and faunistic data on Neotropical Cochylini

SHILAP Revista de Lepidopterología

ISSN: 0300-5267

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Sociedad Hispano-Luso-Americana de

Lepidopterología

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Razowski, J.; Becker, V. O.

Systematic and faunistic data on Neotropical Cochylini, with description of new species. Part 3

(Lepidoptera: Tortricidae)

SHILAP Revista de Lepidopterología, vol. 35, núm. 137, marzo, 2007, pp. 67-86

Sociedad Hispano-Luso-Americana de Lepidopterología

Madrid, España

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67

Systematic and faunistic data on NeotropicalCochylini, with description of new species. Part 3

(Lepidoptera: Tortricidae)

J. Razowski & V. O. Becker

Abstract

16 new species (Maricaona maricaonana Razowski & Becker, sp. n., Phalonidia mayarina Razowski &Becker, sp. n., Saphenista turguinoa Razowski & Becker, sp. n., Saphenista rosariana Razowski & Becker, sp. n.,Saphenista simillima Razowski & Becker, sp. n., Saphenista cubana Razowski & Becker, sp. n., Platphalonidiaholguina Razowski & Becker, sp. n., Platphalonidia remissa Razowski & Becker, sp. n., Lasiothyris guanana Ra-zowski & Becker, sp. n., Lasiothyris subsorbia Razowski & Becker, sp. n., Lasiothyris puertoricana Razowski &Becker, sp. n., Spinipogon virginanus Razowski & Becker, sp. n., Eugnosta fraudulenta Razowski & Becker, sp.n., Lorita insulicola Razowski & Becker, sp. n., Aethes pinara Razowski & Becker, sp. n., Cochylis sierramaes-trae Razowski & Becker, sp. n.) and one new genus (Maricaona Razowski & Becker, gen. n.) are described. Thematerial studied was collected chiefly in Cuba, Puerto Rico, and Ecuador. The data on distribution and morpholo-gy of 3 further (Saphenista semistrigata Forbes, 1931, Mimeugnosta particeps Razowski, 1986, Lorita lepidulana(Forbes, 1931) species are provided.KEY WORDS: Lepidoptera, Tortricidae, Cochylini, distribution, new taxa, Neotropical.

Datos sistemáticos y faunísticos sobre Cochylini Neotropicales, con descripción de nuevas especies. Parte 3(Lepidoptera: Tortricidae)

Resumen

Se describen 16 nuevas especies (Maricaona maricaonana Razowski & Becker, sp. n., Phalonidia mayari-na Razowski & Becker, sp. n., Saphenista turguinoa Razowski & Becker, sp. n., Saphenista rosariana Razowski& Becker, sp. n., Saphenista simillima Razowski & Becker, sp. n., Saphenista cubana Razowski & Becker, sp. n.,Platphalonidia holguina Razowski & Becker, sp. n., Platphalonidia remissa Razowski & Becker, sp. n., Lasio-thyris guanana Razowski & Becker, sp. n., Lasiothyris subsorbia Razowski & Becker, sp. n., Lasiothyris puertor-icana Razowski & Becker, sp. n., Spinipogon virginanus Razowski & Becker, sp. n., Eugnosta fraudulenta Ra-zowski & Becker, sp. n., Lorita insulicola Razowski & Becker, sp. n., Aethes pinara Razowski & Becker, sp. n.,Cochylis sierramaestrae Razowski & Becker, sp. n.) y un nuevo género (Maricaona Razowski & Becker, gen.n.). El material estudiado fue colectado, principalmente, en Cuba, Puerto Rico y Ecuador. Se proporcionan losdatos sobre la distribución y morfología de tres especies adicionales (Saphenista semistrigata Forbes, 1931,Mimeugnosta particeps Razowski, 1986, Lorita lepidulana (Forbes, 1931).PALABRAS CLAVE: Lepidoptera, Tortricidae, Cochylini, distribución, nuevas taxa, Neotropical.

Introduction

The present paper constitutes third part of the series devoted to the systematics and faunistics ofthe Neotropical Cochlini (Tortricinae). First part (RAZOWSKI & BECKER, 2002) contained the de-scriptions of 36 new species and the data on further 26 species from Brazil and Ecuador; part two

SHILAP Revta. lepid., 35 (137), 2007: 67-86 SRLPEF ISSN:0300-5267

67-86 Systematic and faunistic 22/3/07 10:14 Página 67

(RAZOWSKI & BECKER, 2007 [in press]) comprised the description of 32 new species, two newgenera and the remarks on further 19 species from Mexico, Ecuador and Brazil. Now we publish onthe insular cochylines; the material is chiefly from Cuba, Puerto Rico, and Virgin Island. This papercomprises the description of 16 new species and the data on three known species. The cochylinesfrom the studied islands belong to the following eight genera: Maricoana Razowski & Becker, gen.n., Phalonidia Le Marchand, 1933, Saphenista Walsingham, 1914, Platphalonidia Razowski, 1985,Lasiothyris Meyrick, 1917, Spinipogon Razowski, 1967, Lorita Busck, 1939, and Cochylis Tre-itschke, 1829. Similarly as in the continental parts of the Neotropical region Saphenista belongs tothe genera most abundant in species. Wide distribution of Phalonidia, Lasiothyris and Platphaloni-dia in this region is also confirmed.

