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THE COLEOPTERISTS BULLETIN 42(2), 1988

MCTI-Museo Civico di Storia Naturale, Trieste, Italy; MNHN-Museum National #Histoire Naturelle, Paris; MRAC- Musee Royal de 1'Afrique Cen- trale, Tervuren, Belgium; NMBZ-National Museum, Bulawayo, Zimbabwe; NMPC-National Museum, Prague; NMWA-Naturhistorisches Museum, Vi- enna; SAMS - South African Museum, Cape Town; SANC- South African National Collection of Insects, Pretoria; and TMSA-Transvaal Museum, Pre- toria.

TYPE-SPECIES. Discoderes stevensoni ThCry.

DESCRIFTION. Size, less than 7.5 mm in length; flattened, elongate; surface black, rugose or striolate, setose.

Head (Fig. 12) produced between eyes (e.g., Fig. 10); eyes moderately large, diverging dorsally, completely bordered by circumocular groove which is confluent with supra- antenna1 groove; epistoma emarginate distally, laterally depressed to receive basal antennomeres in repose; genal projection obtusely rounded; labrum wider than long, co- riaceous, distally setose. Antennae serrate from antennomere 5.

Pronotum wider than long, widest near middle; anterior margin medially convex; basal margin bisinuate; lateral margins arcuate; laterobasal angles obtusely rounded; one prelateral carina on either side; disc slightly depressed past middle. Scutellum triangular.

Elytra narrower than pronotum at base, as wide as pronotum at apical '13; depressed between humeri and scutellum; lateral margins bisinuate; apices separately rounded. Pygidium with acuminate apical spine.

Thoracic sternites with prosternum with broadly truncate mentonnihe, laterally with propleural depressions; process (Fig. 17) with sides subparallel between procoxae to angulate lateroapical lobes, apex laterally truncate, medially acuminate; mesosternum (Fig. 17) partially hidden beneath prosternal process; metacoxal plates with anterior and posterior margins parallel in median '/2.

Legs with femora feebly fusiform; protibia as in Figure 13; metatibia as in Figure 14. Abdominal sternites with suture between 1 and 2 only vaguely indicated; sutures

between 2, 3, 4 and 5 more or less evenly transverse. Ovipositor with two dense ventral setose "brushes." Metathoracic wing as in Figure 18.

REMARKS. The name Discoderella is feminine and is derived from the fact that the species closely resemble those of Discoderes. The very obvious and broadly produced mentonniere immediately separates Discoderella from Dis- coderes Chevrolat and was also the main character that Th6ry used for erecting another related genus, Discoderoides. Discoderoides can be separated from Dis- coderella by differences in the wing venation and also by the strongly flattened tibiae in most species (see Bellamy 1986a). The following tabulation will serve to clearly separate Discoderella from the look-alike species of Discoderes. Dis- coderes salzrnanni (Solier) is illustrated (Figs. 1-9) for comparison.

Discoderes Discoderella, gen. nov. -

Antennomeres 5-1 0 subequally serrate.

Eyes (Fig. 3) moderately large, convex, mostly visible from above.

Lateral margins of pronotum (Fig. 1) evenly arcuate.

Prosternum (Fig. 2) without mentonniere.

Antennomere 5 subserrate, 6-1 0 subequally serrate.

Eyes (Fig. 12) small, flattened, only vaguely visible from above.

Basolateral margins of pronotum (Fig. 10) with notch.

With mentonniere (Fig. 1 1).

THE COLEOPTERISTS BULLETIN 42(2), 1988 175

Prosternal process (Fig. 4) evenly at- Prosternal process (Fig. 17) with angulate la- tenuate, with small lateral lobes. teroapical lobes, apex with median acu-

minate tooth. Mesepimeron (Fig. 2) strongly swol- Mesepimeron (Fig. 11) not swollen laterally,

len laterally, partially visible from only very slightly visible from above. above.

Northern subsaharan Africa. Southeastern Africa.

The type-species, D. stevensoni, was quite adequately described originally (ThCry 1932: 167) and will only be treated diagnostically to allow a clear sep- aration from D. dilaticollis, sp. nov.

Discoderella stevensoni (ThCry), comb. nov. Figures 10- 1 5

Discoderes stevensoni ThCry 1 932: 1 67; Obenberger 1 934:800.

DESCRIPTION. Size, 5.7-7.2 x 1.9-2.4 mm; surface rugose, black, covered with small widely spaced white setose patches. Head (Fig. 12) produced to subtruncate apex between eyes. Pronoturn 1.25 x wider than long. Male genitalia as in Figure 15.

MATER~AL EXAMINED. Paratypes, 1 male, 1 female (NMBZ): (ZIMBABWE) Penkridge, N. Melsetter Dist., S. Rhodesia, 24.12.1928, R. H. R. Stevenson; 1 female (CLBC): Umtali, S. Rh(odesia), XII. 1932, P. A. Shepard. REMARKS. The original placement of this taxon in Discoderes by ThCry can

be understood because of the very close superficial resemblance to D. salzmanni. This species can be separated from D. dilaticollis, sp. nov., as discussed below.

Discoderella dilaticollis Bellamy, sp. nov. Figures 1 6- 1 8

DESCRIPTION. Size, 7.1 x 2.4 mm; elongate, flattened; surface black, irregularly striolate; sparsely setose on head, pronotum and underside; elytra with slightly denser patches or zig zag fasciae.

Head produced between eyes, with vertex longitudinally grooved, thus bilobed; eyes moderately large, feebly diverging above, disc feebly convex, completely margined by circumocular groove; frons with transverse supra-antenna1 groove confluent with ocular grooves laterally, medially with depressed base of epistoma; epistoma constricted slightly between antennae, apex bisinuately emarginate; gena depressed below eye for antenna in repose, apex with rounded obtuse projection; labrum only slightly visible, sparsely setose distally. Antennae with antennomere 1 geniculate, slightly swollen; 2 longer than 3 or 4; 3 and 4 subequal; 5-10 serrate; 1 1 oblong.

