

Heterotrissocladius hirtape-x Sather, 1975a;39.-Ont.

latilaminus. 8.C., Mont.Heterotrissocladius latilaminas Sather,

1975a: 36.-8.C.maeaeri. Alas.: PA.

Heterotrissocladius mriiiri Brundin, L949:812.-Sweden.

marcidus, B.C., Ont., Qu6., N.Y., Mich.,N.C., S.C., Fla.; PA.

Chironomus marcidus Walker, 1'956a;177.-England.

oliveri. N.W.T., B.C. to Ont., s. to Calif.,N.y.: pA.

Heterotrissocladius oliveri Sether, I975a.:19.-N.W.T.

subpilosus. N.W.T.; PA.Dactylocladius subpilosus Kieffer in

Kieffer & Lundbeck. 1911: 273.-BearIsland.

Genus HYDROBAENUS FriesHydrobaenus Fries, 1830: 177.

Type-species, lugubris Fries (orig. des.).

calvescens. S. Dak.Hydrob aenus c alvescens Saether, 1976:

158.-5. Dak.conformis. N.W.T., Qu6., Labr., Grld.; PA.

Chironomus conformis Holmgren, 1869:42.-Spitsbergen.

Hydrobaenus conformis conformisHolmgren (Srether, 1976: 83).

Hydrobaenus conformislabradorensis Sether. 1976:89.-Qu6., Labr.

Orthocladius natvigi Goetghebuer, 1933:25.-Franz Joseoh Land.


fusistylus. Alas., N.W.T., Alta., Mont., Colo.,Grld.

Orthocladius fusistylus Goetghebuer, 1933:26.-Grld.

Hydrobaenus fusistylus var. fusistylus(Goetghebuer), Sather, 7976:- 94.

Hydrobaenus fusistylus var. octomerusSrether, 7976: 99.

hudsoni. S. Dak.Hydrobaenus hudsoni Sether, 1976:

131.-S. Dak.johannseni. Man., Ont., S. Dak., Minn., Ohio,

N.Y., Md., Ark., N.C., S.C.Orthocladius johannseni Sublette, 1967b:

504 (n. nam. for Orthocladiusnivoriundus Johannsen, 1934: 348,preocc. Fitch, 1847).-N.Y.

Orthocladius domus Sublette, 1966:594.-Md.

Trissocladius hamiltoni Sether, 1969:42.-Man.

laticaudus. Alas.Hydrobaenus laticaudus Sether, 1976:

80.-Alas.martini. N.W.T., N.B.; PA.

Hydrobaenus martini Sether, 1976:134.-N.W.T.

pilipes. Ont., Qu€., S. Dak., Ohio, Ill., Calif.,N. Mex., N.C., S.C., Fla.

Orthocladius pilipes Malloch, 1915b:522.-\1.

pilipodex. S. Dak., Ark., Ala.Hydrobaenus pilipodex Sether, 1976:

144.-5. Dak.scapulapilosus. Alas., Ont.

Hydrobaenus scapulapilosus Sether, L976:124.-Ont.

spinnatis. Ont., Qu6.; PA.Hydrobaenus spinnatis Sather, 1976:

152.-Qu€.travisi. Mass.

Hydrobaenus travisi Sa.ther, 1989b:55.-Mass.

tumidistylus. N.W.T.Hydrobaenus tumidistylus Sether, 1976:

160.-N.W.T.virgo. Ont.

Hydrobaenus virgo Sather, L976:128.-Ont.

Genus KRENOSMITTIA Thienemann &Krtiger

Krenosmittia Thienemann & KrUger, L939:253. Type-species, Smittia gynoceraEdwards, 1937, sensu Thienemann &Krriger (misident. = Spaniotoma

camptophleps Edwards, 1,929) (orig.des.).

?boreoalpina. B.C., Alta.; PA.Smittia boreoalpina Goetghebuer, 1944:

43.-Austria.?camptophleps. B.C.; PA.

Spaniotoma camptophleps Edwards, 1929:353.-Great Britain.

halvorseni. N.W.T., N.B.; PA.Rheosmittia halvorseni Cranston & Sather.

1986: 45.-Norwav.

Genus LIMNOPHYES EatonLimnophyes Eaton, 1875: 60.