The paper is based on the collection of the junior author (with only two exceptions). The holo-types of the new described species and other material studied are preserved in the Becker Collection.It will be eventually transferred to one of the museums in Brazil. One holotype is from the AmericanMuseum of Natural History, New York. Representatives of a few species originally from the BeckerCollection have been kindly donated to the Institute of Systematics and Evolution of Animals, PAS,Kraków, Poland.

AbbreviationsAMNH - American Museum of Natural HistoryBrit. - BritishGS - genitalia slideINRA - Institut National de la Recherche AgronomiqueMaest. - MaestreSier. - SierraStgo - Santiago

[] - the numbers in brackets are the entry numbers of the specimens in the register book of theBecker Collection.

Note - the number given in descriptions of the labial palpi indicate the proportion of their totallength to the horizontal diameter of eye.

Systematic part

Maricaona Razowski & Becker, gen. n.

Type-species: Maricaona maricaonana Razowski & Becker, sp. n.Description: Venation: in forewing R4-R5 connate or strongly approaching to one another at

median cell; R5 to apex; M3-CuA1 stalked to 1/4; CuA2 opposite R1. In hindwing Rr-M1 stalked tobeyond middle; M2 far from base of M2-M3 which are stalked to about 1/4.

Male genitalia: Tegumen short, broad; base of socii rather separate from tegumen, hairy; sociiwithout free lobes but with slender apical processes; vinculum consisting of pair of broad terminallyarms; valva broad with well developed costa; sacculus long, with free termination; transtilla large,extending dorsally, with short median, spiny part; juxta small, simple; aedeagus very broad; caulispostmedian; coecum penis short, broad; cornuti absent.

Female genitalia: Ovipositor short; papilla analis small; apophyses slender; posterior portion ofsterigma short; cup-shaped part broad; colliculum ill-defined; ductus bursae shorter than corpus bur-sae, with numerous longitudinal folds extending laterally to middle of corpus bursae; accessory bursaoriginating just before middle of the latter.

Diagnosis: Facies and transtilla resembling some representatives of Henricus Busck, 2943; sociiwithout free portions, with apical processes similar to Mourecochylis Razowski & Becker, 1983.Tentatively placed in the Phalonidia-group of genera.

Etymology: The generic name refers to the name of the type-locality of the type species.

J. RAZOWSKI & V. O. BECKER

68 SHILAP Revta. lepid., 35 (137), 2007


Maricaona maricaonana Razowski & Becker, sp. n. (Figs 49)

Holotype male: “Puerto Rico, Maricao, 770 m, 12-VII-1987, V. O. Becker”; [67639]; GS 42Mar. Paratype female with identical label, GS 43 Mar.

Description: Wing span 14.5 mm. Head brownish, labial palpus missing; thorax brown-grey,tegula darker than median portion. Forewing slender, expanding terminad; costa straight to middle,then slightly bent; termen moderately oblique, almost straight. Ground colour white preserved inform of large postmedian blotch at costa; proximal half of wing brownish grey with brown, diffusestrigulation; dorsal third of wing brownish white with brown-grey suffusions and spots; grayish suf-fusion beyond end of median cell marked by rust red terminal spots, Cilia grayish. Hindwing grey,paler basally; cilia grey.

Female (wing span ca 16 mm): head white, labial palpus 1.5; thorax whitish; tegula brownish.Colouration darker than in male; brown and rust red spots in proximal and terminal parts of wing; re-fractive spots and groups of erect scales present.

Male genitalia (Figs 1, 2): as described for the genus.Female genitalia (Fig. 37): as described for the genus.Diagnosis: The only species of the genus; cf. its diagnosis.Etymology: The specific epithet refers to the name of the country.

Phalonidia mayarina Razowski & Becker, sp. n. (Fig. 50)

Holotype male: “Cuba: Holguin, Mayari, 400 m, 12-VII-1990, V. O. Becker”; [72538]; GS 147Mar.

Description: Wing span 18.5 mm. Head cream; labial palpus ca 1.3; thorax cream tinged withyellow-brown. Forewing not expanding terminally; costa and termen rather straight. Ground colourcream preserved along edges of markings; remaining surface suffused with brownish yellow; dotsand strigulae brown. Markings: dorsal blotch at 1/3 of dorsum obliquely towards middle of mediancell accompanied by brownish spot at mid-costa; dorsal blotch yellow brown with blackish marks;blackish spot at tornus; subterminal fascia fusing with blotch between end of median cell and tornus.Cilia cream. Hindwing brownish grey; cilia similar.

Male genitalia (Figs 3, 4): Base of socii broad; arms of vinculum membranously connected withone another; median part of transtilla long, slender; valva slender with basal portion of costa stronglyconvex; sacculus short, broad, rounded terminally; aedeagus large, strongly curved, with slenderventral termination; caulis postmedian; cornutus very large.

Female not known.Diagnosis: Facies similar to Saphenista cubana but majarina without strong dorsal blotch;

closely allied with Cochylis parallelana Walsingham, 1879 from California, USA but majarina dis-tinguished by presence of cornutus.

Saphenista turguinoa Razowski & Becker, sp. n. (Fig. 51)

Holotype male: “Cuba: S[an]tiago, Turguino, 470 m, 27/9-VII-1990, V. O. Becker” [73383]; GS138 Mar.