Pronoturn 1.34 x wider than long, widest near middle; anterior margin convex medially; basal margin bisinuate, prescutellar lobe truncate, lateral angles obtusely rounded; lateral margins sinuate past basal angle then broadly arcuate to apex; disc with transverse groove medially just past middle; one arcuate carina before lateral margins on middle l/2; slightly swollen before transverse groove with pair of concentrically striolate rounded feeble elevations. Scutellurn triangular, length subequal to width.

Elytra with base depressed between humeri and scutellum; narrower than pronotum at base, equal to pronotal width at apical l/3; lateral margins subparallel to past humeri, narrowing to middle before widening to apical Y3, then gradually narrowing to finely serrate and separately rounded apices; sutural carinae slightly elevated on apical '/3; pygidium with short acuminate apical spine, partially visible between elytral apices.

Undersidewith prosternum with broad truncate mentonnikre, laterally with propleural depressions, disc flattened, sides compressed between broad convex hypomera; prosternal process with sides subparallel between procoxae to obtuse lateral angles, apex lateroap-


Figs. 1-9. Discoderes salzmanni (Solier). 1, left dorsal view. 2, lateral view. 3, head, frontal view. 4, sternal cavity, ventral view. 5, protibia, ventral view. 6, metatibia, ventral view. 7, metatarsus, distal tarsomeres, ventral view. 8, metathoracic wing, dorsal view. 9, male genitalia, dorsal view.

Figs. 10-1 5. Discodorella stevensoni (Thiry). 10, left dorsal view. 1 1, lateral view. 12, head, frontal view. 13, protibia, ventral view. 14, metatibia, ventral view. 15, male genitalia, dorsal view.


ically truncate with narrow median acuminate tooth; metacoxal plate with anterior and posterior margins subparallel in median l/2, posterior margin dilated on posterolateral angle; basal abdominal sternites laterally expanded, visible from above; sutures between sternites 2, 3 ,4 and 5 more or less transverse; apex of 5 subtruncate, disc with moderate concentration of recumbent testaceous setae.

MATERIAL EXAMINED. Holotype, female (NMPC 26804): Africa mer[idional]. / 13H [red].

REMARKS. The name is the genitive form of the Latin prefix dilat (expand) and suffix coll (neck) for the broad pronotum. Discoderella dilaticollis can immediately be distinguished from D. stevensoni by the wider pronotum and by differences in the setal pattern.

TYPE-SPECIES. Paraxenita tesselata, sp. nov.

DESCRIIYTION. Size less than 9.0 mm in length; elongate; flattened above. Head (Fig. 2 1) feebly longitudinally grooved, frons narrowed between eyes; eyes large,

convex, diverging dorsally; circumocular grooves almost entire, not on genae, confluent with supra-antenna1 groove; frontoclypeus reduced to single carina between antennae, distal margin broadly, shallowly, triangularly emarginate; antenna1 cavities large, deeply excavated, confluent with transverse genal depression beneath eyes; labrum mostly hidden, bilobed, sparsely setose apically; mandibles distally emarginate, with slight arcuate- acuminate tooth. Antennae (Fig. 22) with antennomere 2 elongate, subglobose, as long as 3 and 4 together, 5 subserrate, 6 serrate, 7-10 with lateral serrate portion prolonged; 1 1 oblong, curved.

Pronotum wider than long, widest at middle; anterior margin with slightly convex lobe medially; base broadly arcuate; lateral margins notched at base, subparallel on middle l/2, arcuate to apex; basal angles acute; prelateral carinae, one on either side, slightly arcuate on middle l/2; disc slightly depressed medially and in each laterobasal l/3.

Scutellum triangular, longer than wide, disc depressed. Elytra narrower than pronotum, widest opposite humeri; base depressed between

humeri and suture; lateral margin subparallel to past humeri, then broadly concave to apical Y3, widening slightly before gradually narrowing to broadly separately rounded apices; disc with very feeble longitudinal costa separating depressed sutural l/2 from feebly declivous lateral l/2. Pygidium slightly emarginate around projecting apical spine.

Thoracic sternites with prosternum with mentonni6re entire, feebly convex, disc slightly compressed, process with sides subparallel to triangularly acuminate apex; mesepimeron partially visible from above; metepimeron partially covered by lateral projection of abdominal sternite 1; metacoxal plate longest laterally, distal margin bisinuate, outer apical angle roundly acute, produced into slightly projecting lobe.

Legs with femora narrowly fusiform; tibiae straight, with external edge carinate, explanate before margin; protibiae with one short, slightly curved distal spine; metatibiae (Fig. 23) with setal comb on external margin; tarsomeres 1 4 subequal, each with ventral pulvilli, 4 with pulvillus nearly 2 x larger than on 1-3; 5 as long as 1-4 together, claws broadly appendiculate internally, external portion recurved.

Abdominal sternites with suture between sternites 1 and 2 feebly indicated, evenly transverse laterally, anteriorly convex medially; basal sternites laterally expanded, partially visible from above; remaining sutures transverse; lateral angles obtuse; sternite 5 broadly rounded, with transverse latitudinally carinate plate in female.

Metathoracic wings with venation as in Xenita (see Bellamy 1986b:fig. 54).

C

Figs. 16-1 8. Discodorella dilaticollis, sp. nov. 16, left dorsal view. 17, sternal cavity, ventral view. 18, metathoracic wing, dorsal view.

178 THE COLEOPTERISTS BULLETIN 42(2), 1988

REMARKS. The generic name is designated as feminine and is named for being "close" to Xenita ThCry and can be separated as follows.

Xenita Paraxenita, gen. nov.

Prosternal process with lateral lobes. Prosternal process acuminate. Abdominal sternite 5 not sexually Abdominal sternite 5 sexually dimorphic, fe-

dimorphic. male with transverse carinate plate. Lateral margins of pronotum evenly Pronotum emarginate basolaterally, basal an-

arcuate, basal angles obtuse. gles acute. Kenya, Tanzania. Cameroun.

The only species in Paraxenita is described below with only a morphomet- rical and specific diagnosis used to augment the generic description.

