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4.07 - Extending the principal of Beverton-Holt Life History Invariants for length based assessment of SPR Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History Invariants for length based assessment of SPR. In: ICES World Conference on Stock Assessment Methods for Sustainable Fisheries (WCSAM 2013), 15 - 16 July, Boston, USA.

4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

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Page 1: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

4.07 - Extending the principal of Beverton-Holt Life History Invariants for length based assessment of SPR

Adrian Hordyk & Jeremy Prince*

Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History Invariants for length based assessment of SPR. In: ICES World Conference on Stock Assessment Methods for Sustainable Fisheries (WCSAM 2013), 15 - 16 July, Boston, USA.

Page 2: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Collected high quality biological parameters for range of marine species (Gislason et al. 2010 – Criteria used).

For each species:

• Growth model • Natural mortality (M) • Size-fecundity model or maturity ogive • Length – weight model

Examined patterns life history strategies

Meta-analysis

Page 3: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

SPR at Size

Page 4: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

SPR at Size

Page 5: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

SPR at Size: r- vs. K- strategists

Type I 𝑴

𝒌> 𝟏

Clupeidae

Page 6: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

SPR at Size: r- vs. K- strategists

Type I 𝑴

𝒌> 𝟏

Type II 𝑴

𝒌< 𝟏

Haliotidae

Trachichthyidae

Page 7: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

SPR at Size: r- vs. K- strategists

Type I 𝑴

𝒌> 𝟏

Type II 𝑴

𝒌< 𝟏

Type III 𝑴

𝒌< 𝟏

& 𝑳𝒎

𝑳∞> 𝟎. 𝟖𝟓

Physeteridae

Page 8: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Beverton-Holt Life History Invariants

M/k = 1.5 Lm/L∞= 0.66 M x Agem = 1.65 Fec. ~ Adult Wt.

Page 9: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Meta-analysis & Beverton (1992)

Lm/L∞ = 3/(3+ M/k).

Page 10: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Unfished Length Composition

Page 11: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Length Based SPR Estimation Method

Standardised to L∞

Unfished

(𝑴

𝒌)

: expected unfished length distribution

Page 12: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Standardised to L∞

Fished

(𝒁

𝒌)

selectivity

(F/M)

: expected unfished length distribution

: length frequency of catch (Z = F + M)

SPR & F/M: Calculated from M/k & Lm/L∞

Length Based SPR Estimation Method

Page 13: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Important Assumptions

Length frequency of catch representative of exploited stock Asymptotic selectivity Same growth curve or female length data Knowledge of maturity at size Equilibrium method

Page 14: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Neoplatycephalus richardsoni

Tiger Flathead

Calibration against Stock Assessments

Low

Ok

High

Page 15: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Merluccius productus

Pacific Hake

Calibration against Stock Assessments

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LB-SPR Assessment

Merluccius merluccius

Northern Hake – ICES dataset

Calibration against Stock Assessments

Page 17: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Merluccius merluccius

Northern Hake – ICES dataset

Calibration against Stock Assessments

Page 18: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Conclusion

Meta-analysis

M/k ratio defines life-history strategy & Size composition e.g. tuna are just scaled up anchovy. Conceptual framework for borrowing information from data-rich species.

BH-LHI Only covers a small subset of the species in the meta-analysis. Productivity of K-strategists parameterised by BH-LHI have been over-estimated.

Application Cost-effective estimation of SPR & F/M from length-data, Lm & meta-analysis for Data-poor and small scale fisheries.

Page 19: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Acknowledgements

Funding David and Lucile Packard Foundation Marine Stewardship Council The Nature Conservancy Murdoch University Data & Assistance Kotaro Ono, Sarah Valencia, Keith Sainsbury, Neil Loneragan

Thank you

Page 20: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Atheresthes stomias

Arrowtooth Flounder

Calibration against Stock Assessments

Page 21: 4.07 - Extending the principal of Beverton-Holt Life ... · Adrian Hordyk & Jeremy Prince* Hordyk, A. and Prince, J.D. (2013) Extending the principal of Beverton-Holt Life History

Estimation Model

Estimated parameters: Model input parameters:

F/M

SL50 & SL95

SPR

M/k L∞

CVL∞

L50 & L95

Female parameters

𝑴𝑳𝑬 𝑺𝑳𝟓𝟎 ,𝑺𝑳𝟗𝟓 , 𝑭/𝑴 =𝒂𝒓𝒈 𝒎𝒊𝒏(𝑺𝑳𝟓𝟎, 𝑺𝑳𝟗𝟓, 𝑭/𝑴)

𝑶𝑳𝒍𝒐𝒈𝑷𝑷𝑳𝑶𝑷𝑳

𝑳=𝑳𝒎𝒂𝒙

𝑳=𝑳𝒎𝒊𝒏