Description: Wing span 8 mm. Head cream, frons white; labial palpus 1.3; thorax brownishcream, base of tegula darker. Forewing not expanding terminad; costa weakly convex; termen slight-ly oblique, straight beneath apex. Ground colour cream hardly mixed with brownish yellow, withbrowner strigulation. Markins brownish yellow consisting of costal part of basal blotch, dorsopost-basal marking; median fascia divided into three parts accompanied by tornal marking; subterminalslender fascia reaching tornal third of termen. Cilia whitish cream. Hindwing pale brownish grey,paler basally than on periphery; cilia concolorous with median part of wing.

Male genitalia (Figs 5, 6): Socii large, rather broad; vinculum without lateral processes; valva

SYSTEMATIC AND FAUNISTIC DATA ON NEOTROPICAL COCHYLINI, WITH DESCRIPTION OF NEW SPECIES

SHILAP Revta. lepid., 35 (137), 2007 69


and sacculus simple; median part of transtilla long with slender terminal parts; aedeagus about twicelonger than median part of transtilla; cornutus strong, bent.

Female not known.Diagnosis: Allied with S. multistrigana Walsingham, 1914 from Veracruz, Mexico but tur-

guinoa with short aedeagus and strong cornutus.Etymology: The name refers to the type locality.

Saphenista rosariana Razowski & Becker, sp. n. (Fig. 52)

Holotype male: “Cuba: Pinar Rio, Sierra Rosario, 5-15-VI-1990, 400 m, V. O. Becker”;[71535]; GS 159 Mar.

Description: Wing span 7.5 mm. Head cream, thorax with slight brownish hue; labial palpus ca1.3. Forewing slender, broadest postmedially; costa gradually convex; termen rather straight,oblique. Ground colour cream; suffusions pale brownish cream, dots browner. Markings pale brown-ish cream with slight olive hue consisting of weak dorsopostbasal mark, broad median fascia consist-ing of three elongate spots accompanied by browner spot at mid-costa; distinct, uniformly broad sub-terminal fascia, and slender fascia extending from end of termen towards end of median cell. Ciliacream. Hindwing pale brownish cream; cilia paler.

Variation: Paratypes with paler ground colour and more distinct markings.Male genitalia (Figs 7, 8): Socii fairly large; vinculum simple; sacculus simple, small; median

part of transtilla long, slender; juxta small; aedeagus slender with short ventral termination; cornutuslong, swung. Abdominal scent organ in form of two short lobes of 7th sternite.

Female (Fig. 38): Papilla analis proportionally large; sterigma weakly sclerotized, with shortcup-shaped part; ductus bursae broad, corpus bursae tapering from middle proximally; ductus semi-nalis apical.

Diagnosis: This species is allied with semistrigata and cubana but rosariana differs from themin having well developed abdominal scent organs, longer aedeagus and shorter cornutus. A similarscent organ occurs also with S. imaginaria Razowski & Becker, 1986 from Costa Rica and Brazilian(from Santa Catarina) S. naufraga Razowski & Becker, 1982. S. simillima differs from all thesespecies in its long aedeagus. Female genitalia similar to S. onychina Razowski & Becker, 1986 fromCosta Rica and some allied species but rosariana with ductus bursae shorter and broader. From S.delifrobursa Razowski, 1992 from Costa Rica rosariana differs in apical, not lateral, position of duc-tus seminalis.

Etymology: The specific name refers to the type locality.

Saphenista semistrigata Forbes, 1931

Material examined: Four specimens from Patillas (590 m, VII-1987) and 2 males and one fe-male from Cayey (450 m, 2-VIII-1987), and 2 from Maricao 770 m, 12-VIII-1987, Puerto Rico.

Redescription of genitalia is as follows:Male genitalia (Figs 9, 10): Socii long; vinculum without lateral processes; valva slender; sac-

culus simple, small; median part of transtilla long with long terminal processes directed laterally andslight apical concavity; aedeagus slender with long ventral termination; coecum penis very short;cornutus long, curved. Abdominal scent organ absent.

Female genitalia (Fig. 39): Papilla analis proportionally large; cup-shaped part of sterigma shal-low; ductus bursae sclerotized; with base of accessory ductus well sclerotized, originating medially;corpus bursae slightly longer than ductus bursae with proximal ductus seminalis.

Remarks: Facies similar to S. lacteipalpis (Walsingham, 1891) from St. Vincent, West Indies butsemistrigata with strongly sclerotized ductus bursae and basal part of ductus of accessory bursa, longterminal processes of median part of transtilla, and long aedeagus. For the differences to simillimasee diagnosis of that species.




Saphenista simillima Razowski & Becker, sp. n. (Fig. 53)

Holotype female: “Cuba: Stgo. Sier. Maestra P. Cuba, 31-VII-1990, 1500 m, V. O. Becker”;[73591]; GS 114 Mar. Paratypes, 3 males and 2 females with identical labels.

Description: Wing span 9 mm. Head cream; labial palpus ca 1.5, tinged ochreous basally; thoraxbrownish cream. Forewing slender, not expanding terminad; median portion of costa straight; termenoblique, almost straight. Ground colour whitish cream with some pale brownish cream suffusionsand brownish dots. Markings yellowish brown consisting of spots at base, middle and subterminal(extending in a slender, paler fascia to termen); dorsobasal marking slender, reaching median cell;dorsal blotch and a diffuse blotch at end of median cell pale yellowish brown with some brown dots;some concolorous dots along termen. Cilia cream. Hindwing pale brownish, transparent in basal half;cilia pale brownish.

Variation: Ground colour of forewing varies from cream to ferruginous cream with distinct fer-ruginous strigulation. Markings brownish to pale ferruginous brown.