Paraxenita tesselata Bellamy, sp. nov. Figures 1 9-23

DESCRIPTION. Holotype female. Size, 8.1 x 2.5 mm; colour nitid black. Head with vertex with sparse short black squamiform setae; frons sparsely covered with adpressed white setae between eyes. Pronotum 1.5 x wider than long; surface imbricate, covered with setae as in Figure 19, with dark areas having white recurved setae, light areas with sparse short black squamiform setae. Elytra 2.5 x longer than wide; setal pattern tesselate (Fig. 19) with light and dark areas as pronotum. Underside moderately covered with short adpressed white setae.

MATERIAL EXAMINED. Holotype, female (NMPC 26798): Cameroon S. E. [white labe1]/26H [red label].

REMARKS. The species name is from the Latin tesselat, for "checkered" in reference to the black and white setal pattern on the elytra.

TYPE-SPECIES. Katangiella squamivela, sp. nov.

DESCRIPTION. Size less than 9.0 mm in length; elongate; flattened above, convex below; with concentrations of squamose setae on frons, in large basolateral pronotal depressions and more sparsely, yet generally, on entire ventral surface.

Head (Fig. 26) angularly produced between eyes, apex broadly truncate; eyes moderate, not touching pronotum behind, inner margins moderately diverging dorsally; circumocular groove entire around perimeter of eye, confluent with supra-antenna1 trans- frontal groove, this confluent with medial frontoclypeal depression; frontoclypeus with slightly elevated subparallel lateral carinae between antennae, apex broadly, shallowly emarginate; antenna1 cavities large, separated by slightly larger distance than their width; gena slightly depressed beneath eye for reception of basal antennomeres in repose, margin broadly obtuse; labrum wider than long, with sparse elongate feebly curved setae distally; mandibles recessed within oral cavity, attenuate distally to subtruncate apices; maxillary palpi with segment 3 subequal to 1, each longer than 2. Antennae with antennomeres 5- 10 serrate, 1 1 appendiculate.

Pronotum wider than long, widest before middle; anterior margin strongly convex medially; basal margin bisinuate with median subtruncate prescutellar lobe; lateral margins concave to widest point, then broadly arcuate to anterior margin, carinate from base to apex; basal angles obtuse; disc irregularly depressed. Scutellum small, triangular, length subequal to width, disc slightly depressed.

Elytra narrower than pronotum, widest opposite humeri; base slightly depressed between humeri and suture; humeri moderately elevated, short, oblique; epipleural fold not separated from disc by carina; lateral margins narrowing past humeri, then widening


to before apical l/3 before steeply narrowing to attenuate, separately rounded apices; disc rugose with transverse setal fasciae. Pygidium with apex emarginate around projecting median spine.

Thoracic sternites with prosternum lacking mentonni&re, slightly compressed, disc convex; process with sides subparallel, slightly narrowing to widely acuminate apex; mesepimeron slightly expanded laterally, partially visible from above; metepimeron partially covered by anterolateral projection of abdominal sternite 1; metacoxal plate with posterior margin broadly, shallowly concave; outer apical angle acute, slightly projecting.

Legs with femora flattened, slightly fusiform basally, inner margin with short apically- curved erect setae, inner margin arcuate on apical 2/3 SO that tibiae are only contiguous in repose at either end; tibiae slightly arcuate with short, stout curved distal spine; protibiae with distal angulate appendix; tarsomere 1 slightly longer than 2; 2, 3 and 4 subequal, 1-4 with ventral pulvilli, 5 elongate, claws broadly appendiculate.

Abdominal sternites 1 and 2 laterally expanded, partially visible from above, suture only slightly visible laterally; sutures between 2, 3 and 4 evenly transverse, between 4 and 5 slightly arcuate; posterolateral angles of 2, 3 and 4 obtuse; margin of 5 broadly arcuate to truncate apex.

Metathoracic wings as in Figure 28.

REMARKS. This generic name is designated feminine and is based on the Latin diminutive of the place name, Katanga. The arcuate tibiae and wing venation relate Katangiella to the Strigulia Kerremans- Cupriscobina Bellamy and Holm group of genera. Katangiella is distinct from these and most other African coroebines with two apparently apomorphic characters, the appendiculate antennomere 1 1 and the squamiform setal covering.

The only species in Katangiella is described below with only a morpho- metrical and specific character diagnosis presented to augment the generic description.

Katangiella squamivela Bellamy, sp. nov. Figures 24-28

DESCRIPTION. Holotype male. Size, 8.4 x 2.5 mm; colour black with slight purplish or bluish reflections. Head rugose with very slight narrow longitudinal depression midway between eyes. Pronoturn 1.3 x wider than long; disc coarsely rugose, with slight aeneous reflection; anterior margin with slight finely rugose band, widest medially; prelateral carinae extend from before middle sinuately to past middle, one on either side; discal depressions as follows: 2 small on midline, 1 at base, 1 just past middle, 1 small pair before anterior margin, before lateral Y3, one large pair more or less over basolateral Y2. Elytra nearly 2.5 x longer than wide; surface black, rugose; setal concentrations in pairs: 1 in basal depression, 1 in medial '/2 at basal l/3 and 1 just past middle; 1 pair of transverse fasciae, in lateral '/2, just before apical '13 and 1 zigzag entire fascia at apical '12; lateroapical margin serrate from apical '/4 to just inside sutural angle; squamiform setae slightly more concentrated on visible portion of abdominal tergites 1 and 2. Genitalia as in Figure 27.

MATERIAL EXAMINED. Holotype, male (MRAC): [ZAIRE] Katanga: Kolwezi [S10.43 E25.281 VII. 1953, Dr. Allard.

REMARKS. The specific name is a combination of the Latin squam (scales) and vel (a veil) to describe the covering of squamiform setae.

TYPE-SPECIES. Toxoscelus africanus Kerremans 1 899:274.