Type-species, pusillus Eaton (mon.).

asquamatus. N.Y., Grld.; PA.Limnophyes eltoni var. asquamatus

Andersen, 1937 : 7 Z.-Grld.atomarius. N.Y., Grld.; PA.

Chironomus atomarius Zetterstedt, 1850:3522.-Sweden.

borealis. Y.T. to Grld., N.8., s. to Minn.,Tenn., N.C., S.C.; PA.

Limnophyes borealis Goetghebuer, 1933:29.-Grld.

Limnophyes spatulosus Srether, t975d:1055.-Minn.

carolinensis. Ont., Minn., Tenn., N.C., S.C.Limnophyes carolinensis Srether, I97 5d:

10-50.-s.c.coloradensis. Colo.

Limnophyes coloradensis Sether, 197 5d:1052.-Colo.

eltoni. Grld.: PA.Camptocladius eltoni Edwards, L922:

203.-Bear Island.fumosus. Ont., N.S., N.Y., N.C., S.C., Ga.

Camptocladius fumosus Johannsen, 1905:261.-N.Y.

Limnophyes cristatissimus Sether, 7975d:1032.-S.C.

globifer. Y.T., N.W.T., Alta., Man., Ont.,Grld.: PA.

Camptocladius globifer LundstrOm, 1915:16.-U.S.S.R.

Limnophyes folliculatus Sether, L97 5d:1050.-Man.

groenlandiensis. Grld.Limnophy e s gr o e nl and ie nsis Andersen,

1937. 70.-Grld.hamiltoni. Alas. to N.W.T., B.C., Ont., Calif.,

Grld.Limnophyes hamiltoni Saether, 1969:

101.-B.C.


hastulatus. B.C., Ont., S. Dak., Minn.Limnophyes hastulatus Saether, 1975d.

1054.-S. Dak.immucronatus. Widespread Can. s. to N.

Mex., Nebr., N.C., S.C.; ?PA.Limnophyes immucronatus Sather, 1969:

103.-8.C.Limnophyes hudsoni Sether, 1975d:

1032.-Nebr.margaretae. Man.

Limnophyes margaretae Srether, 797 5d:1053.-Man.

minimus. Idaho, Colo., Ohio, N.Y.; PA.Chironomus minimus Meigen, 1818:

47.-Europe.ninae. N.W"T., Man., Ont.

Limnophyes ninae Snther, 1975d:1048.-Man.

nudiradius. Man., S. Dak.Limnophyes nudiradius Srether, 197 5d:.

1048.-5. Dak.pilicistulus. Y.T., B.C., Oreg., S. Dak.

Limnophyes pilicistulus Sether, 197 5d:1049.-5. Dak.

prolongatus. Y.T., N.W.T., Man., Grld.; PA.C amptocladius pentaplastus v at.

prolongatus Kieffer in Thienemann,1921: 810.-Germany.

pumilio. Grld.; PA.Chironomus pumilio Holmgren, 1869:

41.-Spitsbergen.recisus. S. Dak., Nebr., N. Mex.

Limnophyes recisus Srether, I975d:10-52.-Nebr.

Genus LIPUROMETRIOCNEMUS SetherLipurometriocnemus Sether, 1981b: 13.

Type-species, glabalus Seether (orig.des.).

vixlobatus. Y.T., S.C.Lipur ome triocnemus vixl ob atus Srther,

1982:477.-5.C.

Genus LOPESCLADIUS OliveiraLopescladius Oliveira, 1967: 4I7.

Type-species, minutis s imus Oliveira(orig. des.).

Cordites Brundin, 1966: 428 (nom. nud.).

Subgenus CORDIELLA Coffman & RobackCordiella Coffman & Roback, 1984: 130.

Type-species, hyporheicus Coffman &Roback (orig. des.).

hyporheicus. Pa., Va.Lopescladius hyporheicus Coffman &

Roback, 7984: l3l.-Pa.26

Subgenus LOPESCLADIUS Oliveira

inermis. Kans.Lopescladius inermis Srether, 1983c:

293.-Kans.verruculosus. Ill., Mex.; NT.

Lopes

Documents

A catalog of Nearctic Chironomidaeesc-sec.ca/wp/wp-content/uploads/2017/03/AAFC_catalog_of_nearcti… · Nanocladius Kieffer, 1913d: 31. Type-species, vitellinus Kieffer (orig. des.)