Male genitalia (Figs 11, 12): Base of socii large, terminal parts proportionally short; vinculumsimple; valva moderately broad, curved upwards; sacculus simple, short; median part of transtillalarge; aedeagus slender, bent, with short ventral termination; cornutus moderate, bent.

Female genitalia (Fig. 40): Papilla analis rather slender; cup-shaped part of sterigma short,somewhat asymmetric; ductus bursae in major part sclerotized; base of ductus of accessory bursaslender, sclerotized; corpus bursae small, membranous, minutely spined; ductus seminalis proximal.

Diagnosis: Closely allied with semistrigata but simillima distinguished by brown dorsal blotchof forewing, slender, sclerotized basal part of ductus of accessory bursa, and short ventroterminalpart of aedeagus.

Etymology: The name refers to great genital similarity to semistrigata.

Saphenista cubana Razowski & Becker, sp. n. (Fig. 54)

Holotype male: “Cuba: Stgo. Sier. Maestra P. Cuba 1500 m, 31-VII-1990, V. O. Becker”;[73589]; GS 120 Mar. Paratypes two males and two females, three with same label and one fromHolguin, Pin, Mayari, 640 m, VII-1990.

Diagnosis: Wing span. 13 mm (in paratypes 8-9 mm). Head grayish cream, thorax darker; labialpalpus ca 1.3, whitish. Forewing hardly expanding terminally; costa almost straight; termen weaklyoblique, almost straight. Ground colour cream; suffusions ochreous and pale ochreous brown, dotsbrownish. Postbasal fascia diffuse, yellowish brown; median fascia consisting of concolorous costalblotch and large blackish dorsal blotch; subapical marking weak, brownish yellow, subterminalmarkings indistinct except for blackish median spot. Cilia concolorous with ground colour. Hind-wing whitish tinged with brownish grey, pale basally; cilia similar.

Variation: Ground colour of forewing often whitish, glossy; suffusions more or less dark; mark-ings variable in shape and colour, especially the dorsal blotch.

Male genitalia (Figs 13, 14): Socii large; vinculum simple; valva proportionally broad; sacculusslender, short; median part of transtilla slender with large terminal processes; aedeagus slender withcaulis median; cornutus long, curved near base, not swung. Abdominal scent organ absent.

Female genitalia (Fig. 41): Papilla analis broad; cup-shaped part of sterigma short, concave pos-teriorly; ductus bursae broad, with longitudinal, rather weakly sclerotized folds and anterior wellsclerotized base of accessory bursa; corpus bursae with median broadening and apical ductus semi-nalis.

Diagnosis: Closely allied with semistrigata but cubana distinguished by presence of dark dorsalblotch of forewing and straight, long cornutus; from simillima cubana differs in having broad ductusbursae (all these three species with sclerotized basal part of ductus seminalis).

Etymology: The specific epithet refers to the name of island where there is the type locality.




Platphalonidia holguina Razowski & Becker, sp. n. (Fig. 55)

Holotype male: “Cuba: Holguin Mayari, 400 m, 12-VII-1990, V. O. Becker”, [72540]; GS 128Mar.

Description: Wing span 7 mm. Head white; labial palpus ca 1.5; thorax grayish white. Forewingslightly expanding terminad; costa straight; termen weakly oblique, straight. Ground colour whitish,suffusions brownish. Median fascia yellowish brown, brown subdorsally; subterminal fascia broad,diffuse. Cilia worn, concolorous with ground colour. Hindwing whitish basally, mixed with brownishotherwise; cilia whitish.

Male genitalia (Figs 15, 16): Valva broad to before middle, then slender, curved upwards; ven-tral prominence of sacculus proportionally broad; median part of transtilla broad basally, with out-wardly curved arms of bifurcation; cornutus large.

Female not known.Diagnosis: Allied with P. subolivacea (Walsingham, 1897) from Danish West Indies but hol-

guina with long cornutus, large median part of transtilla and long arms of its bifurcation.Etymology: The specific epithet refers to the type locality.

Platphalonidia remissa Razowski & Becker, sp. n. (Fig. 56)

Holotype male: “Cuba: Holguin Mayari, 400 m, 12-VII-1990, V. O. Becker”; [72030]; GS 129Mar.

Description: Wing span 8.5 mm. Head whitish; labial palpus 1.3, pale brownish cream; thoraxgrayish cream. Forewing weakly expanding terminad; costa straight; termen slightly convex. Groundcolour whitish grey in basal half of wing suffused with grey, in distal third with brownish; dotsbrownish. Markings pale brownish: median fascia convex, subterminal fascia slender. Cilia whitish.Hindwing brownish white basally brownish on periphery; cilia concolorous with wing base.

Male genitalia (Figs 17, 18): Valva broad in basal one-third, then slender, tapering terminally;ventral prominence of sacculus small; median part of transtilla stout, deeply incised apically; cornu-tus large.

Female not known.Diagnosis: Allied with subolivacea and holguina but remissa distinct by broad, short median

part of transtilla and deep bifurcation of its terminal portion.Etymology: The specific name defers to the facies of the moth; Latin: remissa - placid.

Lasiothyris guanana Razowski & Becker, sp. n. (Fig. 57)

Holotype male: “Brit. Virgin Is: Guana I, 0-80 m, 9-23-VII-1987, V. O. Becker & S. E. Miller”;[66728]; GS 137 Mar.