DESCRIPTION. Size less than 8.0 mm in length; elongate; flattened. Head feebly produced between eyes, feebly bilobed; eyes moderate, widely separated;


Figs. 24-28. Katangiella squamivela, gen. et sp. nov. 24, left dorsal view. 25, lateral view. 26, head, frontal view. 27, male genitalia, dorsal view. 28, wing venation, dorsal view.

Figs. 29-34. Figs. 29, 30, 33, 34, Parastrigulia afiicana (Kerremans). Figs. 31, 32, Strigulia asmarica Bellamy. 29, left dorsal view. 30, 3 1, lateral view, pronotum, base of left elytron, and thoracic sternites. 32, 33, head, frontal view. 34, profemur and protibia, dorsal view.

supra-antenna1 grooves and antenna1 cavities widely separated by broad frontoclypeus, which is concavely emarginate distally. Antennae serrate from antennomere 5.

Pronoturn 1.5 x wider than long, widest anterior to middle; anterior margin lobate medially; anterolateral angles produced, obtuse, when viewed from side; lateral margins broadly arcuate; basal margin bisinuate, with median truncate lobe anterior to scutellum; basal angles obtuse. Scutellum triangular, wider than long, basal angles rounded.

EIytra narrower than pronotum at base, sides subparallel past humeri, then narrowed before widening to widest past middle, then narrowing gradually to slightly separately rounded attenuate apices. Pygidium subtruncate, with moderate median projecting spine.


Thoracicsternites with prosternum subtriangular, widest anteriorly, with mentonnihe concave, compressed; disc convex, process strongly narrowed between procoxae to attenuate apex; mesepimeron not visible from above.

Legs as in Strigulia. Abdominal sternites with basal sternites laterally expanded, visible from above.

REMARKS. The name Parastrigulia is feminine and the genus comes nearest to Strigulia Kerremans, hence the name. Parastrigulia can be differentiated by the following characters, based upon comparison with S. asmarica Bellamy, S. bottcheri Thiry and the female holotype of Toxoscelus africanus (BMNH).

Strigulia Parastrigulia, gen. nov.

Frons longer than wide (Fig. 32). Supra-antenna1 groove entire.

Frontoclypeus narrowed to broad canna, distally convex.

Pronotum less than 1.2 x wider than long.

Elytra widest at base. Antennomere 1 1 entire. Mesepimeron strongly dorsally pro-

duced, visible from above (Fig. 3 1).

Abdominal sternite 5 sexually dimorphic.

Frons wider than long (Fig. 33). Supra-antenna1 grooves separated by fronto-

clypeus. Frontoclypeus not carinate, broad between

antenna1 cavities, distally concave. Pronotum 1.5 x wider than long.

Elytra widest past middle. Antennomere 1 1 serrate. Mesepimeron small, not visible from above

(Fig. 30).

Abdominal sternite 5 not sexually dimorphic.

Parastrigulia africana (Kerremans), comb. nov. Figures 29, 30, 33, 34

Toxoscelus africanus Kerremans 1899:274, 1909:36; Obenberger 1934905.

This species was originally described in an otherwise Oriental genus and, indeed, does bear a strong resemblance to species of Toxoscelus. Examination of the female holotype reveals that a more correct placement is in Parastrigulia; P. africana is related to species of the African genus Strigulia and joins the generic complex comprised of these two along with Strigulioides Bellamy, Kerremansia Piringuey and Cupriscobina. These five genera share the character of the strongly arcuate pro- and mesotibiae. Parastrigulia africana also superfi- cially resembles species of Strandietta Obenberger and Cobosietta Bellamy, but differs most noticeably in the form of the first two pairs of legs. Both Strandietta and Cobosietta, along with most species ofDiscoderoides, have the tibiae strongly flattened and lying parallel to the femora in repose.

TYPE-SPECIES. Amorphosoma dentifer Waterhouse.

DESCRIPTION. Size less than 7.0 mm long; elongate; flattened above; elytra with multi- coloured setal fasciae.

Head quadrituberculate, dorsal-most pair between eyes much more strongly produced, with apical fascicle of stout, squamiform setae, frontal tubercles only slightly convex; entire frontovertex moderately covered with recurved setae; eyes large, inner margins diverging dorsally; frontoclypeus with supra-antenna1 grooves separately arcuate dorsally, becoming confluent with slight medial depression on upper part of frontocly-

T H E COLEOPTERISTS BULLETIN 42(2), 1988 183

peus; frontoclypeus compressed between antenna1 cavities, laterally with a very narrow, oblique costa on each side, roundly emarginate medioapically, slightly produced laterally; gena with elongate transverse depression beneath eye for reception of basal antennomeres in repose, with rounded obtuse lobe beneath groove; labrum wider than long, with semi- erect setae on apical %, apical margin sinuate. Antennae with antennomere 2 globose; 3 shorter than 2, subserrate; 4 serrate, longer than 3; 5-10 with width nearly 2 x length, laterally narrowed, slightly arcuate, rounded distally; 1 1 oblong, rounded.

Pronotum wider than long, widest near middle; apical margin slightly arcuate; basal margin trisinuate, with medial prescutellar lobe feebly concave; lateral margins arcuate, marginal carina sinuate; one sinuate costa extends from apical l/3 obliquely discad and anteriad on either side; laterobasal angles obtuse; disc moderately, roundly convex, depressed before one strongly elevated convex tubercle on either side; one slight sinuate carina extends anteriad from just inside basal angle to near middle; surface rugose, sparsely covered with recurved setae. Scutellum with length subequal to width, basal margin arcuate, lateral margin slightly lobate, slightly overlapping elytral surface.

Elytra slightly narrower than pronotum, widest opposite humeri; humeri moderately elevated, oblique; base depressed between each humerus and scutellum; epipleural lobe narrow, elongate, separated from disc by prelateral carina which extends from base to just before apex; sides narrowing beyond humeri before widening again at apical Y2, then gradually narrowing to serrate lateroapical angles; apex of elytron subtruncate except for large median triangular tooth and broadly acute sutural tooth; disc flattened, rugose, moderately setose, apical l/2 with setae concentrated into irregular transverse fasciae.