Description: Wing span 7 mm. Forewing not expanding terminally; costa weakly convex; ter-men moderately oblique, slightly convex. Ground colour white, basal part of costa suffused withbrownish, terminal area of wing with pale ochreous yellow. Markings brownish yellow consisting ofcostal half of median fascia and subterminal fascia. Cilia whitish. Hindwing brownish white; ciliapaler.

Male genitalia (Figs 19, 20): Socii broad; arms of vinculum connected by means of a mem-brane; basal portion of valva broad, terminal portion slender, tapering posteriorly; sacculus simple;median part of transtilla tapering apically; juxta moderate; aedeagus extremely long, with long ven-tral termination; coecum penis broad; cornutus very long.

Female not known.Diagnosis: Allied with L. gravida Razowski, 1986 from Mexico, Vera Cruz and L. luminosa

(Razowski & Becker, 1983) from Santa Catarina, Brazil but gounana with longer aedeagus and slen-




der socius; from L. megapenis Razowski & Becker, 1993 Minas Gerais, Brazil gounana differs main-ly in straight cornutus.

Etymology: the specific name refers to the type locality, Island Guana.

Lasiothyris subsorbia Razowski & Becker, sp. n. (Fig. 58)

Holotype male: “Cuba: Stgo. Sier. Maestr P. Cuba, 1500, 31-VII-1990, V. O. Becker”; [73594];GS 145 Mar.

Description: Wing span 10 mm. Head cream; labial palpus ca 1.3 tinged with pale ochreousproximally; thorax cream ochreous. Forewing slightly expanding terminally, broadest beyond mid-dle; costa straight; termen moderately oblique, hardly convex. Ground colour ochreous cream withslight ferruginous suffusions and somewhat darker strigulation. Markings ferruginous; median fasciafrom before mid-dorsum to mid-costa; subterminal fascia directed towards mid-termen. Cilia concol-orous with ground colour. Hindwing whitish hardly mixed with brownish on periphery; cilia whitish.

Male genitalia (Figs 21, 22): Basal part of socii broad; valva broad basally, slender in posteriorhalf; median part of transtilla broad basally where elongate-triangular, very slender in terminal third;aedeagus large; cornutus missing. Scent organ of seventh sternite with short base and slender arms.

Female not known.Diagnosis: Allied with L. sorbia Razowski & Becker, 1993 from the Federal District of Brazil

but in subsorbia basal portion of median part of transtilla gradually tapering posteriorly, sacculuswith reduced ventral termination, aedeagus longer, and abdominal scent organs preserved.

Etymology: The specific name refers to similarity with sorbia; Latin: sub - beneath.

Lasiothyris puertoricana Razowski & Becker, sp. n. (Fig. 59)

Holotype male: “Puerto Rico: Maricao, 770 m 12-VIII-1987, V. O. Becker”; [67637]; GS 158Mar. Paratype female, same label; GS 157 Mar.

Description: Wing span 9 mm. Head cream; labial palpus ca 1.3; thorax dirty cream. Forewinguniformly broad throughout; costa almost straight; termen oblique. Ground colour cream with indis-tinct pale ochreous suffusions. Markings light ferruginous consisting of remnants of median fasciaand subterminal fascia accompanied by vestiges of median suffusion. Cilia cream. Hindwing withelongate apex, cream white slightly mixed with brownish on periphery; cilia cream.

Female darker than male, with stronger suffusions and markings.Male genitalia (Figs 23, 24): Base of socii large, terminal free parts slender; valva proportional-

ly slender, strongly tapering terminally beyond middle; sacculus to become middle of valva, withventral termination; base of median part of transtilla broad, posterior half slender; aedeagus large;cornutus vestigial. Abdominal scent organ on seventh sternite, short, with small, rounded laterallobes.

Female genitalia (Fig. 42): Cup-shaped part of sterigma large, broadest subterminally; ductusbursae proportionally short, with large mediolateral sac and dense longitudinal folds in proximalhalf; corpus bursae rounded, with minute spinulation.

Diagnosis: Allied with Brazilian L. ficta Razowski & Becker, 1983 but ficta distinguished bysimple sacculus, short median part of transtilla and large lateroterminal processes of abdominal scentorgan.

Etymology: The species name refers to the country of origin of this species.

Spinipogon virginanus Razowski & Becker, sp. n. (Fig. 60)

Holotype male: “Brit. Virgin Is: Guana, X-1989, V. O. Becker”; [71034]; GS 126 Mar. Paratypefemale, same label; GS 127 Mar.

Description: Wing span 9 mm. Head cream; labial palpus 1.3, brownish cream; thorax brownish




cream. Forewing not expanding terminally; costa and termen weakly convex. Ground colour whitishcream with weak yellowish suffusions; dots and strigulae brownish. Markings yellowish brown,browner at costa, mixed with brownish grey at dorsum; median fascia divided into three parts; sub-terminal fascia slender; mediotornal blotch oval, with some brown-blacks dots; dots along termenbrownish yellow. Cilia cream with some pale yellowish brown divisions. Hindwing pale brownishgrey, darker on periphery; cilia cream, tinged with brownish grey in apical portion.

Variation: Female darker than male with broad fascia consisting of pale rust brown costal andmedian spots and large blackish grey dorsal blotch. Hindwing brownish, cilia paler.

Male genitalia (Figs 25, 26): End parts of vinculum fused with one another; valva long, slender;sacculus with wedge-shaped postbasal process; median part of transtilla large, with long apical bifur-cation devoid any spines; juxta large; aedeagus arched, with sharp ventral termination.

Female genitalia (Fig. 43): Ovipositor moderately long; apophyses slender, fairly long; papillaeanales slender; posterior edges of postostial sterigma well sclerotized, anteostial part rounded; cup-shaped part short, deeply concave in middle posteriorly; ductus bursae slender, proportionally long;corpus bursae with very large wart strengthened by numerous spines.

Diagnosis: S. virginana is allied with Brazilian S. atrox Razowski & Becker, 1983 (fromParana) and Costarican S. elaphroterus (Razowski & Becker, 1986) but virginanus characterizes withbroad aedeagus and long bifurcation of transtilla; in elaphroterus termination of sacculus is roundedand in L. studiosus (Razowski & Becker, 1993) from Guerrero, Mexico antrum is shorter and ductusbursae longer.

Etymology: The specific epithet is based on the name of Virgin Island where there is the typelocality.

Mimeugnosta particeps Razowski, 1986

Material examined: A pair from Cuba (Pinar Rio, Vinales, 27-IX-1989). The female was untilnow unknown thus its genitalia (Fig. 44) are described and illustrated. Papilla analis slender; apophy-ses proportionally long, apophyses anteriores with apical broadenings; cup-shaped part of sterigmasclerotized, elongate, with proximal corners rounded; ductus bursae with longitudinal sclerotizedfolds; corpus bursae without larger sclerites; ductus of accessory bursa originating in distal portion ofductus bursae.

Mimeugnosta cf. chascax Razowski, 1994

This species was described from Costa Rica. Our insular specimen is from Santiago, Cuba (Tur-guino, 470 m, 27/9-VII-1990) and shows slight differences to the type (straight distal part of aedea-gus, curved base of cornutus, the presence of rudimentary abdominal scent organs).

Eugnosta fraudulenta Razowski & Becker, sp. n. (Fig. 61)

Holotype male: “Br.[itish] W.[est] Indies”; GS 10555; coll. AMNH. Paratypes: one male labeled“Honduras: Siguatepeque, VI-17-1979, Malaise trap, J. A. Chemsal, M. Kichelbacker, W.W. Mid-dlekauft collectors”, coll. University of Berkeley, California; two males: “Cuba: Holguin, Pin. Ma-yari, 640 m, VII-1990, V. O. Becker”; [72250]; GS 151 and 152 and one female with label “Cuba:Holguin, Pin. Mayari, 640 m, VII-1990, V. O. Becker”; [720228]; GS 150 Mar.

Description: Wing span. 13 mm (paratypes: males 9 mm, female 14 mm). Head brownish white;labial palpus over 2, more brownish grey; thorax pale brownish grey. Forewing not expanding termi-nally; costa rather straight; termen hardly convex, moderately oblique. Ground colour whitish withbrownish grey suffusions and brown dots. Markings in form of dark brown median blotch andbrown-grey, in costal part dark brown, white edged subterminal blotch. Cilia brownish grey. Hind-wing whitish tinged with grey-brown (in female pale brownish grey); cilia whitish.




Male genitalia (Figs 27, 28): Terminal portion of tegumen broadening, rounded; socius slender,long; basal half of valva broad, remaining portion distinctly tapering terminally; sacculus moderate;median part of transtilla long, slender; aedeagus stout; cornutus large.

Female genitalia (Fig. 45): Papilla analis slender, proportionally long; cup-shaped part of sterig-ma short, with lateral corners rounded; ductus bursae slender distally, broad in median and proximalportions where numerous folds and spines; corpus bursae in major part densely spined; both ductusbursae and accessory bursa originating near middle of corpus bursae.

Diagnosis: Similar to Brazilian (from the Minas Gerais) Eugnosta tenacia Razowski & Becker,1994 but fraudulenta with small, apical portion of tegumen (similar to E. bimaculana (Robinson,1869) from the U.S.A.) and much broader aedeagus provided with broad ventral termination.

Etymology: The specific epithet refers to characters of the genitalia; Latin: fraudulenta - delu-sive.

Lorita insulicola Razowski & Becker, sp. n. (Fig. 62)

Holotype male: “Brit. Virgin Is: Guana I: 0 - 80 m, 9-23-VII-1987, V. O. Becker & S. E.Miller,” [66730]; GS 133 Mar. Paratype female, same label, GS 134 Mar.; 3 males from Guadeloupe,Vieux-Habitants, Le Bouchu, 9-I-2006 (coll. ISEZ, donated by Dr. J. Etienne) and one paratype (coll.INRA).

Description: Wing span 7.5 mm. Head cream; labial palpus ca 1.5; thorax cream tinged withbrownish. Forewing uniformly broad throughout; costa almost straight; termen oblique. Groundcolour cream with indistinct yellowish admixture and ochreous cream suffusions; basal one-third ofcosta brownish; some brownish dots along dorsum and in terminal portion of wing. Costal portion ofmedian fascia brownish almost connected with dorsal blotch; the latter yellow-brown, brown-grey atwing edge and proximally; subterminal fascia pale yellowish brown. Cilia cream with brownish suf-fusions. Hindwing cream tinged with brown on periphery; cilia pale brownish cream.

Variation: Female darker than male, with more brownish ferruginous markings and slightlypaler suffusions.

Male genitalia (Figs 29, 30): Terminal part of tegumen and socii typical of the genus; valvaelongate, proportionally slender, tapering from middle to apex, with costa weakly convex at middle;sacculus slender, half the length of valva; median part of transtilla strong; vinculum and aedeaguslong.