Underside with prosternum very short, quite strongly compressed anteroposteriorly, mentonnikre strongly bilobed, process strongly arcuate between procoxae when viewed from side, rounded distally; metasternum with two small ventrally-produced lobes between mesocoxae, forming anterior part of sternal cavity; metacoxal plate with posterior margin bisinuate, with one acute lobe slightly medial to middle, lateroapical angle roundly acute; abdominal sternites with sutures evenly transverse, lateroapical angles slightly obtuse, lateral margin of sternite 5 finely serrate, rounded, apex truncate; surface of abdominal sternites rugose, generally moderately covered with white recurved setae, concentrated into a pair of round patches on sternites 2-5; posterior 2/3 of metasternum and metacoxal plate densely covered with elongate white squamiform setae.

Legs with femora fusiform; protibiae slightly arcuate, outer margin dentate; metatibiae with setal comb on apical 2/3 of outer margin; tarsi with basal four tarsomeres becoming progressively shorter distally, tarsomere 5 narrow, elongate, subequal to 1-4 together, claws bifid, inner teeth short, slightly curving inward.

Male genitalia as in Figure 40; ovipositor with pair of ventral brushes. Metathoracic wing as in Figure 4 1.

REMARKS. T h e name of this new genus is feminine a n d is a combination o f the Greek roots crypt (hidden, concealed) a n d morph (form) because of its very camouflaged appearance.

T h e description o f this genus is the first step in clarifying the heterogeneous Amorphosoma Laporte a n d with its description the following synonymy is proposed.

Cryptomorpha dentifera (Waterhouse), comb. nov. Figures 3 5-4 1

Amorphosoma dentifer Waterhouse 1902:330. Phlocteis dentifera, Obenberger 1 934: 796. Arnorphosoma dentifera (sic), Bellamy 1985: 1 86.

MATERIAL EXAMINED. Holotype (BMNH); 1 ex. (SANC): S O U T H AFRICA: Transvaal, Rustenburg, 12- 1 5 .iii. 1947, A. L. Capener; 1 male, 1 female (CBLC): Tvl., 25 k m E Nelspruit, 1 8/ 1 9.11.1 984, C. L. Bellamy, beating lichen-covered branch of Maytenus heterophylla; 1 female (CLBC), Tvl., 5 k m N Groblersdal,


12.111.1984, C. L. Bellamy; 1 ex. (TMSA): Tvl., Stockpoort, 2.1.1985; 1 male (CLBC): Tvl., Melodie, 2 1 .VII. 1985, C. L. Bellamy, beating Maytenus sp.

GENUS VANROONIA OBENBERGER

Vanroonia Obenberger 1923:28, 1934:8 14. TYPE-SPECIES. Vanroonia coraeboides Obenberger (from original monotypy).

REMARKS. Obenberger (1 923) erected the genus Vanroonia for a new species, V. coraeboides, from Cameroun and placed it near Cisseicoroebus Kerremans and Amorphosoma. Later (1 93 1) he described a second species, V. strandi, from Zambia. In 1958 he revised the genus, providing a key and describing new species from Spanish Guinea, Sierra Leone, New Guinea and the Himalayas. I have compared species of Amorphosoma to those in Vanroonia and feel that Amorphosoma should be restricted to the New World, and the Old World species placed in Vanroonia. The African-Indo-Oriental distributional track of Vanroonia is comparable to that of Sambus Deyrolle, although the latter also has species in Madagascar.

For comparison, Vanroonia strandi Obenberger is illustrated (Figs. 42-48).

Vanroonia coraeboides Obenberger

Vanroonia coraeboides Obenberger 1 923:29, 1 934: 8 14.

MATERIAL EXAMINED. Lectotype (here designated), male (NMPC): [white] Kamerun, Tiko b.Victoria (p)/ [white] 4.1.1943, v.Rothkirch [p]/ [red] TYPUS/ [white] Vanroonia coraeboides m. Type [h] det Dr. Obenberger [p]/ [green] Mus. Nat. Pragae [p] 23646 [h]/ [red] LECTOTYPE [p] Vanroonia coraeboides 8 Obenb. det. [h] C. L. Bellamy [p]; 6 paralectotypes (5, NMPC 23647-5 1; 1, HNBH): same data, except labelled PARALECTOTYPE; 1 male (MRAC): Kamerun, Meyer D.

Vanroonia guineae Obenberger

Vanroonia guineae Obenberger 1 95 8: 5 1 1. Vanroonia leonensis Obenberger 1 9585 1 2, syn. nov.

REMARKS. Examination of the holotypes of V. guineae (NMPC 23666) and V. leonensis (NMPC 23667) reveals the need for this synonymy. Both species were described in the same work (Obenberger 1958) from single specimens. I have examined a third specimen (MRAC) from Ivory Coast and find no sub- stantial differences. This appears then to be a more widespread West African species than originally surmised.

Vanroonia africana (Obenberger), comb. nov.

Coroebus africanus Obenberger 1 928:337.

REMARKS. An examination of the type (NMPC 23707) reveals the need for this new combination, which I had suspected after seeing the original text figure (Obenberger 1928:337, fig. 1). This transfer from Coroebus and those made earlier (see Bellamy 1988) allows a more restricted definition of the genus and its African "relatives." It is curious that this species was not included in Oben- berger's catalogue (1934), although other species described in the same work (1 928) were.


Figs. 35-48. Figs. 35-41. Cryptomorpha dentifera (Waterhouse). Figs. 42-48. Van- roonia strandi Obenberger. 35, 42, left dorsal view. 36, 43, lateral view. 37, 44, head, frontal view. 38, 45, antenna, dorsal view. 39, 46, metatibia, dorsal view. 40, 47, male genitalia, dorsal view. 4 1, 48, wing venation, dorsal view.

TYPE-SPECIES. Meliboeus dermestoides Thomson.

DESCRIPTION. Length less than 10.0 mm, elongate, subcylindrical, subnitid black with faint iridescent reflections, rugose, moderately setose (more densely so on pronoturn), elytra with transverse fasciae.