Female genitalia (Fig. 46): Cup-shaped part of sterigma slightly tapering and rounded proximal-ly, postostial part delicate; ductus bursae short, with median origin of accessory bursa; corpus bursaepiriform, with folds.

Biology: Bread from flowers of Magnifera indica (Anacardiaceae).Distribution: Known from British Virgin Islands and Guadalupe.Diagnosis: Allied with Californian L. baccharivora Pogue, 1988 but insulicola with weak, me-

dian convexity of costa of valva and long, slender aedeagus; from another insular species, lepidu-lana, insulicola differs in slenderer valva with costa weakly convex and in rounded proximal portionof the cup-shaped part of sterigma.

Etymology: The specific epithet refers to insular distribution; Latin: insula - island.

Lorita lepidulana (Forbes, 1931)

Material examined: One pair from Patillas, Puerto Rico (590 m, VIII-1987); one female fromSantiago, Turguino, Cuba (470 m, 27/9-VII-1990), and one male from St. Thomas, Virgin Island,USA (300 m, 25-30-VII-1987).

L. lepidulana was described from Puerto Rico. The genitalia of the Puerto Rican specimens arecharacterized as follows.




Male genitalia (Figs 31, 32): Socii forming a pair of rigid, slender processes; vinculum armslarge fused with one another ventrally; valva broad with costa convex before middle, dorsocaudalportion forming a small lobe and numerous specialized scales of disc; sacculus simple, longer thanhalf the length of valva; median part of transtilla strong; juxta oval; aedeagus typical of the genus.

Female genitalia (Fig. 47): Postostial part of sterigma slender, cup-shaper part large, withoutdorsal concavity as with L. scarificata (Meyrick, 1917); ductus bursae twice longer than cup-shapedportion of sterigma, with longitudinal sclerites and postmedian origin of accessory bursa; corpus bur-sae with folds and minute thorns.

Aethes pinara Razowski & Becker, sp. n. (Fig. 63)

Holotype male: “Cuba: Pinar Rio, Sierra Rosario, 4-6-X-1989, 400 m, V. O. Becker”; [70680];GS 153 Mar. Paratype female, same label; GS 154 Mar.

Description: Wing span 8 mm. Head and thorax white; labial palpus 1.5. Forewing not expand-ing terminally; costa bent near middle, otherwise straight; termen straight, oblique. Ground colourwhitish; base of costa suffused with brownish. Markings brown-yellow with a few brown dots: dor-sobasal blotch slender; median fascia consisting of slender costal part and postmedian spot at dor-sum; subapical spot vestigial; terminal fascia weak. Cilia white. Hindwing whitish tinged with yel-lowish terminally. Cilia white.

Variation: Female (9 mm) with ground colour yellowish white and markings stronger ferrugi-nous brown.

Male genitalia (Figs 33, 34): Valva broad proximally, tapering from the end of sacculus, withweakly oblique caudal edge, terminating in a short, curved thorn like process; ventral edge of saccu-lus almost straight; median part of transtilla slender, tapering terminad; aedeagus large, simple; onespine like cornutus and a large thorny plate in vesica.

Female genitalia (Fig. 48): Sterigma proportionally short, with pair of submedian lobes posteri-orly; ductus bursae in major part distinctly sclerotized; basal portion of ductus of accessory bursasclerotized, curved, originating in basal portion of ductus bursae; corpus bursae with posterior lobe.

Diagnosis: Closely allied with two Mexican species A. ignobilis Razowski, 1994 from Durangoand A. sonorae (Walsingham, 1884) from Sonora but pinara with almost straight caudal edge of val-va, long aedeagus and cornutus; Jamaican A. olibra Razowski, 1994 differs from all the mentionedspecies in having slender distal half of valva and uniformly broad median part of transtilla.

Etymology: The specific name refers to the name of the type locality.

Cochylis sierraemaestrae Razowski & Becker, sp. n. (Fig. 64)

Holotype male: “Cuba: Stgo. Sier. Maestra P. Cuba, 31-VII-1990, 1500 m, V. O. Becker”;[73593]; GS 123 Mar.

Description: Wing span 11 mm. Head cream; labial palpus ca 1.5 tinged with ferruginous to be-yond middle; thorax cream ferruginous proximally. Forewing somewhat expanding terminally; costaalmost straight; termen weakly oblique, straight. Ground colour yellowish cream suffused with fer-ruginous especially at wing base and beyond end of median cell; brown dots along costa and dorsum.Markings: broad chestnut brown fascia from mid-dorsum to mid-costa and much paler slender sub-terminal fascia almost reaching termen. Cilia cream. Hindwing slender, pale brownish grey; ciliasimilar.

Male genitalia (Figs 35, 36): Socii broad; arms of vinculum slender; valva broadest, stronglyconvex postbasally; sacculus slender, with very large, slightly curved process; median part oftranstilla large, hook-shaped; aedeagus very long, curved, with slender ventral termination; coecumpenis long.

Female not known.Diagnosis: Allied with Cochylis caulocatax Razowski, 1984 from Venezuela and the U.S.A. but




sierraemaestrae with larger, slightly curved process of sacculus and longer aedeagus with reducedlateral process.

Etymology: the specific name refers to the range Sierra Maestra.

Acknowledgements

The authors thank Dr. Jean Etienne, for a permission to include his material of Lorita insulicolain this paper and Mr M. Kopec, Kraków who made the genitalia slides, photographs and arrangedthem in plates.

BIBLIOGRAPHY

RAZOWSKI, J., 1994. - Synopsis of the Neotropical Cochylini (Lepidoptera: Tortricidae).– Acta zool. cracov.,37(2): 121-320.

RAZOWSKI, J. & BECKER V.O., 2002. - Systematic and faunistic data on Neotropical Cochylini (Lepidoptera:Tortricidae), with description of new species. Part 1.– Acta zool. cracov., 45(4): 287-316.

RAZOWSKI, J. & BECKER, V. O., 2007 [in press]. - Systematic and faunistic data on Neotropical Cochylini(Lepidoptera: Tortricidae), with description of new species. Part 2. - Acta zool. cracov.

J. R. V. O. B.Institute of Systematics and Evolution of Animals Research AssociatePolish Academy of Sciences Departamento de ZoologiaSlawkowska 17 Universidade de BrasiliaPL-31-016 Kraków, Caixa Postal 04525POLONIA / POLAND 70919-970, Brasilia, DFE-mail: [email protected] BRASIL / BRAZIL

E-mail:[email protected]

(Recibido para publicatión / Received for publication 20-II-2007)(Revisado y acceptado / Revised and accepted 9-III-2007)






Figs. 1-10.– Male genitalia: 1-2. Maricaona maricaonana Razowski & Becker, sp. n., holotype; 3-4.Phalonidia mayarina Razowski & Becker, sp. n., holotype; 5-6. Saphenista turginoa Razowski & Becker, sp.n., holotype; 7-8. Saphenista rosariana Razowski & Becker, sp. n., holotype; 9-10. Saphenista semistrigataForbes, 1931, Porto Rico: Patillas.

1 3

2

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7

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Figs. 11-22.– Male genitalia: 11-12. Saphenista simillima Razowski & Becker, sp. n., paratype; 13-14. Saphenistacubana Razowski & Becker, sp. n., holotype; 15-16. Platphalonidia holguina Razowski & Becker, sp. n.,holotype; 17-18. Platphalonidia remissa Razowski & Becker, sp. n., holotype; 19-20. Lasiothyris guananaRazowski & Becker, sp. n., holotype; 21-22. Lasiothyris subsorbia Razowski & Becker, sp. n., holotype.

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Figs. 23-32. - Male genitalia: 23-24. Lasiothyris puertoricana Razowski & Becker, sp. n., holotype; 25-26.Spinipogon virginanus Razowski & Becker, sp. n., holotype; 27-28. Mimeugnosta particeps Razowski, 1986,Cuba, Vinales; 29-30. Lorita insulicola Razowski & Becker, sp. n., holotype; 31-32. Lorita lepidulana (Forbes,1931), Puerto Rico: Patillas.

2325

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Figs 33-38. - Male and female genitalia: 33-34. Aethes pinara Razowski & Becker, sp. n., paratype; 35-36.Cochylis sierramaesrae Razowski & Becker, sp. n., holotype; 37. Maricaona maricaonana Razowski &Becker, sp. n., paratype; 38. Saphenista rosariana Razowski & Becker, sp. n., paratype.

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Figs 39-42.– Female genitalia: 39. Saphenista semistrigata Forbes, 1931, Puerto Rico, Patillas; 40. Saphenistasimillima Razowski & Becker, sp. n., holotype; 41. Saphenista cubana Razowski & Becker, sp. n., paratype;42. Lasiothyris puertoricana Razowski & Becker, sp. n., paratype.

3940

41

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Figs 43-45.– Female genitalia: 43. Spinipogon virginanus Razowski & Becker, sp. n., paratype; 44.Mimeugnosta particeps Razowski, 1986, Cuba, Vinales; 45. Eugnosta fraudulenta Razowski & Becker, sp. n.,paratype; 46. Lorita insulicola Razowski & Becker, sp. n., paratype.

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Figs 47-48.– Female genitalia: 47. Lorita lepidulana (Forbes, 1931), Puerto Rico, Patillas; 48. Aethes pinaraRazowski & Becker, sp. n., holotype.

47 48




Figs 49-56.– Adults. 49. Maricaona maricaonana Razowski & Becker, sp. n., holotype; 50. Phalonidiamayarina Razowski & Becker, sp. n., holotype; 51. Saphenista turguinoa Razowski & Becker, sp. n., holotype;52. Saphenista rosariana Razowski & Becker, sp. n., holotype; 53. Saphenista simillima Razowski & Becker,sp. n., holotype; 54. Saphenista cubana Razowski & Becker, sp. n., holotype; 55. Platphalonidia holguinaRazowski & Becker, sp. n., holotype; 56. Platphalonidia remissa Razowski & Becker, sp. n., holotype.

49 50

51 52

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Figs 57-64.– Adults. 57. Lasiothyris guanana Razowski & Becker, sp. n., holotype; 58. Lasiothyris subsorbiaRazowski & Becker, sp. n., holotype; 59. Lasiothyris puertoricana Razowski & Becker, sp. n., holotype; 60.Spinipogon virginanus Razowski & Becker, sp. n., holotype; 61. Eugnosta fraudulenta Razowski & Becker, sp.n., holotype; 62. Lorita insulicola Razowski & Becker, sp. n., holotype; 63. Aethes pinara Razowski & Becker,sp. n., holotype; 64. Cochylis sierramaestae Razowski & Becker, sp. n., holotype.

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