Head with frontovertex produced, longitudinally depressed, feebly bilobed between eyes; eyes small, very flat, inner margins subparallel; ocular grooves broad, extending along inner margins from past dorsal pole to past ventral pole of eyes; supra-antenna1 grooves short, separate; frontoclypeus broad, short between antenna1 cavities, emarginate distally; gena lobed beneath eye. Antennae serrate from antennomere 5.

Pronotum wider than long, widest past middle; anterior margin arcuate; posterior margin bisinuate on either side of subtruncate median lobe; basal angles obtuse; lateral margins carinate, subparallel, widening slightly from base to before apex, then arcuate to anterior margin; disc convex, strongly so mediobasally. Scutellum cordiform, basolateral lobes extending slightly over elytral margin.

Elytra slightly narrower than pronotum, widest opposite humeri; sides subparallel past humeri, narrowing slightly to near middle then widening to near apical '13 before becoming gradually attenuate to separately rounded serrate apices; epipleuron narrow, ventrally deflexed, extending almost to apex; disc depressed basally, otherwise transversely convex. Pygidium partially visible from above, extending beyond elytral apices, lateroapical margin subtriangular, serrate.

Thoracic sternites with prosternum with short deflexed bilobed mentonni$re, lobes broadly rounded; disc laterally compressed; process with sides subparallel, narrowing to attenuate apex; hypomeron transversely carinate, anterolateral portion narrowly depressed for distal antennomeres in repose; laterobasally depressed for forelegs in repose, basal notch confluent to notch at base of epipleuron to receive apex of mesofemur in repose; metacoxal plate quite narrow medially, broader laterally, posterior margin broadly shallowly emarginate, lateroapical angle roundly acute, projecting.

Legs with femora fusiform; tibiae arcuate on external margin; tarsomere 5 very slender, longer than 1 4 together; claws bifid.

Abdominal sternites 1 + 2 longer than 3 + 4 + 5; suture between 1 and 2 vague but indicated across entire width; other sutures transverse; lateroapical margin of sternite 5 roundly attenuate.

Metathoracic wing with radial cell open distally; radiomedian crossvein absent; 1 stA 1 and 2dA1 separate, short.

Genitalia with parameres partially fused, with membranous lobes lateroapically. Ovi- positor with two ventral opposing setose brushes.

REMARKS. The species of Kamosiella are related to those of Kamosia; the new name is feminine. The strongly flattened eyes relate Kamosiella to Pseu- dokamosia ThCry, and are clearly a derived character state (S. Caveney, pers. comm.). Otherwise, Kamosiella seems nearer to Meliboeus Deyrolle than to Kamosia Kerremans. Kamosia indigacea (Obenberger) is illustrated (Figs. 49, 5 1, 53, 55, 57) for comparison.

Kamosia Kamosiella, gen. nov.

Head (Fig. 49) not or only slightly produced, only feebly longitudinally grooved between eyes.

Eyes (Figs. 5 1, 5 3) large, moderately to strongly convex.

Antennal cavities small, widely separated by about 2 x their width.

Gena with acute to angularly obtuse tooth (Fig. 53).

Pygidium trispinose apically (Fig. 55).

Abdominal sternites 1 +2=3 +4+ 5 in length.

Tarsi (Fig. 57) with tarsomere 5

Head (Fig. 50) produced, bilobed between eyes.

Eyes (Figs. 52, 54) small, flat to feebly convex.

Antennal cavities large, separated by about 1 x width.

Gena with rounded obtuse tooth (Fig. 54).

Pygidium triangularly attenuate (Fig. 56).

Abdominal sternites with 1 + 2 > 3 + 4 + 5 in length.

Tarsi (Fig. 58) with tarsomere 5 very narrow,


more robust, claws with outer claws small, reduced, outer teeth subparal- teeth widely separated, inner teeth lel, inner teeth contiguous distally. nearly continguous distally.

Generally throughout subsaharan Southern Africa, apparently associated only Africa, with many putative host with Rhus spp.; not anthophilous. plant genera, e.g., Combreturn and anthophilous adult habits.

Kamosiella dermestoides (Thomson), comb. nov. Figures 50, 52, 54, 56, 68

Meliboeus dermestoides Thomson 1 878:8 1. Karnosia dermestoides, Thkry 1 926: 59; Obenberger 1 934:877. Amorphosoma originaria PCringuey 1908:303, syn. nov. Kamosia originaria, Obenberger 1 9 34: 8 7 7.

I have examined the types of both species [dermestoides (MNHN) and originaria (SAMS)], and a number of additional specimens, and find very little

d1if"erence witfiin ffiem.

Kamosiella jactuosula (Pkringuey), comb. nov.

Demostis jactuosula Pkringuey 1 908: 305. Phlocteis jactuosula, Oben berger 1 9 34: 7 9 8. Kamosia jactuosula, Bellamy 1 985 : 1 86.

I have examined the damaged type (SAMS) and a few other specimens (CLBC, GHNC, TMSA). This species differs from K. dermestoides by being generally larger, with the elytral setal pattern broader and more dense and by the shape of the male genitalia. This species appears to be distributed along the tropical forest belt of southeastern Africa from Natal to Mozambique.

GENUS HOLMERIKA BELLAMY, GEN. NOV.

TYPE-SPECIES. Holmerika orientalis, sp. nov.

DESCRIPTION. Size, length less than 10.0 mm; subcylindrical; elongate; nitid black, portions with bright cupreous or aeneous reflections; surface rugose with bicolourous setal covering.

Head (Fig. 62) produced between widely separated convex eyes; frontovertex longitudinally grooved; antenna1 cavities small, separated by nearly 2 x width, each bordered above by short arcuate groove which is confluent with ocular groove; eyes with inner margins slightly diverging dorsally, with deep groove along inner margin to a point opposite apex; frontoclypeus distally emarginate, depressed for basal antennomeres laterally; gena with acutely rounded lobe beneath each eye; labrum very short, distally setose; mandibles with apices attenuate. Antennae (Fig. 63) with antennomere 1 geniculate, fusiform distally; 2 globose, longer than 3; 3 slightly swollen distally; 4 triangularly serrate; 5-10 broadly serrate, length to width ratio increasing distad with 10 subequal to 9; 1 1 oblong, curved.

Pronotum wider than long, widest anterior to middle; apical margin feebly arcuate medially; basal margin bisinuate on either side of slightly emarginate median prescutellar lobe; laterobasal angles obtuse; lateral margins arcuate, carinate, crenulate from basal l/3 to before apex; disc convex anteromedially, otherwise depressed near basolateral portion; either side with very feebly elevated prehumeral carina extending from basolateral angle anteriad. Scutellum cordiform, widest basally; basal margin arcuate; apex attenuate.

Elytra with width subequal to that of pronotum, slightly wider before apical '/3; humeri


Figs. 49-58. Figs. 49, 5 1, 53, 55, 57. Kamosia indigacea (Obenberger). Figs. 50, 52, 54, 56, 58. Kamosiella dermestoides (Thomson). 49, 50, head, dorsal view. 5 1, 52, head, right frontal view. 53, 54, head, pronotum, lateral view. 55, 56, pygidium, dorsal view. 57, 58, metatarsi, lateral view.

elevated, oblique; disc depressed between humeri and scutellum; lateral margins carinate; epipleuron broad basally, then narrowed, separated from disc by carina, ventrally deflexed to opposite suture between abdominal sternites 4 and 5; sides narrowing slightly past opposite humeri, before widening before apical l/3, then attenuate to slightly separately rounded apices; lateroapical margins serrulate from middle, becoming serrate to dentate apically. Pygidium not projecting past elytral apex.

Thoracic sternites with prosternum with acutely bilobed mentonnikre lobes angulate to disc; process ventrally convex, attenuate apically; hypomeron with feebly elevated carina dividing disc on middle portion.

Legs with femora robust, slightly depressed on inner face for tibiae in repose; tibiae arcuate basally; protibiae with outer edge feebly crenulate; metatibiae (Fig. 64) with setal "comb" on apical Y2; tarsomeres 1 4 together slightly longer than 5, 1-4 decreasing in length distally, with ventral pulvilli increasing in width distally; 5 elongate, fusiforrn distally; claws bifid, inner teeth short, almost contiguous apically.

Abdominal sternites with sutures arcuate. Metathoracic wings as in Figure 65. Male genitalia as in Figures 66 and 67. Ovipositor with two opposing ventral setal

"brushes."


REMARKS. The name Holmerika is masculine and is erected to honour Professor Erik Holm, my mentor. The superficial resemblance to species of Coroebus Gory and Laporte and Katonius Thkry caused some initial confusion, on my part, about the placement of the following two new species. However, the presence of the apomorphic ovipositor type and open radial cell in the wing (Fig. 65) helped place it within the Meliboeus group (for a discussion of these tribal characters see Bellamy 1988). Within Holmerika the formation of the transverse carinae on the hypomera is the most rudimentary and lack of any excavation on the hypomera (Fig. 61) for the antennae suggest Holmerika as a link between the Meliboeus-group and Asymades Kerremans. Holmerika comes nearest to Asymades, differing by having the pronotum more or less evenly convex, at least on the anteromedial portion with the basolateral areas evenly depressed, while Asymades has the pronotum uneven and partially tuberculate and is more than twice as large as either species of Holmerika. Asyrnades borana (Obenberger) and A. transvalensis Kerremans are illustrated (Figs. 68-71) for comparison, while the two new species of Holmerika are diagnostically described below.

Holmerika orientalis Bellamy, sp. nov. Figures 60-6 5, 67

DESCRIPTION. Size 5.6-6.0 x 1.7-1.8 mm; surface rugose; shining black with faint blue or purple reflections on head and pronotum, with areas of bright aeneous reflection on scutellum and elytra; medial subglabrous portion of elytra deep blue; moderately setose on frons; pronotum generally with elongate testaceous setal covering, with small patches of short white setae intermixed; elytra with base, lateral '/2 from past humeri to middle, and most of apical l/3 covered with elongate adpressed testaceous setae, with small white setal patches as in Figure 60. Head (Fig. 62) produced between eyes, apex feebly convex when viewed from above; disc of frontoclypeus with cupreous reflection, distal margin broadly shallowly emarginate. Pronotum slightly more than 1.4 x wider than long; disc regularly convex anteromedially. Elytra with apicolateral margin finely senate. Metacoxal plate biarcuately emarginate. Male genitalia as in Figure 67.

MATERIAL EXAMINED. Holotype, male (TMSA): SOUTH AFRICA, Natal, Ndumu Game Reserve, S26.5 1-55 E 32.12-20, 5-8.XI.1984, 50-100 m, C. L. Bellamy; 1 male paratype (CLBC): same data; 1 female paratype (MRAC): S. L. Edouard, r. Rwindi (1,000 m), 25-IV-36, L. Lippens, Parc Nat. Albert; 1 female paratype (NMWA): Zanzibar.

REMARKS. This species is named for its eastern African localities. It can be separated from H. mulanje, sp. nov., by differences in colour, setation, shape of frontovertex, frontoclypeus and metacoxal emargination and male genitalia.

Holmerika mulanje Bellamy, sp. nov. Figures 5 9, 66

DESCRIPTION. Size, 5.8 x 2.0 mm; surface rugose; shining black with faint blue or purple reflections on head and pronotum, with areas of bright cupreous reflection on scutellum and elytra; frons moderately setose; pronotum generally with elongate brunneous setal covering, with small patches of short white setae intermixed; elytra with base and most of apical l/3 covered with elongate adpressed brunneous setae, with small white setal patches as in Figure 59. Head bilobed between eyes; frontoclypeus broadly and arcuately roundly emarginate distally. Pronotum slightly more than 1.6 x as wide as long; disc slightly depressed anteriad to median convexity. Elytra with apicolateral margin dentate. Metacoxal plate evenly arcuately emarginate. Male genitalia as in Figure 66.

190 T H E COLEOPTERISTS BULLETIN 42(2), 1 988

Figs. 59-7 1. Figs. 59,66, Holmerika mulanje, sp. nov. Figs. 60-65,67, H. orientalis, sp. nov. Figs. 68, 69, Asymades borana (Obenberger). Figs. 70, 71. A. transvalensis Kerremans. 59, 60, 68, left dorsal view. 6 1, 69, lateral view. 62, 70, head, frontal view. 63, antenna, dorsal view. 64, metatibia, ventral view. 65, 7 1, wing, dorsal view. 66, 67, male genitalia, dorsal view.

MATERIAL EXAMINED. Holotype, male (TMSA): S O U T H E R N MALAWI: Mabuka, Lauderdale/Mt. Mulanje, SE 16 35 Ba, 30.XI-2.XII. 1983, E. Holm.

REMARKS. This species is named for the primary type-locality, Mt. Mulanje. I t differs from H. orientalis as mentioned under that species.


Asymades Kerremans 1 893: 1 1 8; Obenberger 1 934:8 1 2. TYPE-SPECIES. Asymades transvalensis Kerremans (from original monotypy). Zavattaria Obenberger 1940:652, syn. nov. TYPE-SPECIES. Zavattaria borana Obenberger (from original monotypy).

REMARKS. An examination of two specimens of Z. borana Obenberger shows this taxon to be congeneric with Asymades, one of several genera it was compared to at the time of the original description. With this synonymy the two species of Asymades repeat a pattern observed within the two species of Chal- cophlocteis Obenberger (see Bellamy 1986a), that of an East African species being a replacement to a southern African species.

Asymades borana (Obenberger), comb. nov. Figures 68, 69

Zavattaria borana Obenberger 1940:654.

MATERIAL EXAMINED. Holotype (MCTI): SOMALIA: Borana, Javello, 30.iv. 1937; 1 female (ANSP): KENYA, Kibwezi, Br. E. Africa, XII- 17- 192 1, R. Dummer.

REMARKS. This species is immediately separated from A. transvalensis by the bituberculate pronotum, by having setal tufts on the head and by the widely separated areas of assumed distribution.

I would like to express my gratitude to those that assisted me in the prep: aration of this paper: firstly to Prof. E. Holm of this department, for providing the working space, support and many helpful suggestions on this and many other projects; to G. H. Nelson and R. L. Westcott for their critical reading of an earlier draft and many suggestions towards its improvement and to those who provided type and other material from collections in their care, D. Azuma (ANSP), E. R. Peacock (BMNH), A. Descarpentries and R.-P. Dechambre (MNHN), J. Decelle (MRAC), R. G. Oberprieler (SANC), D. Hancock (NMBZ), S. Biljr (NMPC), H. Schonmann (NMWA), V. B. Whitehead (SAMS), S. En- drody-Younga (TMSA); and G. Alberti (MCTI).

BELLAMY, C. L. 1985. Some nomenclatorial changes in the subfamily Agrilinae (Co- leoptera, Buprestidae). J. Ent. Soc. Sth. Afr. 48: 185-188. . 1986a. Studies in the African Agrilinae, Coraebini IV. Navors. Nas. Mus., Bloemfontein 5:93-167. . 19863. Rectifications and descriptions of new taxa in the Coraebini (Coleoptera, Buprestidae, Agrilinae). J. Ent. Soc. Sth. Afr. 49:239-249. . 1988. Studies in the African Agrilinae, Coroebini VI. J. Ent. Soc. Sth. Afr. 50: 101-1 14. , AND E. HOLM. 1985a. Studies in the African Agrilinae, Coraebini I (Coleoptera, Buprestidae). J. Ent. Soc. Sth. Afr. 48: 12 1-1 28. , AND - . 19856. Studies in the African Agrilinae, Coraebini I1 (Coleoptera, Buprestidae). J. Ent. Soc. Sth. Afr. 48: 129-1 34.

KERREMANS, C. 1893. Essai de groupement des buprestides. Ann. Soc. Ent. Belg. 37: 94-1 22.


. 1899. Buprestides de 1'Afrique Cquatoriale et de Madagascar. Ann. Soc. Ent. Belg. 43:256-298. . 1909. Catalogue raisonnk de la faune entomologique du Congo Belge. Coleop- tera. Buprestidae. Ann. Mus. Congo Belge, Zoologic, series 111, tome I, fasc. 2, pp. 1 4 4 .

OBENBERGER, J. 1923. Une sCrie de nouveaux genres de Buprestides. Acta Ent. Mus. Nat. Pragae 1 : 1344 . . 1928. Opuscula Buprestologica I. Beitrage zur Kenntnis der Buprestiden (Col.). Arch. Naturg. 92(A), 9-1 l(1926): 1-350. . 193 1. Studien iiber die aethiopischen Buprestiden I. Folia Zool. Hydrobiol. 2: 175-20 1. . 1934. Buprestidae 4. Coleopterorum Catalogus, part 143:782-934. . 1940. Coleoptera, Buprestidae. In: Mission biologics nel Paese dei Borana, Raccolte zoologiche 2, Parte 1 (1 939):64 1-663. . 1958. Revision des espGces du genre Vanroonia Obenb. (Col. Bupr.). Acta Ent. Mus. Nat. Pragae 32:503-5 12.

PERINGUEY, L. 1908. South African coleopterous fauna. Ann. So. Afr. Mus. 5:301- 3 14.

T H ~ R Y , A. 1926. Remarques sur les genres Chrysaspis Kerr., Steraspis Sol. et Kamosia Kerr., avec descriptions d'espGces nouvelles. Rev. Zool. Africaine 14:54-59. . 1932. Buprestides nouveaux d'Afrique (4e note). Bull. Soc. Sci. Nat. Maroc 11:144-175.

THOMSON, J. 1878. Typi buprestidarum musaei Thomsoniani. E. Deyrolle, Paris. WATERHOUSE, C. 0. 1902. Descriptions of two new Coleoptera of the family Bupres-

tidae. Ann. Mag. Nat. Hist. (7) 10:330-332.

(Received 23 December 1987; accepted 30 March 1988)

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